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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.990.55542" ID-GBIF-Dataset="1cede2a7-3880-480c-8401-1015e5440145" ID-PMC="PMC7674391" ID-Pensoft-Pub="1313-2970-990-1" ID-Pensoft-UUID="9F51F78CD53F5005A305DE65494002C4" ID-PubMed="33269011" ID-ZooBank="7B73642C278D40F89091B26213C9A704" ModsDocID="1313-2970-990-1" checkinTime="1605112396458" checkinUser="pensoft" docAuthor="Shimizu, So, Broad, Gavin R. &amp; Maeto, Kaoru" docDate="2020" docId="B13DF4CFE4EB582886247C38DB2A3468" docLanguage="en" docName="ZooKeys 990: 1-144" docOrigin="ZooKeys 990" docSource="http://dx.doi.org/10.3897/zookeys.990.55542" docTitle="Enicospilus matsumurai Shimizu 2020, sp. nov." docType="treatment" docUuid="DF52C899-7459-4CC9-8C1C-A2A2FBD46170" docUuidSource="ZooBank" docVersion="5" id="9F51F78CD53F5005A305DE65494002C4" lastPageNumber="1" masterDocId="9F51F78CD53F5005A305DE65494002C4" masterDocTitle="Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1668169845364" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Integrative taxonomy and analysis of species richness patterns of nocturnal Darwin wasps of the genus Enicospilus Stephens (Hymenoptera, Ichneumonidae, Ophioninae) in Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shimizu, So</mods:namePart>
<mods:affiliation>Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan &amp; DC and Overseas Challenge Program for Young Researchers, Japan Society for the Promotion of Science, Tokyo, Japan &amp; Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5202-4552</mods:nameIdentifier>
<mods:nameIdentifier type="email">parasitoidwasp.sou@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Broad, Gavin R.</mods:namePart>
<mods:affiliation>Depertment of Life Sciences, the Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-7223-5333</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maeto, Kaoru</mods:namePart>
<mods:affiliation>Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Rokkodaicho 1 - 1, Nada, Kobe, Hyogo 657 - 8501, Japan</mods:affiliation>
</mods:name>
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<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>990</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>144</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.990.55542</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.990.55542</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-990-1</mods:identifier>
<mods:identifier type="ZooBank">7B73642C278D40F89091B26213C9A704</mods:identifier>
<mods:identifier type="Pensoft-UUID">9F51F78CD53F5005A305DE65494002C4</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="170296366" LSID="urn:lsid:zoobank.org:act:DF52C899-7459-4CC9-8C1C-A2A2FBD46170" httpUri="http://treatment.plazi.org/id/B13DF4CFE4EB582886247C38DB2A3468" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="B13DF4CF-E4EB-5828-8624-7C38DB2A3468" authority="Shimizu" authorityName="Shimizu" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus matsumurai" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="matsumurai" status="sp. nov.">Enicospilus matsumurai Shimizu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">Figure 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific name is derived from one of the greatest Japanese entomologists,
<normalizedToken originalValue="Shônen">Shonen</normalizedToken>
Matsumura.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type series">
<paragraph pageId="0" pageNumber="1">Type series.</paragraph>
<paragraph pageId="0" pageNumber="1">Total of 12 specimens (8♀♀4♂♂): Japan (8♀♀4♂♂).</paragraph>
<paragraph pageId="0" pageNumber="1">
HT: ♀, Mt. Hikosan, Fukuoka Pref.,
<normalizedToken originalValue="Kyûshû">Kyushu</normalizedToken>
, JAPAN, 22.IX.1983, M.T.
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
leg. (LT) (NIAES).
</paragraph>
<paragraph pageId="0" pageNumber="1">
PT: 1♀, Tarumi, Matsuyama, Ehime Pref., Shikoku, JAPAN, 18.IX.1948, M. Miyatake leg. (EUM); 3♀♀, Matsuyama, Ehime Pref., Shikoku, JAPAN, 18.X.1954 (1♀), 19.X.1954 (1♀), 26.XI.1954 (1♀), S. Ueda leg. (EUM); 1♂, Matsuyama, Ehime Pref., Shikoku, JAPAN, 13.I.1952, T. Ishikawa leg. (EUM); 1♀, Teragawa, Hongawa,
<normalizedToken originalValue="Kôchi">Kochi</normalizedToken>
Pref., Shikoku, JAPAN, 22.VII.1989, I. Yamashita leg. (LT) (NIAES); 1♀, Jigokudani,
<normalizedToken originalValue="Kôchi">Kochi</normalizedToken>
City,
<normalizedToken originalValue="Kôchi">Kochi</normalizedToken>
Pref., Shikoku, JAPAN, 7.VIII.1992, I. Yamashita leg. (LT) (NIAES);1♀,
<normalizedToken originalValue="Kôchi">Kochi</normalizedToken>
Pref., Shikoku, JAPAN (MNHA); 1♂, Mt. Hikosan, Fukuoka Pref.,
<normalizedToken originalValue="Kyûshû">Kyushu</normalizedToken>
, JAPAN, 11.X.1980, K. Konishi leg. (LT) (NIAES); 1♂, Nagasaki Pref.,
<normalizedToken originalValue="Kyûshû">Kyushu</normalizedToken>
, JAPAN, V.1935 (NIAES-TIC); 1♂, Mt. Kurodake, Mts.
<normalizedToken originalValue="Kujû">Kuju</normalizedToken>
,
<normalizedToken originalValue="Ōita">Ōita</normalizedToken>
Pref.,
<normalizedToken originalValue="Kyûshû">Kyushu</normalizedToken>
, JAPAN, 3.X.1983, N. Koda &amp; R. Noda leg. (LT) (NIAES).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Eastern Palaearctic region.</paragraph>
<paragraph pageId="0" pageNumber="1">
JAPAN: [Shikoku] Ehime and
<normalizedToken originalValue="Kôchi">Kochi</normalizedToken>
; [
<normalizedToken originalValue="Kyûshû">Kyushu</normalizedToken>
] Fukuoka, Nagasaki and
<normalizedToken originalValue="Ōita">Ōita</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="bionomics">
<paragraph pageId="0" pageNumber="1">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is similar to
<taxonomicName lsidName="E. flavicaput" pageId="0" pageNumber="1" rank="species" species="flavicaput">
<emphasis italics="true" pageId="0" pageNumber="1">E. flavicaput</emphasis>
</taxonomicName>
and some of the type series were misidentified as that species, but
<taxonomicName lsidName="E. matsumurai" pageId="0" pageNumber="1" rank="species" species="matsumurai">
<emphasis italics="true" pageId="0" pageNumber="1">E. matsumurai</emphasis>
</taxonomicName>
sp. nov. is distinguishable from
<taxonomicName lsidName="E. flavicaput" pageId="0" pageNumber="1" rank="species" species="flavicaput">
<emphasis italics="true" pageId="0" pageNumber="1">E. flavicaput</emphasis>
</taxonomicName>
by the shape and position of the central sclerite of the fore wing fenestra (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27F</figureCitation>
), sculpture of the mesosoma (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
), and wing venation (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27F</figureCitation>
). This species also resembles
<taxonomicName lsidName="E. kunigamiensis" pageId="0" pageNumber="1" rank="species" species="kunigamiensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. kunigamiensis</emphasis>
</taxonomicName>
sp. nov. but is distinguishable by several characters (cf. 'Differential
<normalizedToken originalValue="diagnosis">diagnosis'</normalizedToken>
under
<taxonomicName lsidName="E. kunigamiensis" pageId="0" pageNumber="1" rank="species" species="kunigamiensis">
<emphasis italics="true" pageId="0" pageNumber="1">E. kunigamiensis</emphasis>
</taxonomicName>
sp. nov.). In addition, the large size of this species is also a useful diagnostic character.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Female (n = 8). Body length 28.0-30.5 (HT: ca. 30.5) mm.</paragraph>
<paragraph pageId="0" pageNumber="1">
Head with GOI = 2.5-2.9 (HT: 2.8) (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27D</figureCitation>
). Lower face 0.7
<normalizedToken originalValue="×">x</normalizedToken>
as wide as high, strongly shiny, finely punctate with setae (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27B</figureCitation>
). Clypeus 1.3-1.5
<normalizedToken originalValue="×">x</normalizedToken>
(HT: 1.5) as wide as high, sparsely finely punctate with setae, very weakly convex but almost flat in profile, and ventral margin subacute (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27B, D</figureCitation>
). Malar space 0.2
<normalizedToken originalValue="×">x</normalizedToken>
as long as basal mandibular width (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27B, D</figureCitation>
). Mandible moderately twisted by 50-55° (HT: ca. 50°), moderately long, evenly narrowed, outer surface smooth (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27B, D</figureCitation>
). Upper tooth of mandible 1.3-1.4
<normalizedToken originalValue="×">x</normalizedToken>
(HT: 1.4) as long as lower (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27B</figureCitation>
). Frons, vertex and gena strongly shiny with fine setae. Posterior ocellus touching eye (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27B-D</figureCitation>
). Ventral end of occipital carina joining oral carina. Antennae with 59-60 (HT: 60) flagellomeres; first flagellomere 2.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as second; 20th flagellomere 1.7-1.8
<normalizedToken originalValue="×">x</normalizedToken>
(HT: 1.7) as long as wide.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Mesosoma entirely weakly to moderately shiny with setae (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Pronotum diagonally striate (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Mesoscutum 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as maximum width, moderately shiny and finely punctate to smooth with setae, and evenly rounded in profile (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Notauli absent (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Scutellum moderately convex, smooth, with lateral longitudinal carinae along anterior 0.8-1.0 of scutellum. Epicnemium densely punctate with setae. Epicnemial carina strong, straight, strongly inclined to anterior, dorsal end not reaching anterior margin of mesopleuron (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Mesopleuron entirely longitudinally striate (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Submetapleural carina broadened anteriorly (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Metapleuron densely punctate with rather coarse diagonal striae (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
). Propodeum evenly rounded in profile; anterior transverse carina complete; anterior area longitudinally striate; spiracular area smooth; posterior area coarsely rugose; propodeal spiracle elliptical and not joining pleural carina by ridge (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Wings (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27F</figureCitation>
). Fore wing length 19.5-21.5 (HT: ca. 21.0) mm with AI = 0.7-0.8 (HT: 0.8), CI = 0.2-0.3 (HT: 0.2), DI = 0.3, ICI = 0.7-0.8 (HT: 0.8), SDI = 1.2-1.3 (HT: 1.3), SI = 0.1, SRI = 0.2; vein 1m-cu&amp;M slightly sinuous; vein 2r&amp;RS slightly sinuous and angulate; vein RS evenly curved; fenestra and sclerites of discosubmarginal cell as in Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27F</figureCitation>
; proximal sclerite subquadrate, not confluent with distal sclerite, strongly pigmented; central sclerite linear and rather weakly pigmented, subparallel to 2r&amp;RS, positioned in anterodistal part of fenestra; distal sclerite vestigial or absent; proximal corner of marginal cell evenly setose; posterodistal corner of second discal cell 100-110° (HT: ca. 110°) and of subbasal cell 80-85° (HT: ca. 85°); vein 1cu-a antefurcal to M&amp;RS by 0.2
<normalizedToken originalValue="×">x</normalizedToken>
length of 1cu-a. Hind wing with NI = 2.6-2.9 (HT: 2.6); vein RS basally slightly bowed and slightly evenly curved; vein RA with 8-9 (HT: 9) uniform hamuli.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Legs. Hind leg with coxa in profile 1.7-1.8
<normalizedToken originalValue="×">x</normalizedToken>
(HT: 1.7) as long as deep; basitarsus 1.8-1.9
<normalizedToken originalValue="×">x</normalizedToken>
(HT: 1.9) as long as second tarsomere; fourth tarsomere 2.8-3.3
<normalizedToken originalValue="×">x</normalizedToken>
(HT: 2.8) as long as wide; tarsal claw simply pectinate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Metasoma with DMI = 1.3-1.4 (HT: 1.4), PI = 3.1-3.5 (HT: 3.1), THI = 4.3-4.7 (HT: 4.7); thyridium oval, rather small, not strongly impressed; ovipositor sheath not longer than posterior depth of metasoma (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27A</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
Colour (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Enicospilus matsumurai Shimizu, sp. nov. ♀ (HT) from Japan A habitus B head, frontal view C head, dorsal view D head, lateral view E mesosoma, lateral view F central part of fore wing." figureDoi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1">27</figureCitation>
). Entirely red-brown except for apex of mandible black. Wings weakly infuscate; sclerites pigmented and amber; veins red-brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">Male (n = 4). Very similar to female except body length 22.0-30.5 mm; antennae with 55-63 flagellomeres.</paragraph>
<caption doi="10.3897/zookeys.990.55542.figure27" httpUri="https://binary.pensoft.net/fig/473793" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
<taxonomicName authorityName="Shimizu" authorityYear="2020" class="Insecta" family="Ichneumonidae" genus="Enicospilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enicospilus matsumurai" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="matsumurai">
<emphasis italics="true" pageId="0" pageNumber="1">Enicospilus matsumurai</emphasis>
</taxonomicName>
Shimizu, sp. nov. ♀ (HT) from Japan
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
head, frontal view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
head, dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
head, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
mesosoma, lateral view
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
central part of fore wing.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>