481 lines
80 KiB
XML
481 lines
80 KiB
XML
<document ID-CLB-Dataset="276892" ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.109702" ID-GBIF-Dataset="9fe0bff4-b6c2-415d-ba49-0f1e074a9710" ID-Pensoft-Pub="1313-2970-1184-1" ID-Pensoft-UUID="C2F3B4D92BF558A0AB92E0607CB93262" ID-ZooBank="3D745E43FB6C48A8B089DDAEE1A422BB" ModsDocID="1313-2970-1184-1" checkinTime="1699693871686" checkinUser="pensoft" docAuthor="Yamada, Aiki, Nguyen, Dai Dac & Eguchi, Katsuyuki" docDate="2023" docId="B0BFD6587E1157F69104A2110758231D" docLanguage="en" docName="ZooKeys 1184: 1-17" docOrigin="ZooKeys 1184" docPubDate="2023-11-10" docSource="http://dx.doi.org/10.3897/zookeys.1184.109702" docTitle="Eburopone easoana Yamada & Nguyen & Eguchi 2023, sp. nov." docType="treatment" docUuid="73CBF0E9-4D91-4EAD-952B-F324E03D370E" docUuidSource="ZooBank" docVersion="3" id="C2F3B4D92BF558A0AB92E0607CB93262" lastPageNumber="1" masterDocId="C2F3B4D92BF558A0AB92E0607CB93262" masterDocTitle="First discovery of the ant genus Eburopone Borowiec, 2016 (Hymenoptera, Formicidae, Dorylinae) in the Oriental realm, with description of a new species from Vietnam" masterLastPageNumber="17" masterPageNumber="1" pageNumber="1" updateTime="1699701014511" updateUser="ExternalLinkService">
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<mods:title id="C325A199671CCF2AD93C58E81773D098">First discovery of the ant genus Eburopone Borowiec, 2016 (Hymenoptera, Formicidae, Dorylinae) in the Oriental realm, with description of a new species from Vietnam</mods:title>
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<mods:roleTerm id="1062775F220B1BF7FFCBF8542C409214">Author</mods:roleTerm>
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<mods:namePart id="DB1A08DCE904D2ED087CA296D8CB4325">Yamada, Aiki</mods:namePart>
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<mods:nameIdentifier id="13A0721D5610E5D9C9AEF6967CDE5F32" type="ORCID">https://orcid.org/0000-0002-2141-0661</mods:nameIdentifier>
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<mods:affiliation id="FF635D9809CFF44F23F2FA38942C7164">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397, Japan</mods:affiliation>
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<mods:nameIdentifier id="FCBC50F32527C67D65A5CFF4464CD142" type="email">aiki.ymd@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CE65713A5FEB9E9F7DED6FF6BF814336">Nguyen, Dai Dac</mods:namePart>
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<mods:nameIdentifier id="009BA55681D7ED5E0CBFA0B4F5776539" type="ORCID">https://orcid.org/0009-0006-8024-2553</mods:nameIdentifier>
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<mods:affiliation id="EC277DBB12A77A40FD609DB25B92E25C">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397, Japan & Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Cau Giay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="C805EAC8C2860DD7F5290DAF56008E2C">Eguchi, Katsuyuki</mods:namePart>
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<mods:nameIdentifier id="24ECF19291C5673BA77CC5F5A2A337C1" type="ORCID">https://orcid.org/0000-0002-1054-1295</mods:nameIdentifier>
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<mods:affiliation id="F8AE769C3086A59AB3F767A33C3A13D8">Systematic Zoology Laboratory, Department of Biological Sciences, Graduate School of Science, Tokyo Metropolitan University, 1 - 1 Minami-Osawa, Hachioji, Tokyo, 192 - 0397, Japan & Department of International Health and Medical Anthropology, Institute of Tropical Medicine, Nagasaki University, 1 - 12 - 4 Sakamoto, Nagasaki, Nagasaki, 852 - 8523, Japan</mods:affiliation>
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<mods:title id="0D2F677873C24A78AE2CB61C582EB656">ZooKeys</mods:title>
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<mods:date id="5D2ACF3586651B78816205BE70A66483">2023</mods:date>
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<mods:number id="31651E5BF6B6E6F3CC974BA8C1B15C6F">2023-11-10</mods:number>
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<mods:number id="3A92AA061A2569114193B13DBF63CF66">1184</mods:number>
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<treatment id="B0BFD6587E1157F69104A2110758231D" ID-GBIF-Taxon="217559630" LSID="urn:lsid:zoobank.org:act:73CBF0E9-4D91-4EAD-952B-F324E03D370E" httpUri="http://treatment.plazi.org/id/B0BFD6587E1157F69104A2110758231D" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection id="90130CD90B8901EA128957BE155B58F4" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="8893668B5740DE94D5DDCBDD3F8E219F" pageId="0" pageNumber="1">
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<taxonomicName id="C7EE88DAACF34726EA338321747D485A" LSID="https://zoobank.org/73CBF0E9-4D91-4EAD-952B-F324E03D370E" authority="Yamada & Nguyen & Eguchi, 2023" authorityName="Yamada & Nguyen & Eguchi" authorityYear="2023" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone easoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="easoana" status="sp. nov.">Eburopone easoana</taxonomicName>
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<taxonomicNameLabel id="4EE9C80C9874A0DB4F7BB2008FE05149" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="79BCAFD0A7573FEEEB412E0AE3E4C645" pageId="0" pageNumber="1" type="description">
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<paragraph id="5CBFC2AB2C3057361E4FB78F968F6270" pageId="0" pageNumber="1">
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<figureCitation id="2C7D7ACC121177E44BE9BD3EFDDA3B3F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Eburopone easoana sp. nov. worker, holotype, colony Eg 17 ix 19 - 297 A head in full-face view B habitus in lateral view C habitus in dorsal view. Abbreviation: fl - frontal line." figureDoi="10.3897/zookeys.1184.109702.figure2" httpUri="https://binary.pensoft.net/fig/934722" pageId="0" pageNumber="1">Figs 2</figureCitation>
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<figureCitation id="765B5BAAA1F88DE1EAA1E5E4447143D0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">, 3</figureCitation>
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<figureCitation id="4DAF553D52A278F2DFA3F350EF519460" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">, 4</figureCitation>
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<figureCitation id="8E99C1D31950A725A32E17555BB23C22" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">, 5</figureCitation>
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<figureCitation id="D98EDC0D5120E3E251D3766B55AB8F5A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Abdominal segments IV-VII morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype A abdominal segments IV-VII in dorsal view B abdominal segments IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow C cinctus and presclerites of abdominal segment IV in dorsal view D abdominal tergite VII (pygidium) in dorsoposterior view." figureDoi="10.3897/zookeys.1184.109702.figure6" httpUri="https://binary.pensoft.net/fig/934726" pageId="0" pageNumber="1">, 6</figureCitation>
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<figureCitation id="58DFDD8C6C8FB640B0A04804354482DA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of dealate queens of Eburopone easoana sp. nov., colony Eg 17 ix 19 - 297, paratypes (all except D are of the same individual) A head in full-face view B habitus in lateral view C habitus in dorsal view D abdominal segment IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow." figureDoi="10.3897/zookeys.1184.109702.figure7" httpUri="https://binary.pensoft.net/fig/934727" pageId="0" pageNumber="1">, 7</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9970A4445423E453671C14EBA9CFB3DD" pageId="0" pageNumber="1" type="type material">
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<paragraph id="301173554939A6C10615C5C33121D990" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="DA3E50973B447E187BACA3EC839DE7FF" pageId="0" pageNumber="1">
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<emphasis id="263EB4643D0F89A18632A76DB138BE40" bold="true" pageId="0" pageNumber="1">
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<emphasis id="73CEBD342EBBB5CB90AA6CA746E0D3FA" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Vietnam • worker; Vietnam, Dak Lak Province, Ea Kar District, Ea So Nature Reserve; 12.9676°N, 108.5230°E, 478 m alt.; 17 Sept. 2019; K. Eguchi leg.; colony code Eg17ix19-297; IEBR.
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</paragraph>
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<paragraph id="A357031BAA2A4686C9F1E58C2AAE015D" pageId="0" pageNumber="1">
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<emphasis id="98EE975C4F8AB932B1334FE775B9838D" bold="true" pageId="0" pageNumber="1">
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<emphasis id="A5F63BF717A175BF1928302720455691" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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6 workers and 2 dealate queens from the same colony as the holotype; IEBR, MCZC, MHNG, MNHAH.
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</paragraph>
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</subSubSection>
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<subSubSection id="03E02A4E638BE1499FC7B4D79F5C3EB7" pageId="0" pageNumber="1" type="type material">
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<paragraph id="40492F7C6FD6D647132314C85E51CFD2" pageId="0" pageNumber="1">Non-type material examined.</paragraph>
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<paragraph id="951CBF9467187CCD7A732F7C6F7DF71D" pageId="0" pageNumber="1">Two workers from the same colony as the type series, used for SEM and dissecting observations (one of these was used also for DNA barcoding).</paragraph>
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</subSubSection>
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<subSubSection id="A01D5348948B0B6E2CA174EB95AD967F" pageId="0" pageNumber="1" type="dna barcode">
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<paragraph id="825A47F5F4D4D6ECCA6899C37AEB79B4" pageId="0" pageNumber="1">DNA barcode.</paragraph>
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<paragraph id="452DC6475A31DD4DDB5E6D671496D7D4" pageId="0" pageNumber="1">A partial mitochondrial COI sequence of 658 bp length determined from a non-type worker (the same colony as the type series) is deposited at GenBank: accession number, LC776907.</paragraph>
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</subSubSection>
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<subSubSection id="9FDBC7DAF89E47009E8DBF4787E88642" pageId="0" pageNumber="1" type="worker diagnosis">
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<paragraph id="DDFB309F75B62F2F586EA1615D56FC07" pageId="0" pageNumber="1">Worker diagnosis.</paragraph>
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<paragraph id="B42814DD8761626709617E027475470B" pageId="0" pageNumber="1">Body rather bicolored: abdominal segments III-VII distinctively paler-colored than most surfaces of cranium and mesosoma. Frontal line distinct, extending a little beyond mid-length of cranium. Occipital corner in lateral view strongly produced posteriad to form conspicuous angle. Anterior (frontoclypeal) margins of torulo-posttorular complex not forming conspicuous lobes protruding over anterior clypeal margin in full-face view. Mandibles when closed in full-face view forming only a little space between anterior clypeal margin and mandibles. Promesonotal suture faint and inconspicuous. Petiole in dorsal view almost as long as or slightly longer than wide, with weakly convex lateral margins. Subpetiolar process in lateral view rounded lobate, with anterobasal margin weakly emarginate; posteroventral slope gentle, weakly concave. Abdominal segment III in dorsal view subtrapezoidal, strongly wider posteriorly, distinctly wider than long.</paragraph>
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</subSubSection>
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<subSubSection id="45B8B11DC390A92600FA6705F5571ECB" pageId="0" pageNumber="1" type="description">
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<paragraph id="7B674D568297A20D33C4BD486F7B5266" pageId="0" pageNumber="1">Worker measurements (in mm) and indices.</paragraph>
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<paragraph id="104341259CF225B99249CEFC71BDEA32" pageId="0" pageNumber="1">
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<emphasis id="674092F05C76AC88E188540529FF772F" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: HL 0.63, HW 0.55, SL 0.31, WL 0.86, DML 0.73, PH 0.31, PW 0.37, MFL 0.43, PTH 0.26, PTL 0.29, PTW 0.28, A3L 0.35, A3W 0.46, A4L 0.58, A4W 0.62, CI 87, SI 50, DMI 43, DMI2 85, LMI 36, MFI 79, LPI 111, DPI 97, DA3I 131, DA4I 107.
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</paragraph>
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<paragraph id="685BD14091DFB5EC84DB3739E539D529" pageId="0" pageNumber="1">
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<emphasis id="8B1FD7A911A84460D46F9D3F1C470533" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(
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<emphasis id="F394089E790941287F623761B496D85B" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 6): HL 0.63-0.65, HW 0.55-0.57, SL 0.31-0.33, WL 0.89-0.92, DML 0.74-0.80, PH 0.32-0.35, PW 0.37-0.40, MFL 0.44-0.47, PTH 0.27-0.29, PTL 0.30-0.33, PTW 0.29-0.32, A3L 0.37-0.43, A3W 0.47-0.51, A4L 0.56-0.64, A4W 0.64-0.66, CI 86-88, SI 48-51, DMI 42-43, DMI2 84-87, LMI 36-38, MFI 80-83, LPI 111-114, DPI 95-98, DA3I 118-130, DA4I 103-112.
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</paragraph>
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</subSubSection>
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<subSubSection id="8BEA5B8FF381F75060EF6659FB153D87" pageId="0" pageNumber="1" type="description">
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<paragraph id="79A7B38DE1EBBE59CEB3F5D49CE951C8" pageId="0" pageNumber="1">Queen measurements (in mm) and indices.</paragraph>
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<paragraph id="6A1E87D8EBCA7BE6681E7CF5E191F18D" pageId="0" pageNumber="1">
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<emphasis id="3B355C0A02C9F74C0F5D7CB7BA29B088" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(
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<emphasis id="565FE8BB507C3AFD2C9D419CEED4AD9C" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 2): HL 0.70-0.72, HW 0.58-0.61, EL 0.19-0.20, OL 0.05-0.06, SL 0.35-0.36, WL 1.18-1.20, DML 1.07-1.08, PW 0.57-0.58, MFL 0.51-0.53, PTH 0.31, PTL 0.35-0.36, PTW 0.35-0.37, A3L 0.48-0.49, A3W 0.56-0.58, A4L 0.82, A4W 0.73-0.75, CI 84, SI 49-50, EI 28, OI 7-8, DMI 48, DMI2 90-91, MFI 87-88, LPI 111-117, DPI 100-101, DA3I 115-119, DA4I 89-91.
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</paragraph>
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</subSubSection>
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<subSubSection id="56017AB0D6F9F2393BA8808D15CCC282" pageId="0" pageNumber="1" type="description">
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<paragraph id="4413C0E982E992BE974BCA6161B6A398" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="EB015C3488898E0B9E6880C4731C4397" pageId="0" pageNumber="1">
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<emphasis id="28ADA894CFD6B509F0818EC32A1D20A9" bold="true" pageId="0" pageNumber="1">
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Worker.
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<emphasis id="FBDA801C1E212B65064AF2E7873EC1FA" italics="true" pageId="0" pageNumber="1">Body coloration</emphasis>
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.
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</emphasis>
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Body rather bicolored: cranium, mesosoma, and petiole, mostly dark reddish brown; antennae, anterior part of cranium, mandibles, legs, and abdominal segments III-VII paler brownish to yellowish (Fig.
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<figureCitation id="C7A40405D0F89577F4A0B5C6A67643FF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Eburopone easoana sp. nov. worker, holotype, colony Eg 17 ix 19 - 297 A head in full-face view B habitus in lateral view C habitus in dorsal view. Abbreviation: fl - frontal line." figureDoi="10.3897/zookeys.1184.109702.figure2" httpUri="https://binary.pensoft.net/fig/934722" pageId="0" pageNumber="1">2A-C</figureCitation>
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). Pleural endophragmal pit conspicuously blackish.
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</paragraph>
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||
<caption id="07556BE94A3822E0639B27D0026A879A" doi="10.3897/zookeys.1184.109702.figure2" httpUri="https://binary.pensoft.net/fig/934722" pageId="0" pageNumber="1" start="Figure 2" startId="F2">
|
||
<paragraph id="85F9CA7FCCF3B1716876E0A0D5912E38" pageId="0" pageNumber="1">
|
||
<emphasis id="2B2C4E63D7A955BFA364E5282093E109" bold="true" pageId="0" pageNumber="1">Figure 2.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="E5E0BB8224541B7A618A4D714C8B6196" authorityName="Yamada & Nguyen & Eguchi" authorityYear="2023" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone easoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="easoana">
|
||
<emphasis id="31D8BAED45280497E3A5AE92B164D778" italics="true" pageId="0" pageNumber="1">Eburopone easoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. worker, holotype, colony Eg17ix19-297
|
||
<emphasis id="BB17EF6623E22E287F12AF45336C76DF" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
head in full-face view
|
||
<emphasis id="B34EC467AE649C909B4C4E7321DF5770" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
habitus in lateral view
|
||
<emphasis id="8377E800FB1E0E51927E36E625693898" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
habitus in dorsal view. Abbreviation: fl - frontal line.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B286330C7E3DE9FE8FA4EECDA9CCC207" pageId="0" pageNumber="1">
|
||
<emphasis id="07346BDC964A77E194A59ECA1C8FC716" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="E9D0A645A6C2C8E023DBE2574DC99AAD" italics="true" pageId="0" pageNumber="1">Head structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Cranium in full-face view subrectangular, 1.14-1.17
|
||
<normalizedToken id="DEC58443817F4F4772779B9A28158074" originalValue="×">x</normalizedToken>
|
||
longer than wide (CI, 86-88), with convex lateral margins; posterior margin widely weakly concave. Occipital corner in lateral view strongly produced posteriad to form conspicuous angle. Occipital carina (oc, Fig.
|
||
<figureCitation id="F799BEF91B4F4B47DA2BED69EB2B3EE4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3A, B</figureCitation>
|
||
) distinct and completely encircles occiput, forming inconspicuous and slightly undulate carina dorsally and a conspicuous flange lateroventrally. Frontal line (fl, Figs
|
||
<figureCitation id="01F7740A13BACDBE21BF76488C05B2CA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Eburopone easoana sp. nov. worker, holotype, colony Eg 17 ix 19 - 297 A head in full-face view B habitus in lateral view C habitus in dorsal view. Abbreviation: fl - frontal line." figureDoi="10.3897/zookeys.1184.109702.figure2" httpUri="https://binary.pensoft.net/fig/934722" pageId="0" pageNumber="1">2A</figureCitation>
|
||
,
|
||
<figureCitation id="C96D28D8AFCCB6E44EBFE3043014CAE9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3C, D</figureCitation>
|
||
) distinct, extending a little beyond mid-length of cranium. Torulo-posttorular complex (tptc, Fig.
|
||
<figureCitation id="412337D3D1BEC65D88A71215A65B337A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3C-E</figureCitation>
|
||
) in full-face view arrow-like shaped with narrowed median part; anterior (frontoclypeal) margins not forming conspicuous lobes protruding over anterior clypeal margin in full-face view; maximal width of posterior arrowhead-like part a little shorter than major diameter of antennal socket. Anterior clypeal margin evenly weakly concave in full-face view. Lateroclypeal teeth (lct, Fig.
|
||
<figureCitation id="0D59D0BEB4F4CA46DC0BB5E60C657554" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3A, C, D</figureCitation>
|
||
) large, rounded lobate, strongly protruding latero-anteriad to form anteriormost points of cranium, lateroventrally with a weak protrusion (black arrows in Fig.
|
||
<figureCitation id="3C2AC67AE60F5267B1EFA37A799F1910" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3A, C</figureCitation>
|
||
). Parafrontal ridges (pfr, Fig.
|
||
<figureCitation id="ED635B6D00A59087B2FEA466F5AB4DED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3A, C-E</figureCitation>
|
||
) conspicuous and strongly elevated, turning at posteriorly to incompletely surround antennal sockets, and forming well-marginated subtriangular mound-like lobes; its lateral margins evenly rounded arc-like in full-face view; minimal distance from medialmost end of parafrontal ridge to margin of torulo-posttorular complex in full-face view as long as major diameter of antennal socket. Eye and ocelli completely absent. Antenna 12-merous; scape short, just reaching approximately mid-length of cranium when laid backward; antennomere XII distinctively longer than summed length of three preceding antennomeres IX-XI. Mandible subtriangular, with rounded obtuse basal angle and blunt apex, when closed in full-face view forming only a little space between anterior clypeal margin and mandibles (basal angles nearly reaching center line of cranium when mandibles closed); masticatory margin virtually edentate but with a series of feeble inconspicuous denticles. Postgenal ridge (pgr, Fig.
|
||
<figureCitation id="BB6E1016162743C3CE4A54BE0BE6CC71" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4A</figureCitation>
|
||
) externally recognizable as a broad cross-ribbed furrow, ending a little before the level of occipital carina. Hypostomal teeth (hyt, Fig.
|
||
<figureCitation id="3C78B365C8B5C2D2B46355BBA97E06C9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4A</figureCitation>
|
||
) large and conspicuous with rounded apex in ventral view. Atalar acetabulum (ala, Figs
|
||
<figureCitation id="E05E78DFF400F78898E5A64176C01B65" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3C</figureCitation>
|
||
,
|
||
<figureCitation id="C513A4AD356C640E75D4427F879B6864" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4A</figureCitation>
|
||
) strongly produced anterolaterad. Labrum subrectangular with distal margin weakly bilobed with a small median notch; lateral labral process (lbrp, Fig.
|
||
<figureCitation id="D87A7218C40C18995D62444D126410B8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4B</figureCitation>
|
||
) not visible in frontal view, strongly produced distad and truncated tooth-like. Maxillary palps bi-merous; palpomere II elongate, much longer than palpomere I (Fig.
|
||
<figureCitation id="C3EF3519D03BD6A49FFA9E0DD7673E1E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4C</figureCitation>
|
||
). Labial palps tri-merous; palpomere III distinctly shorter than preceding palpomeres; palpomere II strongly curved basally (Fig.
|
||
<figureCitation id="AAF65BF711DF7CE0ADA4322C06D06D51" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="EDEEF99708900E32646E237CA7E0C322" doi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1" start="Figure 3" startId="F3">
|
||
<paragraph id="BBD1E21A439E3D73A325798FBDEC9C1C" pageId="0" pageNumber="1">
|
||
<emphasis id="0F6A2D0F213938E6FF3619DCE9489F8E" bold="true" pageId="0" pageNumber="1">Figure 3.</emphasis>
|
||
Cranium morphology of
|
||
<taxonomicName id="0E421F19A32551269BDEB917368F0A55" authorityName="Yamada & Nguyen & Eguchi" authorityYear="2023" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone easoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="easoana">
|
||
<emphasis id="702CFFECE3CFEFD7D1DA1CD2415A72FD" italics="true" pageId="0" pageNumber="1">Eburopone easoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. workers, colony Eg17ix19-297, holotype (
|
||
<emphasis id="DBD0728AC516A207B33C8D21A3FEDDD4" bold="true" pageId="0" pageNumber="1">A-C, E</emphasis>
|
||
) and nontype (
|
||
<emphasis id="0756276314E780CEE8912B71BA8C42A2" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
)
|
||
<emphasis id="9A57F6B94D020F02EFC8E13CCBE08DE7" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
head in lateral view
|
||
<emphasis id="448316712FD47743F1DD2497419597D5" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
occipital corners in posterior view
|
||
<emphasis id="9600FE6D1C5D88BA41DC3BB221574222" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
anterior part of cranium in ventrofrontal view
|
||
<emphasis id="42423A76764B690D610DEE5880252AC7" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains)
|
||
<emphasis id="29F6FFA2809AB4EF8515B351A8929087" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="4CE36B369D03856E5B894418ECE425BB" doi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
|
||
<paragraph id="3C4989F79663F0513A0EB76392541870" pageId="0" pageNumber="1">
|
||
<emphasis id="7178600B21C5783E28F9FC83CA087FA2" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
|
||
Cranial venter and mouthparts morphology of
|
||
<taxonomicName id="425640D8F9613F3BF8A27890BA05B873" authorityName="Yamada & Nguyen & Eguchi" authorityYear="2023" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone easoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="easoana">
|
||
<emphasis id="56C7F4F9D4B8620EAB051EB3E8771316" italics="true" pageId="0" pageNumber="1">Eburopone easoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. workers, colony Eg17ix19-297, nontypes
|
||
<emphasis id="DE27AF70B7A0FACE28321C1C932EF4CF" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed
|
||
<emphasis id="830340A6CF47B6524D97BA690ACF42B3" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface
|
||
<emphasis id="04DD7F3C6A1DF5E0C2F79E8F7398E083" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows
|
||
<emphasis id="37A84C595049A5296F9886C5795FCAA1" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6DB1419EDC6468766D3694ECB1E6E46A" pageId="0" pageNumber="1">
|
||
<emphasis id="E134F2A88DD621EA48216280B00B20B1" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="5DF5768FDE64142C5D1C60965B769CA7" italics="true" pageId="0" pageNumber="1">Mesosomal structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Mesosoma with evenly and slightly convex dorsum in lateral view (Fig.
|
||
<figureCitation id="EC8DD19D14560319383B2D4450B053DE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A</figureCitation>
|
||
); lateral margins in dorsal view almost parallel, weakly and widely compressed laterally around mesopleura (Fig.
|
||
<figureCitation id="FE6FEBA9E9A88129297AA000E0C58638" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5B</figureCitation>
|
||
). Pronotal flange only weakly marginated posteriorly from collar. Pronotum and mesopleuron unfused in terms of absence of sclerotized fusion, with dorsal part of pronotomesopleural junction apparently connected by a narrow intersegmental membrane (Fig.
|
||
<figureCitation id="BEDA1BCBA81EEE9B7CCA2EC042EC62F8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A, C</figureCitation>
|
||
). Promesonotal suture inconspicuous, only faintly recognizable as a shallow groove (pmns, Fig.
|
||
<figureCitation id="BA34F657C9438D9CE4CAF604ADCC0E44" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5D</figureCitation>
|
||
; may be hardly recognizable under normal optical observation as in Fig.
|
||
<figureCitation id="0287F65A409474F0B0671DFE5955A0C6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5B</figureCitation>
|
||
, but the weak suture can be recognizable by changing angle of observation or lighting condition). Concavity surrounding pleural endophragmal pit (epp, Fig.
|
||
<figureCitation id="551CCFFE557118E355188F7FDD56534C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A, C</figureCitation>
|
||
) deep and conspicuous. Propodeal declivity only weakly marginate dorsally, almost immarginate laterally; dorsal margin weakly arched anteriad in dorsal view (Fig.
|
||
<figureCitation id="8A69B597598A987FB087EB7CFC2CA287" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A, B</figureCitation>
|
||
). Posterodorsal corner of propodeum in lateral view blunt, only weakly angulated. Propodeal lobe evenly rounded in lateral view. Pretarsal claws simple without teeth.
|
||
</paragraph>
|
||
<caption id="77E9A10E8FCE12146300870F1E7B19E1" doi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
|
||
<paragraph id="C6CF9D3797EAFB4B25B38028AC4BD946" pageId="0" pageNumber="1">
|
||
<emphasis id="2759E900EDEDBB812FB54FB391EEB197" bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
|
||
Mesosoma and abdominal segments II-III morphology of
|
||
<taxonomicName id="3A619CFFFCF9B29836605037220AD7EE" authorityName="Yamada & Nguyen & Eguchi" authorityYear="2023" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone easoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="easoana">
|
||
<emphasis id="9152281EBDE9E37918D51A8202B23B29" italics="true" pageId="0" pageNumber="1">Eburopone easoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. workers, colony Eg17ix19-297, holotype (
|
||
<emphasis id="977F64F1B6755C73B0D18B2F57415EE1" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
|
||
) and non-type (
|
||
<emphasis id="21A281E2ACC66C9E968AAFA35A68786E" bold="true" pageId="0" pageNumber="1">C, D</emphasis>
|
||
)
|
||
<emphasis id="36B5B53F5458F60AE4F2843F3F13F1A0" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
mesosoma and abdominal segment II-III in lateral view
|
||
<emphasis id="2BE563948645E340023665E05C0C79CA" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
mesosoma and abdominal segment II-III in dorsal view
|
||
<emphasis id="B3CC630E1BE5A3A52355637EDD1C8244" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view
|
||
<emphasis id="11237AA806ACD6C082713967D4FC938F" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FC3351F650D440184FC470D9C07988C8" pageId="0" pageNumber="1">
|
||
<emphasis id="D30423002066DF833AB65A6762F8472C" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="F17F7BD5BD6DFFD1CA0E1A1CFFB7A1DE" italics="true" pageId="0" pageNumber="1">Metasomal structure</emphasis>
|
||
.
|
||
</emphasis>
|
||
Petiole (abdominal segment II) in dorsal view 1.02-1.05
|
||
<normalizedToken id="F716EFAF624E462D229D39BC3D72F6A7" originalValue="×">x</normalizedToken>
|
||
longer than wide (length measured from anterodorsal corner to posterior margin of posterior petiolar peduncle; DPI, 95-98), with weakly convex lateral margins; posterior margin of posterior petiolar peduncle slightly arched anteriad (Fig.
|
||
<figureCitation id="BC08A1DC2B508C23FB3C445F367A3C50" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5B</figureCitation>
|
||
). Dorsal outline of petiole in lateral view roundly convex; anterolateral petiolar carina dorsally protrude to form a stout prominence with acute apex. Subpetiolar process in lateral view rounded lobate, with anterobasal margin weakly emarginate; posteroventral slope gentle, weakly concave (Fig.
|
||
<figureCitation id="FC39B4FF1CB50271CA289AE8359B67F0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A</figureCitation>
|
||
). Abdominal segment III in dorsal view subtrapezoidal, strongly wider posteriorly, 1.18-1.31
|
||
<normalizedToken id="546E4F91CC0C660AB61348D2EFD89C28" originalValue="×">x</normalizedToken>
|
||
wider than long (DA3I, 118-131; Fig.
|
||
<figureCitation id="ACC78530E85BE7F4541B5161868D2BFC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5B</figureCitation>
|
||
); lateral margins only feebly evenly convex; 1.59-1.62
|
||
<normalizedToken id="90E3DDD6AB4E89746FA08DC588EE76E2" originalValue="×">x</normalizedToken>
|
||
wider than petiole. Abdominal posttergite III subrectangular in lateral view, with almost vertical anterior face; distinctly larger in height than fused poststernite III (Fig.
|
||
<figureCitation id="36C628FAACCFBA88419AE17B7559E096" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A</figureCitation>
|
||
). Abdominal segment IV in dorsal view 1.03-1.12
|
||
<normalizedToken id="C0DD309943A9E778F995D9ECBE39CC1C" originalValue="×">x</normalizedToken>
|
||
wider than long (DA4I, 103-112), with strongly convex lateral margins (Fig.
|
||
<figureCitation id="6FC0C803FFEFF836044DDE94F6A3088B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Abdominal segments IV-VII morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype A abdominal segments IV-VII in dorsal view B abdominal segments IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow C cinctus and presclerites of abdominal segment IV in dorsal view D abdominal tergite VII (pygidium) in dorsoposterior view." figureDoi="10.3897/zookeys.1184.109702.figure6" httpUri="https://binary.pensoft.net/fig/934726" pageId="0" pageNumber="1">6A</figureCitation>
|
||
). Putative glandular patch near posterior edge of abdominal poststernite IV feebly recognizable as oval pale area contrasted by little darker-colored proximate surrounding (black arrow in Fig.
|
||
<figureCitation id="5D63B8C05E97FC763F4D6934FB3D5F5F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Abdominal segments IV-VII morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype A abdominal segments IV-VII in dorsal view B abdominal segments IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow C cinctus and presclerites of abdominal segment IV in dorsal view D abdominal tergite VII (pygidium) in dorsoposterior view." figureDoi="10.3897/zookeys.1184.109702.figure6" httpUri="https://binary.pensoft.net/fig/934726" pageId="0" pageNumber="1">6B</figureCitation>
|
||
; no specialized structure such as gland opening recognizable on the external sternal surface).
|
||
</paragraph>
|
||
<caption id="F02DA3775765C56E129F03E09ECDFF0D" doi="10.3897/zookeys.1184.109702.figure6" httpUri="https://binary.pensoft.net/fig/934726" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
|
||
<paragraph id="E0BA04C80FF720E56179DAFF5357638D" pageId="0" pageNumber="1">
|
||
<emphasis id="F4C03ECA9EA7A230959ED9934B295295" bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
|
||
Abdominal segments IV-VII morphology of
|
||
<taxonomicName id="EFAB0B8D86F49F93CF88FF7D082B53E0" authorityName="Yamada & Nguyen & Eguchi" authorityYear="2023" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone easoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="easoana">
|
||
<emphasis id="344B22C0BA120B9DAAD7CAC64B5A1229" italics="true" pageId="0" pageNumber="1">Eburopone easoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. workers, colony Eg17ix19-297, holotype
|
||
<emphasis id="86848415CC40AE2D117C6C75384CA30F" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
abdominal segments IV-VII in dorsal view
|
||
<emphasis id="5BE6AF1A3EB8C327167ED4F60C8F34C2" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
abdominal segments IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow
|
||
<emphasis id="794C815E8EB3B9310975B96F6DD6ADB2" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
cinctus and presclerites of abdominal segment IV in dorsal view
|
||
<emphasis id="02EFB4F147F2FBB298FFCA976A5AF8A5" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
abdominal tergite VII (pygidium) in dorsoposterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="52903AD9354D0F02CE991F7D84F97A63" pageId="0" pageNumber="1">
|
||
<emphasis id="F670AE8F1B2836B860EB6630E429EFF3" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="8F46F797D7629AFBF4AAC2F31C7FE43C" italics="true" pageId="0" pageNumber="1">Setation and sculpture</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body setation relatively sparse. Anterolateral surfaces of cranium coarsely longitudinally rugose (Fig.
|
||
<figureCitation id="F0B3103E77BCAA292D249220EE67CC71" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Eburopone easoana sp. nov. worker, holotype, colony Eg 17 ix 19 - 297 A head in full-face view B habitus in lateral view C habitus in dorsal view. Abbreviation: fl - frontal line." figureDoi="10.3897/zookeys.1184.109702.figure2" httpUri="https://binary.pensoft.net/fig/934722" pageId="0" pageNumber="1">2A</figureCitation>
|
||
); other areas of cranium basically smooth except having sparse seta-bearing foveae; posterolateral surfaces very smooth with few setae and sculpturing (Fig.
|
||
<figureCitation id="1CDB14982EFD1A54B6D1F10DD9F9A4EA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cranium morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A-C, E) and nontype (D) A head in lateral view B occipital corners in posterior view C anterior part of cranium in ventrofrontal view D scanning electron microscopic (SEM) photograph of anterior part of cranium in dorsofrontal view, antennae and mandibles removed (left antennal bulbus remains) E torulo-posttorular complex and parafrontal ridges in posterior vertical view against its posterior face. Black arrows in A and C indicate a weak protrusion on lateroclypeal teeth. Abbreviations: ala - atalar acetabulum; oc - occipital carina; fl - frontal line; lct - lateroclypeal teeth; pfr - parafrontal ridge; tptc - torulo-posttorular complex." figureDoi="10.3897/zookeys.1184.109702.figure3" httpUri="https://binary.pensoft.net/fig/934723" pageId="0" pageNumber="1">3A</figureCitation>
|
||
). Frontal face of labrum reticulate (Fig.
|
||
<figureCitation id="513B314610681C4FA7916A771F23BA45" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4A</figureCitation>
|
||
). Proximal face of maxillary stipes mostly smooth. Dorsum of mesosoma and petiole smooth and shiny with sparse seta-bearing foveae; pronotal flange, lateral surfaces of mesosoma and petiole shiny but superficially reticulate-imbricate (Fig.
|
||
<figureCitation id="5FDBC5A72049084EA95463569B782501" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A-C</figureCitation>
|
||
); propodeal declivity and anterior face of petiole smooth. Abdominal postsclerites III and IV smooth, except sparse seta-bearing foveae. Cinctus of abdominal segment IV almost smooth, with coarse short longitudinal ribs near its anterior margin (Fig.
|
||
<figureCitation id="FD5E116E979F0A6A6566ED22FA6A5FDE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Abdominal segments IV-VII morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype A abdominal segments IV-VII in dorsal view B abdominal segments IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow C cinctus and presclerites of abdominal segment IV in dorsal view D abdominal tergite VII (pygidium) in dorsoposterior view." figureDoi="10.3897/zookeys.1184.109702.figure6" httpUri="https://binary.pensoft.net/fig/934726" pageId="0" pageNumber="1">6C</figureCitation>
|
||
). Presclerites of abdominal segment IV reticulate-imbricate. Abdominal tergites V-VI largely smooth with superficially imbricate anterior face. Abdominal tergite VII (pygidium) coarsely areolate-reticulate except superficially imbricate anterior face, armed with numerous stout chaetae arranged in two or three irregular oblique-longitudinal rows along each posterolateral margin (Fig.
|
||
<figureCitation id="81B711C630351B728FA672DAD7C54BBA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Abdominal segments IV-VII morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype A abdominal segments IV-VII in dorsal view B abdominal segments IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow C cinctus and presclerites of abdominal segment IV in dorsal view D abdominal tergite VII (pygidium) in dorsoposterior view." figureDoi="10.3897/zookeys.1184.109702.figure6" httpUri="https://binary.pensoft.net/fig/934726" pageId="0" pageNumber="1">6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D66AC685EDABE0E5C23D4E64C798E5BF" pageId="0" pageNumber="1">
|
||
<emphasis id="F29BF4101F42E50113CBB418B206EF09" bold="true" pageId="0" pageNumber="1">Dealate queen.</emphasis>
|
||
Largely similar to the worker except for some queen-specific features. Eyes and ocelli large and conspicuous (Fig.
|
||
<figureCitation id="526C9AED4571F639F2FE8A819491C72B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of dealate queens of Eburopone easoana sp. nov., colony Eg 17 ix 19 - 297, paratypes (all except D are of the same individual) A head in full-face view B habitus in lateral view C habitus in dorsal view D abdominal segment IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow." figureDoi="10.3897/zookeys.1184.109702.figure7" httpUri="https://binary.pensoft.net/fig/934727" pageId="0" pageNumber="1">7A, B</figureCitation>
|
||
); eyes circular with ~ 20 ommatidia at maximum diameter in lateral view; minimal distance between margins of median and lateral ocellus as long as major diameter of median ocellus; minimal distance between margins of lateral ocelli ~ 2
|
||
<normalizedToken id="EC8CEFAA1E4391A5FB8DABECDE9AA82E" originalValue="×">x</normalizedToken>
|
||
major diameter of median ocellus. Mesosoma with fully developed flight sclerites and wing basal remnants (Fig.
|
||
<figureCitation id="4C56CA2E1D8A63865C0FB4B485DBC523" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of dealate queens of Eburopone easoana sp. nov., colony Eg 17 ix 19 - 297, paratypes (all except D are of the same individual) A head in full-face view B habitus in lateral view C habitus in dorsal view D abdominal segment IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow." figureDoi="10.3897/zookeys.1184.109702.figure7" httpUri="https://binary.pensoft.net/fig/934727" pageId="0" pageNumber="1">7C</figureCitation>
|
||
); pronotum dorsolaterally coarsely foveolate-reticulate; notauli weakly present as coarse groove; parapsidal lines distinct; mesopleuron almost smooth; oblique mesopleural sulcus deep and conspicuous. Petiole in dorsal view almost as long as wide (DPI, 100-101); abdominal segment III in dorsal view 1.15-1.19
|
||
<normalizedToken id="993FC79F29B97F92FD992498453A9201" originalValue="×">x</normalizedToken>
|
||
wider than long (DA3I, 115-119); abdominal segment IV in dorsal view 1.10-1.12
|
||
<normalizedToken id="02131D633859510F75FBCFD5C589A5B2" originalValue="×">x</normalizedToken>
|
||
longer than wide (DA4I, 89-91). Putative glandular patch near posterior edge of abdominal poststernite IV feebly recognizable as in worker (black arrow in Fig.
|
||
<figureCitation id="0481262C54FB396AEDA50DBBCBFF6CBD" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus of dealate queens of Eburopone easoana sp. nov., colony Eg 17 ix 19 - 297, paratypes (all except D are of the same individual) A head in full-face view B habitus in lateral view C habitus in dorsal view D abdominal segment IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow." figureDoi="10.3897/zookeys.1184.109702.figure7" httpUri="https://binary.pensoft.net/fig/934727" pageId="0" pageNumber="1">7D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="21BBE87EE6A3C53002D58B8F0375D1DB" doi="10.3897/zookeys.1184.109702.figure7" httpUri="https://binary.pensoft.net/fig/934727" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
|
||
<paragraph id="035679434CFB9987BC0FD3F9F0ECF862" pageId="0" pageNumber="1">
|
||
<emphasis id="92BFF26686C5795BFE186F139EEEBE8C" bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
|
||
Habitus of dealate queens of
|
||
<taxonomicName id="108EFF9698FEB72E01DCDA105D169482" authorityName="Yamada & Nguyen & Eguchi" authorityYear="2023" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone easoana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="easoana">
|
||
<emphasis id="4FA4326C6004C8C2463F5E33E692B615" italics="true" pageId="0" pageNumber="1">Eburopone easoana</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., colony Eg17ix19-297, paratypes (all except D are of the same individual)
|
||
<emphasis id="27D3BA7671DEEECAB81AAB4E81BC1238" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
head in full-face view
|
||
<emphasis id="2EF78C35EABAE56B6DCBB9D6B740F3AD" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
habitus in lateral view
|
||
<emphasis id="5BDC6BB76FBB4834DC849F147C9014AF" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
habitus in dorsal view
|
||
<emphasis id="2C34FC3C306A130DC32CD10350C844B7" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
abdominal segment IV-VII in ventral view, with the putative glandular patch of poststernite IV indicated by a black arrow.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A41E7E2DB5D4405CE3BA6328197F7152" pageId="0" pageNumber="1">
|
||
<emphasis id="21EF8712EF9098119765983DE0A0BB51" bold="true" pageId="0" pageNumber="1">Male.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8EDFC0546F2C2C3083DDA3394C6586F6" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="635755FCC855DC635A1061FDA267B6B2" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="52E4F2DACB35284A77F5144A99DFD270" pageId="0" pageNumber="1">
|
||
The specific epithet is named after the type locality, Ea So Nature Reserve:
|
||
<emphasis id="38F7A3D8B5D80E9A204011C1DC92776E" italics="true" pageId="0" pageNumber="1">easo</emphasis>
|
||
combined with the Latin feminine suffix -
|
||
<emphasis id="0CF9C04F88A9B539BC284A0BD90B0F2A" italics="true" pageId="0" pageNumber="1">ana</emphasis>
|
||
, adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="396C01B1613E1CF2447DEA71740975E0" pageId="0" pageNumber="1" type="habitat">
|
||
<paragraph id="AF0B48DD1E14316980C4DC7D3A4EEB9D" pageId="0" pageNumber="1">Habitat.</paragraph>
|
||
<paragraph id="722D2BDD94F11C67441BF179EF4364F3" pageId="0" pageNumber="1">The collecting site of the type colony series is primarily covered with a relatively disturbed secondary evergreen forest. The collector (K. Eguchi) did not record the exact microhabitat and collecting situation of the colony fragment.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DDA4B055509E1B1311776D2B80112B4" pageId="0" pageNumber="1" type="differences from e. wroughtoni">
|
||
<paragraph id="8D5F98944519EEF6D6DE9FB9ADCFC25C" pageId="0" pageNumber="1">
|
||
Differences from
|
||
<taxonomicName id="61C82917E2463A007DF5D92959561524" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
|
||
<emphasis id="BA3B325D6D1B79892B26A89A56168B09" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3C17EAFDFC50CD827DF7C55EC1BF3128" pageId="0" pageNumber="1">
|
||
The worker of
|
||
<taxonomicName id="292281D24ACC72617512F36BF9621677" lsidName="E. easoana" pageId="0" pageNumber="1" rank="species" species="easoana">
|
||
<emphasis id="5E91F208F35A5F5BC2285A55309469F7" italics="true" pageId="0" pageNumber="1">E. easoana</emphasis>
|
||
</taxonomicName>
|
||
is morphologically easily distinguished from the only other valid congener
|
||
<taxonomicName id="0F87F9F65FBEF1CD2FF85AD4860E036C" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
|
||
<emphasis id="735D96B4ECD22719CD840053D5926B7D" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
|
||
</taxonomicName>
|
||
from southern Africa (see also the account of
|
||
<taxonomicName id="6325C84FB8434715E4E8FD04F8647D77" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
|
||
<emphasis id="9586AE95DE81BACA0966CB5C865C763E" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
|
||
</taxonomicName>
|
||
below) by the combination of following characteristics: i) frontal line distinct, extending a little beyond mid-length of cranium; ii) anterior (frontoclypeal) margins of torulo-posttorular complex not forming conspicuous lobes protruding over anterior clypeal margin in full-face view (anterior clypeal margin evenly weakly concave in full-face view); iii) mandibles when closed in full-face view forming only a little space between anterior clypeal margin and mandibles (basal angles nearly reaching center line of cranium when mandibles closed); iv) promesonotal suture faint and inconspicuous; v) abdominal segment III in dorsal view distinctly wider than long, with lateral margins only feebly convex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A0650786AC6D082E84D69C9D8E20429E" pageId="0" pageNumber="1" type="morphological remarks">
|
||
<paragraph id="EFA673BAD3D0831585299F38A1A0B4B5" pageId="0" pageNumber="1">Morphological remarks.</paragraph>
|
||
<paragraph id="716DE679706B0D1817FC1E3BDD6B8F3B" pageId="0" pageNumber="1">
|
||
The morphology of workers of
|
||
<taxonomicName id="95CFB5BC44DDBF7A829E134E91D6453C" lsidName="E. easoana" pageId="0" pageNumber="1" rank="species" species="easoana">
|
||
<emphasis id="88F95C30277016468380BBA170EBE073" italics="true" pageId="0" pageNumber="1">E. easoana</emphasis>
|
||
</taxonomicName>
|
||
is generally consistent with the concept of
|
||
<taxonomicName id="A38B04B5A3639A961C675ABE79EDE859" authorityName="Borowiec" authorityYear="2016" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9FEE837038F369D872127E2554D7C181" italics="true" pageId="0" pageNumber="1">Eburopone</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="14F66F0E107CAF230BC1B839D8EC9897" DOI="https://doi.org/10.3897/zookeys.608.9427" author="Borowiec, ML" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 280" refId="B6" refString="Borowiec, ML, 2016. Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae). ZooKeys 608: 1 - 280, DOI: https://doi.org/10.3897/zookeys.608.9427" title="Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)." url="https://doi.org/10.3897/zookeys.608.9427" volume="608" year="2016">Borowiec (2016)</bibRefCitation>
|
||
, except for the number of labial palpomeres.
|
||
<bibRefCitation id="23AED3A355A53FE516FCC74AAB591109" DOI="https://doi.org/10.3897/zookeys.608.9427" author="Borowiec, ML" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 280" refId="B6" refString="Borowiec, ML, 2016. Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae). ZooKeys 608: 1 - 280, DOI: https://doi.org/10.3897/zookeys.608.9427" title="Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)." url="https://doi.org/10.3897/zookeys.608.9427" volume="608" year="2016">Borowiec (2016)</bibRefCitation>
|
||
stated that both the maxillary and labial palps of
|
||
<taxonomicName id="A62D084FDBC9E868750DE28189085C4F" authorityName="Borowiec" authorityYear="2016" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6DD0C92A20E46D627ACF696878F01E36" italics="true" pageId="0" pageNumber="1">Eburopone</emphasis>
|
||
</taxonomicName>
|
||
are bi-merous in workers. However,
|
||
<taxonomicName id="1EBA940190A47EF722853104AF38CC76" lsidName="E. easoana" pageId="0" pageNumber="1" rank="species" species="easoana">
|
||
<emphasis id="5410763FB935B28B09FAEDF9F559BF17" italics="true" pageId="0" pageNumber="1">E. easoana</emphasis>
|
||
</taxonomicName>
|
||
has a bi-merous maxillary and tri-merous labial palps (Fig.
|
||
<figureCitation id="64BC7FC60EFB2EFAF773938E6C4B430A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cranial venter and mouthparts morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, nontypes A scanning electron microscopic (SEM) photograph of cranium in ventral view, mandibles removed B transmission light microscopic (TLM) photograph of labrum in frontal view, partly focus-stacked to emphasize lateral labral processes on caudal surface C TLM photograph of right maxilla in ventral view, with maxillary palpomeres indicated by black arrows D TLM photograph of labium in oblique lateral view, left side, with labial palpomeres indicated by black arrows. Abbreviations: oc - occipital carina; ala - atalar acetabulum; hyt - hypostomal teeth; lbrp - lateral labral process; pgr - postgenal ridge." figureDoi="10.3897/zookeys.1184.109702.figure4" httpUri="https://binary.pensoft.net/fig/934724" pageId="0" pageNumber="1">4C, D</figureCitation>
|
||
), indicating the inter-specific variation of the character within the genus. According to
|
||
<bibRefCitation id="BAC9463FAA4CABE8D6643F9AEB8C9084" DOI="https://doi.org/10.3897/zookeys.608.9427" author="Borowiec, ML" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 280" refId="B6" refString="Borowiec, ML, 2016. Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae). ZooKeys 608: 1 - 280, DOI: https://doi.org/10.3897/zookeys.608.9427" title="Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)." url="https://doi.org/10.3897/zookeys.608.9427" volume="608" year="2016">Borowiec (2016)</bibRefCitation>
|
||
, in dorylines, the labial palps of workers have, as a rule, fewer palpomeres than the maxillary palps of the same individual, except for the New World army ants and
|
||
<taxonomicName id="B2C844FAC47904C7379977D565938D20" authorityName="Mayr" authorityYear="1887" class="Hexapoda" family="Formicidae" genus="Acanthostichus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acanthostichus" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="72EB61656F775000A9CF30696C6D719C" italics="true" pageId="0" pageNumber="1">Acanthostichus</emphasis>
|
||
</taxonomicName>
|
||
Mayr, 1887. Therefore,
|
||
<taxonomicName id="A5B959D9D642D78AE8EE507B94ACC674" lsidName="E. easoana" pageId="0" pageNumber="1" rank="species" species="easoana">
|
||
<emphasis id="3A5885FB40FAD1858E8D519508D55CEF" italics="true" pageId="0" pageNumber="1">E. easoana</emphasis>
|
||
</taxonomicName>
|
||
represents a new exception against this rule. The known palp formula of
|
||
<taxonomicName id="945135CEA898F20B29D2E3DBCB776D8B" authorityName="Borowiec" authorityYear="2016" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E818944FC748021F047E54D268BF6FD0" italics="true" pageId="0" pageNumber="1">Eburopone</emphasis>
|
||
</taxonomicName>
|
||
worker is updated to 2, 2 or 2, 3.
|
||
</paragraph>
|
||
<paragraph id="A997FE1B98D9B1244ECA0D4AA3AAB2D1" pageId="0" pageNumber="1">
|
||
<bibRefCitation id="8489044BA0540244542F5CB20B6A222D" DOI="https://doi.org/10.3897/zookeys.608.9427" author="Borowiec, ML" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 280" refId="B6" refString="Borowiec, ML, 2016. Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae). ZooKeys 608: 1 - 280, DOI: https://doi.org/10.3897/zookeys.608.9427" title="Generic revision of the ant subfamily Dorylinae (Hymenoptera, Formicidae)." url="https://doi.org/10.3897/zookeys.608.9427" volume="608" year="2016">Borowiec (2016)</bibRefCitation>
|
||
described the condition of pronotomesopleural junction in
|
||
<taxonomicName id="AE5E8E1C625F4BB0E14CB8EF10F397A5" authorityName="Borowiec" authorityYear="2016" class="Insecta" family="Formicidae" genus="Eburopone" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eburopone" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B1BAA9769459A5F39C25285CE790FEA7" italics="true" pageId="0" pageNumber="1">Eburopone</emphasis>
|
||
</taxonomicName>
|
||
workers "pronotomesopleural suture visible as groove but not unfused". However, the pronotomesopleural junction of the
|
||
<taxonomicName id="9729870C547F422F57EF68585F8BDBD9" lsidName="E. easoana" pageId="0" pageNumber="1" rank="species" species="easoana">
|
||
<emphasis id="AE76D2E4BDCC8F07322B0730C4111058" italics="true" pageId="0" pageNumber="1">E. easoana</emphasis>
|
||
</taxonomicName>
|
||
workers can be interpretable as unfused in terms of the absence of sclerotized fusion, since the
|
||
<normalizedToken id="C8CC187C6DEBF44288A0B7E117B0E28B" originalValue="“groove”">"groove"</normalizedToken>
|
||
between the two sclerites seems membranous (Fig.
|
||
<figureCitation id="106D5FEA0AFBED4236ABD2F00AB21B50" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesosoma and abdominal segments II-III morphology of Eburopone easoana sp. nov. workers, colony Eg 17 ix 19 - 297, holotype (A, B) and non-type (C, D) A mesosoma and abdominal segment II-III in lateral view B mesosoma and abdominal segment II-III in dorsal view C scanning electron microscopic (SEM) photograph of pronotomesopleural junction in lateral view D SEM photograph of promesonotum in dorsal view. Abbreviations: epp - pleural endophragmal pit; pmns - promesonotal suture." figureDoi="10.3897/zookeys.1184.109702.figure5" httpUri="https://binary.pensoft.net/fig/934725" pageId="0" pageNumber="1">5A, C</figureCitation>
|
||
). Based on examination of the specimen images on AntWeb, the pronotomesopleural junction of
|
||
<taxonomicName id="1AE360A46F23D7184CB7C1678422B10D" lsidName="E. wroughtoni" pageId="0" pageNumber="1" rank="species" species="wroughtoni">
|
||
<emphasis id="A6BF240AB369D3A731F740FEE6B03D24" italics="true" pageId="0" pageNumber="1">E. wroughtoni</emphasis>
|
||
</taxonomicName>
|
||
also seems to be in the same condition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |