194 lines
23 KiB
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194 lines
23 KiB
XML
<document id="7768BDE255B354BECE43E6ABB8DE244D" ID-DOI="10.24199/j.mmv.2013.70.04" ID-ISSN="1447-2554" ID-Zenodo-Dep="12208349" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718988993683" checkinUser="felipe" docAuthor="O’Loughlin, P. Mark, Mackenzie, Melanie & VandenSpiegel, Didier" docDate="2013" docId="B049879AFFC5FFE8592EFCAEFE19F8E1" docLanguage="en" docName="MemMusVictoria.70.37-50.pdf" docOrigin="Memoirs of Museum Victoria 70" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-70-2013/pages-37-50/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Pannychia taylorae O'Loughlin 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="45" masterDocId="4C70FFE2FFC3FFE05A07FFA8FFE6FF8F" masterDocTitle="New sea cucumber species from the seamounts on the Southwest Indian Ocean Ridge (Echinodermata: Holothuroidea: Aspidochirotida, Elasipodida, Dendrochirotida)" masterLastPageNumber="50" masterPageNumber="37" pageNumber="43" updateTime="1718995370909" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="93A573FDF5F83D90CA5D6F97B003C2AE">New sea cucumber species from the seamounts on the Southwest Indian Ocean Ridge (Echinodermata: Holothuroidea: Aspidochirotida, Elasipodida, Dendrochirotida)</mods:title>
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<mods:namePart id="F3CC810E720701A18A2572862411DDEB">O’Loughlin, P. Mark</mods:namePart>
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<mods:namePart id="3075081F337F66542066AD8BBE710607">Mackenzie, Melanie</mods:namePart>
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<mods:namePart id="C0E8C01A7B6EE4664446FCC55C6DFC7D">VandenSpiegel, Didier</mods:namePart>
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<mods:title id="27D9D873E5566D1FBDC2ABC280011F0B">Memoirs of Museum Victoria</mods:title>
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<mods:date id="DD9E494EDD786C938AB3BFAF9D6F791B">2013</mods:date>
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<paragraph id="385F368CFFC5FFE6592EFCAEFB20FC93" blockId="6.[809,1222,774,796]" box="[809,1222,774,796]" pageId="6" pageNumber="43">
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<heading id="631781E0FFC5FFE6592EFCAEFB20FC93" box="[809,1222,774,796]" fontSize="9" level="2" pageId="6" pageNumber="43" reason="5">
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<taxonomicName id="FFE04D0FFFC5FFE6592EFCAEFB95FC93" ID-CoL="6TMBQ" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2013" box="[809,1139,774,796]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="6" pageNumber="43" phylum="Echinodermata" rank="species" species="taylorae" status="sp. nov.">
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<emphasis id="0A94EA9EFFC5FFE6592EFCAEFC10FC93" bold="true" box="[809,1014,774,796]" italics="true" pageId="6" pageNumber="43">Pannychia taylorae</emphasis>
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O’Loughlin
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<taxonomicNameLabel id="11A757E5FFC5FFE65E7EFCAFFB20FC93" box="[1145,1222,775,796]" pageId="6" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
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<uri id="4C713A8EFFC5FFE65EBAFC9AFA47FCCA" box="[1213,1441,818,837]" pageId="6" pageNumber="43">urn:lsid:zoobank.org:act:</uri>
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<paragraph id="385F368CFFC5FFE6592EFCE5FB2AFCEF" blockId="6.[809,1441,818,864]" box="[809,1228,845,864]" pageId="6" pageNumber="43">
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<uuid id="4C460C59FFC5FFE6592EFCE5FB2AFCEF" box="[809,1228,845,864]" pageId="6" pageNumber="43">77705FFB-3331-44AC-9266-BDD53F760E49</uuid>
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<paragraph id="385F368CFFC5FFE6592EFCD0FBC6FC02" blockId="6.[809,1056,887,910]" box="[809,1056,887,910]" pageId="6" pageNumber="43">
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<figureCitation id="A0DB2A09FFC5FFE6592EFCD0FC6EFC02" box="[809,904,888,910]" captionStart="Figure 1" captionStartId="2.[140,199,1830,1849]" captionTargetBox="[140,1434,832,1802]" captionTargetId="figure-334@2.[140,1440,819,1812]" captionTargetPageId="2" captionText="Figure 1. RRS James Cook cruise map showing the Southwest Indian Ocean Ridge and the locations of the five seamounts that were visited during NERC JC066. The hydrothermal vent system studied during JC067 is indicated by the filled circle near Middle of What seamount. The sea cucumbers described in this study were collected from theAtlantis Bank, northeast on the Ridge,and the Coral Seamount,southwest on the Ridge." figureDoi="http://doi.org/10.5281/zenodo.12208351" httpUri="https://zenodo.org/record/12208351/files/figure.png" pageId="6" pageNumber="43">Figures 1</figureCitation>
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,
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<figureCitation id="A0DB2A09FFC5FFE65993FCDFFC44FC02" box="[916,930,887,909]" captionStart="Figure 5" captionStartId="7.[147,207,1060,1079]" captionTargetBox="[233,1361,227,1046]" captionTargetId="figure-9@7.[233,1361,227,1046]" captionTargetPageId="7" captionText="Figure 5. Photos of live holotype specimen of Pannychia taylorae O’Loughlin sp. nov. (NHMUK 2013.5). a, photo of live holotype specimen (in situ on Coral Seamount; taken by ROV during cruise JC066; copyright AD Rogers University of Oxford/NERC); b, ventral view of live holotype (with two commensal polynoid specimens) (photo taken by David Shale and used with permission)." figureDoi="http://doi.org/10.5281/zenodo.12208361" httpUri="https://zenodo.org/record/12208361/files/figure.png" pageId="6" pageNumber="43">5</figureCitation>
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,
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<figureCitation id="A0DB2A09FFC5FFE659ABFCDFFC5CFC02" box="[940,954,887,909]" captionStart="Figure 6" captionStartId="7.[147,207,1827,1846]" captionTargetBox="[291,1305,1179,1809]" captionTargetId="figure-70@7.[291,1305,1179,1809]" captionTargetPageId="7" captionText="Figure 6. Photos of holotype of Pannychia taylorae O’Loughlin sp. nov. (preserved, 180 mm long, NHMUK 2013.5). Ventral view of holotype with oral end below and midventral tube feet more numerous posteriorly. Insert with ventral view of damaged oral region showing non-retractile tentacles." figureDoi="http://doi.org/10.5281/zenodo.12208363" httpUri="https://zenodo.org/record/12208363/files/figure.png" pageId="6" pageNumber="43">6</figureCitation>
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,
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<figureCitation id="A0DB2A09FFC5FFE659C2FCD0FC34FC02" box="[965,978,888,909]" captionStart="Figure 7" captionStartId="8.[140,200,1435,1454]" captionTargetBox="[140,1440,228,1418]" captionTargetId="figure-130@8.[140,1440,227,1418]" captionTargetPageId="8" captionText="Figure 7. SEM images of ossicles from holotype of Pannychia taylorae O’Loughlin sp. nov. (NHMUK 2013.5). a, tentacle rods with outer surface spines (scale bar 50 µm; small plate ossicle probable contaminant from body wall); b, wheels from dorsal body wall (scale bars 100 µm); c, wheels and small plates from dorsal papilla (scale bars 50 µm); d, small concave plates from ventral body wall (scale bars 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.12208365" httpUri="https://zenodo.org/record/12208365/files/figure.png" pageId="6" pageNumber="43">7</figureCitation>
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, table 1
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</paragraph>
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<paragraph id="385F368CFFC5FFE6592EFC0BFB4FFC63" blockId="6.[809,1440,931,1004]" pageId="6" pageNumber="43">
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<emphasis id="0A94EA9EFFC5FFE6592EFC0BFC3EFC39" box="[809,984,931,950]" italics="true" pageId="6" pageNumber="43">Material examined</emphasis>
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.
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<typeStatus id="E75B882EFFC5FFE659E2FC0BFBDAFC39" box="[997,1084,931,950]" pageId="6" pageNumber="43" type="holotype">Holotype</typeStatus>
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. Southwest Indian Ocean Ridge,
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<emphasis id="0A94EA9EFFC5FFE65F6CFC0BFA46FC39" box="[1387,1440,931,950]" italics="true" pageId="6" pageNumber="43">Coral</emphasis>
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Seamount,
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<geoCoordinate id="5DD4504BFFC5FFE65994FC16FC3EFC5E" box="[915,984,958,977]" degrees="41.38" direction="south" orientation="latitude" pageId="6" pageNumber="43" precision="555" value="-41.38">41.38ºS</geoCoordinate>
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<geoCoordinate id="5DD4504BFFC5FFE659E6FC16FBCDFC5E" box="[993,1067,958,977]" degrees="42.85" direction="east" orientation="longitude" pageId="6" pageNumber="43" precision="555" value="42.85">42.85ºE</geoCoordinate>
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,
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<quantity id="FF189B69FFC5FFE65E30FC16FB66FC5E" box="[1079,1152,958,977]" metricMagnitude="3" metricUnit="m" metricValue="1.286" pageId="6" pageNumber="43" unit="m" value="1286.0">1286 m</quantity>
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, JC066, specimen no. JC066– 204, ROV,
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<date id="4C5E104CFFC5FFE6598AFC71FBE6FC63" box="[909,1024,985,1004]" pageId="6" pageNumber="43" value="2011-11-13">13 Nov 2011</date>
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, NHMUK 2013.5.
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</paragraph>
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<paragraph id="385F368CFFC5FFE6592EFBACFC77FAAD" blockId="6.[809,1441,1027,1726]" pageId="6" pageNumber="43">
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<emphasis id="0A94EA9EFFC5FFE6592EFBACFC4FFB96" box="[809,937,1028,1049]" italics="true" pageId="6" pageNumber="43">Description.</emphasis>
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Body wall thick, soft; body with low elevation, rounded anteriorly and posteriorly, slight brim marginally, body
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<quantity id="FF189B69FFC5FFE65963FB96FC5FFBDB" box="[868,953,1086,1108]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="43" unit="mm" value="180.0">180 mm</quantity>
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long, up to
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<quantity id="FF189B69FFC5FFE65E3EFB96FB64FBDB" box="[1081,1154,1086,1108]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="43" unit="mm" value="45.0">45 mm</quantity>
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wide; numerous dorsal and lateral thin papillae evident in photo of live animal, few remaining on preserved
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<typeStatus id="E75B882EFFC5FFE65E2BFBD1FB61FB00" box="[1068,1159,1145,1167]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
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, up to about
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<quantity id="FF189B69FFC5FFE65F1FFBD1FA85FB00" box="[1304,1379,1145,1167]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="43" unit="mm" value="15.0">15 mm</quantity>
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long; mouth subventral, anus dorsoposterior; damaged orally, 16 of 20 tentacles remaining; single series of 19 tube feet on each lateroventral ambulacrum, median ventral ambulacrum with 23 smaller tube feet in paired series posteriorly, more scattered anteriorly.
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</paragraph>
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<paragraph id="385F368CFFC5FFE65948FA81FC0CF9EA" blockId="6.[809,1441,1027,1726]" pageId="6" pageNumber="43">Ossicles in dorsal and ventral body wall and dorsal papillae abundant wheels and small concave oval to round plates; wheels up to 328 µm diameter, 9–15 outer spokes, rounded triangular teeth between the bases of the spokes, central wheel hub with 5–7 spokes, predominantly 6 never 4, hub covered by a membrane; small plates about 56 µm long, up to about 70 µm long, irregularly oval to round, slightly lobed margin, slightly concave, about 15 perforations, typically 11 marginally with 4 centrally, sometimes 2 larger centrally. Tentacle ossicles rarely branched, curved rods with thick spines on outer surface, rods up to 350 µm long.</paragraph>
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<paragraph id="385F368CFFC5FFE65948F9C5FC31F931" blockId="6.[809,1441,1027,1726]" pageId="6" pageNumber="43">Colour. Live: body pale blue, tentacle and tube foot ends pale brown. Preserved: body off-white, tentacle and tube feet ends pale brown.</paragraph>
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<paragraph id="385F368CFFC5FFE6592EF97CFA47F965" blockId="6.[809,1441,1748,1770]" box="[809,1441,1748,1770]" pageId="6" pageNumber="43">
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<emphasis id="0A94EA9EFFC5FFE6592EF97CFC4CF966" box="[809,938,1748,1769]" italics="true" pageId="6" pageNumber="43">Distribution.</emphasis>
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Southwest Indian Ocean,
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<emphasis id="0A94EA9EFFC5FFE65EA1F97CFB39F966" box="[1190,1247,1748,1769]" italics="true" pageId="6" pageNumber="43">Coral</emphasis>
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Seamount,
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<quantity id="FF189B69FFC5FFE65F56F97CFA7AF965" box="[1361,1436,1748,1770]" metricMagnitude="3" metricUnit="m" metricValue="1.286" pageId="6" pageNumber="43" unit="m" value="1286.0">1286 m</quantity>
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.
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</paragraph>
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<paragraph id="385F368CFFC5FFE6592EF8A9FAA9F8E0" blockId="6.[809,1440,1793,1903]" pageId="6" pageNumber="43">
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<emphasis id="0A94EA9EFFC5FFE6592EF8A9FC78F899" box="[809,926,1793,1814]" italics="true" pageId="6" pageNumber="43">Etymology.</emphasis>
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Named for Michelle Taylor (Department of Zoology, University of Oxford) in appreciation of Michelle’s tireless and efficient work in organizing the biological science team and processing the collections for voyage JC066.
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</paragraph>
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<caption id="6C9F6604FFC4FFE75A94FB8CFBFFFBE3" ID-DOI="http://doi.org/10.5281/zenodo.12208361" ID-Zenodo-Dep="12208361" httpUri="https://zenodo.org/record/12208361/files/figure.png" pageId="7" pageNumber="44" startId="7.[147,207,1060,1079]" targetBox="[233,1361,227,1046]" targetPageId="7" targetType="figure">
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<paragraph id="385F368CFFC4FFE75A94FB8CFBFFFBE3" blockId="7.[147,1448,1060,1132]" pageId="7" pageNumber="44">
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Figure 5. Photos of live holotype specimen of
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<taxonomicName id="FFE04D0FFFC4FFE75836FB8CFCB2FBB8" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2013" box="[561,852,1060,1079]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="44" phylum="Echinodermata" rank="species" species="taylorae" status="sp. nov.">
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<emphasis id="0A94EA9EFFC4FFE75836FB8CFD02FBB8" box="[561,740,1060,1079]" italics="true" pageId="7" pageNumber="44">Pannychia taylorae</emphasis>
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O’Loughlin
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</taxonomicName>
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<taxonomicNameLabel id="11A757E5FFC4FFE7595DFB8DFC78FBB7" box="[858,926,1061,1080]" pageId="7" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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(NHMUK 2013.5). a, photo of live holotype specimen (
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<emphasis id="0A94EA9EFFC4FFE75F91FB8DFF52FBDE" italics="true" pageId="7" pageNumber="44">in situ</emphasis>
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on
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<emphasis id="0A94EA9EFFC4FFE75AD3FB97FEEFFBDD" box="[212,265,1087,1106]" italics="true" pageId="7" pageNumber="44">Coral</emphasis>
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Seamount; taken by ROV during cruise JC066; copyright AD Rogers University of Oxford/NERC); b, ventral view of live holotype (with two commensal polynoid specimens) (photo taken by David Shale and used with permission).
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</paragraph>
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</caption>
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<paragraph id="385F368CFFC4FFE75A94F88BFF0FF8E4" blockId="7.[147,1447,1827,1899]" pageId="7" pageNumber="44">
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Figure 6. Photos of holotype of
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<taxonomicName id="FFE04D0FFFC4FFE75BB0F88BFD31F8B9" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2013" box="[439,727,1827,1846]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="7" pageNumber="44" phylum="Echinodermata" rank="species" species="taylorae" status="sp. nov.">
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<emphasis id="0A94EA9EFFC4FFE75BB0F88BFD80F8B9" box="[439,614,1827,1846]" italics="true" pageId="7" pageNumber="44">Pannychia taylorae</emphasis>
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O’Loughlin
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</taxonomicName>
|
||
<taxonomicNameLabel id="11A757E5FFC4FFE758DAF88CFCC5F8B8" box="[733,803,1828,1847]" pageId="7" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(preserved, 180 mm long, NHMUK 2013.5). Ventral view of holotype with oral end below and midventral tube feet more numerous posteriorly. Insert with ventral view of damaged oral region showing non-retractile tentacles.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6C9F6604FFCBFFE85A8BFA33FAC3FA6D" ID-DOI="http://doi.org/10.5281/zenodo.12208365" ID-Zenodo-Dep="12208365" httpUri="https://zenodo.org/record/12208365/files/figure.png" pageId="8" pageNumber="45" startId="8.[140,200,1435,1454]" targetBox="[140,1440,228,1418]" targetPageId="8" targetType="figure">
|
||
<paragraph id="385F368CFFCBFFE85A8BFA33FAC3FA6D" blockId="8.[140,1440,1435,1506]" pageId="8" pageNumber="45">
|
||
Figure 7. SEM images of ossicles from holotype of
|
||
<taxonomicName id="FFE04D0FFFCBFFE85851FA33FC95FA21" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2013" box="[598,883,1435,1454]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="45" phylum="Echinodermata" rank="species" species="taylorae" status="sp. nov.">
|
||
<emphasis id="0A94EA9EFFCBFFE85851FA33FCE2FA21" box="[598,772,1435,1454]" italics="true" pageId="8" pageNumber="45">Pannychia taylorae</emphasis>
|
||
O’Loughlin
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="11A757E5FFCBFFE8597FFA33FC5AFA21" box="[888,956,1435,1454]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(NHMUK 2013.5). a, tentacle rods with outer surface spines (scale bar 50 µm; small plate ossicle probable contaminant from body wall); b, wheels from dorsal body wall (scale bars 100 µm); c, wheels and small plates from dorsal papilla (scale bars 50 µm); d, small concave plates from ventral body wall (scale bars 10 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="385F368CFFCBFFE85A8BF9DAFE19F8E1" blockId="8.[140,771,1650,1903]" pageId="8" pageNumber="45">
|
||
<emphasis id="0A94EA9EFFCBFFE85A8BF9DAFF0AF908" box="[140,236,1650,1671]" italics="true" pageId="8" pageNumber="45">Remarks.</emphasis>
|
||
The single
|
||
<typeStatus id="E75B882EFFCBFFE85B65F9DBFE68F907" box="[354,398,1651,1672]" pageId="8" pageNumber="45">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="FFE04D0FFFCBFFE85811F9DBFCE4F907" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2013" box="[534,770,1650,1672]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="45" phylum="Echinodermata" rank="species" species="taylorae" status="sp. nov.">
|
||
<emphasis id="0A94EA9EFFCBFFE85811F9DBFD63F908" box="[534,645,1650,1672]" italics="true" pageId="8" pageNumber="45">P. taylorae</emphasis>
|
||
O’Loughlin
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="11A757E5FFCBFFE85A8BF938FF07F92A" box="[140,225,1680,1701]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is closest in its morphological characters to the Antarctic Admiralty Seamount specimens that we have examined, but we judge that they are not conspecific. These specimens represent a species of
|
||
<taxonomicName id="FFE04D0FFFCBFFE85805F94EFD89F974" authorityName="Theel" authorityYear="1882" box="[514,623,1766,1787]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="45" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="0A94EA9EFFCBFFE85805F94EFD89F974" box="[514,623,1766,1787]" italics="true" pageId="8" pageNumber="45">Pannychia</emphasis>
|
||
</taxonomicName>
|
||
that is quite distinct morphologically and genetically from
|
||
<taxonomicName id="FFE04D0FFFCBFFE85880F8ABFCE4F898" authorityName="Theel" authorityYear="1882" box="[647,770,1794,1816]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="45" phylum="Echinodermata" rank="species" species="moseleyi">
|
||
<emphasis id="0A94EA9EFFCBFFE85880F8ABFCE4F898" box="[647,770,1794,1816]" italics="true" pageId="8" pageNumber="45">P. moseleyi</emphasis>
|
||
</taxonomicName>
|
||
(Davey
|
||
<emphasis id="0A94EA9EFFCBFFE85ADEF888FEE3F8BB" box="[217,261,1823,1845]" italics="true" pageId="8" pageNumber="45">et al</emphasis>
|
||
., in preparation). In table 1 we detail the significant morphological diagnostic differences between
|
||
<taxonomicName id="FFE04D0FFFCBFFE8588DF895FEE5F8E0" authority="O'Loughlin" authorityName="O'Loughlin" authorityYear="2013" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="45" phylum="Echinodermata" rank="species" species="taylorae" status="sp. nov.">
|
||
<emphasis id="0A94EA9EFFCBFFE8588DF895FCE4F8DE" box="[650,770,1852,1874]" italics="true" pageId="8" pageNumber="45">P. taylorae</emphasis>
|
||
O’Loughlin
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="11A757E5FFCBFFE85B0EF8F1FEB0F8E1" box="[265,342,1881,1902]" pageId="8" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="FFE04D0FFFCBFFE85B81F8F2FE1FF8E1" authorityName="Theel" authorityYear="1882" box="[390,505,1881,1903]" class="Holothuroidea" family="Laetmogonidae" genus="Pannychia" kingdom="Animalia" order="Elasipodida" pageId="8" pageNumber="45" phylum="Echinodermata" rank="species" species="moseleyi">
|
||
<emphasis id="0A94EA9EFFCBFFE85B81F8F2FE1FF8E1" box="[390,505,1881,1903]" italics="true" pageId="8" pageNumber="45">P. moseleyi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |