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<document id="889747473B1250EA5EEA1F05CCAF7F4B" ID-DOI="10.11646/zootaxa.5028.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5453967" ID-ZooBank="C678F0FE-1B62-4F34-8A66-449CF9806B50" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630655426570" checkinUser="plazi" docAuthor="Santos, Bruna S., Marques, Mariana P., Bauer, Aaron M. &amp; Ceríaco, Luis M. P." docDate="2021" docId="B007F528FFFAFFF716D7FF36FE7D5B62" docLanguage="en" docName="zootaxa.5028.1.1.pdf" docOrigin="Zootaxa 5028 (1)" docSource="http://zoobank.org/c678f0fe-1b62-4f34-8a66-449cf9806b50" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Philothamnus A.Smith 1847" docType="treatment" docVersion="4" lastPageNumber="35" masterDocId="4C3E8D50FFD8FFD51640FFA1FFD2597B" masterDocTitle="Herpetological results of Francisco Newtons Zoological Expedition to Angola (1903 1906): a taxonomic revision and new records of a forgotten collection" masterLastPageNumber="80" masterPageNumber="1" pageNumber="35" updateTime="1699281765633" updateUser="plazi">
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<mods:title id="0551AB82DC03FB9600851D6ACD1CD89D">Herpetological results of Francisco Newtons Zoological Expedition to Angola (1903 1906): a taxonomic revision and new records of a forgotten collection</mods:title>
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<mods:namePart id="2081BBC11D42CE2287BF4A24465A0475">Santos, Bruna S.</mods:namePart>
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<mods:namePart id="F91774A56C5EE15401062487F3C2407C">Marques, Mariana P.</mods:namePart>
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<mods:namePart id="878058410A09C2F46E77C538E9893147">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="6415EE8CC8C16D82EDE654C8FCC40101">Ceríaco, Luis M. P.</mods:namePart>
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<treatment id="B007F528FFFAFFF716D7FF36FE7D5B62" ID-DOI="http://doi.org/10.5281/zenodo.5495383" ID-Zenodo-Dep="5495383" LSID="urn:lsid:plazi:treatment:B007F528FFFAFFF716D7FF36FE7D5B62" httpUri="http://treatment.plazi.org/id/B007F528FFFAFFF716D7FF36FE7D5B62" lastPageNumber="35" pageId="34" pageNumber="35">
<subSubSection id="70B417B5FFFAFFF716D7FF36FD2C59CA" box="[151,766,151,178]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="3811443EFFFAFFF716D7FF36FD2C59CA" blockId="34.[151,766,151,178]" box="[151,766,151,178]" pageId="34" pageNumber="35">
<heading id="6359F352FFFAFFF716D7FF36FD2C59CA" bold="true" box="[151,766,151,178]" fontSize="11" level="2" pageId="34" pageNumber="35" reason="4">
<emphasis id="0ADA982CFFFAFFF716D7FF36FD2C59CA" bold="true" box="[151,766,151,178]" pageId="34" pageNumber="35">
<taxonomicName id="FFAE3FBDFFFAFFF716D7FF36FE9059C9" authorityName="A.Smith" authorityYear="1847" box="[151,322,151,178]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="0ADA982CFFFAFFF716D7FF36FE9059C9" bold="true" box="[151,322,151,178]" italics="true" pageId="34" pageNumber="35">Philothamnus</emphasis>
</taxonomicName>
cf.
<emphasis id="0ADA982CFFFAFFF7172FFF36FDE359C9" bold="true" box="[367,561,151,178]" italics="true" pageId="34" pageNumber="35">heterolepitdotus</emphasis>
(Günther, 1863)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="70B417B5FFFAFFF716D7FF7EFD645866" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="3811443EFFFAFFF716D7FF7EFD645866" blockId="34.[151,1437,223,537]" pageId="34" pageNumber="35">
<emphasis id="0ADA982CFFFAFFF716D7FF7EFE175982" bold="true" box="[151,453,223,249]" pageId="34" pageNumber="35">
Material (
<specimenCount id="2EA88FB7FFFAFFF71750FF7EFE6A5982" box="[272,440,223,249]" pageId="34" pageNumber="35" type="generic">one specimen</specimenCount>
):
</emphasis>
One adult specimen (MHNCUP/REP 198, formerly UP-MHNFCP-017411;
<figureCitation id="A09558BBFFFAFFF7136BFF7EFA5D5981" box="[1323,1423,223,250]" captionStart="FIGURE S" captionStartId="79.[151,247,1069,1093]" captionTargetBox="[170,1416,164,1044]" captionTargetId="figure-18@79.[170,1416,164,1044]" captionTargetPageId="79" captionText="FIGURE S75. Specimen of Philotamnus cf. heterolepidotus (MHNCUP/REP 198), from unknown locality. Photo by Bruna Santos." figureDoi="http://doi.org/10.5281/zenodo.5454133" httpUri="https://zenodo.org/record/5454133/files/figure.png" pageId="34" pageNumber="35">Fig. S75</figureCitation>
), collected from unknown locality, date unknown.
</paragraph>
</subSubSection>
<subSubSection id="70B417B5FFFAFFF71687FE86FE7D5B62" pageId="34" pageNumber="35" type="discussion">
<paragraph id="3811443EFFFAFFF71687FE86FE7D5B62" blockId="34.[151,1437,223,537]" pageId="34" pageNumber="35">
<emphasis id="0ADA982CFFFAFFF71687FE86FE80583A" bold="true" box="[199,338,295,321]" pageId="34" pageNumber="35">Comments:</emphasis>
<taxonomicName id="FFAE3FBDFFFAFFF71718FE86FD75583A" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[344,679,295,321]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="heterolepidotus">
<emphasis id="0ADA982CFFFAFFF71718FE86FD75583A" box="[344,679,295,321]" italics="true" pageId="34" pageNumber="35">Philothamnus heterolepidotus</emphasis>
</taxonomicName>
is known from sub-Saharan Africa, with a distribution across
<collectingCountry id="40B904AEFFFAFFF71319FE86FF0F581D" name="Sierra Leone" pageId="34" pageNumber="35">Sierra Leone</collectingCountry>
to
<collectingCountry id="40B904AEFFFAFFF71741FEEAFE98581E" box="[257,330,331,357]" name="Sudan" pageId="34" pageNumber="35">Sudan</collectingCountry>
, western
<collectingCountry id="40B904AEFFFAFFF717F3FEEAFDC0581E" box="[435,530,331,357]" name="Ethiopia" pageId="34" pageNumber="35">Ethiopia</collectingCountry>
and
<collectingCountry id="40B904AEFFFAFFF71409FEEDFD40581D" box="[585,658,332,358]" name="Kenya" pageId="34" pageNumber="35">Kenya</collectingCountry>
and south to the
<collectingCountry id="40B904AEFFFAFFF71512FEEAFB61581E" box="[850,1203,331,357]" name="Democratic Republic of the Congo" pageId="34" pageNumber="35">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="40B904AEFFFAFFF712FFFEEAFAC7581E" box="[1215,1301,331,357]" name="Zambia" pageId="34" pageNumber="35">Zambia</collectingCountry>
and
<collectingCountry id="40B904AEFFFAFFF7130AFEEAFA4E581E" box="[1354,1436,331,357]" name="Angola" pageId="34" pageNumber="35">Angola</collectingCountry>
(
<bibRefCitation id="5C3F39CFFFFAFFF716DFFECEFE5258F2" author="Marques, M. P. &amp; Ceriaco, L. M. P. &amp; Blackburn, D. C. &amp; Bauer, A. M." box="[159,384,367,393]" pageId="34" pageNumber="35" pagination="1 - 501" refId="ref38562" refString="Marques, M. P., Ceriaco, L. M. P., Blackburn, D. C. &amp; Bauer, A. M. (2018) Diversity and distribution of the amphibians and terrestrial reptiles of Angola atlas of historical and bibliographic records (1840 - 2017). Proceedings of the California Academy of Sciences, Series 4, 65 (Supplement 2), 1 - 501." type="journal article" year="2018">
Marques
<emphasis id="0ADA982CFFFAFFF71748FED1FEEC58F2" box="[264,318,367,393]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
2018
</bibRefCitation>
). Ferreira cited
<specimenCount id="2EA88FB7FFFAFFF71469FED1FD1F58F2" box="[553,717,367,394]" pageId="34" pageNumber="35" type="generic">two specimens</specimenCount>
with the same temporal formula as this specimen, 1+1 one from “Cambondo” as
<emphasis id="0ADA982CFFFAFFF7170EFE32FDA458D6" box="[334,630,403,429]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="FFAE3FBDFFFAFFF71717FE32FDA458D6" baseAuthorityName="Leach" baseAuthorityYear="1819" box="[343,630,403,429]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="irregularis">Philothamnus irregularis</taxonomicName>
</emphasis>
” and another without locality info as “
<taxonomicName id="FFAE3FBDFFFAFFF71262FE32FAFA58D6" authority="(Ferreira 1906)" authorityName="Pfeffer" authorityYear="1893" baseAuthorityName="Ferreira" baseAuthorityYear="1906" box="[1058,1320,403,429]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="guntheri">
<emphasis id="0ADA982CFFFAFFF71262FE32FAFA58D6" box="[1058,1320,403,429]" italics="true" pageId="34" pageNumber="35">Philothamnus Guntheri</emphasis>
</taxonomicName>
” (
<bibRefCitation id="5C3F39CFFFFAFFF71302FE32FF0158A9" author="Ferreira, J. B." pageId="34" pageNumber="35" pagination="159 - 171" refId="ref37198" refString="Ferreira, J. B. (1906) Algumas especies novas ou pouco conhecidas de amphibios e reptis de Angola (Coleccao Newton - 1903 - 1904). Jornal de Sciencias Mathematicas Physicas e Naturaes, Segunda Serie, 7 (26), 159 - 171." type="journal article" year="1906">Ferreira 1906</bibRefCitation>
), rendering it difficult to assign the aforementioned specimen to either. After reviewing its morphology, which shows three supralabials in contact with the eye, we found it to closely resemble
<emphasis id="0ADA982CFFFAFFF71277FE7DFADA588E" box="[1079,1288,475,501]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="FFAE3FBDFFFAFFF71277FE7DFAD6588E" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[1079,1284,475,501]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="heterolepidotus">P. heterolepidotus</taxonomicName>
,
</emphasis>
to which we here tentatively assign it.
</paragraph>
</subSubSection>
</treatment>
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