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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.179.63994" ID-GBIF-Dataset="26cdc4ef-8ff1-44b7-8882-083ba5531c94" ID-PMC="PMC8275566" ID-Pensoft-Pub="1314-2003-179-91" ID-Pensoft-UUID="441C694B2C9D529395AF32225D065E47" ID-PubMed="34285638" ModsDocID="1314-2003-179-91" checkinTime="1625591121449" checkinUser="pensoft" docAuthor="Qin, Ying, Chen, Hailing, Deng, Zhenhai &amp; Liu, Yan" docDate="2021" docId="AF8E39B6855655B68975809069C885A8" docLanguage="en" docName="PhytoKeys 179: 91-97" docOrigin="PhytoKeys 179" docPubDate="2021-07-05" docSource="http://dx.doi.org/10.3897/phytokeys.179.63994" docTitle="Aphyllorchis yachangensis Ying Qin &amp; Yan Liu 2021, sp. nov." docType="treatment" docVersion="4" id="441C694B2C9D529395AF32225D065E47" lastPageNumber="91" masterDocId="441C694B2C9D529395AF32225D065E47" masterDocTitle="Aphyllorchis yachangensis (Orchidaceae), a new holomycotrophic orchid from China" masterLastPageNumber="97" masterPageNumber="91" pageNumber="91" updateTime="1668140377344" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Aphyllorchis yachangensis (Orchidaceae), a new holomycotrophic orchid from China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Qin, Ying</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2683-206X</mods:nameIdentifier>
<mods:affiliation>Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, Guangxi, 541006, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Hailing</mods:namePart>
<mods:affiliation>Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, Guangxi, 541006, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deng, Zhenhai</mods:namePart>
<mods:affiliation>Management Center of Yachang Orchid National Nature Reserve, Baise, Guangxi, 533200, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Yan</mods:namePart>
<mods:affiliation>Guangxi Institute of Botany, Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences, Guilin, Guangxi, 541006, China</mods:affiliation>
<mods:nameIdentifier type="email">gxibly@163.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-07-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>179</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>91</mods:start>
<mods:end>97</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.179.63994</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.179.63994</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-179-91</mods:identifier>
<mods:identifier type="Pensoft-UUID">441C694B2C9D529395AF32225D065E47</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="183392481" LSID="urn:lsid:plazi:treatment:AF8E39B6855655B68975809069C885A8" httpUri="http://treatment.plazi.org/id/AF8E39B6855655B68975809069C885A8" lastPageNumber="91" pageId="0" pageNumber="91">
<subSubSection pageId="0" pageNumber="91" type="nomenclature">
<paragraph pageId="0" pageNumber="91">
<taxonomicName LSID="AF8E39B6-8556-55B6-8975-809069C885A8" authority="Ying Qin &amp; Yan Liu" authorityName="Ying Qin &amp; Yan Liu" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphyllorchis yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Tracheophyta" rank="species" species="yachangensis" status="sp. nov.">Aphyllorchis yachangensis Ying Qin &amp; Yan Liu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="91">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="description">
<paragraph pageId="0" pageNumber="91">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Aphyllorchis yachangensis A flowering habit B inflorescence C bract D flower in side view E flower in front view F column in top view G column in ventral view H lip, dorsal sepal, lateral sepals, petals, column and ovary I column in side view J lip in side view K lip in top view L pollinaria M anther cap in top view N rhizome and roots. Photographed by Ying Qin from Ying Qin et al. QY 20190719001 (holotype: IBK!)." figureDoi="10.3897/phytokeys.179.63994.figure1" httpUri="https://binary.pensoft.net/fig/562064" pageId="0" pageNumber="91">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Aphyllorchis yachangensis (an individual with dark colors in the same environment with Ying Qin et al. QY 20190719001) A flowering habit B bud C flower in back view D flower in front view. Photographed by Ying Qin from Ying Qin et al. QYYC 20200703009 (paratype: IBK!)." figureDoi="10.3897/phytokeys.179.63994.figure2" httpUri="https://binary.pensoft.net/fig/562065" pageId="0" pageNumber="91">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="diagnosis">
<paragraph pageId="0" pageNumber="91">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="91">
<taxonomicName authorityName="Ying Qin &amp; Yan Liu" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphyllorchis yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Tracheophyta" rank="species" species="yachangensis">
<emphasis italics="true" pageId="0" pageNumber="91">Aphyllorchis yachangensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. caudata" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="91">A. caudata</emphasis>
</taxonomicName>
but differs from the latter mainly by its hypochile with two wings of 1-1.4
<normalizedToken originalValue="×">x</normalizedToken>
0.4-0.6 mm (vs. 2-3
<normalizedToken originalValue="×">x</normalizedToken>
ca. 4 mm), epichile adaxially smooth (vs. densely papillose), lateral lobes triangular-ovate (vs. semicircular), sepals acute at apex (vs. long cuspidate) and column clavate (vs. approximately cylindrical). Detailed morphological comparisons between
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="yachangensis">
<emphasis italics="true" pageId="0" pageNumber="91">A. yachangensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. caudata" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="91">A. caudata</emphasis>
</taxonomicName>
are provided in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison of Aphyllorchis yachangensis and A. caudata." httpUri="http://table.plazi.org/id/3F47E3C082718BA1D1FEC18A4B7ED0C1" pageId="0" pageNumber="91" tableUuid="3F47E3C082718BA1D1FEC18A4B7ED0C1">1</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="3F47E3C082718BA1D1FEC18A4B7ED0C1" httpUri="http://table.plazi.org/id/3F47E3C082718BA1D1FEC18A4B7ED0C1" pageId="0" pageNumber="91" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Table 1.</emphasis>
Morphological comparison of
<taxonomicName authorityName="Ying Qin &amp; Yan Liu" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphyllorchis yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Tracheophyta" rank="species" species="yachangensis">
<emphasis italics="true" pageId="0" pageNumber="91">Aphyllorchis yachangensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. caudata" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="91">A. caudata</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="91">
<table pageId="0" pageNumber="91">
<tr pageId="0" pageNumber="91">
<th colspan="1" pageId="0" pageNumber="91" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="91" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="yachangensis">
<emphasis italics="true" pageId="0" pageNumber="91">A. yachangensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="91" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. caudata" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="caudata">
<emphasis italics="true" pageId="0" pageNumber="91">A. caudata</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Plant size</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">107-113 cm</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">100 cm</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Inflorescence</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">42-47 cm</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">rachis to 50 cm</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Bracts</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
lanceolate,
<emphasis bold="true" pageId="0" pageNumber="91">12-30</emphasis>
<normalizedToken originalValue="×">x</normalizedToken>
2.8-6.8 mm
</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
narrowly lanceolate,
<emphasis bold="true" pageId="0" pageNumber="91">40-48</emphasis>
<normalizedToken originalValue="×">x</normalizedToken>
5-6 mm
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Dorsal sepal</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
lanceolate, 25-40
<normalizedToken originalValue="×">x</normalizedToken>
7.9-10.7 mm,
<emphasis bold="true" pageId="0" pageNumber="91">apex acute</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
linear-lanceolate or lanceolate, 30-35
<normalizedToken originalValue="×">x</normalizedToken>
ca. 8.0 mm,
<emphasis bold="true" pageId="0" pageNumber="91">apex long cuspidate</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Lateral sepals</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
lanceolate, 6.1-8.2 mm wide,
<emphasis bold="true" pageId="0" pageNumber="91">apex acute</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
linear-lanceolate or lanceolate, 6-7 mm wide,
<emphasis bold="true" pageId="0" pageNumber="91">apex long cuspidate</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Petals</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
lanceolate,
<emphasis bold="true" pageId="0" pageNumber="91">22-27</emphasis>
<normalizedToken originalValue="×">x</normalizedToken>
5.6-7.2 mm
</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
lanceolate,
<emphasis bold="true" pageId="0" pageNumber="91">ca. 20</emphasis>
<normalizedToken originalValue="×">x</normalizedToken>
6-7 mm
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Hypochile</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="91">6.6-7.1</emphasis>
mm long, with 2 wings;
<emphasis bold="true" pageId="0" pageNumber="91">
wings semicircular, 1-1.4
<normalizedToken originalValue="×">x</normalizedToken>
0.4-0.6 mm
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="91">2-3</emphasis>
mm long, with 2 wings;
<emphasis bold="true" pageId="0" pageNumber="91">
wings ligulate, 2-3
<normalizedToken originalValue="×">x</normalizedToken>
ca. 4 mm
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Epichile</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="91">1.9-2.2</emphasis>
cm long,
<emphasis bold="true" pageId="0" pageNumber="91">adaxially smooth</emphasis>
;
<emphasis bold="true" pageId="0" pageNumber="91">lateral lobes triangular-ovate</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="91">ca. 1.2</emphasis>
cm long,
<emphasis bold="true" pageId="0" pageNumber="91">adaxially densely papillose; lateral lobes semicircular</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Column</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
1.4-1.6 cm long,
<emphasis bold="true" pageId="0" pageNumber="91">clavate (apex obviously inflated)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">
1.1-1.4 cm long,
<emphasis bold="true" pageId="0" pageNumber="91">approximately cylindrical</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="91">
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">Flowering period</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">late June and July</td>
<td colspan="1" pageId="0" pageNumber="91" rowspan="1">July and August</td>
</tr>
</table>
</paragraph>
<caption doi="10.3897/phytokeys.179.63994.figure1" httpUri="https://binary.pensoft.net/fig/562064" pageId="0" pageNumber="91" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Figure 1.</emphasis>
<taxonomicName authorityName="Ying Qin &amp; Yan Liu" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphyllorchis yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Tracheophyta" rank="species" species="yachangensis">
<emphasis italics="true" pageId="0" pageNumber="91">Aphyllorchis yachangensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="91">A</emphasis>
flowering habit
<emphasis bold="true" pageId="0" pageNumber="91">B</emphasis>
inflorescence
<emphasis bold="true" pageId="0" pageNumber="91">C</emphasis>
bract
<emphasis bold="true" pageId="0" pageNumber="91">D</emphasis>
flower in side view
<emphasis bold="true" pageId="0" pageNumber="91">E</emphasis>
flower in front view
<emphasis bold="true" pageId="0" pageNumber="91">F</emphasis>
column in top view
<emphasis bold="true" pageId="0" pageNumber="91">G</emphasis>
column in ventral view
<emphasis bold="true" pageId="0" pageNumber="91">H</emphasis>
lip, dorsal sepal, lateral sepals, petals, column and ovary
<emphasis bold="true" pageId="0" pageNumber="91">I</emphasis>
column in side view
<emphasis bold="true" pageId="0" pageNumber="91">J</emphasis>
lip in side view
<emphasis bold="true" pageId="0" pageNumber="91">K</emphasis>
lip in top view
<emphasis bold="true" pageId="0" pageNumber="91">L</emphasis>
pollinaria
<emphasis bold="true" pageId="0" pageNumber="91">M</emphasis>
anther cap in top view
<emphasis bold="true" pageId="0" pageNumber="91">N</emphasis>
rhizome and roots. Photographed by Ying Qin from
<emphasis italics="true" pageId="0" pageNumber="91">Ying Qin</emphasis>
et al.
<emphasis italics="true" pageId="0" pageNumber="91">QY20190719001</emphasis>
(holotype: IBK!).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="materials_examined">
<paragraph pageId="0" pageNumber="91">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="91">
<materialsCitation collectingDate="2019-07-19" country="China" county="Baise" elevation="1862" location="Yachang Orchid National Nature Reserve" municipality="Leye county" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<collectingCountry name="China">China</collectingCountry>
.
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
,
<collectingCounty>Baise</collectingCounty>
city,
<collectingMunicipality>Leye county</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AF8E39B6855655B68975809069C885A8:F873B5A1D39FFFB23E555FA2BD53FD21" country="China" county="Baise" municipality="Leye county" name="Yachang Orchid National Nature Reserve" stateProvince="Guangxi">Yachang Orchid National Nature Reserve</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.8619999999999997" unit="m" value="1862.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.8619999999999997" unit="m" value="1862.0">1862 m</elevation>
</quantity>
,
<collectingDate value="2019-07-19">19 July 2019</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="91">Ying Qin</emphasis>
et al.
<emphasis italics="true" pageId="0" pageNumber="91">QY20190719001</emphasis>
(
<emphasis bold="true" italics="true" pageId="0" pageNumber="91">
<typeStatus>holotype</typeStatus>
</emphasis>
: IBK! isotype: IBK!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="description">
<paragraph pageId="0" pageNumber="91">Description.</paragraph>
<paragraph pageId="0" pageNumber="91">Herb holomycotrophic, leafless, 107-113 cm tall. Rhizome ca. 10.8 cm long, 4.9-6.2 mm in diameter, densely noded; internodes 2.4-12 mm long. Roots spreading, stout, 3.4-6.4 mm in diameter. Stem 61-71 cm tall, 4.5-7.3 mm in diameter, yellow-green, usually with many dark purple stripes and spots, with 1.4-4 cm long sheaths; the uppermost sheath lanceolate; other sheaths tubular. Inflorescence terminal, racemose, 42-47 cm long, with 19-23 well-spaced flowers; rachis sparsely glandular puberulent, yellow-green, usually with many dark purple stripes and spots. Bracts lanceolate, shorter than pedicel and ovary, 1.2-3 cm long, 2.8-6.8 mm wide, yellow-green, usually with many dark purple stripes and spots on abaxial surface, sparsely glandular puberulent. Ovary cylindrical, yellow-green, usually with many dark purple stripes and spots, sparsely glandular puberulent, 2.2-2.8 mm in diameter, including pedicel to 2.3-3.5 cm long. Sepals yellow-green, usually with many dark purple stripes and spots on abaxial surface, abaxially sparsely glandular puberulent; dorsal sepal lanceolate, slightly concave, 2.5-4 cm long, 7.9-10.7 mm wide, apex acute, reflexed; lateral sepals lanceolate, slightly oblique at base, 2.5-4 cm long, 6.1-8.2 mm wide, apex acute, reflexed. Petals lanceolate, reflexed, 2.2-2.7 cm long, 5.6-7.2 mm wide, yellow-green, pale purple to dark purple at base, apex acute. Lip 2-partite; hypochile narrow, concave, 6.6-7.1 mm long, 3.6-4.5 mm wide, yellow or reddish brown, with 2 unobvious wings at base; wings semicircular, 1-1.4 mm wide at base, 0.4-0.6 mm tall; epichile ca. 1.9-2.2 cm long, 1.3-1.5 cm wide when flattened, 3-lobed, yellow with pale purple, or purple to dark purple, adaxially smooth, margin slightly erose; lateral lobes triangular-ovate, 8.2-8.9 mm long, 3.6-4.3 mm wide; mid-lobe narrowly triangular, 1.1-1.3 cm long, 5.6-5.9 mm wide, acute at apex, margin slightly involute. Column clavate, arcuate, 1.4-1.6 cm long, 1.8-3.3 mm in diameter in lower part, 4.3-6.1 mm in diameter in upper part, yellow-green, usually purple from middle to base; stigma ovate, slightly concave; staminodes 2, on both sides of apex, yellow-white to pale yellow-green; pollinia 2, ovate, granular-farinaceous, 4 mm long, yellow; anther cap ovate, slightly laterally compressed, 4 mm long, pale yellow.</paragraph>
<caption doi="10.3897/phytokeys.179.63994.figure2" httpUri="https://binary.pensoft.net/fig/562065" pageId="0" pageNumber="91" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">Figure 2.</emphasis>
<taxonomicName authorityName="Ying Qin &amp; Yan Liu" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphyllorchis yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Tracheophyta" rank="species" species="yachangensis">
<emphasis italics="true" pageId="0" pageNumber="91">Aphyllorchis yachangensis</emphasis>
</taxonomicName>
(an individual with dark colors in the same environment with
<emphasis italics="true" pageId="0" pageNumber="91">Ying Qin</emphasis>
et al.
<emphasis italics="true" pageId="0" pageNumber="91">QY20190719001</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="91">A</emphasis>
flowering habit
<emphasis bold="true" pageId="0" pageNumber="91">B</emphasis>
bud
<emphasis bold="true" pageId="0" pageNumber="91">C</emphasis>
flower in back view
<emphasis bold="true" pageId="0" pageNumber="91">D</emphasis>
flower in front view. Photographed by Ying Qin from
<emphasis italics="true" pageId="0" pageNumber="91">Ying Qin</emphasis>
et al.
<emphasis italics="true" pageId="0" pageNumber="91">QYYC20200703009</emphasis>
(paratype: IBK!).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="distribution">
<paragraph pageId="0" pageNumber="91">Distribution, habitat and ecology.</paragraph>
<paragraph pageId="0" pageNumber="91">
<taxonomicName authorityName="Ying Qin &amp; Yan Liu" authorityYear="2021" class="Liliopsida" family="Orchidaceae" genus="Aphyllorchis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphyllorchis yachangensis" order="Asparagales" pageId="0" pageNumber="91" phylum="Tracheophyta" rank="species" species="yachangensis">
<emphasis italics="true" pageId="0" pageNumber="91">Aphyllorchis yachangensis</emphasis>
</taxonomicName>
was discovered in the Yachang Orchid National Nature Reserve, Leye county, Baise city, Guangxi Province, China. The holotype subpopulation is distributed in a subtropical evergreen and deciduous broad-leaved mixed forest, and is found growing with
<taxonomicName class="Monocotyledoneae" family="Gramineae" genus="Chimonobambusa" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Chimonobambusa quadrangularis" order="Poales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="quadrangularis">
<emphasis italics="true" pageId="0" pageNumber="91">Chimonobambusa quadrangularis</emphasis>
</taxonomicName>
(Franceschi) Makino (
<taxonomicName authorityName="Makino" authorityYear="1914" class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Poales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Gramineae</taxonomicName>
),
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Lithocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Lithocarpus glaber" order="Fagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="glaber">
<emphasis italics="true" pageId="0" pageNumber="91">Lithocarpus glaber</emphasis>
</taxonomicName>
(Thunberg) Nakai (
<taxonomicName class="Dicotyledoneae" family="Fagaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Fagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Fagaceae</taxonomicName>
),
<taxonomicName class="Cladi" family="Magnoliaceae" genus="Manglietia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Manglietia fordiana" order="Magnoliales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="fordiana">
<emphasis italics="true" pageId="0" pageNumber="91">Manglietia fordiana</emphasis>
</taxonomicName>
Oliver (
<taxonomicName class="Cladi" family="Magnoliaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Magnoliales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Magnoliaceae</taxonomicName>
),
<taxonomicName class="Dicotyledoneae" family="Juglandaceae" genus="Juglans" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Juglans mandshurica" order="Juglandales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="mandshurica">
<emphasis italics="true" pageId="0" pageNumber="91">Juglans mandshurica</emphasis>
</taxonomicName>
Maximowicz (
<taxonomicName authorityName="Linnaeus" authorityYear="1753" class="Dicotyledoneae" family="Juglandaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Juglandales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Juglandaceae</taxonomicName>
),
<taxonomicName authorityName="T. S. Ying" authorityYear="1979" baseAuthorityName="Gagnep." class="Dicotyledoneae" family="Berberidaceae" genus="Dysosma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Dysosma majorensis" order="Ranunculales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="majorensis">
<emphasis italics="true" pageId="0" pageNumber="91">Dysosma majorensis</emphasis>
</taxonomicName>
(Gagnep.) Ying (
<taxonomicName class="Dicotyledoneae" family="Berberidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Ranunculales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Berberidaceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Liliaceae" genus="Ophiopogon" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ophiopogon bockianus" order="Liliales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="bockianus">
<emphasis italics="true" pageId="0" pageNumber="91">Ophiopogon bockianus</emphasis>
</taxonomicName>
Diels (
<taxonomicName class="Magnoliopsida" family="Liliaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Liliales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Liliaceae</taxonomicName>
),
<taxonomicName authorityName="Hance" authorityYear="1883" class="Magnoliopsida" family="Liliaceae" genus="Disporopsis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Disporopsis fuscopicta" order="Liliales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="fuscopicta">
<emphasis italics="true" pageId="0" pageNumber="91">Disporopsis fuscopicta</emphasis>
</taxonomicName>
Hance (
<taxonomicName class="Magnoliopsida" family="Liliaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Liliales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Liliaceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Orchidaceae" genus="Calanthe" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calanthe brevicornu" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="brevicornu">
<emphasis italics="true" pageId="0" pageNumber="91">Calanthe brevicornu</emphasis>
</taxonomicName>
Lindley (
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Orchidaceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Orchidaceae" genus="Goodyera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Goodyera biflora" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="biflora">
<emphasis italics="true" pageId="0" pageNumber="91">Goodyera biflora</emphasis>
</taxonomicName>
(Lindley) J. D. Hooker (
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Orchidaceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="G. velutina" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="species" species="velutina">
<emphasis italics="true" pageId="0" pageNumber="91">G. velutina</emphasis>
</taxonomicName>
Maximowicz ex Regel (
<taxonomicName class="Magnoliopsida" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="91" phylum="Angiospermae" rank="family">Orchidaceae</taxonomicName>
), etc.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="phenology">
<paragraph pageId="0" pageNumber="91">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="91">Flowering in late June and July, capsules not seen.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="etymology">
<paragraph pageId="0" pageNumber="91">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="91">The specific epithet is derived from the type locality, Yachang Orchid National Nature Reserve.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="chinese name">
<paragraph pageId="0" pageNumber="91">Chinese name.</paragraph>
<paragraph pageId="0" pageNumber="91">
<normalizedToken originalValue="雅长无叶兰">雅长无叶兰</normalizedToken>
(Ya Chang Wu Ye Lan)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="91" type="materials_examined">
<paragraph pageId="0" pageNumber="91">Additional specimens examined</paragraph>
<paragraph pageId="0" pageNumber="91">
<emphasis bold="true" pageId="0" pageNumber="91">
(
<typeStatus>paratypes</typeStatus>
).
</emphasis>
<materialsCitation collectingDate="2018-07-28" country="China" county="Baise" elevation="1862" location="Yachang Orchid National Nature Reserve" municipality="Leye county" specimenCount="1" stateProvince="Guangxi">
<collectingCountry name="China">China</collectingCountry>
:
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
:
<collectingCounty>Baise</collectingCounty>
city,
<collectingMunicipality>Leye county</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AF8E39B6855655B68975809069C885A8:73E77B88B78EB876393EC303089C1715" country="China" county="Baise" municipality="Leye county" name="Yachang Orchid National Nature Reserve" stateProvince="Guangxi">Yachang Orchid National Nature Reserve</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.8619999999999997" unit="m" value="1862.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.8619999999999997" unit="m" value="1862.0">1862 m</elevation>
</quantity>
,
<collectingDate value="2018-07-28">28 July 2018</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="91">Ying Qin</emphasis>
et al.
<emphasis italics="true" pageId="0" pageNumber="91">YC3639</emphasis>
(IBK!)
</materialsCitation>
;
<materialsCitation collectingDate="2020-07-03" country="China" county="Baise" elevation="1859" location="Yachang Orchid National Nature Reserve" municipality="Leye county" specimenCount="1" stateProvince="Guangxi">
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
:
<collectingCounty>Baise</collectingCounty>
city,
<collectingMunicipality>Leye county</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:AF8E39B6855655B68975809069C885A8:EAA32DBBCC34784AB5B9079E3369E99E" country="China" county="Baise" municipality="Leye county" name="Yachang Orchid National Nature Reserve" stateProvince="Guangxi">Yachang Orchid National Nature Reserve</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.859" unit="m" value="1859.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.859" unit="m" value="1859.0">1859 m</elevation>
</quantity>
,
<collectingDate value="2020-07-03">03 July 2020</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="91">Ying Qin</emphasis>
et al.
<emphasis italics="true" pageId="0" pageNumber="91">QYYC20200703009</emphasis>
(IBK!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>