135 lines
17 KiB
XML
135 lines
17 KiB
XML
<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="AF634E30201B5FA89D59A15E71291C04" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina billeti Fischer 1898" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:title id="28287F3BF486CD2596218AD664D5A0F5">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:namePart id="F3F77B23F1EDAF1776BDF27A6EDCFDFB">Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier id="1C9C416F9A698B5FBF0BCD043DFD9469" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation id="B894A299CECC173AA4917A09276E8CB1">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:roleTerm id="47CB0A2EF2EC8A2339B82FCFE19325E1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FF63C6B2A4906DEBDAA5479CB9298F24">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="311007A5C4912F644DA89C0DE0ED177F">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="09B3F2E1D8679201EA894210092DB252" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="65BE49381983222BDAFB34F30E11304E">Panha, Somsak</mods:namePart>
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<mods:affiliation id="2F5A0260983BBA4953BB81DE0A049591">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="E846D88FAF843F8658D39F766B947CAC" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title id="3D3A0F083ED8F7236BB2E3DE8FF1D7EB">ZooKeys</mods:title>
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<mods:date id="48DB58C79213CB28F15D94B9042100F9">2022</mods:date>
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<mods:number id="E1706E8A1F8ED269B2B535752B6312D2">2022-08-25</mods:number>
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<mods:number id="7D5B332EFCDB4020593490C89B730D4F">1119</mods:number>
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<mods:identifier id="6C7E475F5501A4760AB4F93315E0DE49" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<treatment id="AF634E30201B5FA89D59A15E71291C04" ID-GBIF-Taxon="199838997" LSID="urn:lsid:plazi:treatment:AF634E30201B5FA89D59A15E71291C04" httpUri="http://treatment.plazi.org/id/AF634E30201B5FA89D59A15E71291C04" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupininae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="6CB7EED29BE5346227E75C394657A870" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="80C2DD572CDBD3C2FE647A0119D54121" pageId="0" pageNumber="1">
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<taxonomicName id="746B3F293DB6CF83511518B2AAE0AB8E" ID-CoL="4QJQW" LSID="AF634E30-201B-5FA8-9D59-A15E71291C04" authority="Fischer, 1898" authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti">Pupina billeti Fischer, 1898</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="541A162ED77BEDFC3BAA405CB932FBC3" pageId="0" pageNumber="1" type="description">
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<paragraph id="A3324329DB76A6D55885B9F87143F903" pageId="0" pageNumber="1">
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<figureCitation id="5CE505DFE71EC168D4E8FAA21975C404" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">Figs 21S</figureCitation>
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<figureCitation id="D7F2D067BA15A83608A47D759A2410C3" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D)." figureDoi="10.3897/zookeys.1119.85400.figure26" httpUri="https://binary.pensoft.net/fig/735387" pageId="0" pageNumber="1">, 26C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9913AA56E5947EDDF2161D92BE9ECF9F" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="8CEA2AA0F7D9FD9CCA2D7B628709721A" pageId="0" pageNumber="1">
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<taxonomicName id="3C59153644E85D7904A77481D1F3D3F1" authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti">Pupina billeti</taxonomicName>
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Fischer, 1898: 333, 334, pl. 18, figs 38-41. Type locality: Rochers calcaires
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<normalizedToken id="92DB3C3FB9E42C09968CFFAE721A651D" originalValue="Déo-Ma-Phuc">Deo-Ma-Phuc</normalizedToken>
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[limestone areas around Ma Phuc Pass, Tra Linh District, Cao Bang Province, Vietnam].
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<bibRefCitation id="3B5F9D3FEA85FDBBCA922B17EC78E14A" author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 431, Bac-Kan, Tonkin [Bac Kan Province, Vietnam].
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</paragraph>
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<paragraph id="F149C391DF62C96C2A815FCDED1239DF" pageId="0" pageNumber="1">
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<taxonomicName id="442F68B8632D19FE85975364F566A219" authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti" subGenus="Tylotoechus">Pupina (Tylotoechus) billeti</taxonomicName>
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-
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<bibRefCitation id="26B1CB0C7BA8DAD332D10FB131E60F67" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 309.
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</paragraph>
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</subSubSection>
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<subSubSection id="12E0ABEAF31641D2523F722550A44710" pageId="0" pageNumber="1" type="type material">
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<paragraph id="0E7A0FF3C898263E3B3021BCE7B361CE" pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph id="151807BCE79DE1C5A8748891B4572197" pageId="0" pageNumber="1">
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<emphasis id="C5F515216ECC4A142188EB79771BB5E3" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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MNHN-IM-2000-35841 (Figs
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<figureCitation id="552AD8FADF4EE863FD17A88C7C3F8420" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21S</figureCitation>
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,
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<figureCitation id="E2508248FEAB2A0CA20FE5C321B2D72E" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. A Pupina bensoni sp. nov., paratype CUMZ 12046 / 1 from Khao Wong Cave, Uthai Thani B Pupina hungerfordiana, specimen NHMUK 91.3.14.686 - 7 from Hsaddan Koo C Pupina billeti, holotype MNHN-IM- 2000 - 35841 D Pupina verneaui, syntype MNHN-IM- 2000 - 35843 from Ha-Giang, Tonkin E, F Pupina peguensis E syntype of Pupina blanfordi NHMUK 1888.12.4.100 from Pegu and F specimen NHMUK ex. Cuming coll. from Lao Mountains, Camboja. Photo: P. Maestrati, MNHN (C, D)." figureDoi="10.3897/zookeys.1119.85400.figure26" httpUri="https://binary.pensoft.net/fig/735387" pageId="0" pageNumber="1">26C</figureCitation>
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) from Deo-Ma-Phuc.
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</paragraph>
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</subSubSection>
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<subSubSection id="BFE2DC06602308BA9C667B2EFF62888A" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="82E44A30D9937B7227DAB30699DE5F19" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="BC889FF7E102690D3671ABF3F5390009" pageId="0" pageNumber="1">Shell ovate; last whorl ca. 70% of shell height. Apertural lip extremely thickened; with a furrow between inner and outer peristomes; inner peristome thickened and cord-like; parietal callus distinct. Both parietal and columellar teeth extremely thickened; parietal tooth covering posterior canal; columellar tooth next to slit-like anterior canal.</paragraph>
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</subSubSection>
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<subSubSection id="1325CBB4159360FBC97DDC80D9EC1DA3" pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph id="27263332BC4B18C2052DBA5EBAB22E27" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph id="5558CD471FC9717D6DB22F78D1062032" pageId="0" pageNumber="1">
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<taxonomicName id="50620E05C7B67786FC6036A10A2474D4" authorityName="Fischer" authorityYear="1898" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti">
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<emphasis id="5D65F1E0D92B2EE2C661F73C51340C2F" italics="true" pageId="0" pageNumber="1">Pupina billeti</emphasis>
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</taxonomicName>
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can be distinguished from all other species in the
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<taxonomicName id="F17695F85C5C3F3D39A28EE0DC043E42" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="artata">
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<emphasis id="5B756BDD91C0DBCE23A4C04941C1CE78" italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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species group from mainland Southeast Asia by having the thickest parietal and columellar teeth and apertural lip, and a distinct parietal callus.
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</paragraph>
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</subSubSection>
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<subSubSection id="2A45B5BBA1C159C5B72C971F589252EC" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="26B3E9B15C85AD54F54FE0A85E2DEE2F" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="DDB297FA7A7CF38FFAB8590FE08E69BC" pageId="0" pageNumber="1">
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Northern Vietnam (
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<bibRefCitation id="89D6BF07D9C3296E1371143C049FBD2E" author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D7389D5B88BF90868BB9867911A89B98" pageId="0" pageNumber="1" type="remarks">
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<paragraph id="C8872072D5AAC69F617168E7E99F75C1" pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph id="F5A31119A899443477AF80E1430AA889" pageId="0" pageNumber="1">
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As
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<taxonomicName id="D880D32B576A2459D272E5593A7D496F" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. billeti" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="billeti">
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<emphasis id="494A8E354B1229060F87B850AE32B0F5" italics="true" pageId="0" pageNumber="1">P. billeti</emphasis>
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</taxonomicName>
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was described based on a single specimen as explicitly stated in the original description, that specimen is the holotype fixed by monotypy (
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<bibRefCitation id="6B09F3A99A58890B562366B70BFDC539" author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
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: Art. 73.1.2).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |