treatments-xml/data/AF/3D/87/AF3D87B5A23EFF832A8F787FFDB64874.xml

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<mods:title id="52FBA37966C07D66CFE7F946B48A3F3F">Species of Ancistrus (Siluriformes, Loricariidae) from Ecuador, with the description of a new species from the Amazon River Basin</mods:title>
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<treatment id="AF3D87B5A23EFF832A8F787FFDB64874" ID-DOI="http://doi.org/10.5281/zenodo.6489404" ID-GBIF-Taxon="154508035" ID-Zenodo-Dep="6489404" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874" httpUri="http://treatment.plazi.org/id/AF3D87B5A23EFF832A8F787FFDB64874" lastPageId="11" lastPageNumber="222" pageId="6" pageNumber="217">
<subSubSection id="6F8E6528A23EFF8E2A8F787FFE1F4C9B" pageId="6" pageNumber="217" type="nomenclature">
<paragraph id="272B36A3A23EFF8E2A8F787FFE4D4C7A" blockId="6.[151,491,1619,1679]" box="[151,491,1619,1646]" pageId="6" pageNumber="217">
<heading id="7C6381CFA23EFF8E2A8F787FFE4D4C7A" bold="true" box="[151,491,1619,1646]" fontSize="11" level="1" pageId="6" pageNumber="217" reason="1">
<taxonomicName id="E0944D20A23EFF8E2A8F787FFE4D4C7A" ID-CoL="69QG4" authority="(Cope, 1872)" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[151,491,1619,1646]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="217" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A23EFF8E2A8F787FFE4D4C7A" bold="true" box="[151,491,1619,1646]" pageId="6" pageNumber="217">
<emphasis id="15E0EAB1A23EFF8E2A8F787FFEE04C79" bold="true" box="[151,326,1619,1645]" italics="true" pageId="6" pageNumber="217">Ancistrus alga</emphasis>
(
<bibRefCitation id="43054B52A23EFF8E2B4D7878FE454C7A" author="Cope, E. D." box="[341,483,1620,1646]" pageId="6" pageNumber="217" pagination="250 - 294" refId="ref13970" refString="Cope, E. D. (1872) On the fishes of the Ambyiacu river. Proceedings of the Academy of Natural Sciences of Philadelphia, 23, 250 - 294, pls. 3 - 16. [for 1871]" type="journal article" year="1872">Cope, 1872</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="272B36A3A23EFF8E2A8F785AFE1F4C9B" blockId="6.[151,491,1619,1679]" box="[151,441,1654,1679]" pageId="6" pageNumber="217">
<tableCitation id="6A160318A23EFF8E2A8F785AFF504C9B" box="[151,246,1654,1679]" captionStart="TABLE 2" captionStartId="10.[151,239,152,175]" captionText="TABLE 2. Morphometric data of Ancistrus alga. Measurements are expressed as percentage of standard length. N=11" pageId="6" pageNumber="217">Tables 2</tableCitation>
, 
<tableCitation id="6A160318A23EFF8E2B1B785AFEB74C9B" box="[259,273,1654,1679]" captionStart="TABLE 5" captionStartId="21.[151,239,1049,1072]" captionText="TABLE 5. Comparative morphometric data for the males of three species. Measurements are expressed as percentage of standard length." pageId="6" pageNumber="217">5</tableCitation>
<figureCitation id="BFAF2A26A23EFF8E2B0F785AFE254C9B" box="[279,387,1654,1679]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[204,1390,193,1949]" captionTargetId="figure@5.[204,1390,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Females of Ancistrus species, lateral view. From top to bottom: A. shuar, MEPN 17983, 93.9 mm SL. A. alga, MEPN 15169, 160.9 mm SL. A. clementinae, MEPN 5996, 100.1 mm SL. and A. malacops, MEPN 7804, 93.4 mm SL." httpUri="https://zenodo.org/record/2612129/files/figure.png" pageId="6" pageNumber="217">Figures 2</figureCitation>
, 
<figureCitation id="BFAF2A26A23EFF8E2B96785BFE384C9B" box="[398,414,1655,1679]" captionStart="FIGURE 3" captionStartId="7.[151,250,1606,1628]" captionTargetBox="[151,1436,589,1585]" captionTargetId="figure@7.[151,1436,589,1585]" captionTargetPageId="7" captionText="FIGURE 3. Ventral view of the head of females to show the presence (arrows) or absence of soft, fleshy tentacles on snout. A) A. clementinae, MEPN 5996, 100.1 mm SL. B) A. malacops, MEPN 7804, 93.4 mm SL. C) A. alga, MEPN 15169, 160.9 mm SL, D) A. shuar, MEPN 17983, 93.9 mm SL." httpUri="https://zenodo.org/record/2612131/files/figure.png" pageId="6" pageNumber="217">3</figureCitation>
, 
<figureCitation id="BFAF2A26A23EFF8E2BB3785AFE1F4C9B" box="[427,441,1654,1679]" captionStart="FIGURE 5" captionStartId="9.[151,250,1988,2010]" captionTargetBox="[177,1414,193,1976]" captionTargetId="figure@9.[177,1414,193,1976]" captionTargetPageId="9" captionText="FIGURE 5. Ancistrus alga, MEPN 14853, 115.1 mm SL, male. From top to bottom, lateral, dorsal, and ventral view." httpUri="https://zenodo.org/record/2612135/files/figure.png" pageId="6" pageNumber="217">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6F8E6528A23EFF8E2A8F7892FA3A4D06" pageId="6" pageNumber="217" type="reference_group">
<paragraph id="272B36A3A23EFF8E2A8F7892FA3A4CC0" blockId="6.[151,1436,1726,1932]" box="[151,1436,1726,1748]" pageId="6" pageNumber="217">
<taxonomicName id="E0944D20A23EFF8E2A8F7892FE784CC0" ID-CoL="69QG4" authority="Cope, 1872" authorityName="Cope" authorityYear="1872" box="[151,478,1726,1748]" class="Actinopterygii" family="Loricariidae" genus="Chaetostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="217" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A23EFF8E2A8F7892FEFA4CC0" box="[151,348,1726,1748]" italics="true" pageId="6" pageNumber="217">Chaetostomus alga</emphasis>
<bibRefCitation id="43054B52A23EFF8E2B7C7892FE784CC0" author="Cope, E. D." box="[356,478,1726,1748]" pageId="6" pageNumber="217" pagination="250 - 294" refId="ref13970" refString="Cope, E. D. (1872) On the fishes of the Ambyiacu river. Proceedings of the Academy of Natural Sciences of Philadelphia, 23, 250 - 294, pls. 3 - 16. [for 1871]" type="journal article" year="1872">Cope, 1872</bibRefCitation>
</taxonomicName>
. Proceedings of the Academy of Natural Sciences of Philadelphia, 
<date id="532A1063A23EFF8E2E817893FB6B4CC0" box="[1177,1229,1726,1748]" pageId="6" pageNumber="217" value="1923-05">v. 23</date>
:287, Pl. 15 (fig. 3).
</paragraph>
<paragraph id="272B36A3A23EFF8E2ADF78F1FA3A4D06" blockId="6.[151,1436,1726,1932]" pageId="6" pageNumber="217">
Ambyiacu (Ampiyacu?) River, 
<collectingCountry id="5F837633A23EFF8E281E78F2FD914CE7" box="[518,567,1758,1779]" name="Peru" pageId="6" pageNumber="217">Peru</collectingCountry>
. 
<typeStatus id="F82F8801A23EFF8E285978F1FD044CE7" box="[577,674,1757,1779]" pageId="6" pageNumber="217" type="syntype">Syntypes</typeStatus>
: ANSP 16461-62 (2). 
<taxonomicName id="E0944D20A23EFF8E2A8F78D0FDBC4D06" ID-CoL="69QGV" authority="Cope, 1872" authorityName="Cope" authorityYear="1872" box="[151,538,1788,1810]" class="Actinopterygii" family="Loricariidae" genus="Chaetostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="217" phylum="Chordata" rank="species" species="tectirostris">
<emphasis id="15E0EAB1A23EFF8E2A8F78D0FE3C4D06" box="[151,410,1788,1810]" italics="true" pageId="6" pageNumber="217">Chaetostomus tectirostris</emphasis>
<bibRefCitation id="43054B52A23EFF8E2BB978D0FDBC4D06" author="Cope, E. D." box="[417,538,1788,1810]" pageId="6" pageNumber="217" pagination="250 - 294" refId="ref13970" refString="Cope, E. D. (1872) On the fishes of the Ambyiacu river. Proceedings of the Academy of Natural Sciences of Philadelphia, 23, 250 - 294, pls. 3 - 16. [for 1871]" type="journal article" year="1872">Cope, 1872</bibRefCitation>
</taxonomicName>
. Proceedings of the Academy of Natural Sciences of Philadelphia 
<date id="532A1063A23EFF8E2EDA78D1FB534D06" box="[1218,1269,1788,1810]" pageId="6" pageNumber="217" value="1923-05">v. 23</date>
:288, Pl. 15 (fig.
</paragraph>
</subSubSection>
<subSubSection id="6F8E6528A23EFF8E2ADF7936FE874D98" pageId="6" pageNumber="217" type="description">
<paragraph id="272B36A3A23EFF8E2ADF7936FA3A4D7A" blockId="6.[151,1436,1726,1932]" pageId="6" pageNumber="217">
2). Ambyiacu (Ampiyacu?) River, 
<collectingCountry id="5F837633A23EFF8E28307937FDFC4D24" box="[552,602,1819,1840]" name="Peru" pageId="6" pageNumber="217">Peru</collectingCountry>
. 
<typeStatus id="F82F8801A23EFF8E287B7936FD624D24" box="[611,708,1818,1840]" pageId="6" pageNumber="217" type="syntype">Syntypes</typeStatus>
: ANSP 8298 (1), 8300 (1). 
<taxonomicName id="E0944D20A23EFF8E2A8F7916FE544D5B" authority="Isbrucker, 1980: 66" authorityName="Isbrucker" authorityPageNumber="66" authorityYear="1980" box="[151,498,1849,1871]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="217" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A23EFF8E2A8F7916FE8C4D5B" box="[151,298,1849,1871]" italics="true" pageId="6" pageNumber="217">Ancistrus alga</emphasis>
<bibRefCitation id="43054B52A23EFF8E2B287915FE744D5B" author="Isbrucker, I. J. H." box="[304,466,1849,1871]" pageId="6" pageNumber="217" refId="ref14645" refString="Isbrucker, I. J. H. (1980) Classification and catalogue of the mailed Loricariidae (Pisces, Siluriformes). Ferslagen en Technische Gegevens, Instituut voor Taxonomische Zoologie, Universiteit van Amsterdam, 22,1 - 181." type="book" year="1980">Isbrücker, 1980</bibRefCitation>
:66
</taxonomicName>
. Ortega &amp; Vari, 1986:16. Burgess, 1989:436. Isbrücker, 2001:25, 26. 
<bibRefCitation id="43054B52A23EFF8E2EA97915FAF04D5B" author="Isbrucker, I. J. H." box="[1201,1366,1849,1871]" pageId="6" pageNumber="217" pagination="11 - 30" refId="ref14747" refString="Isbrucker, I. J. H. (2002) Nomenclator of the 108 genera with 692 species of the mailed catfishes, family Loricariidae Rafinesque, 1815 (Teleostei, Ostariophysi). Cat Chat, Journal of the catfish study group (UK), 3 (1), 11 - 30." type="journal article" year="2002">Isbrücker, 2002</bibRefCitation>
:11. 
<taxonomicName id="E0944D20A23EFF8E2A8F7975FDE74D7A" authority="Isbrucker, 1980: 73" authorityName="Isbrucker" authorityPageNumber="73" authorityYear="1980" box="[151,577,1880,1902]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="217" phylum="Chordata" rank="species" species="tectirostris">
<emphasis id="15E0EAB1A23EFF8E2A8F7975FEC94D7A" box="[151,367,1881,1902]" italics="true" pageId="6" pageNumber="217">Ancistrus tectirostris</emphasis>
<bibRefCitation id="43054B52A23EFF8E2B637974FD854D7A" author="Isbrucker, I. J. H." box="[379,547,1880,1902]" pageId="6" pageNumber="217" refId="ref14645" refString="Isbrucker, I. J. H. (1980) Classification and catalogue of the mailed Loricariidae (Pisces, Siluriformes). Ferslagen en Technische Gegevens, Instituut voor Taxonomische Zoologie, Universiteit van Amsterdam, 22,1 - 181." type="book" year="1980">Isbrücker, 1980</bibRefCitation>
:73
</taxonomicName>
. Ortega &amp; Vari, 1986:16. Burgess, 1989:437. Isbrücker, 2001:25, 26. 
<bibRefCitation id="43054B52A23EFF8E2F2F7974FA3A4D7A" author="Isbrucker, I. J. H." box="[1335,1436,1880,1902]" pageId="6" pageNumber="217" pagination="11 - 30" refId="ref14747" refString="Isbrucker, I. J. H. (2002) Nomenclator of the 108 genera with 692 species of the mailed catfishes, family Loricariidae Rafinesque, 1815 (Teleostei, Ostariophysi). Cat Chat, Journal of the catfish study group (UK), 3 (1), 11 - 30." type="journal article" year="2002">Isbrücker,</bibRefCitation>
</paragraph>
<paragraph id="272B36A3A23EFF8E2ADF795AFE874D98" blockId="6.[151,1436,1726,1932]" box="[199,289,1910,1932]" pageId="6" pageNumber="217">2002:12.</paragraph>
</subSubSection>
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<paragraph id="272B36A3A23EFF8E2A8F7995FA3A4DE2" blockId="6.[151,1436,1977,2038]" pageId="6" pageNumber="217">
<emphasis id="15E0EAB1A23EFF8E2A8F7995FE254DC6" bold="true" box="[151,387,1977,2002]" pageId="6" pageNumber="217">Material examined.</emphasis>
All from 
<collectingCountry id="5F837633A23EFF8E2BE37995FDFD4DC6" box="[507,603,1977,2002]" name="Ecuador" pageId="6" pageNumber="217">Ecuador</collectingCountry>
. 
<emphasis id="15E0EAB1A23EFF8E287F7995FCE44DC6" bold="true" box="[615,834,1977,2002]" pageId="6" pageNumber="217">
<collectingRegion id="E550F841A23EFF8E287F7995FCE44DC6" box="[615,834,1977,2002]" country="Ecuador" name="Orellana" pageId="6" pageNumber="217">Orellana province</collectingRegion>
</emphasis>
: MEPN-460, 
<quantity id="E06C9B46A23EFF8E29FD7996FBDE4DC5" box="[997,1144,1978,2002]" metricMagnitude="0" metricUnit="m" metricValue="1.1100999999999999" pageId="6" pageNumber="217" unit="mm" value="1110.1">1, 110.1 mm</quantity>
SL, Quebrada Cotoyacu, 
<geoCoordinate id="42A05064A23EFF8E2A8F79F2FEBD4DE2" box="[151,283,2014,2038]" direction="south" orientation="latitude" pageId="6" pageNumber="217" precision="15" value="-1.0086111">01°00’31”S</geoCoordinate>
<geoCoordinate id="42A05064A23EFF8E2B3F79F1FE104DE2" box="[295,438,2013,2038]" direction="west" orientation="longitude" pageId="6" pageNumber="217" precision="15" value="-75.43667">75°26’12”W</geoCoordinate>
, R. Barriga &amp; K. Galacatos, 
<date id="532A1063A23EFF8E290D79F1FC114DE2" box="[789,951,2013,2038]" pageId="6" pageNumber="217" value="1996-06-15">15 June 1996</date>
. MEPN-829, 1, 119.0 mm SL, Quebrada
</paragraph>
<paragraph id="272B36A3A23FFF8F2A8F7EBBFA32480C" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="218">
Tambococha, 
<geoCoordinate id="42A05064A23FFF8F2B2B7EBBFE1E4AA4" box="[307,440,151,176]" direction="south" orientation="latitude" pageId="7" pageNumber="218" precision="15" value="-0.9594444">00°57’34”S</geoCoordinate>
<geoCoordinate id="42A05064A23FFF8F2BA77EBBFDE94AA4" box="[447,591,151,176]" direction="west" orientation="longitude" pageId="7" pageNumber="218" precision="15" value="-75.44056">75°26’26”W</geoCoordinate>
, R. Barriga &amp; K. Galacatos, 
<date id="532A1063A23FFF8F29887EBBFBFC4AA4" box="[912,1114,151,176]" pageId="7" pageNumber="218" value="1996-02-26">26 February 1996</date>
. 
<materialsCitation id="97FC3CFEA23FFF8F2E7D7EB4FB244AEC" ID-GBIF-Occurrence="2029174818" collectingDate="1981-10-28" collectorName="D. Stewart &amp; R. Barriga &amp; Ibarra" latitude="-0.7480556" location="rio Tiputini" longLatPrecision="21" longitude="-75.79472" municipality="Tributario" pageId="7" pageNumber="218" specimenCount="1" stateProvince="Napo">
MEPN-1308, 5, 60.0– 
<quantity id="E06C9B46A23FFF8F2F477EBBFF654AC0" metricMagnitude="-1" metricUnit="m" metricValue="1.258" pageId="7" pageNumber="218" unit="mm" value="125.8">125.8 mm</quantity>
SL, 
<collectingMunicipality id="C74FACD9A23FFF8F2AE17E90FECF4AC1" box="[249,361,188,213]" pageId="7" pageNumber="218">Tributario</collectingMunicipality>
norteño del 
<location id="224B6078A23FFF8F2BEE7E90FDDF4AC1" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874:224B6078A23FFF8F2BEE7E90FDDF4AC1" box="[502,633,188,213]" latitude="-0.7480556" longLatPrecision="21" longitude="-75.79472" municipality="Tributario" name="rio Tiputini" pageId="7" pageNumber="218" stateProvince="Napo">río Tiputini</location>
, a unas 1.5 horas en canoa motorizada, aguas arriba de la boca en el río 
<collectingRegion id="E550F841A23FFF8F2A8F7ECCFF714AE3" box="[151,215,224,247]" country="Ecuador" name="Napo" pageId="7" pageNumber="218">Napo</collectingRegion>
, 
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, 
<collectorName id="8A615375A23FFF8F28097ECCFD294AEC" box="[529,655,224,248]" pageId="7" pageNumber="218">D. Stewart</collectorName>
, 
<collectorName id="8A615375A23FFF8F28857ECCFCB24AEC" box="[669,788,223,248]" pageId="7" pageNumber="218">R. Barriga</collectorName>
&amp; M. 
<collectorName id="8A615375A23FFF8F29707EF3FC084AEC" box="[872,942,223,248]" pageId="7" pageNumber="218">Ibarra</collectorName>
, 
<date id="532A1063A23FFF8F29A47ECCFB244AEC" box="[956,1154,223,248]" pageId="7" pageNumber="218" value="1981-10-28">
<collectingDate id="436EE98BA23FFF8F29A47ECCFB244AEC" box="[956,1154,223,248]" pageId="7" pageNumber="218" value="1981-10-28">28 October 1981</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="97FC3CFEA23FFF8F2E887EF3FEF64B54" ID-GBIF-Occurrence="2029173570" collectingDate="1998-03-08" collectorName="R. Barriga" latitude="-1.2011112" location="rio Nashino" longLatPrecision="21" longitude="-75.81334" municipality="Rio Gabaron" pageId="7" pageNumber="218" specimenCount="1" stateProvince="Napo">
MEPN-2544, 7, 105.4– 
<quantity id="E06C9B46A23FFF8F2A8F7F28FEAF4B08" box="[151,265,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="1.4809999999999999" pageId="7" pageNumber="218" unit="mm" value="148.1">148.1 mm</quantity>
SL, 
<collectingMunicipality id="C74FACD9A23FFF8F2B587F28FE764B09" box="[320,464,260,285]" pageId="7" pageNumber="218">Río Gabarón</collectingMunicipality>
afluente del 
<location id="224B6078A23FFF8F287B7F28FD4C4B09" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874:224B6078A23FFF8F287B7F28FD4C4B09" box="[611,746,260,285]" latitude="-1.2011112" longLatPrecision="21" longitude="-75.81334" municipality="Rio Gabaron" name="rio Nashino" pageId="7" pageNumber="218" stateProvince="Napo">río Nashiño</location>
, pozo OBE, bloque 31, 
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<geoCoordinate id="42A05064A23FFF8F2E9D7F28FAB24B09" box="[1157,1300,260,285]" direction="west" orientation="longitude" pageId="7" pageNumber="218" precision="15" value="-75.81334">75°48’48”W</geoCoordinate>
, 
<collectorName id="8A615375A23FFF8F2F387F29FA3E4B09" box="[1312,1432,260,285]" pageId="7" pageNumber="218">R. Barriga</collectorName>
, 0 
<date id="532A1063A23FFF8F2ABE7F04FEF64B54" box="[166,336,295,320]" pageId="7" pageNumber="218" value="1998-03-08">
<collectingDate id="436EE98BA23FFF8F2ABE7F04FEF64B54" box="[166,336,295,320]" pageId="7" pageNumber="218" value="1998-03-08">8 March 1998</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="97FC3CFEA23FFF8F2B797F0BFC124B71" ID-GBIF-Occurrence="2029175602" collectingDate="2006-06-20" collectorName="de Chiroisla &amp; R. Barriga &amp; Jose Pinargote" latitude="-0.7097222" location="rio Tiputini" longLatPrecision="21" longitude="-75.93166" municipality="Laguna Muyuna" pageId="7" pageNumber="218" specimenCount="1" stateProvince="Napo">
MEPN-14853, 
<quantity id="E06C9B46A23FFF8F280A7F04FD0D4B2B" box="[530,683,295,320]" metricMagnitude="0" metricUnit="m" metricValue="1.1151" pageId="7" pageNumber="218" unit="mm" value="1115.1">1, 115.1 mm</quantity>
SL, 
<collectingMunicipality id="C74FACD9A23FFF8F28F47F04FC0B4B54" box="[748,941,296,320]" pageId="7" pageNumber="218">Laguna Muyuna</collectingMunicipality>
, km 12.5 
<collectorName id="8A615375A23FFF8F2E357F0BFB1D4B54" box="[1069,1211,295,320]" pageId="7" pageNumber="218">de Chiroisla</collectorName>
en el 
<location id="224B6078A23FFF8F2F087F04FA3E4B54" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874:224B6078A23FFF8F2F087F04FA3E4B54" box="[1296,1432,295,320]" latitude="-0.7097222" longLatPrecision="21" longitude="-75.93166" municipality="Laguna Muyuna" name="rio Tiputini" pageId="7" pageNumber="218" stateProvince="Napo">río Tiputini</location>
, 
<geoCoordinate id="42A05064A23FFF8F2A8F7F60FEBD4B71" box="[151,283,332,357]" direction="south" orientation="latitude" pageId="7" pageNumber="218" precision="15" value="-0.7097222">00°42’35”S</geoCoordinate>
<geoCoordinate id="42A05064A23FFF8F2B3D7F60FE174B71" box="[293,433,332,357]" direction="west" orientation="longitude" pageId="7" pageNumber="218" precision="15" value="-75.93166">75°55’54”W</geoCoordinate>
, 
<collectorName id="8A615375A23FFF8F2BD87F61FD914B71" box="[448,567,332,357]" pageId="7" pageNumber="218">R. Barriga</collectorName>
&amp; 
<collectorName id="8A615375A23FFF8F28787F60FCAE4B71" box="[608,776,332,357]" pageId="7" pageNumber="218">José Pinargote</collectorName>
, 
<date id="532A1063A23FFF8F290D7F60FC124B71" box="[789,948,332,357]" pageId="7" pageNumber="218" value="2006-06-20">
<collectingDate id="436EE98BA23FFF8F290D7F60FC124B71" box="[789,948,332,357]" pageId="7" pageNumber="218" value="2006-06-20">20 June 2006</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="97FC3CFEA23FFF8F29D87F60FB9D4B9C" ID-GBIF-Occurrence="2029173182" collectingDate="1963-03" latitude="-1.5758333" location="Rio Curaray" longLatPrecision="21" longitude="-76.35667" municipality="Pavacachi" pageId="7" pageNumber="218" specimenCount="1" stateProvince="Pastaza">
<emphasis id="15E0EAB1A23FFF8F29D87F60FB2B4B71" bold="true" box="[960,1165,332,357]" pageId="7" pageNumber="218">
<collectingRegion id="E550F841A23FFF8F29D87F60FB2B4B71" box="[960,1165,332,357]" country="Ecuador" name="Pastaza" pageId="7" pageNumber="218">Pastaza province</collectingRegion>
</emphasis>
: MEPN-10348, 1, 
<quantity id="E06C9B46A23FFF8F2F737F60FF654B93" metricMagnitude="-2" metricUnit="m" metricValue="7.69" pageId="7" pageNumber="218" unit="mm" value="76.9">76.9 mm</quantity>
SL, 
<collectingMunicipality id="C74FACD9A23FFF8F2AEF7F43FECD4B9C" box="[247,363,367,392]" pageId="7" pageNumber="218">Pavacachi</collectingMunicipality>
, 
<location id="224B6078A23FFF8F2B6E7F5CFDA64B9C" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874:224B6078A23FFF8F2B6E7F5CFDA64B9C" box="[374,512,368,392]" latitude="-1.5758333" longLatPrecision="21" longitude="-76.35667" municipality="Pavacachi" name="Rio Curaray" pageId="7" pageNumber="218" stateProvince="Pastaza">Río Curaray</location>
, 
<geoCoordinate id="42A05064A23FFF8F28137F5CFD294B9C" box="[523,655,368,392]" direction="south" orientation="latitude" pageId="7" pageNumber="218" precision="15" value="-1.5758333">01°34’33”S</geoCoordinate>
<geoCoordinate id="42A05064A23FFF8F288E7F43FC834B9C" box="[662,805,367,392]" direction="west" orientation="longitude" pageId="7" pageNumber="218" precision="15" value="-76.35667">76°21’24”W</geoCoordinate>
, G. 
<collectingRegion id="E550F841A23FFF8F29567F5CFC004B93" box="[846,934,368,391]" country="Panama" name="Herrera" pageId="7" pageNumber="218">Herrera</collectingRegion>
, 
<date id="532A1063A23FFF8F29A97F43FB9D4B9C" box="[945,1083,367,392]" pageId="7" pageNumber="218" value="1963-03">
<collectingDate id="436EE98BA23FFF8F29A97F43FB9D4B9C" box="[945,1083,367,392]" pageId="7" pageNumber="218" value="1963-03">March 1963</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="97FC3CFEA23FFF8F2E5E7F43FDE84BC4" ID-GBIF-Occurrence="2029173934" collectingDate="1998-05-24" collectorName="R. Barriga &amp; D. Stewart" county="Rio Aguarico" latitude="-0.8833333" location="Puerto" longLatPrecision="21" longitude="-75.22833" municipality="rio Yanayacu" pageId="7" pageNumber="218" specimenCount="1" stateProvince="Sucumbios">
<emphasis id="15E0EAB1A23FFF8F2E5E7F43FA914B9C" bold="true" box="[1094,1335,367,392]" pageId="7" pageNumber="218">
<collectingRegion id="E550F841A23FFF8F2E5E7F43FA914B9C" box="[1094,1335,367,392]" country="Ecuador" name="Sucumbios" pageId="7" pageNumber="218">Sucumbíos province</collectingRegion>
</emphasis>
: MEPN- 2273, 
<quantity id="E06C9B46A23FFF8F2AC47FB8FECC4BB8" box="[220,362,404,428]" metricMagnitude="0" metricUnit="m" metricValue="1.1099" pageId="7" pageNumber="218" unit="mm" value="1109.9">1, 109.9 mm</quantity>
SL, 
<collectingCounty id="CE4A4E2FA23FFF8F2B867FB8FD9F4BB9" box="[414,569,404,429]" pageId="7" pageNumber="218">Río Aguarico</collectingCounty>
, tributario norteño del 
<collectingMunicipality id="C74FACD9A23FFF8F29207FB8FC6D4BB9" box="[824,971,404,429]" pageId="7" pageNumber="218">río Yanayacu</collectingMunicipality>
at 
<location id="224B6078A23FFF8F29F47FB9FB934BB8" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874:224B6078A23FFF8F29F47FB9FB934BB8" box="[1004,1077,405,428]" county="Rio Aguarico" latitude="-0.8833333" longLatPrecision="21" longitude="-75.22833" municipality="rio Yanayacu" name="Puerto" pageId="7" pageNumber="218" stateProvince="Sucumbios">Puerto</location>
<collectingRegion id="E550F841A23FFF8F2E247FB8FBD44BB9" box="[1084,1138,404,429]" country="Ecuador" name="Loja" pageId="7" pageNumber="218">Loja</collectingRegion>
, 
<geoCoordinate id="42A05064A23FFF8F2E657FB8FAA44BB9" box="[1149,1282,404,429]" direction="south" orientation="latitude" pageId="7" pageNumber="218" precision="15" value="-0.8833333">00°53’00”S</geoCoordinate>
<geoCoordinate id="42A05064A23FFF8F2F107FB8FA314BB9" box="[1288,1431,404,429]" direction="west" orientation="longitude" pageId="7" pageNumber="218" precision="15" value="-75.22833">75°13’42”W</geoCoordinate>
, 
<collectorName id="8A615375A23FFF8F2A8F7F94FEAA4BC4" box="[151,268,439,464]" pageId="7" pageNumber="218">R. Barriga</collectorName>
&amp; 
<collectorName id="8A615375A23FFF8F2B287F94FE0C4BC4" box="[304,426,440,464]" pageId="7" pageNumber="218">D. Stewart</collectorName>
, 
<date id="532A1063A23FFF8F2BAE7F94FDE84BC4" box="[438,590,440,464]" pageId="7" pageNumber="218" value="1998-05-24">
<collectingDate id="436EE98BA23FFF8F2BAE7F94FDE84BC4" box="[438,590,440,464]" pageId="7" pageNumber="218" value="1998-05-24">24 May 1998</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="97FC3CFEA23FFF8F28427F9BFB654BE0" ID-GBIF-Occurrence="2029174227" collectingDate="1993-05-01" collectorName="R. Barriga &amp; E. Medina" county="Laguna Imuya" latitude="-0.60194445" location="Transturi" longLatPrecision="21" longitude="-75.24083" municipality="rio Lagartococha" pageId="7" pageNumber="218" specimenCount="1" stateProvince="Sucumbios">
MEPN-13896, 
<quantity id="E06C9B46A23FFF8F291D7F94FC324BDB" box="[773,916,440,464]" metricMagnitude="0" metricUnit="m" metricValue="1.1314" pageId="7" pageNumber="218" unit="mm" value="1131.4">1, 131.4 mm</quantity>
SL, 
<collectingCounty id="CE4A4E2FA23FFF8F29D17F94FBC94BC4" box="[969,1135,440,464]" pageId="7" pageNumber="218">Laguna Imuya</collectingCounty>
, junto al 
<collectingMunicipality id="C74FACD9A23FFF8F2ECE7F94FA3E4BC4" box="[1238,1432,439,464]" pageId="7" pageNumber="218">río Lagartococha</collectingMunicipality>
, campamento 
<location id="224B6078A23FFF8F2B2C7FF0FE3B4BE1" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874:224B6078A23FFF8F2B2C7FF0FE3B4BE1" box="[308,413,476,501]" county="Laguna Imuya" latitude="-0.60194445" longLatPrecision="21" longitude="-75.24083" municipality="rio Lagartococha" name="Transturi" pageId="7" pageNumber="218" stateProvince="Sucumbios">Transturi</location>
, 
<geoCoordinate id="42A05064A23FFF8F2BB67FF0FD944BE1" box="[430,562,476,501]" direction="south" orientation="latitude" pageId="7" pageNumber="218" precision="15" value="-0.60194445">00°36’07”S</geoCoordinate>
<geoCoordinate id="42A05064A23FFF8F28587FF0FD694BE1" box="[576,719,476,501]" direction="west" orientation="longitude" pageId="7" pageNumber="218" precision="15" value="-75.24083">75°14’27”W</geoCoordinate>
, 
<collectorName id="8A615375A23FFF8F28FA7FF1FCFB4BE1" box="[738,861,476,501]" pageId="7" pageNumber="218">R. Barriga</collectorName>
&amp; 
<collectorName id="8A615375A23FFF8F29967FF1FBAB4BE1" box="[910,1037,476,501]" pageId="7" pageNumber="218">E. Medina</collectorName>
, 0 
<date id="532A1063A23FFF8F2E377FF0FB654BE0" box="[1071,1219,476,501]" pageId="7" pageNumber="218" value="1993-05-01">
<collectingDate id="436EE98BA23FFF8F2E377FF0FB654BE0" box="[1071,1219,476,501]" pageId="7" pageNumber="218" value="1993-05-01">1 May 1993</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="97FC3CFEA23FFF8F2ECD7FF0FA36480C" ID-GBIF-Occurrence="2029174819" collectingDate="1994-10-30" collectorName="R. Barriga" latitude="-0.5833333" location="Zancudococha" longLatPrecision="21" longitude="-75.5" municipality="Canal" pageId="7" pageNumber="218" specimenCount="1" stateProvince="Sucumbios">
MEPN-15169, 2, 112.14– 
<quantity id="E06C9B46A23FFF8F2AEA7FD3FEC54803" box="[242,355,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="1.609" pageId="7" pageNumber="218" unit="mm" value="160.9">160.9 mm</quantity>
SL, 
<collectingMunicipality id="C74FACD9A23FFF8F2B807FD3FE7F480C" box="[408,473,511,536]" pageId="7" pageNumber="218">Canal</collectingMunicipality>
de la laguna 
<location id="224B6078A23FFF8F28777FD3FCB3480C" LSID="urn:lsid:plazi:treatment:AF3D87B5A23EFF832A8F787FFDB64874:224B6078A23FFF8F28777FD3FCB3480C" box="[623,789,511,536]" latitude="-0.5833333" longLatPrecision="21" longitude="-75.5" municipality="Canal" name="Zancudococha" pageId="7" pageNumber="218" stateProvince="Sucumbios">Zancudococha</location>
, 
<geoCoordinate id="42A05064A23FFF8F29387FD3FC01480C" box="[800,935,511,536]" direction="south" orientation="latitude" pageId="7" pageNumber="218" precision="15" value="-0.5833333">00°35’00”S</geoCoordinate>
, 
<geoCoordinate id="42A05064A23FFF8F29AA7FD3FBE4480C" box="[946,1090,511,536]" direction="west" orientation="longitude" pageId="7" pageNumber="218" precision="15" value="-75.5">75°30’00”W</geoCoordinate>
, 
<collectorName id="8A615375A23FFF8F2E557C2CFB63480C" box="[1101,1221,511,536]" pageId="7" pageNumber="218">R. Barriga</collectorName>
, 
<date id="532A1063A23FFF8F2EC87C2CFA36480C" box="[1232,1424,511,536]" pageId="7" pageNumber="218" value="1994-10-30">
<collectingDate id="436EE98BA23FFF8F2EC87C2CFA36480C" box="[1232,1424,511,536]" pageId="7" pageNumber="218" value="1994-10-30">30 October 1994</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="73EB662BA23FFF8F2A8F786AFDC74CB5" httpUri="https://zenodo.org/record/2612131/files/figure.png" pageId="7" pageNumber="218" startId="7.[151,250,1606,1628]" targetBox="[151,1436,589,1585]" targetPageId="7">
<paragraph id="272B36A3A23FFF8F2A8F786AFDC74CB5" blockId="7.[151,1435,1606,1697]" pageId="7" pageNumber="218">
<emphasis id="15E0EAB1A23FFF8F2A8F786AFEB44C48" bold="true" box="[151,274,1606,1628]" pageId="7" pageNumber="218">FIGURE 3.</emphasis>
Ventral view of the head of females to show the presence (arrows) or absence of soft, fleshy tentacles on snout. A) 
<taxonomicName id="E0944D20A23FFF8F2A8F7845FE8B4C6A" authorityName="Rendahl" authorityYear="1937" box="[151,301,1640,1662]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="clementinae">
<emphasis id="15E0EAB1A23FFF8F2A8F7845FE8B4C6A" box="[151,301,1640,1662]" italics="true" pageId="7" pageNumber="218">A. clementinae</emphasis>
</taxonomicName>
, MEPN 5996, 100.1 mm SL. B) 
<taxonomicName id="E0944D20A23FFF8F28997845FD584C6A" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[641,766,1640,1662]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A23FFF8F28997845FD584C6A" box="[641,766,1640,1662]" italics="true" pageId="7" pageNumber="218">A. malacops</emphasis>
</taxonomicName>
, MEPN 7804, 93.4 mm SL. C) 
<taxonomicName id="E0944D20A23FFF8F2E5C7845FB284C6A" authorityName="Isbrucker" authorityYear="1980" box="[1092,1166,1640,1662]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A23FFF8F2E5C7845FB284C6A" box="[1092,1166,1640,1662]" italics="true" pageId="7" pageNumber="218">A. alga</emphasis>
</taxonomicName>
, MEPN 15169, 160.9 mm SL, D) 
<taxonomicName id="E0944D20A23FFF8F2AFA78A0FE914CB5" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[226,311,1675,1697]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A23FFF8F2AFA78A0FE914CB5" box="[226,311,1675,1697]" italics="true" pageId="7" pageNumber="218">A. shuar</emphasis>
</taxonomicName>
, MEPN 17983, 93.9 mm SL.
</paragraph>
</caption>
<subSubSection id="6F8E6528A23FFF802ADF78E1FC714BE1" lastPageId="8" lastPageNumber="219" pageId="7" pageNumber="218" type="diagnosis">
<paragraph id="272B36A3A23FFF802ADF78E1FC714BE1" blockId="7.[151,1436,1741,2018]" lastBlockId="8.[151,1437,151,501]" lastPageId="8" lastPageNumber="219" pageId="7" pageNumber="218">
<emphasis id="15E0EAB1A23FFF8F2ADF78E1FEE74CF2" bold="true" box="[199,321,1741,1766]" pageId="7" pageNumber="218">Diagnosis.</emphasis>
<taxonomicName id="E0944D20A23FFF8F2B5178E2FE484CF2" authorityName="Isbrucker" authorityYear="1980" box="[329,494,1741,1766]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A23FFF8F2B5178E2FE484CF2" box="[329,494,1741,1766]" italics="true" pageId="7" pageNumber="218">Ancistrus alga</emphasis>
</taxonomicName>
can be distinguished from other species that inhabit Amazonian Andean piedmont rivers, except 
<taxonomicName id="E0944D20A23FFF8F2B5878DEFE654D1E" baseAuthorityName="Günther" baseAuthorityYear="1869" box="[320,451,1777,1802]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="15E0EAB1A23FFF8F2B5878DEFE654D1E" box="[320,451,1777,1802]" italics="true" pageId="7" pageNumber="218">A. latifrons</emphasis>
</taxonomicName>
and 
<taxonomicName id="E0944D20A23FFF8F281D78DEFD304D1E" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[517,662,1777,1802]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A23FFF8F281D78DEFD304D1E" box="[517,662,1777,1802]" italics="true" pageId="7" pageNumber="218">A. malacops</emphasis>
</taxonomicName>
, by its mandibular ramus length which fits 3.1–4.0 times in the interorbital width. In 
<taxonomicName id="E0944D20A23FFF8F2B9F793AFDA14D3A" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[391,519,1813,1838]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="bufonius">
<emphasis id="15E0EAB1A23FFF8F2B9F793AFDA14D3A" box="[391,519,1813,1838]" italics="true" pageId="7" pageNumber="218">A. bufonius</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F280E793AFD1C4D39" baseAuthorityName="Regan" baseAuthorityYear="1904" box="[534,698,1814,1837]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="marcapatae">
<emphasis id="15E0EAB1A23FFF8F280E793AFD1C4D39" box="[534,698,1814,1837]" italics="true" pageId="7" pageNumber="218">A. marcapatae</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F28D0793AFCF34D39" baseAuthorityName="Regan" baseAuthorityYear="1904" box="[712,853,1814,1837]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="montanus">
<emphasis id="15E0EAB1A23FFF8F28D0793AFCF34D39" box="[712,853,1814,1837]" italics="true" pageId="7" pageNumber="218">A. montanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F297B793AFB894D3A" baseAuthorityName="Regan" baseAuthorityYear="1912" box="[867,1071,1813,1838]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="heterorhynchus">
<emphasis id="15E0EAB1A23FFF8F297B793AFB894D3A" box="[867,1071,1813,1838]" italics="true" pageId="7" pageNumber="218">A. heterorhynchus</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2E25793AFB614D3A" box="[1085,1223,1813,1838]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="boliviana">
<emphasis id="15E0EAB1A23FFF8F2E25793AFB614D3A" box="[1085,1223,1813,1838]" italics="true" pageId="7" pageNumber="218">A. boliviana</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2ECD793AFA334D3A" authorityName="Pearson" authorityYear="1924" box="[1237,1429,1813,1838]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="megalostomus">
<emphasis id="15E0EAB1A23FFF8F2ECD793AFA334D3A" box="[1237,1429,1813,1838]" italics="true" pageId="7" pageNumber="218">A. megalostomus</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2A8F7916FF5C4D46" authorityName="Eigenmann" authorityYear="1928" box="[151,250,1849,1874]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="occloi">
<emphasis id="15E0EAB1A23FFF8F2A8F7916FF5C4D46" box="[151,250,1849,1874]" italics="true" pageId="7" pageNumber="218">A. occloi</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2B117916FEC94D45" authorityName="in Isbrucker" baseAuthorityName="Isbrucker" baseAuthorityYear="2001" box="[265,367,1850,1873]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="greeni">
<emphasis id="15E0EAB1A23FFF8F2B117916FEC94D45" box="[265,367,1850,1873]" italics="true" pageId="7" pageNumber="218">A. greeni</emphasis>
</taxonomicName>
and 
<taxonomicName id="E0944D20A23FFF8F2BBF7916FDA14D46" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[423,519,1849,1874]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A23FFF8F2BBF7916FDA14D46" box="[423,519,1849,1874]" italics="true" pageId="7" pageNumber="218">A. shuar</emphasis>
</taxonomicName>
the mandibular ramus length fits fewer than 3.0 times in the interorbital width. It differs further except from 
<taxonomicName id="E0944D20A23FFF8F2BD47972FDEC4D62" baseAuthorityName="Günther" baseAuthorityYear="1869" box="[460,586,1885,1910]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="15E0EAB1A23FFF8F2BD47972FDEC4D62" box="[460,586,1885,1910]" italics="true" pageId="7" pageNumber="218">A. latifrons</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F28427972FCBC4D62" authorityName="Pearson" authorityYear="1924" box="[602,794,1885,1910]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="megalostomus">
<emphasis id="15E0EAB1A23FFF8F28427972FCBC4D62" box="[602,794,1885,1910]" italics="true" pageId="7" pageNumber="218">A. megalostomus</emphasis>
</taxonomicName>
and 
<taxonomicName id="E0944D20A23FFF8F294B7972FC154D62" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[851,947,1885,1910]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A23FFF8F294B7972FC154D62" box="[851,947,1885,1910]" italics="true" pageId="7" pageNumber="218">A. shuar</emphasis>
</taxonomicName>
by its interorbital width which fits 1.8–2.3 times in head length. In 
<taxonomicName id="E0944D20A23FFF8F2BA879AEFD974D8E" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[432,561,1921,1946]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="bufonius">
<emphasis id="15E0EAB1A23FFF8F2BA879AEFD974D8E" box="[432,561,1921,1946]" italics="true" pageId="7" pageNumber="218">A. bufonius</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F285979AEFD684D8E" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[577,718,1921,1946]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A23FFF8F285979AEFD684D8E" box="[577,718,1921,1946]" italics="true" pageId="7" pageNumber="218">A. malacops</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F28C679AEFCE44D8E" baseAuthorityName="Steindachner" baseAuthorityYear="1876" box="[734,834,1921,1946]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="jelskii">
<emphasis id="15E0EAB1A23FFF8F28C679AEFCE44D8E" box="[734,834,1921,1946]" italics="true" pageId="7" pageNumber="218">A. jelskii</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F294A79AEFC514D8D" baseAuthorityName="Regan" baseAuthorityYear="1904" box="[850,1015,1922,1945]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="marcapatae">
<emphasis id="15E0EAB1A23FFF8F294A79AEFC514D8D" box="[850,1015,1922,1945]" italics="true" pageId="7" pageNumber="218">A. marcapatae</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2E1079AEFB304D8D" baseAuthorityName="Regan" baseAuthorityYear="1904" box="[1032,1174,1922,1945]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="montanus">
<emphasis id="15E0EAB1A23FFF8F2E1079AEFB304D8D" box="[1032,1174,1922,1945]" italics="true" pageId="7" pageNumber="218">A. montanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2EBE79AEFAD24D8E" baseAuthorityName="Regan" baseAuthorityYear="1912" box="[1190,1396,1921,1946]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="heterorhynchus">
<emphasis id="15E0EAB1A23FFF8F2EBE79AEFAD24D8E" box="[1190,1396,1921,1946]" italics="true" pageId="7" pageNumber="218">A. heterorhynchus</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2F9C79AEFEA74DAA" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="boliviana">
<emphasis id="15E0EAB1A23FFF8F2F9C79AEFEA74DAA" italics="true" pageId="7" pageNumber="218">A. boliviana</emphasis>
</taxonomicName>
, 
<taxonomicName id="E0944D20A23FFF8F2B17798AFED54DAA" authorityName="Eigenmann" authorityYear="1928" box="[271,371,1957,1982]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="occloi">
<emphasis id="15E0EAB1A23FFF8F2B17798AFED54DAA" box="[271,371,1957,1982]" italics="true" pageId="7" pageNumber="218">A. occloi</emphasis>
</taxonomicName>
and 
<taxonomicName id="E0944D20A23FFF8F2BB4798AFDB44DA9" authorityName="in Isbrucker" baseAuthorityName="Isbrucker" baseAuthorityYear="2001" box="[428,530,1958,1981]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="greeni">
<emphasis id="15E0EAB1A23FFF8F2BB4798AFDB44DA9" box="[428,530,1958,1981]" italics="true" pageId="7" pageNumber="218">A. greeni</emphasis>
</taxonomicName>
, interorbital width fits more than 2.35 times in head length. In 
<taxonomicName id="E0944D20A23FFF8F2EC5798AFACD4DAA" authorityName="Fowler" authorityYear="1943" box="[1245,1387,1957,1982]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="lineolatus">
<emphasis id="15E0EAB1A23FFF8F2EC5798AFACD4DAA" box="[1245,1387,1957,1982]" italics="true" pageId="7" pageNumber="218">A. lineolatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="E0944D20A23FFF8F2A8F79E6FE954DF6" authorityName="Fowler" authorityYear="1945" box="[151,307,1993,2018]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="tamboensis">
<emphasis id="15E0EAB1A23FFF8F2A8F79E6FE954DF6" box="[151,307,1993,2018]" italics="true" pageId="7" pageNumber="218">A. tamboensis</emphasis>
</taxonomicName>
, it fits less than 1.7 times in head length. 
<taxonomicName id="E0944D20A23FFF8F291979E6FC024DF6" authorityName="Isbrucker" authorityYear="1980" box="[769,932,1993,2018]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A23FFF8F291979E6FC024DF6" box="[769,932,1993,2018]" italics="true" pageId="7" pageNumber="218">Ancistrus alga</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="E0944D20A23FFF8F2ECA79E6FAE84DF6" baseAuthorityName="Günther" baseAuthorityYear="1869" box="[1234,1358,1993,2018]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="15E0EAB1A23FFF8F2ECA79E6FAE84DF6" box="[1234,1358,1993,2018]" italics="true" pageId="7" pageNumber="218">A. latifrons</emphasis>
</taxonomicName>
and 
<taxonomicName id="E0944D20A23FFF802F9C79E6FEB14AA4" baseAuthorityName="Gunther" baseAuthorityYear="1864" class="Actinopterygii" family="Loricariidae" genus="Xenocara" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="8" lastPageNumber="219" order="Siluriformes" pageId="7" pageNumber="218" phylum="Chordata" rank="species" species="hoplogenys">
<emphasis id="15E0EAB1A23FFF802F9C79E6FEB14AA4" italics="true" lastPageId="8" lastPageNumber="219" pageId="7" pageNumber="218">A. hoplogenys</emphasis>
</taxonomicName>
(last one according the image available at ACSI image database) by the color pattern, its body and fins are dark brown or black with very small white dots 
<emphasis id="15E0EAB1A230FF8028F87E91FD584AC0" box="[736,766,189,212]" italics="true" pageId="8" pageNumber="219">vs.</emphasis>
uniform black, and uniform brown yellowish, respectively. The shape of caudal fin of 
<taxonomicName id="E0944D20A230FF802BC97ECCFD824AEC" authorityName="Isbrucker" authorityYear="1980" box="[465,548,223,248]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A230FF802BC97ECCFD824AEC" box="[465,548,223,248]" italics="true" pageId="8" pageNumber="219">A. alga</emphasis>
</taxonomicName>
is lunate 
<emphasis id="15E0EAB1A230FF80288F7ECDFD134AEC" box="[663,693,225,248]" italics="true" pageId="8" pageNumber="219">vs.</emphasis>
oblique truncate in 
<taxonomicName id="E0944D20A230FF8029867ECCFB994AEC" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[926,1087,223,248]" class="Actinopterygii" family="Loricariidae" genus="Xenocara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="hoplogenys">
<emphasis id="15E0EAB1A230FF8029867ECCFB994AEC" box="[926,1087,223,248]" italics="true" pageId="8" pageNumber="219">A. hoplogenys</emphasis>
</taxonomicName>
. Among the species caught in eastern 
<collectingCountry id="5F837633A230FF802AF57F28FEEC4B09" box="[237,330,260,285]" name="Ecuador" pageId="8" pageNumber="219">Ecuador</collectingCountry>
(
<taxonomicName id="E0944D20A230FF802B427F28FE454B09" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[346,483,260,285]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A230FF802B427F28FE454B09" box="[346,483,260,285]" italics="true" pageId="8" pageNumber="219">A. malacops</emphasis>
</taxonomicName>
and 
<taxonomicName id="E0944D20A230FF8028017F28FDD14B09" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[537,631,260,285]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A230FF8028017F28FDD14B09" box="[537,631,260,285]" italics="true" pageId="8" pageNumber="219">A. shuar</emphasis>
</taxonomicName>
), it is easily recognized by having a massive head and anterior region of the body; cleithral width 34.3%–36.2% SL 
<emphasis id="15E0EAB1A230FF80289E7F05FD024B54" box="[646,676,297,320]" italics="true" pageId="8" pageNumber="219">vs.</emphasis>
27.9–31.8% SL in 
<taxonomicName id="E0944D20A230FF80299B7F04FBB64B54" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[899,1040,295,320]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A230FF80299B7F04FBB64B54" box="[899,1040,295,320]" italics="true" pageId="8" pageNumber="219">A. malacops</emphasis>
</taxonomicName>
and 30.3%–32.5% SL in 
<taxonomicName id="E0944D20A230FF802F2E7F04FA334B54" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[1334,1429,295,320]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A230FF802F2E7F04FA334B54" box="[1334,1429,295,320]" italics="true" pageId="8" pageNumber="219">A. shuar</emphasis>
</taxonomicName>
. Interorbital width 17.5%–20.5% SL 
<emphasis id="15E0EAB1A230FF8028247F61FDFC4B70" box="[572,602,333,356]" italics="true" pageId="8" pageNumber="219">vs.</emphasis>
13.4%–16.4% SL in 
<taxonomicName id="E0944D20A230FF80294F7F60FC424B71" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[855,996,332,357]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A230FF80294F7F60FC424B71" box="[855,996,332,357]" italics="true" pageId="8" pageNumber="219">A. malacops</emphasis>
</taxonomicName>
and 14.4%–17.0% SL in 
<taxonomicName id="E0944D20A230FF802F0B7F60FAD54B71" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[1299,1395,332,357]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A230FF802F0B7F60FAD54B71" box="[1299,1395,332,357]" italics="true" pageId="8" pageNumber="219">A. shuar</emphasis>
</taxonomicName>
. In males, the values are higher, cleithral width 35.3%–36.2% SL 
<emphasis id="15E0EAB1A230FF80297E7F5DFC234B9C" box="[870,901,369,392]" italics="true" pageId="8" pageNumber="219">vs.</emphasis>
27.8%–30.2% SL in 
<taxonomicName id="E0944D20A230FF802E667F5CFAAD4B9C" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[1150,1291,367,392]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A230FF802E667F5CFAAD4B9C" box="[1150,1291,367,392]" italics="true" pageId="8" pageNumber="219">A. malacops</emphasis>
</taxonomicName>
and 30.7%– 32.5% SL in 
<taxonomicName id="E0944D20A230FF802B327FB8FE214BB9" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[298,391,404,429]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A230FF802B327FB8FE214BB9" box="[298,391,404,429]" italics="true" pageId="8" pageNumber="219">A. shuar</emphasis>
</taxonomicName>
. Interorbital width 20.1%–20.5% SL 
<emphasis id="15E0EAB1A230FF8029367FB9FCEA4BB8" box="[814,844,405,428]" italics="true" pageId="8" pageNumber="219">vs.</emphasis>
13.5%–15.7% SL in 
<taxonomicName id="E0944D20A230FF802E257FB8FB614BB9" baseAuthorityName="Cope" baseAuthorityYear="1872" box="[1085,1223,404,429]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="malacops">
<emphasis id="15E0EAB1A230FF802E257FB8FB614BB9" box="[1085,1223,404,429]" italics="true" pageId="8" pageNumber="219">A. malacops</emphasis>
</taxonomicName>
and 15.4%–17.0% SL in 
<taxonomicName id="E0944D20A230FF802AC67F94FE994BC4" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[222,319,439,464]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="shuar">
<emphasis id="15E0EAB1A230FF802AC67F94FE994BC4" box="[222,319,439,464]" italics="true" pageId="8" pageNumber="219">A. shuar</emphasis>
</taxonomicName>
(
<tableCitation id="6A160318A230FF802B577F9BFE014BC4" box="[335,423,439,464]" captionStart="TABLE 5" captionStartId="21.[151,239,1049,1072]" captionText="TABLE 5. Comparative morphometric data for the males of three species. Measurements are expressed as percentage of standard length." pageId="8" pageNumber="219">Table 5</tableCitation>
). Also, the color pattern of 
<taxonomicName id="E0944D20A230FF8028FD7F94FC9E4BC4" authorityName="Isbrucker" authorityYear="1980" box="[741,824,439,464]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A230FF8028FD7F94FC9E4BC4" box="[741,824,439,464]" italics="true" pageId="8" pageNumber="219">A. alga</emphasis>
</taxonomicName>
, black or dark brown with very minute white dots on body and fins is distinctive when compared with other species (
<figureCitation id="BFAF2A26A230FF8029467FF0FC0B4BE0" box="[862,941,476,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[204,1390,193,1949]" captionTargetId="figure@5.[204,1390,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Females of Ancistrus species, lateral view. From top to bottom: A. shuar, MEPN 17983, 93.9 mm SL. A. alga, MEPN 15169, 160.9 mm SL. A. clementinae, MEPN 5996, 100.1 mm SL. and A. malacops, MEPN 7804, 93.4 mm SL." httpUri="https://zenodo.org/record/2612129/files/figure.png" pageId="8" pageNumber="219">Figs. 2</figureCitation>
, 
<figureCitation id="BFAF2A26A230FF8029A17FF0FC6F4BE1" box="[953,969,476,501]" captionStart="FIGURE 5" captionStartId="9.[151,250,1988,2010]" captionTargetBox="[177,1414,193,1976]" captionTargetId="figure@9.[177,1414,193,1976]" captionTargetPageId="9" captionText="FIGURE 5. Ancistrus alga, MEPN 14853, 115.1 mm SL, male. From top to bottom, lateral, dorsal, and ventral view." httpUri="https://zenodo.org/record/2612135/files/figure.png" pageId="8" pageNumber="219">5</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="73EB662BA230FF802A8F7B08FA9A4F2E" box="[151,1340,1316,1338]" httpUri="https://zenodo.org/record/2612133/files/figure.png" pageId="8" pageNumber="219" startId="8.[151,250,1316,1338]" targetBox="[169,1417,553,1294]" targetPageId="8">
<paragraph id="272B36A3A230FF802A8F7B08FA9A4F2E" blockId="8.[151,1340,1316,1338]" box="[151,1340,1316,1338]" pageId="8" pageNumber="219">
<emphasis id="15E0EAB1A230FF802A8F7B08FEB44F2D" bold="true" box="[151,274,1316,1338]" pageId="8" pageNumber="219">FIGURE 4.</emphasis>
<emphasis id="15E0EAB1A230FF802B017B08FE1A4F2E" box="[281,444,1316,1338]" italics="true" pageId="8" pageNumber="219">
<taxonomicName id="E0944D20A230FF802B017B08FE1F4F2E" authorityName="Provenzano &amp; Barriga-Salazar" authorityYear="2018" box="[281,441,1316,1338]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="219" phylum="Chordata" rank="species" species="shuar">Ancistrus shuar</taxonomicName>
,
</emphasis>
MECN-DP-1637, 96.5 mm SL, male, paratype when alive (Photo courtesy Jorge Brito).
</paragraph>
</caption>
<subSubSection id="6F8E6528A230FF832ADF7B4DFDB64874" lastPageId="11" lastPageNumber="222" pageId="8" pageNumber="219" type="description">
<paragraph id="272B36A3A230FF802ADF7B4DFC9C4C3A" blockId="8.[151,1437,1377,1942]" pageId="8" pageNumber="219">
<emphasis id="15E0EAB1A230FF802ADF7B4DFEFF4F6E" bold="true" box="[199,345,1377,1402]" pageId="8" pageNumber="219">Description.</emphasis>
Morphometric data given in 
<tableCitation id="6A160318A230FF8028B97B4DFD594F6D" box="[673,767,1377,1402]" captionStart="TABLE 2" captionStartId="10.[151,239,152,175]" captionText="TABLE 2. Morphometric data of Ancistrus alga. Measurements are expressed as percentage of standard length. N=11" pageId="8" pageNumber="219">Tables 2</tableCitation>
and 
<tableCitation id="6A160318A230FF80292E7B4DFCE14F6E" box="[822,839,1377,1402]" captionStart="TABLE 5" captionStartId="21.[151,239,1049,1072]" captionText="TABLE 5. Comparative morphometric data for the males of three species. Measurements are expressed as percentage of standard length." pageId="8" pageNumber="219">5</tableCitation>
. Body robust, depressed anteriorly and progressively become compressed posteriorly. Caudal peduncle compressed, deep and, robust. Dorsal profile of body from tip of snout through dorsal-fin origin gently convex, then descending in straight line to adipose-fin origin. From this point to caudal-fin origin straight and horizontal. Ventral profile of body flat and straight or slightly concave. Ventral surface of head and belly naked until anal-fin origin. Urogenital papilla is not usually visible, in some mature males opening is partially visible just posterior to anus (
<figureCitation id="BFAF2A26A230FF8028D97839FCB64C39" box="[705,784,1557,1582]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[204,1390,193,1949]" captionTargetId="figure@5.[204,1390,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Females of Ancistrus species, lateral view. From top to bottom: A. shuar, MEPN 17983, 93.9 mm SL. A. alga, MEPN 15169, 160.9 mm SL. A. clementinae, MEPN 5996, 100.1 mm SL. and A. malacops, MEPN 7804, 93.4 mm SL." httpUri="https://zenodo.org/record/2612129/files/figure.png" pageId="8" pageNumber="219">Figs. 2</figureCitation>
, 
<figureCitation id="BFAF2A26A230FF8029047839FC8A4C3A" box="[796,812,1557,1582]" captionStart="FIGURE 5" captionStartId="9.[151,250,1988,2010]" captionTargetBox="[177,1414,193,1976]" captionTargetId="figure@9.[177,1414,193,1976]" captionTargetPageId="9" captionText="FIGURE 5. Ancistrus alga, MEPN 14853, 115.1 mm SL, male. From top to bottom, lateral, dorsal, and ventral view." httpUri="https://zenodo.org/record/2612135/files/figure.png" pageId="8" pageNumber="219">5</figureCitation>
).
</paragraph>
<paragraph id="272B36A3A230FF802ADF7815FE424D82" blockId="8.[151,1437,1377,1942]" pageId="8" pageNumber="219">
Head massive, very wide and depressed. In dorsal and lateral view, this species shows a massive head compared with the other two species. Snout partially naked with or without fleshy cylindrical tentacles branched or not, its contour semicircular. In males, naked area wider but does not reach nares or orbits. Females have only a narrow naked strip on snout border. Nostrils juxtaposed and closer to eyes than tip of snout. Eyes in dorsolateral position, orbits not raised and without odontodes. Interorbital space is broad and flat. Supraoccipital flat without ridges, posterior border straight and truncate. Movable hypertrophied cheek odontodes well developed, specimens can have from eight to twelve. The size of these odontodes also is variable in each specimen, the longest odontode nearly extends to pectoral-fin origin independently of size of specimen. Anteriorly, base of movable hypertrophied odontodes has covering plates. Opercular bone has an exposed surface easily visible externally, its lateral margin carries odontodes (
<figureCitation id="BFAF2A26A230FF802B737951FE1C4D81" box="[363,442,1917,1942]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[204,1390,193,1949]" captionTargetId="figure@5.[204,1390,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Females of Ancistrus species, lateral view. From top to bottom: A. shuar, MEPN 17983, 93.9 mm SL. A. alga, MEPN 15169, 160.9 mm SL. A. clementinae, MEPN 5996, 100.1 mm SL. and A. malacops, MEPN 7804, 93.4 mm SL." httpUri="https://zenodo.org/record/2612129/files/figure.png" pageId="8" pageNumber="219">Figs. 2</figureCitation>
, 
<figureCitation id="BFAF2A26A230FF802BDE7951FE704D82" box="[454,470,1917,1942]" captionStart="FIGURE 5" captionStartId="9.[151,250,1988,2010]" captionTargetBox="[177,1414,193,1976]" captionTargetId="figure@9.[177,1414,193,1976]" captionTargetPageId="9" captionText="FIGURE 5. Ancistrus alga, MEPN 14853, 115.1 mm SL, male. From top to bottom, lateral, dorsal, and ventral view." httpUri="https://zenodo.org/record/2612135/files/figure.png" pageId="8" pageNumber="219">5</figureCitation>
).
</paragraph>
<caption id="73EB662BA231FF812A8F79E8FA9A4DCE" box="[151,1340,1988,2010]" httpUri="https://zenodo.org/record/2612135/files/figure.png" pageId="9" pageNumber="220" startId="9.[151,250,1988,2010]" targetBox="[177,1414,193,1976]" targetPageId="9">
<paragraph id="272B36A3A231FF812A8F79E8FA9A4DCE" blockId="9.[151,1340,1988,2010]" box="[151,1340,1988,2010]" pageId="9" pageNumber="220">
<emphasis id="15E0EAB1A231FF812A8F79E8FEB44DCD" bold="true" box="[151,274,1988,2010]" pageId="9" pageNumber="220">FIGURE 5.</emphasis>
<taxonomicName id="E0944D20A231FF812B0179E8FE0A4DCE" authorityName="Isbrucker" authorityYear="1980" box="[281,428,1988,2010]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="220" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A231FF812B0179E8FE0A4DCE" box="[281,428,1988,2010]" italics="true" pageId="9" pageNumber="220">Ancistrus alga</emphasis>
</taxonomicName>
, MEPN 14853, 115.1 mm SL, male. From top to bottom, lateral, dorsal, and ventral view.
</paragraph>
</caption>
<caption id="73EB662BA232FF822A8F7EB4FA254ABB" box="[151,1411,151,175]" pageId="10" pageNumber="221">
<paragraph id="272B36A3A232FF822A8F7EB4FA254ABB" blockId="10.[151,1411,151,175]" box="[151,1411,151,175]" pageId="10" pageNumber="221">
<emphasis id="15E0EAB1A232FF822A8F7EB4FEAF4ABB" bold="true" box="[151,265,152,175]" pageId="10" pageNumber="221">TABLE 2.</emphasis>
Morphometric data of 
<taxonomicName id="E0944D20A232FF8228197EB4FD3D4ABB" authorityName="Isbrucker" authorityYear="1980" box="[513,667,151,175]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="221" phylum="Chordata" rank="species" species="alga">
<emphasis id="15E0EAB1A232FF8228197EB4FD3D4ABB" box="[513,667,151,175]" italics="true" pageId="10" pageNumber="221">Ancistrus alga</emphasis>
</taxonomicName>
. Measurements are expressed as percentage of standard length. N=11
</paragraph>
</caption>
<paragraph id="272B36A3A232FF822A857EE0FAB04E08" pageId="10" pageNumber="221">
<table id="5594C403A23200772A857EE0FA894E08" box="[157,1327,204,1052]" colsContinueIn="15.[158,1325,235,1116]" gridcols="5" gridrows="21" pageId="10" pageNumber="221">
<tr id="99A434E1A23200772A857EE0FA894AF6" box="[157,1327,204,226]" gridrow="0" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857EE0FE3B4AF6" box="[157,413,204,226]" gridcol="0" gridrow="0" pageId="10" pageNumber="221">Character</th>
<th id="DA755D9DA232007728A37EE0FCA84AF6" box="[699,782,204,226]" gridcol="1" gridrow="0" pageId="10" pageNumber="221">Average</th>
<th id="DA755D9DA232007729917EE0FC1F4AF6" box="[905,953,204,226]" gridcol="2" gridrow="0" pageId="10" pageNumber="221">STD</th>
<th id="DA755D9DA23200772E377EE0FBC64AF6" box="[1071,1120,204,226]" gridcol="3" gridrow="0" pageId="10" pageNumber="221">Min.</th>
<th id="DA755D9DA23200772EEE7EE0FA894AF6" box="[1270,1327,204,226]" gridcol="4" gridrow="0" pageId="10" pageNumber="221">Max.</th>
</tr>
<tr id="99A434E1A23200772A857ED9FA894B1F" box="[157,1327,245,267]" gridrow="1" pageId="10" pageNumber="221" rowspan-1="1" rowspan-2="1">
<th id="DA755D9DA23200772A857ED9FE3B4B1F" box="[157,413,245,267]" gridcol="0" gridrow="1" pageId="10" pageNumber="221">SL (mm)</th>
<td id="DA755D9DA23200772E377ED9FBC64B1F" box="[1071,1120,245,267]" gridcol="3" gridrow="1" pageId="10" pageNumber="221">60.7</td>
<td id="DA755D9DA23200772EEE7ED9FA894B1F" box="[1270,1327,245,267]" gridcol="4" gridrow="1" pageId="10" pageNumber="221">160.9</td>
</tr>
<tr id="99A434E1A23200772A857F32FA894B20" box="[157,1327,286,308]" gridrow="2" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857F32FE3B4B20" box="[157,413,286,308]" gridcol="0" gridrow="2" pageId="10" pageNumber="221">Head length</th>
<td id="DA755D9DA232007728A37F32FCA84B20" box="[699,782,286,308]" gridcol="1" gridrow="2" pageId="10" pageNumber="221">39.4</td>
<td id="DA755D9DA232007729917F32FC1F4B20" box="[905,953,286,308]" gridcol="2" gridrow="2" pageId="10" pageNumber="221">2.3</td>
<td id="DA755D9DA23200772E377F32FBC64B20" box="[1071,1120,286,308]" gridcol="3" gridrow="2" pageId="10" pageNumber="221">36.3</td>
<td id="DA755D9DA23200772EEE7F32FA894B20" box="[1270,1327,286,308]" gridcol="4" gridrow="2" pageId="10" pageNumber="221">43.9</td>
</tr>
<tr id="99A434E1A23200772A857F64FA894B4A" box="[157,1327,328,350]" gridrow="3" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857F64FE3B4B4A" box="[157,413,328,350]" gridcol="0" gridrow="3" pageId="10" pageNumber="221">Predorsal length</th>
<td id="DA755D9DA232007728A37F64FCA84B4A" box="[699,782,328,350]" gridcol="1" gridrow="3" pageId="10" pageNumber="221">48.7</td>
<td id="DA755D9DA232007729917F64FC1F4B4A" box="[905,953,328,350]" gridcol="2" gridrow="3" pageId="10" pageNumber="221">2.3</td>
<td id="DA755D9DA23200772E377F64FBC64B4A" box="[1071,1120,328,350]" gridcol="3" gridrow="3" pageId="10" pageNumber="221">45.7</td>
<td id="DA755D9DA23200772EEE7F64FA894B4A" box="[1270,1327,328,350]" gridcol="4" gridrow="3" pageId="10" pageNumber="221">52.9</td>
</tr>
<tr id="99A434E1A23200772A857F5DFA894B93" box="[157,1327,369,391]" gridrow="4" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857F5DFE3B4B93" box="[157,413,369,391]" gridcol="0" gridrow="4" pageId="10" pageNumber="221">Postdorsal length</th>
<td id="DA755D9DA232007728A37F5DFCA84B93" box="[699,782,369,391]" gridcol="1" gridrow="4" pageId="10" pageNumber="221">31.6</td>
<td id="DA755D9DA232007729917F5DFC1F4B93" box="[905,953,369,391]" gridcol="2" gridrow="4" pageId="10" pageNumber="221">1.2</td>
<td id="DA755D9DA23200772E377F5DFBC64B93" box="[1071,1120,369,391]" gridcol="3" gridrow="4" pageId="10" pageNumber="221">29.5</td>
<td id="DA755D9DA23200772EEE7F5DFA894B93" box="[1270,1327,369,391]" gridcol="4" gridrow="4" pageId="10" pageNumber="221">33.4</td>
</tr>
<tr id="99A434E1A23200772A857FB6FA894BA4" box="[157,1327,410,432]" gridrow="5" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857FB6FE3B4BA4" box="[157,413,410,432]" gridcol="0" gridrow="5" pageId="10" pageNumber="221">Interdorsal length</th>
<td id="DA755D9DA232007728A37FB6FCA84BA4" box="[699,782,410,432]" gridcol="1" gridrow="5" pageId="10" pageNumber="221">16.8</td>
<td id="DA755D9DA232007729917FB6FC1F4BA4" box="[905,953,410,432]" gridcol="2" gridrow="5" pageId="10" pageNumber="221">0.8</td>
<td id="DA755D9DA23200772E377FB6FBC64BA4" box="[1071,1120,410,432]" gridcol="3" gridrow="5" pageId="10" pageNumber="221">15.2</td>
<td id="DA755D9DA23200772EEE7FB6FA894BA4" box="[1270,1327,410,432]" gridcol="4" gridrow="5" pageId="10" pageNumber="221">18.2</td>
</tr>
<tr id="99A434E1A23200772A857FE8FA894BCE" box="[157,1327,452,474]" gridrow="6" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857FE8FE3B4BCE" box="[157,413,452,474]" gridcol="0" gridrow="6" pageId="10" pageNumber="221">Preanal length</th>
<td id="DA755D9DA232007728A37FE8FCA84BCE" box="[699,782,452,474]" gridcol="1" gridrow="6" pageId="10" pageNumber="221">70.2</td>
<td id="DA755D9DA232007729917FE8FC1F4BCE" box="[905,953,452,474]" gridcol="2" gridrow="6" pageId="10" pageNumber="221">0.8</td>
<td id="DA755D9DA23200772E377FE8FBC64BCE" box="[1071,1120,452,474]" gridcol="3" gridrow="6" pageId="10" pageNumber="221">68.9</td>
<td id="DA755D9DA23200772EEE7FE8FA894BCE" box="[1270,1327,452,474]" gridcol="4" gridrow="6" pageId="10" pageNumber="221">71.2</td>
</tr>
<tr id="99A434E1A23200772A857FC1FA894817" box="[157,1327,493,515]" gridrow="7" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857FC1FE3B4817" box="[157,413,493,515]" gridcol="0" gridrow="7" pageId="10" pageNumber="221">Postanal length</th>
<td id="DA755D9DA232007728A37FC1FCA84817" box="[699,782,493,515]" gridcol="1" gridrow="7" pageId="10" pageNumber="221">28.4</td>
<td id="DA755D9DA232007729917FC1FC1F4817" box="[905,953,493,515]" gridcol="2" gridrow="7" pageId="10" pageNumber="221">1.0</td>
<td id="DA755D9DA23200772E377FC1FBC64817" box="[1071,1120,493,515]" gridcol="3" gridrow="7" pageId="10" pageNumber="221">26.7</td>
<td id="DA755D9DA23200772EEE7FC1FA894817" box="[1270,1327,493,515]" gridcol="4" gridrow="7" pageId="10" pageNumber="221">30.0</td>
</tr>
<tr id="99A434E1A23200772A857C3AFA894838" box="[157,1327,534,556]" gridrow="8" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857C3AFE3B4838" box="[157,413,534,556]" gridcol="0" gridrow="8" pageId="10" pageNumber="221">Thoracic length</th>
<td id="DA755D9DA232007728A37C3AFCA84838" box="[699,782,534,556]" gridcol="1" gridrow="8" pageId="10" pageNumber="221">24.0</td>
<td id="DA755D9DA232007729917C3AFC1F4838" box="[905,953,534,556]" gridcol="2" gridrow="8" pageId="10" pageNumber="221">0.9</td>
<td id="DA755D9DA23200772E377C3AFBC64838" box="[1071,1120,534,556]" gridcol="3" gridrow="8" pageId="10" pageNumber="221">22.4</td>
<td id="DA755D9DA23200772EEE7C3AFA894838" box="[1270,1327,534,556]" gridcol="4" gridrow="8" pageId="10" pageNumber="221">25.8</td>
</tr>
<tr id="99A434E1A23200772A857C6CFA894842" box="[157,1327,576,598]" gridrow="9" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857C6CFE3B4842" box="[157,413,576,598]" gridcol="0" gridrow="9" pageId="10" pageNumber="221">Abdominal length</th>
<td id="DA755D9DA232007728A37C6CFCA84842" box="[699,782,576,598]" gridcol="1" gridrow="9" pageId="10" pageNumber="221">21.8</td>
<td id="DA755D9DA232007729917C6CFC1F4842" box="[905,953,576,598]" gridcol="2" gridrow="9" pageId="10" pageNumber="221">0.8</td>
<td id="DA755D9DA23200772E377C6CFBC64842" box="[1071,1120,576,598]" gridcol="3" gridrow="9" pageId="10" pageNumber="221">20.4</td>
<td id="DA755D9DA23200772EEE7C6CFA894842" box="[1270,1327,576,598]" gridcol="4" gridrow="9" pageId="10" pageNumber="221">22.8</td>
</tr>
<tr id="99A434E1A23200772A857C45FA89486B" box="[157,1327,617,639]" gridrow="10" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857C45FE3B486B" box="[157,413,617,639]" gridcol="0" gridrow="10" pageId="10" pageNumber="221">Dorsal fin base</th>
<td id="DA755D9DA232007728A37C45FCA8486B" box="[699,782,617,639]" gridcol="1" gridrow="10" pageId="10" pageNumber="221">25.2</td>
<td id="DA755D9DA232007729917C45FC1F486B" box="[905,953,617,639]" gridcol="2" gridrow="10" pageId="10" pageNumber="221">1.1</td>
<td id="DA755D9DA23200772E377C45FBC6486B" box="[1071,1120,617,639]" gridcol="3" gridrow="10" pageId="10" pageNumber="221">23.1</td>
<td id="DA755D9DA23200772EEE7C45FA89486B" box="[1270,1327,617,639]" gridcol="4" gridrow="10" pageId="10" pageNumber="221">26.6</td>
</tr>
<tr id="99A434E1A23200772A857CBEFA8948BC" box="[157,1327,658,680]" gridrow="11" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857CBEFE3B48BC" box="[157,413,658,680]" gridcol="0" gridrow="11" pageId="10" pageNumber="221">Dorsal spine length</th>
<td id="DA755D9DA232007728A37CBEFCA848BC" box="[699,782,658,680]" gridcol="1" gridrow="11" pageId="10" pageNumber="221">31.5</td>
<td id="DA755D9DA232007729917CBEFC1F48BC" box="[905,953,658,680]" gridcol="2" gridrow="11" pageId="10" pageNumber="221">1.1</td>
<td id="DA755D9DA23200772E377CBEFBC648BC" box="[1071,1120,658,680]" gridcol="3" gridrow="11" pageId="10" pageNumber="221">29.8</td>
<td id="DA755D9DA23200772EEE7CBEFA8948BC" box="[1270,1327,658,680]" gridcol="4" gridrow="11" pageId="10" pageNumber="221">33.4</td>
</tr>
<tr id="99A434E1A23200772A857C90FA8948C6" box="[157,1327,700,722]" gridrow="12" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857C90FE3B48C6" box="[157,413,700,722]" gridcol="0" gridrow="12" pageId="10" pageNumber="221">Pectoral spine length</th>
<td id="DA755D9DA232007728A37C90FCA848C6" box="[699,782,700,722]" gridcol="1" gridrow="12" pageId="10" pageNumber="221">35.3</td>
<td id="DA755D9DA232007729917C90FC1F48C6" box="[905,953,700,722]" gridcol="2" gridrow="12" pageId="10" pageNumber="221">1.9</td>
<td id="DA755D9DA23200772E377C90FBC648C6" box="[1071,1120,700,722]" gridcol="3" gridrow="12" pageId="10" pageNumber="221">31.9</td>
<td id="DA755D9DA23200772EEE7C90FA8948C6" box="[1270,1327,700,722]" gridcol="4" gridrow="12" pageId="10" pageNumber="221">38.9</td>
</tr>
<tr id="99A434E1A23200772A857CC9FA8948EF" box="[157,1327,741,763]" gridrow="13" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857CC9FE3B48EF" box="[157,413,741,763]" gridcol="0" gridrow="13" pageId="10" pageNumber="221">Pelvic spine length</th>
<td id="DA755D9DA232007728A37CC9FCA848EF" box="[699,782,741,763]" gridcol="1" gridrow="13" pageId="10" pageNumber="221">26.8</td>
<td id="DA755D9DA232007729917CC9FC1F48EF" box="[905,953,741,763]" gridcol="2" gridrow="13" pageId="10" pageNumber="221">1.2</td>
<td id="DA755D9DA23200772E377CC9FBC648EF" box="[1071,1120,741,763]" gridcol="3" gridrow="13" pageId="10" pageNumber="221">24.4</td>
<td id="DA755D9DA23200772EEE7CC9FA8948EF" box="[1270,1327,741,763]" gridcol="4" gridrow="13" pageId="10" pageNumber="221">28.3</td>
</tr>
<tr id="99A434E1A23200772A857D22FA894930" box="[157,1327,782,804]" gridrow="14" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857D22FE3B4930" box="[157,413,782,804]" gridcol="0" gridrow="14" pageId="10" pageNumber="221">Cleithral width</th>
<td id="DA755D9DA232007728A37D22FCA84930" box="[699,782,782,804]" gridcol="1" gridrow="14" pageId="10" pageNumber="221">35.3</td>
<td id="DA755D9DA232007729917D22FC1F4930" box="[905,953,782,804]" gridcol="2" gridrow="14" pageId="10" pageNumber="221">0.6</td>
<td id="DA755D9DA23200772E377D22FBC64930" box="[1071,1120,782,804]" gridcol="3" gridrow="14" pageId="10" pageNumber="221">34.3</td>
<td id="DA755D9DA23200772EEE7D22FA894930" box="[1270,1327,782,804]" gridcol="4" gridrow="14" pageId="10" pageNumber="221">36.2</td>
</tr>
<tr id="99A434E1A23200772A857D14FA89495A" box="[157,1327,824,846]" gridrow="15" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857D14FE3B495A" box="[157,413,824,846]" gridcol="0" gridrow="15" pageId="10" pageNumber="221">Caudal peduncle depth</th>
<td id="DA755D9DA232007728A37D14FCA8495A" box="[699,782,824,846]" gridcol="1" gridrow="15" pageId="10" pageNumber="221">10.9</td>
<td id="DA755D9DA232007729917D14FC1F495A" box="[905,953,824,846]" gridcol="2" gridrow="15" pageId="10" pageNumber="221">0.3</td>
<td id="DA755D9DA23200772E377D14FBC6495A" box="[1071,1120,824,846]" gridcol="3" gridrow="15" pageId="10" pageNumber="221">10.0</td>
<td id="DA755D9DA23200772EEE7D14FA89495A" box="[1270,1327,824,846]" gridcol="4" gridrow="15" pageId="10" pageNumber="221">11.2</td>
</tr>
<tr id="99A434E1A23200772A857D4DFA894963" box="[157,1327,865,887]" gridrow="16" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857D4DFE3B4963" box="[157,413,865,887]" gridcol="0" gridrow="16" pageId="10" pageNumber="221">Head depth</th>
<td id="DA755D9DA232007728A37D4DFCA84963" box="[699,782,865,887]" gridcol="1" gridrow="16" pageId="10" pageNumber="221">21.2</td>
<td id="DA755D9DA232007729917D4DFC1F4963" box="[905,953,865,887]" gridcol="2" gridrow="16" pageId="10" pageNumber="221">1.5</td>
<td id="DA755D9DA23200772E377D4DFBC64963" box="[1071,1120,865,887]" gridcol="3" gridrow="16" pageId="10" pageNumber="221">18.8</td>
<td id="DA755D9DA23200772EEE7D4DFA894963" box="[1270,1327,865,887]" gridcol="4" gridrow="16" pageId="10" pageNumber="221">24.5</td>
</tr>
<tr id="99A434E1A23200772A857DA6FA8949B4" box="[157,1327,906,928]" gridrow="17" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857DA6FE3B49B4" box="[157,413,906,928]" gridcol="0" gridrow="17" pageId="10" pageNumber="221">Snout length</th>
<td id="DA755D9DA232007728A37DA6FCA849B4" box="[699,782,906,928]" gridcol="1" gridrow="17" pageId="10" pageNumber="221">23.5</td>
<td id="DA755D9DA232007729917DA6FC1F49B4" box="[905,953,906,928]" gridcol="2" gridrow="17" pageId="10" pageNumber="221">1.6</td>
<td id="DA755D9DA23200772E377DA6FBC649B4" box="[1071,1120,906,928]" gridcol="3" gridrow="17" pageId="10" pageNumber="221">21.9</td>
<td id="DA755D9DA23200772EEE7DA6FA8949B4" box="[1270,1327,906,928]" gridcol="4" gridrow="17" pageId="10" pageNumber="221">26.4</td>
</tr>
<tr id="99A434E1A23200772A857D98FA8949DE" box="[157,1327,948,970]" gridrow="18" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857D98FE3B49DE" box="[157,413,948,970]" gridcol="0" gridrow="18" pageId="10" pageNumber="221">Interorbital width</th>
<td id="DA755D9DA232007728A37D98FCA849DE" box="[699,782,948,970]" gridcol="1" gridrow="18" pageId="10" pageNumber="221">19.7</td>
<td id="DA755D9DA232007729917D98FC1F49DE" box="[905,953,948,970]" gridcol="2" gridrow="18" pageId="10" pageNumber="221">0.9</td>
<td id="DA755D9DA23200772E377D98FBC649DE" box="[1071,1120,948,970]" gridcol="3" gridrow="18" pageId="10" pageNumber="221">17.5</td>
<td id="DA755D9DA23200772EEE7D98FA8949DE" box="[1270,1327,948,970]" gridcol="4" gridrow="18" pageId="10" pageNumber="221">20.5</td>
</tr>
<tr id="99A434E1A23200772A857DF1FA8949E7" box="[157,1327,989,1011]" gridrow="19" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857DF1FE3B49E7" box="[157,413,989,1011]" gridcol="0" gridrow="19" pageId="10" pageNumber="221">Orbital diameter</th>
<td id="DA755D9DA232007728A37DF1FCA849E7" box="[699,782,989,1011]" gridcol="1" gridrow="19" pageId="10" pageNumber="221">6.1</td>
<td id="DA755D9DA232007729917DF1FC1F49E7" box="[905,953,989,1011]" gridcol="2" gridrow="19" pageId="10" pageNumber="221">0.4</td>
<td id="DA755D9DA23200772E377DF1FBC649E7" box="[1071,1120,989,1011]" gridcol="3" gridrow="19" pageId="10" pageNumber="221">5.4</td>
<td id="DA755D9DA23200772EEE7DF1FA8949E7" box="[1270,1327,989,1011]" gridcol="4" gridrow="19" pageId="10" pageNumber="221">7.0</td>
</tr>
<tr id="99A434E1A23200772A857A2AFA894E08" box="[157,1327,1030,1052]" gridrow="20" pageId="10" pageNumber="221">
<th id="DA755D9DA23200772A857A2AFE3B4E08" box="[157,413,1030,1052]" gridcol="0" gridrow="20" pageId="10" pageNumber="221">Mandibular ramus length</th>
<td id="DA755D9DA232007728A37A2AFCA84E08" box="[699,782,1030,1052]" gridcol="1" gridrow="20" pageId="10" pageNumber="221">5.5</td>
<td id="DA755D9DA232007729917A2AFC1F4E08" box="[905,953,1030,1052]" gridcol="2" gridrow="20" pageId="10" pageNumber="221">0.3</td>
<td id="DA755D9DA23200772E377A2AFBC64E08" box="[1071,1120,1030,1052]" gridcol="3" gridrow="20" pageId="10" pageNumber="221">5.1</td>
<td id="DA755D9DA23200772EEE7A2AFA894E08" box="[1270,1327,1030,1052]" gridcol="4" gridrow="20" pageId="10" pageNumber="221">6.0</td>
</tr>
</table>
</paragraph>
<paragraph id="272B36A3A232FF822ADF7A71FAE54F82" blockId="10.[151,1437,1117,2006]" pageId="10" pageNumber="221">
Mouth oval or rounded. Upper lip narrow, usually cover premaxilla and only external surface is visible, edge is almost horizontal and with very minute undulations. The internal surface papillose. Lower lip broad, its border with very minute undulations. Lower lip surface papillose. Papillae smaller near border of lip increasing in size near lower jaws. Papillae of anterior lip have similar size from those near lower jaws. Maxillary barbels evident and free. Upper and lower jaws with similar length. The hemimandibles are straight, placed horizontally or forming a 
<emphasis id="15E0EAB1A232FF822F937AC2FA3A4F11" box="[1419,1436,1262,1285]" italics="true" pageId="10" pageNumber="221">V</emphasis>
between them. Teeth numerous and minute. Between 65 to 80 teeth in each hemimandible. Premaxillary and dentary teeth with same size. Teeth incisor 
<typeStatus id="F82F8801A232FF82289F7B1AFD1C4F5A" box="[647,698,1334,1358]" pageId="10" pageNumber="221">type</typeStatus>
, asymmetrically bifid, medial cusp longer and wider than lateral cusp. Medial cusp rounded or straight truncated, lateral cusp pointed. Tooth apex curved toward interior of mouth. Tooth apex yellowish, stalk whitish. Premaxillary and dentary without posterior papillae or ornamentation.
</paragraph>
<paragraph id="272B36A3A232FF822ADF7B8DFD564CCD" blockId="10.[151,1437,1117,2006]" pageId="10" pageNumber="221">Lateral line plates 23–24. Post-anal plates 11–12. Six plates between dorsal-fin and adipose-fin, all plates without keels. Origin dorsal-fin is anterior to vertical passing through pelvic-fin origin. Dorsal-fin with one spine, followed by seven branched rays; when depressed reaching adipose-fin origin. Adipose-fin well developed and always present. Spine of adipose-fin wide, and curved. Pectoral-fin with one spine, and six branched rays. When depressed, spine of pectoral-fin reaches half, (females and males) of pelvic-fin spine length. Spine and first branched ray of pectoral-fin with same length. Distal region of pectoral-fin spine with enlarged odontodes, more developed in males, and a small fleshy prominence. Pelvic-fin with one spine and five branched rays; its posterior margin surpassed anal-fin base when depressed. Anal-fin with one flexible spine and four branched rays. Caudalfin rays i,14,i. Posterior border of caudal-fin concave.</paragraph>
<paragraph id="272B36A3A232FF832ADF78C9FC3C4AEC" blockId="10.[151,1437,1117,2006]" lastBlockId="11.[151,1437,151,608]" lastPageId="11" lastPageNumber="222" pageId="10" pageNumber="221">
<emphasis id="15E0EAB1A232FF822ADF78C9FE104CEA" bold="true" box="[199,438,1765,1790]" pageId="10" pageNumber="221">Sexual dimorphism.</emphasis>
Adult males have a wide naked area on snout, with developed (long) and branched fleshy tentacles. Tentacles are arranged in the following pattern: Along snout edge a row of tentacles, long and conical are present, those at snout tip are longest and branched. Dorsal midline with two or three tentacles very long, branched or not. Anterior the nares, the dorsal midline row of tentacles split in three or two tentacles on each side. The tentacles near midline can show ramifications, while the laterals are just conical. On each side of head, from snout edge to near eyes, another row of three or four, conical tentacles is present, passing in front of cheek odontodes (
<figureCitation id="BFAF2A26A232FF822A877991FF424DC2" box="[159,228,1981,2006]" captionStart="FIGURE 5" captionStartId="9.[151,250,1988,2010]" captionTargetBox="[177,1414,193,1976]" captionTargetId="figure@9.[177,1414,193,1976]" captionTargetPageId="9" captionText="FIGURE 5. Ancistrus alga, MEPN 14853, 115.1 mm SL, male. From top to bottom, lateral, dorsal, and ventral view." httpUri="https://zenodo.org/record/2612135/files/figure.png" pageId="10" pageNumber="221">Fig. 5</figureCitation>
). Some cataloged lots have adult males with small or very small fleshy tentacles. Females possess a narrow naked anterior stripe on snout, as in females of the other species analyzed. On the naked strip, a single row of small, conical fleshy tentacles is present, from one side to the other side. The two central tentacles a bit longer, and occasionally, conical tentacles have bifid or branched tip (
<figureCitation id="BFAF2A26A233FF8329387EF3FCD64AEC" box="[800,880,223,248]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[204,1390,193,1949]" captionTargetId="figure@5.[204,1390,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Females of Ancistrus species, lateral view. From top to bottom: A. shuar, MEPN 17983, 93.9 mm SL. A. alga, MEPN 15169, 160.9 mm SL. A. clementinae, MEPN 5996, 100.1 mm SL. and A. malacops, MEPN 7804, 93.4 mm SL." httpUri="https://zenodo.org/record/2612129/files/figure.png" pageId="11" pageNumber="222">Figs. 2</figureCitation>
, 
<figureCitation id="BFAF2A26A233FF8329647ECCFC2A4AEC" box="[892,908,224,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,1606,1628]" captionTargetBox="[151,1436,589,1585]" captionTargetId="figure@7.[151,1436,589,1585]" captionTargetPageId="7" captionText="FIGURE 3. Ventral view of the head of females to show the presence (arrows) or absence of soft, fleshy tentacles on snout. A) A. clementinae, MEPN 5996, 100.1 mm SL. B) A. malacops, MEPN 7804, 93.4 mm SL. C) A. alga, MEPN 15169, 160.9 mm SL, D) A. shuar, MEPN 17983, 93.9 mm SL." httpUri="https://zenodo.org/record/2612131/files/figure.png" pageId="11" pageNumber="222">3</figureCitation>
).
</paragraph>
<paragraph id="272B36A3A233FF832ADF7F28FD264BB9" blockId="11.[151,1437,151,608]" pageId="11" pageNumber="222">
<emphasis id="15E0EAB1A233FF832ADF7F28FEB64B09" bold="true" box="[199,272,260,285]" pageId="11" pageNumber="222">Color.</emphasis>
Head, and dorsolateral body surface dark brown to black with very small dots, white or whitish, sometimes faded or difficult to observe. Belly slightly paler than sides of the body; with very small dots, white or whitish, generally more evident. All fins uniform black, with minute white dots spaced on interradial membranes. In juveniles, the dots on dorsal-fin membrane are relatively larger. Dorsal-fin with black spot between bases of spine and the first branched ray (
<figureCitation id="BFAF2A26A233FF83281E7FB8FDF34BB8" box="[518,597,404,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[204,1390,193,1949]" captionTargetId="figure@5.[204,1390,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Females of Ancistrus species, lateral view. From top to bottom: A. shuar, MEPN 17983, 93.9 mm SL. A. alga, MEPN 15169, 160.9 mm SL. A. clementinae, MEPN 5996, 100.1 mm SL. and A. malacops, MEPN 7804, 93.4 mm SL." httpUri="https://zenodo.org/record/2612129/files/figure.png" pageId="11" pageNumber="222">Figs. 2</figureCitation>
, 
<figureCitation id="BFAF2A26A233FF83287A7FB8FDD44BB9" box="[610,626,404,429]" captionStart="FIGURE 5" captionStartId="9.[151,250,1988,2010]" captionTargetBox="[177,1414,193,1976]" captionTargetId="figure@9.[177,1414,193,1976]" captionTargetPageId="9" captionText="FIGURE 5. Ancistrus alga, MEPN 14853, 115.1 mm SL, male. From top to bottom, lateral, dorsal, and ventral view." httpUri="https://zenodo.org/record/2612135/files/figure.png" pageId="11" pageNumber="222">5</figureCitation>
).
</paragraph>
<paragraph id="272B36A3A233FF832ADF7F9BFDB64874" blockId="11.[151,1437,151,608]" pageId="11" pageNumber="222">
<emphasis id="15E0EAB1A233FF832ADF7F9BFDAC4BC4" bold="true" box="[199,522,439,464]" pageId="11" pageNumber="222">Geographical distribution.</emphasis>
The species inhabits the northeastern part of the country, in 
<collectingRegion id="E550F841A233FF832EF37F9BFACE4BC4" box="[1259,1384,439,464]" country="Ecuador" name="Sucumbios" pageId="11" pageNumber="222">Sucumbíos</collectingRegion>
and 
<collectingRegion id="E550F841A233FF832A8F7FF0FF514BE1" box="[151,247,476,501]" country="Ecuador" name="Orellana" pageId="11" pageNumber="222">Orellana</collectingRegion>
provinces, in systems that drain to the 
<collectingRegion id="E550F841A233FF8328A67FF1FD5D4BE0" box="[702,763,477,500]" country="Ecuador" name="Napo" pageId="11" pageNumber="222">Napo</collectingRegion>
River. There is one sample (juvenile) that comes from the Curaray River, in 
<collectingRegion id="E550F841A233FF832B797C2CFD84480C" box="[353,546,511,536]" country="Ecuador" name="Pastaza" pageId="11" pageNumber="222">Pastaza province</collectingRegion>
; apparently this locality may be the southern limit. On the other hand, there are no samples from the 
<collectingRegion id="E550F841A233FF832B8B7C09FDE64829" box="[403,576,548,573]" country="Ecuador" name="Napo" pageId="11" pageNumber="222">Napo province</collectingRegion>
, these results seems to indicate that the species has a preference for low altitudes in the drainages (
<figureCitation id="BFAF2A26A233FF832BA57C6BFDA44874" box="[445,514,583,608]" captionStart="FIGURE 8" captionStartId="14.[151,250,1963,1985]" captionTargetBox="[151,1435,563,1937]" captionTargetId="figure@14.[151,1435,563,1937]" captionTargetPageId="14" captionText="FIGURE 8. Map of Ecuador showing the geographical distribution of the species. A. shuar, star. A. alga, square. A. clementinae, circle. A. malacops, triangle. Symbols may represent more than one lot." httpUri="https://zenodo.org/record/2612141/files/figure.png" pageId="11" pageNumber="222">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>