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<document id="67B6148F36FC5511F1309516FF6D7469" ID-CLB-Dataset="57636" ID-DOI="10.5852/ejt.2023.877.2151" ID-GBIF-Dataset="74111f75-19b1-4944-b6d0-09db245266fd" ID-ISSN="2118-9773" ID-Zenodo-Dep="8110586" ID-ZooBank="65E8C3F7-5EA4-4013-A09E-37C96B4929FB" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1688395635639" checkinUser="felipe" docAuthor="Haran, Julien M., Marvaldi, Adriana E., Benoit, Laure, Oberlander, Kenneth, Stals, Riaan &amp; Oberprieler, Rolf G." docDate="2023" docId="AF3A7125FFE4FF99FD896712FAD907A1" docLanguage="en" docName="ejt-877_1-89.pdf" docOrigin="European Journal of Taxonomy 877 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.877.2151" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cryptolarynx namaquanus Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler 2023, sp. nov." docType="treatment" docUuid="F889226B-4182-43BB-A918-3DE821DEC07E" docUuidSource="ZooBank" docVersion="12" lastPageNumber="30" masterDocId="5303095DFFFFFF84FF99651EFFAF016C" masterDocTitle="Revision of the enigmatic South African Cryptolaryngini (Coleoptera, Curculionidae), with description of a new genus and twenty-two new species" masterLastPageNumber="89" masterPageNumber="1" pageNumber="28" updateTime="1699533238137" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="67BCFB72B88E64CF203E75ACA037D61D">Revision of the enigmatic South African Cryptolaryngini (Coleoptera, Curculionidae), with description of a new genus and twenty-two new species</mods:title>
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<mods:namePart id="3D9ECEE05429A8767B3875B384BA15F9">Haran, Julien M.</mods:namePart>
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<mods:affiliation id="CE00F0D2124A993D7533191C3E2B30F4">División Entomología, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, Buenos Aires, Argentina.</mods:affiliation>
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<mods:affiliation id="4E55442A74A7019E5D6CCAAA5FC5073A">Schweickerdt Herbarium, Department of Plant and Soil Sciences, Plant Sciences Complex, University of Pretoria, Private Bag X 20, Hatfield 0024, South Africa. &amp; Kenneth.</mods:affiliation>
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<mods:affiliation id="2D56C5D2035CEF9B46933C31900529EC">Australian National Insect Collection, G. P. O. Box 1700, Canberra, A. C. T. 2601, Australia.</mods:affiliation>
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7.
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<emphasis id="15E71C21FFE4FF9FFDB76715FCDC034A" bold="true" box="[558,883,523,550]" italics="true" pageId="27" pageNumber="28">Cryptolarynx namaquanus</emphasis>
Haran
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<figureCitation id="BFA8DCB6FFE4FF9FFDF4674CFD620300" box="[621,717,594,620]" captionStart="Fig" captionStartId="9.[190,233,1845,1871]" captionTargetId="figure-13@10.[308,1280,265,2019]" captionTargetPageId="10" captionText="Fig. 1 (Part I) (next page). Habitus of males of species of Cryptolarynx Van Schalkwyk, 1966 (dorsal view). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113555" httpUri="https://zenodo.org/record/8113555/files/figure.png" pageId="27" pageNumber="28">Figs 1G</figureCitation>
,
<figureCitation id="BFA8DCB6FFE4FF9FFD40674CFD500300" box="[729,767,594,620]" captionStart="Fig" captionStartId="15.[189,232,1787,1813]" captionTargetId="figure-13@16.[201,1387,265,2019]" captionTargetPageId="16" captionText="Fig. 2 (Part I) (next page). Male genitalia of species of Cryptolarynx Van Schalkwyk, 1966. Penis in dorsal and lateral view (left), tegmen in dorsal view (middle) and details of apex of a parameroid lobe and spiculum gastrale in dorsal view (right). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113561" httpUri="https://zenodo.org/record/8113561/files/figure.png" pageId="27" pageNumber="28">2G</figureCitation>
,
<figureCitation id="BFA8DCB6FFE4FF9FFC92674CFC9E0300" box="[779,817,594,620]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetId="figure-13@20.[189,1398,300,1983]" captionTargetPageId="20" captionText="Fig. 3 (Part I) (next page). Habitus of males of species of Cryptolarynx Van Schalkwyk, 1966 (lateral view). A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113565" httpUri="https://zenodo.org/record/8113565/files/figure.png" pageId="27" pageNumber="28">3G</figureCitation>
,
<figureCitation id="BFA8DCB6FFE4FF9FFCA7674CFCCA0300" box="[830,869,594,620]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetId="figure-12@26.[226,1361,265,2019]" captionTargetPageId="26" captionText="Fig. 4 (Part I) (next page). Head of males of species of Cryptolarynx Van Schalkwyk, 1966. A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113569" httpUri="https://zenodo.org/record/8113569/files/figure.png" pageId="27" pageNumber="28">4G</figureCitation>
,
<figureCitation id="BFA8DCB6FFE4FF9FFCE9674CFC380301" box="[880,919,594,621]" captionStart="Fig" captionStartId="31.[189,232,1822,1848]" captionTargetBox="[191,1387,271,1758]" captionTargetId="figure-12@31.[188,1397,262,1759]" captionTargetPageId="31" captionText="Fig. 5 (Part I). Ventrites of males of species of Cryptolarynx Van Schalkwyk, 1966. A. C. vitis (Marshall, 1957). B. C. subglaber Haran sp. nov. C. C. squamulatus Haran sp. nov. D. C. muellerae Haran sp. nov. E. C. hirtulus Haran sp. nov. F. C. robustus Haran sp. nov. G. C. namaquanus Haran sp. nov. H. C. carinatus Haran sp. nov. I. C. variabilis Haran sp. nov. J. C. estriatus (Marshall, 1957), with details of a cuticular carina (white arrows). K. C. pyrophilus Haran sp. nov. L. C. pilipes Haran sp. nov. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113573" httpUri="https://zenodo.org/record/8113573/files/figure.png" pageId="27" pageNumber="28">5G</figureCitation>
,
<figureCitation id="BFA8DCB6FFE4FF9FFC3A674CFC680300" box="[931,967,594,620]" captionStart="Fig" captionStartId="45.[189,232,1582,1608]" captionTargetBox="[195,1393,274,1677]" captionTargetId="figure-13@46.[189,1398,265,1692]" captionTargetPageId="46" captionText="Fig. 8 (next page). Structural details and measurements of Cryptolaryngini Van Schalkwyk, 1966. A. Cryptolarynx spinicornis Haran sp. nov., cephalic view, arrows indicating width of forehead (top) and of epifrons (bottom). B. Head of C. pyrophilus Haran sp. nov., lateral view, arrows indicating distance between eye and scrobe. C. Head of C. pyrophilus Haran sp. nov., in cephalic view, arrows showing prothoracic scales near eyes (top), position of scape when antennae in repose (middle) and position of clubs near procoxae when in repose (bottom). D. C. namaquanus Haran sp. nov., dorsal view, arrows indicate length and width dimensions of pronotum and elytra as measured.E. Metafemora and metatarsus of C. variabilis Haran sp. nov. (left) and C. carinatus Haran sp. nov. (right), arrows indicating position of proximal inner carina in latter species (highlighted with a white line in C. variabilis Haran sp. nov.) and short apical mucro. F. Metatarsus of Hadrocryptolarynx major Haran gen. et sp. nov., arrow indicating elongate segment 2 (longer than wide). G. Hindleg of C. vitis (Marshall, 1957), arrow indicating apical mucro. H. Antenna of C. spinicornis Haran sp. nov., arrows pointing at inner tooth of funicle segments 2 and 4. I. Protibia of C. armatus Haran sp. nov., arrow pointing at inner preapical expansion proximally of mucro. J. Penis of C. oberprieleri Haran sp. nov., dorsal view, black arrows indicating length of penis body and temones and white arrows indicating width of penis body as measured; bottom arrow indicating copulatory sclerite. K. Ovipositor, Crypolarynx variabilis Haran sp. nov. L. Sternite VIII, same species. M. Spermatheca, same species. N. Ovipositor, Hadrocryptolarynx major Haran gen. et sp. nov. O. Sternite VIII, same species. P. Spermatheca, same species. Q. Right maxilla, same species. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.8113581" httpUri="https://zenodo.org/record/8113581/files/figure.png" pageId="27" pageNumber="28">8D</figureCitation>
</paragraph>
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<emphasis id="15E71C21FFE4FF9FFF24678DFE6603C1" bold="true" box="[189,457,659,685]" pageId="27" pageNumber="28">Differential diagnosis</emphasis>
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<taxonomicName id="E093BBB0FFE4FF9FFF2467DCFE5403B0" authority="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler, 2023" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[189,507,706,732]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="namaquanus" status="sp. nov.">
<emphasis id="15E71C21FFE4FF9FFF2467DCFE5403B0" box="[189,507,706,732]" italics="true" pageId="27" pageNumber="28">Cryptolarynx namaquanus</emphasis>
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<taxonomicNameLabel id="0ED4A15AFFE4FF9FFD9E67DDFDC803B1" box="[519,615,707,733]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from other species of
<taxonomicName id="E093BBB0FFE4FF9FFB1067DCFA8A03B0" authorityName="Van Schalkwyk" authorityYear="1966" box="[1161,1317,706,732]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="15E71C21FFE4FF9FFB1067DCFA8A03B0" box="[1161,1317,706,732]" italics="true" pageId="27" pageNumber="28">Cryptolarynx</emphasis>
</taxonomicName>
by its wide epifrons (subequal to length of scapes), its narrow forehead (narrower than width of eyes) and its proximally cylindrical metatibiae. Its broad parameroid lobes of the male genitalia, bearing only very short setae, and the shape of the spiculum are also unique among the species of the genus.
<taxonomicName id="E093BBB0FFE4FF9FFB436632FEFC0205" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="namaquanus">
<emphasis id="15E71C21FFE4FF9FFB436632FEFC0205" italics="true" pageId="27" pageNumber="28">Cryptolarynx namaquanus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="E093BBB0FFE4FF9FFDA16651FD630205" authority="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler, 2023" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[568,716,847,873]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="15E71C21FFE4FF9FFDA16651FD630205" box="[568,716,847,873]" italics="true" pageId="27" pageNumber="28">C. variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0ED4A15AFFE4FF9FFD4F6651FC9C0205" box="[726,819,847,873]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="E093BBB0FFE4FF9FFCF66651FBAD0205" authority="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler, 2023" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[879,1026,847,873]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="carinatus" status="sp. nov.">
<emphasis id="15E71C21FFE4FF9FFCF66651FBAD0205" box="[879,1026,847,873]" italics="true" pageId="27" pageNumber="28">C. carinatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0ED4A15AFFE4FF9FFB926651FBC90205" box="[1035,1126,847,873]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
, but in the male it can be easily distinguished from these by its proximally cylindrical metatibiae (bearing an inner carina in
<taxonomicName id="E093BBB0FFE4FF9FFF266688FEFC02C3" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[191,339,917,944]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="15E71C21FFE4FF9FFF266688FEFC02C3" box="[191,339,917,944]" italics="true" pageId="27" pageNumber="28">C. variabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="E093BBB0FFE4FF9FFE0A6688FD8802C3" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[403,551,918,944]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="15E71C21FFE4FF9FFE0A6688FD8802C3" box="[403,551,918,944]" italics="true" pageId="27" pageNumber="28">C. carinatus</emphasis>
</taxonomicName>
) and the conformations of its male genitalia. Uncorrected p-distances between
<taxonomicName id="E093BBB0FFE4FF9FFEBE66A7FE4F02BE" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[295,480,953,979]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="namaquanus">
<emphasis id="15E71C21FFE4FF9FFEBE66A7FE4F02BE" box="[295,480,953,979]" italics="true" pageId="27" pageNumber="28">C. namaquanus</emphasis>
</taxonomicName>
and these two species were found to span 12.815.1% for
<emphasis id="15E71C21FFE4FF9FFB0066A7FB6302BF" box="[1177,1228,953,979]" italics="true" pageId="27" pageNumber="28">COI</emphasis>
and 3.74.2% for
<emphasis id="15E71C21FFE4FF9FFF7E66C2FEB6029A" box="[231,281,988,1014]" italics="true" pageId="27" pageNumber="28">EF1</emphasis>
with
<taxonomicName id="E093BBB0FFE4FF9FFEC566C2FE41029A" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[348,494,988,1014]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="15E71C21FFE4FF9FFEC566C2FE41029A" box="[348,494,988,1014]" italics="true" pageId="27" pageNumber="28">C. variabilis</emphasis>
</taxonomicName>
and 9.310.4% for
<emphasis id="15E71C21FFE4FF9FFD4166C2FCA4029A" box="[728,779,988,1014]" italics="true" pageId="27" pageNumber="28">COI</emphasis>
and 3.4% for
<emphasis id="15E71C21FFE4FF9FFC2866C2FC4B029A" box="[945,996,988,1014]" italics="true" pageId="27" pageNumber="28">EF1</emphasis>
with
<taxonomicName id="E093BBB0FFE4FF9FFBBF66C2FB170299" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[1062,1208,988,1014]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="15E71C21FFE4FF9FFBBF66C2FB170299" box="[1062,1208,988,1014]" italics="true" pageId="27" pageNumber="28">C. carinatus</emphasis>
</taxonomicName>
(Supp. file 1).
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFE4FF9FFF246103FC8005C0" pageId="27" pageNumber="28" type="etymology">
<paragraph id="272CC033FFE4FF9FFF246103FEEB055B" blockId="27.[189,324,1053,1079]" box="[189,324,1053,1079]" pageId="27" pageNumber="28">
<heading id="7C64775FFFE4FF9FFF246103FEEB055B" bold="true" box="[189,324,1053,1079]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="15E71C21FFE4FF9FFF246103FEEB055B" bold="true" box="[189,324,1053,1079]" pageId="27" pageNumber="28">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFE4FF9FFF246155FC8005C0" blockId="27.[188,1399,1099,1197]" pageId="27" pageNumber="28">
The species name
<taxonomicName id="E093BBB0FFE4FF9FFE0B6152FD870509" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[402,552,1100,1125]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="namaquanus">
<emphasis id="15E71C21FFE4FF9FFE0B6152FD870509" box="[402,552,1100,1125]" italics="true" pageId="27" pageNumber="28">namaquanus</emphasis>
</taxonomicName>
refers to the area where this species was found, the Namaqualand region of the
<collectingRegion id="E5570ED1FFE4FF9FFE956171FD8305E5" box="[268,556,1135,1161]" country="South Africa" name="Northern Cape" pageId="27" pageNumber="28">Northern Cape province</collectingRegion>
and part of the traditional home of the Nama people (Namaqua). The specific epithet is an adjective in the masculine form.
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFE4FF9FFF2461CDFB77075D" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="272CC033FFE4FF9FFF2461CDFE050581" blockId="27.[189,426,1235,1261]" box="[189,426,1235,1261]" pageId="27" pageNumber="28">
<heading id="7C64775FFFE4FF9FFF2461CDFE050581" bold="true" box="[189,426,1235,1261]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="15E71C21FFE4FF9FFF2461CDFE050581" bold="true" box="[189,426,1235,1261]" pageId="27" pageNumber="28">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFE4FF9FFF73601CFCA904C5" blockId="27.[189,1399,1282,1449]" pageId="27" pageNumber="28">
<materialsCitation id="97FBCA6EFFE4FF9FFF73601CFCA904C5" ID-GBIF-Occurrence="4145879419" collectingDate="2019-08-29" collectionCode="SAMC" collectorName="J. Haran" country="South Africa" county="Kamieskroon" latitude="-30.17" location="Namaqua National Park" longLatPrecision="73" longitude="17.793" pageId="27" pageNumber="28" specimenCode="JHAR02535_0101" specimenCount="1" specimenCount-male="1" stateProvince="Northern Cape Province" typeStatus="holotype">
<emphasis id="15E71C21FFE4FF9FFF73601CFEF30470" bold="true" box="[234,348,1282,1308]" pageId="27" pageNumber="28">
<typeStatus id="F8287E91FFE4FF9FFF73601CFEF30470" box="[234,348,1282,1308]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
REPUBLIC OF SOUTH AFRICA •
<emphasis id="15E71C21FFE4FF9FFDF16038FDD1042E" bold="true" box="[616,638,1318,1346]" pageId="27" pageNumber="28">
<specimenCount id="31950BBAFFE4FF9FFDF16038FDD1042E" box="[616,638,1318,1346]" pageId="27" pageNumber="28" type="male"></specimenCount>
</emphasis>
; “
<collectingCountry id="5F8480A3FFE4FF9FFD0E603BFB9D0453" box="[663,1074,1317,1343]" name="South Africa" pageId="27" pageNumber="28">REPUBLIC OF SOUTH AFRICA</collectingCountry>
.
<emphasis id="15E71C21FFE4FF9FFBA5603BFAC00453" bold="true" box="[1084,1391,1317,1343]" pageId="27" pageNumber="28">
<collectingRegion id="E5570ED1FFE4FF9FFBA5603BFAC00453" box="[1084,1391,1317,1343]" country="South Africa" name="Northern Cape" pageId="27" pageNumber="28">Northern Cape Province</collectingRegion>
</emphasis>
,
<collectingCounty id="CE4DB8BFFFE4FF9FFF246056FEF4040E" box="[189,347,1352,1378]" pageId="27" pageNumber="28">Kamieskroon</collectingCounty>
[
<locationDeviation id="369ED911FFE4FF9FFEF36056FE41040F" bearing="315.0" box="[362,494,1352,1379]" distance="15000.0" pageId="27" pageNumber="28">
<quantity id="E06B6DD6FFE4FF9FFEF36056FE1A040E" box="[362,437,1352,1379]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="28" unit="km" value="15.0">15 km</quantity>
NW
</locationDeviation>
,
<location id="224C96E8FFE4FF9FFE606057FCB9040E" LSID="urn:lsid:plazi:treatment:AF3A7125FFE4FF99FD896712FAD907A1:224C96E8FFE4FF9FFE606057FCB9040E" box="[505,790,1352,1379]" country="South Africa" county="Kamieskroon" latitude="-30.17" longLatPrecision="73" longitude="17.793" name="Namaqua National Park" pageId="27" pageNumber="28" stateProvince="Northern Cape Province">Namaqua National Park</location>
,
<location id="224C96E8FFE4FF9FFCB86056FB87040E" LSID="urn:lsid:plazi:treatment:AF3A7125FFE4FF99FD896712FAD907A1:224C96E8FFE4FF9FFCB86056FB87040E" box="[801,1064,1352,1378]" country="South Africa" county="Kamieskroon" latitude="-30.17" longLatPrecision="73" longitude="17.793" municipality="Namaqua National Park" name="Skilpad Flower Camp" pageId="27" pageNumber="28" stateProvince="Northern Cape Province">Skilpad Flower Camp</location>
].
<date id="532DE6F3FFE4FF9FFBA36056FB64040E" box="[1082,1227,1352,1378]" pageId="27" pageNumber="28" value="2019-08-29">
<collectingDate id="43691F1BFFE4FF9FFBA36056FB64040E" box="[1082,1227,1352,1378]" pageId="27" pageNumber="28" value="2019-08-29">29.viii.2019</collectingDate>
</date>
.
<collectorName id="8A66A5E5FFE4FF9FFB4F6057FA97040F" box="[1238,1336,1353,1379]" pageId="27" pageNumber="28">J. Haran</collectorName>
leg.” “
<geoCoordinate id="42A7A6F4FFE4FF9FFF5E6072FEED04EA" box="[199,322,1388,1414]" degrees="30.170" direction="south" orientation="latitude" pageId="27" pageNumber="28" precision="55" value="-30.17">30.170° S</geoCoordinate>
<geoCoordinate id="42A7A6F4FFE4FF9FFED66072FE6604EA" box="[335,457,1388,1414]" degrees="17.793" direction="east" orientation="longitude" pageId="27" pageNumber="28" precision="55" value="17.793">17.793° E</geoCoordinate>
at base of
<taxonomicName id="E093BBB0FFE4FF9FFDC56075FCAC04E9" box="[604,771,1387,1413]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="obtusa">
<emphasis id="15E71C21FFE4FF9FFDC56075FCAC04E9" box="[604,771,1387,1413]" italics="true" pageId="27" pageNumber="28">Oxalis obtusa</emphasis>
</taxonomicName>
.
<specimenCode id="77356848FFE4FF9FFC8F6075FC5904EA" box="[790,1014,1387,1414]" pageId="27" pageNumber="28">JHAR02535_0101</specimenCode>
. Cirad-CBGP coll.” “Holotype.
<taxonomicName id="E093BBB0FFE4FF9FFF246091FE5804C5" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[189,503,1423,1449]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="namaquanus">
<emphasis id="15E71C21FFE4FF9FFF246091FE5804C5" box="[189,503,1423,1449]" italics="true" pageId="27" pageNumber="28">Cryptolarynx namaquanus</emphasis>
</taxonomicName>
. Haran 2023”;
<collectionCode id="418258F6FFE4FF9FFD356091FCAC04C5" box="[684,771,1423,1449]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="27" pageNumber="28" type="Museum">SAMC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="272CC033FFE4FF9FFF7360CEFB77075D" blockId="27.[189,1398,1488,1585]" pageId="27" pageNumber="28">
<materialsCitation id="97FBCA6EFFE4FF9FFF7360CEFEBF075D" ID-GBIF-Occurrence="4145879346" collectingDate="2019-08-29" collectionCode="SAMC" collectorName="J. Haran" country="South Africa" county="Kamieskroon" latitude="-30.17" location="Northern Cape" longLatPrecision="73" longitude="17.793" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Northern Cape" typeStatus="paratype">
<emphasis id="15E71C21FFE4FF9FFF7360CEFEC80486" bold="true" box="[234,359,1488,1514]" pageId="27" pageNumber="28">
<typeStatus id="F8287E91FFE4FF9FFF7360CEFEC80486" box="[234,359,1488,1514]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="5F8480A3FFE4FF9FFF2460EAFDFC0762" box="[189,595,1524,1550]" name="South Africa" pageId="27" pageNumber="28">REPUBLIC OF SOUTH AFRICA</collectingCountry>
<emphasis id="15E71C21FFE4FF9FFDEB60EAFC9E0761" bold="true" box="[626,817,1523,1550]" pageId="27" pageNumber="28">
<collectingRegion id="E5570ED1FFE4FF9FFDEB60EAFC9E0761" box="[626,817,1523,1550]" country="South Africa" name="Northern Cape" pageId="27" pageNumber="28">Northern Cape</collectingRegion>
</emphasis>
<specimenCount id="31950BBAFFE4FF9FFCD260EAFCD60762" box="[843,889,1524,1550]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="31950BBAFFE4FF9FFC1E60EAFC1B0762" box="[903,948,1524,1550]" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="418258F6FFE4FF9FFF246309FEBF075D" box="[189,272,1559,1585]" collectionName="South Africa, Cape Town, Iziko Museum of Capetown (formerly South African Museum)" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="27" pageNumber="28" type="Museum">SAMC</collectionCode>
</materialsCitation>
<materialsCitation id="97FBCA6EFFE4FF9FFEB06309FB77075D" ID-GBIF-Occurrence="4145879343" box="[297,1240,1559,1585]" collectingDate="2019-08-29" collectionCode="CBGP" collectorName="J. Haran" country="South Africa" county="Kamieskroon" latitude="-30.17" location="Namaqua National Park" longLatPrecision="73" longitude="17.793" pageId="27" pageNumber="28" specimenCount="10" specimenCount-male="1" stateProvince="Northern Cape Province" typeStatus="paratype">
<specimenCount id="31950BBAFFE4FF9FFEB06309FEF8075D" box="[297,343,1559,1585]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="31950BBAFFE4FF9FFEFD6309FEDC075D" box="[356,371,1559,1585]" pageId="27" pageNumber="28">9</specimenCount>
specs (preserved in ethanol); same collection data as for holotype;
<collectionCode id="418258F6FFE4FF9FFB116309FB7E075D" box="[1160,1233,1559,1585]" pageId="27" pageNumber="28">CBGP</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFE4FF99FF246346FE5704A1" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="272CC033FFE4FF9FFF246346FED1071E" blockId="27.[189,382,1624,1650]" box="[189,382,1624,1650]" pageId="27" pageNumber="28">
<emphasis id="15E71C21FFE4FF9FFF246346FEE1071E" bold="true" box="[189,334,1624,1650]" pageId="27" pageNumber="28">Description</emphasis>
(
<specimenCount id="31950BBAFFE4FF9FFEC66346FED9071E" box="[351,374,1624,1650]" pageId="27" pageNumber="28" type="male"></specimenCount>
)
</paragraph>
<paragraph id="272CC033FFE4FF9FFF246399FDD207CD" blockId="27.[189,637,1670,1697]" box="[189,637,1670,1697]" pageId="27" pageNumber="28">
<smallCapsWord id="21CA56EFFFE4FF9FFF246399FEC107F3" baselines="1690,1690" box="[189,366,1671,1697]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="27" pageNumber="28">MEASUREMENTS</smallCapsWord>
. Body length
<quantity id="E06B6DD6FFE4FF9FFD8C6399FDD607CD" box="[533,633,1671,1697]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
.
</paragraph>
<paragraph id="272CC033FFE2FF99FF246417FD7800BC" blockId="29.[187,1400,265,465]" pageId="29" pageNumber="30">
<smallCapsWord id="21CA56EFFFE2FF99FF246417FEB7004D" baselines="284,284" box="[189,280,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="29" pageNumber="30">COLOUR</smallCapsWord>
<smallCapsWord id="21CA56EFFFE2FF99FE876411FEE4004E" baselines="285" box="[286,331,271,290]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="29" pageNumber="30">AND</smallCapsWord>
<smallCapsWord id="21CA56EFFFE2FF99FEC86410FE6D004D" baselines="284" box="[337,450,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="vestiture" pageId="29" pageNumber="30">VESTITURE</smallCapsWord>
. Body integument black, antennae and tarsi reddish. Dorsal vestiture (pronotum + elytra) consisting of overlapping, recumbent, parallel-sided clothing scales, 2 × as long as wide, truncate at apex; colour of scales varying from evenly greyish or creamy-white to brown or black; pale scales concentrated in two longitudinal stripes on pronotum and on elytral interstriae 4 as well as forming a pair of whitish spots surrounded by black scales at apical ⅔ of elytra; scales on interstriae recumbent, in lateral view not distinct from rest of vestiture.
</paragraph>
<paragraph id="272CC033FFE2FF99FF2464E6FD150268" blockId="29.[188,1401,503,772]" pageId="29" pageNumber="30">
<smallCapsWord id="21CA56EFFFE2FF99FF2464E6FF53037C" baselines="523,523" box="[189,252,504,530]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="29" pageNumber="30">HEAD</smallCapsWord>
. Forehead as wide as epifrons near antennal insertions, slightly narrower than width of an eye, scales recumbent. Eyes convex, in dorsal view exceeding outline of head, surrounded by a ring of short pale scales, on forehead directed towards occiput; distance between eye and scrobe as large as or slightly smaller than width of antennal club. Epifrons wide, distance between antennal insertions as large as length of scape, scales at most 2× as long as wide, recumbent, subcontiguous. Frons with pairs of erect lateral setae. Epistome without median seta. Antennal funicles with segments 12 elongate, subequal, about 3× as long as wide; 34 as long as wide, compressed, slightly angular on inside; 57 globular, isodiametric; 7 sometimes wider than long.
</paragraph>
<paragraph id="272CC033FFE2FF99FF246635FE8B020B" blockId="29.[187,1398,811,871]" pageId="29" pageNumber="30">
<smallCapsWord id="21CA56EFFFE2FF99FF246635FE99022F" baselines="830,830" box="[189,310,811,837]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Pronotum" pageId="29" pageNumber="30">PRONOTUM</smallCapsWord>
. Transverse (W:L ratio 1.11.2), widest near midlength, sides arcuate; apex and base subequal in width.
</paragraph>
<paragraph id="272CC033FFE2FF99FF246691FB6702C4" blockId="29.[189,1224,910,937]" box="[189,1224,910,937]" pageId="29" pageNumber="30">
<smallCapsWord id="21CA56EFFFE2FF99FF246691FEBD02CB" baselines="930,930" box="[189,274,911,937]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Elytra" pageId="29" pageNumber="30">ELYTRA</smallCapsWord>
. Broadly ovate, isodiametric (W:L ratio 1), sides convex, widest near midlength.
</paragraph>
<paragraph id="272CC033FFE2FF99FF2466CEFAF80560" blockId="29.[188,1399,975,1037]" pageId="29" pageNumber="30">
<smallCapsWord id="21CA56EFFFE2FF99FF2466CEFF590284" baselines="995,995" box="[189,246,976,1002]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="29" pageNumber="30">LEGS</smallCapsWord>
. Protibiae with outer margin straight, inner margin slightly bisinuate; metatibiae with apical mucro and inner setal fringe, setae shorter than segment 5 of metatarsus. Tarsi with segment 2 isodiametric.
</paragraph>
<paragraph id="272CC033FFE2FF99FF24612DFB150503" blockId="29.[188,1399,1075,1136]" pageId="29" pageNumber="30">
<smallCapsWord id="21CA56EFFFE2FF99FF24612DFE830527" baselines="1094,1094" box="[189,300,1075,1101]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="29" pageNumber="30">ABDOMEN</smallCapsWord>
. Ventrites 1 and 5 slightly concave medially; ventrites 14 with creamy-white plumose scales, almost concealing integument, intermixed with long suberect setae, each apically bifid.
</paragraph>
<paragraph id="272CC033FFE2FF99FF246189FAB80457" blockId="29.[187,1399,1174,1340]" pageId="29" pageNumber="30">
<smallCapsWord id="21CA56EFFFE2FF99FF246189FEEC05C3" baselines="1194,1194" box="[189,323,1175,1201]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Terminalia" pageId="29" pageNumber="30">TERMINALIA</smallCapsWord>
. Body of penis moderately elongate (W:L ratio 0.5), 2× as short as temones, sides subparallel in basal half, converging in distal half; curvature in profile weak and regular, dorsoventrally slightly thickened at apex. Copulatory sclerite weakly sclerotised or not discerned in examined specimens. Parameroid lobes separate, divided by very deep median notch, each lobe broad, rounded at apex, bearing a series of very short setae. Spiculum gastrale with basal arms short and feebly curved.
</paragraph>
<paragraph id="272CC033FFE2FF99FF24607CFE1F0410" blockId="29.[189,432,1378,1404]" box="[189,432,1378,1404]" pageId="29" pageNumber="30">
<heading id="7C64775FFFE2FF99FF24607CFE1F0410" bold="true" box="[189,432,1378,1404]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="15E71C21FFE2FF99FF24607CFE1F0410" bold="true" box="[189,432,1378,1404]" pageId="29" pageNumber="30">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFE2FF99FF24608EFE5704A1" blockId="29.[189,1398,1424,1485]" pageId="29" pageNumber="30">Males can be distinguished from females by the shape of ventrites 1 and 5 (concave in male, flat or slightly convex in female).</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFE2FF99FF2460EDFEB50732" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="272CC033FFE2FF99FF2460EDFEE00761" blockId="29.[189,335,1523,1549]" box="[189,335,1523,1549]" pageId="29" pageNumber="30">
<heading id="7C64775FFFE2FF99FF2460EDFEE00761" bold="true" box="[189,335,1523,1549]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="15E71C21FFE2FF99FF2460EDFEE00761" bold="true" box="[189,335,1523,1549]" pageId="29" pageNumber="30">Life history</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFE2FF99FF24633FFEB50732" blockId="29.[189,1398,1569,1630]" pageId="29" pageNumber="30">
Adults of
<taxonomicName id="E093BBB0FFE2FF99FEAB633CFE440757" authority="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler, 2023" authorityName="Haran &amp; Marvaldi &amp; Benoit &amp; Oberlander &amp; Stals &amp; Oberprieler" authorityYear="2023" box="[306,491,1570,1596]" class="Insecta" family="Curculionidae" genus="Cryptolarynx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="namaquanus" status="sp. nov.">
<emphasis id="15E71C21FFE2FF99FEAB633CFE440757" box="[306,491,1570,1596]" italics="true" pageId="29" pageNumber="30">C. namaquanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0ED4A15AFFE2FF99FE6B633CFDE30750" box="[498,588,1570,1596]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
were found in monospecific stands of
<taxonomicName id="E093BBB0FFE2FF99FB8B633FFB1B0757" box="[1042,1204,1569,1595]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="obtusa">
<emphasis id="15E71C21FFE2FF99FB8B633FFB1B0757" box="[1042,1204,1569,1595]" italics="true" pageId="29" pageNumber="30">Oxalis obtusa</emphasis>
</taxonomicName>
, in the month of August.
</paragraph>
</subSubSection>
<subSubSection id="6F8993B8FFE2FF99FF24639BFAD907A1" box="[189,1398,1669,1742]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="272CC033FFE2FF99FF24639BFEF907F3" blockId="29.[189,342,1669,1695]" box="[189,342,1669,1695]" pageId="29" pageNumber="30">
<heading id="7C64775FFFE2FF99FF24639BFEF907F3" bold="true" box="[189,342,1669,1695]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="15E71C21FFE2FF99FF24639BFEF907F3" bold="true" box="[189,342,1669,1695]" pageId="29" pageNumber="30">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="272CC033FFE2FF99FF2463ADFAD907A1" blockId="29.[189,1398,1715,1742]" box="[189,1398,1715,1742]" pageId="29" pageNumber="30">
All specimens collected were found at the
<typeStatus id="F8287E91FFE2FF99FD0F63AAFD6707A2" box="[662,712,1716,1742]" pageId="29" pageNumber="30">type</typeStatus>
locality, the only one thus far known for the species (
<figureCitation id="BFA8DCB6FFE2FF99FA8F63ADFAC707A1" box="[1302,1384,1715,1741]" captionStart="Fig" captionStartId="77.[189,232,1963,1989]" captionTargetBox="[366,1201,265,1898]" captionTargetId="figure-11@77.[361,1226,265,1922]" captionTargetPageId="77" captionText="Fig. 13. Distribution maps of species of Cryptolarynx Van Schalkwyk, 1966 and Hadrocryptolarynx Haran gen. nov." figureDoi="http://doi.org/10.5281/zenodo.8110596" httpUri="https://zenodo.org/record/8110596/files/figure.png" pageId="29" pageNumber="30">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>