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<document id="513CFEFC64BBB8E692C52C41F22ED62E" ID-DOI="http://dx.doi.org/10.3897/zookeys.715.15913" ID-GBIF-Dataset="894fe765-e2af-4695-9f90-fcf45b0c4a2c" ID-PMC="PMC5740430" ID-Pensoft-Pub="1313-2970-715-17" ID-PubMed="29302230" ID-ZooBank="3587B83BD0D4494393CA09B16BBF231B" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-715-17" ModsDocOrigin="ZooKeys 715" ModsDocTitle="Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E.aipo-group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1510239869137" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sumoked, Bob &amp; Balke, Michael" docDate="2017" docId="AF01991AB054EE2520043A9494B9B319" docLanguage="en" docName="ZooKeys 715: 17-37" docOrigin="ZooKeys 715" docSource="http://dx.doi.org/10.3897/zookeys.715.15913" docTitle="Exocelina talaki Balke 1998" docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="FD28FFA11471A342006BFF903075FF81" masterDocTitle="Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E. aipo-group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="37" masterPageNumber="17" pageNumber="31" updateTime="1701317016017" updateUser="plazi">
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<mods:title id="FC248CC67AF98BE559B7C8B3388761C4">Descriptions of two new species and one new subspecies from the Exocelinaokbapensis-group, and notes on the E. aipo-group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="1EA2F8CA5EB475FD8D3F3A3D5E8ECB51">Shaverdo, Helena</mods:namePart>
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<mods:namePart id="2169BDD74453B0EC348B237519D679E5">Sumoked, Bob</mods:namePart>
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<mods:namePart id="CF2FF0FF539175E3E6356E03E09B9B38">Balke, Michael</mods:namePart>
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<mods:date id="7827F730010194E3DEA40EB9CCBEB0BC">2017</mods:date>
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<treatment id="AF01991AB054EE2520043A9494B9B319" ID-GBIF-Taxon="137255798" LSID="urn:lsid:plazi:treatment:AF01991AB054EE2520043A9494B9B319" httpUri="http://treatment.plazi.org/id/AF01991AB054EE2520043A9494B9B319" lastPageId="15" lastPageNumber="32" pageId="14" pageNumber="31">
<subSubSection id="0C285FAB9F7EBBCA604AC85E4DFBADD2" pageId="14" pageNumber="31" type="nomenclature">
<paragraph id="905E26D8DE2DB86E059A321BD84C3CC2" pageId="14" pageNumber="31">
4.
<taxonomicName id="6FEF2F0DA6720A49330651E934069A42" ID-CoL="3DLRL" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina talaki" order="Coleoptera" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="talaki">Exocelina talaki (Balke, 1998)</taxonomicName>
Figs 9, 11
</paragraph>
</subSubSection>
<subSubSection id="909DE2C42DC6B6CCEB281019901A56D9" pageId="14" pageNumber="31" type="reference_group">
<paragraph id="58E901CB1CBDC0A8954D6C55C0B789A5" pageId="14" pageNumber="31">
<taxonomicName id="FAC21581694AC0A734FDDDA45CE4687E" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) talaki" order="Coleoptera" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="talaki" subGenus="Papuadytes">Copelatus (Papuadytes) talaki</taxonomicName>
Balke, 1998: 337;
<bibRefCitation id="5FA7511FBE7BEA0520FB14CB25E16413" author="Nilsson, AN" journalOrPublisher="Stenstrup, Apollo Books" pageId="17" pageNumber="34" title="Dytiscidae. World catalogue of insects. Vol. 3." year="2001">Nilsson 2001</bibRefCitation>
: 77 (catalogue).
</paragraph>
<paragraph id="3F99EB66E31AAF6BB5350AB662819F87" pageId="14" pageNumber="31">
<taxonomicName id="21D1D29B9C98727E80B32130CD758E83" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes talaki" order="Coleoptera" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="talaki">Papuadytes talaki</taxonomicName>
(Balke, 1998):
<bibRefCitation id="F4BBF99471241DF791A592EE411280C0" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="17" pageNumber="34" pagination="55 - 74" title="World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
: 56 (comb.n.).
</paragraph>
<paragraph id="F4E4D27A0F9B4F57A22B809A098D7282" pageId="14" pageNumber="31">
<taxonomicName id="C64B3E4CF567971C792FE3A7AEF98D6C" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina talaki" order="Coleoptera" pageId="14" pageNumber="31" phylum="Arthropoda" rank="species" species="talaki">Exocelina talaki</taxonomicName>
(Balke, 1998):
<bibRefCitation id="8A65722D9E697899A6002ADE56D4D02E" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="17" pageNumber="34" pagination="33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
: 34 (comb.n.).
</paragraph>
</subSubSection>
<subSubSection id="5649F3DB3B1201A495DE5CF3D25FCD22" pageId="14" pageNumber="31" type="type locality">
<paragraph id="23B487EE3ACD6785BFBD1B4C0944BBF8" pageId="14" pageNumber="31">Type locality.</paragraph>
<paragraph id="45A8ECCFB6481FA919611F254CDB5D67" pageId="14" pageNumber="31">
Papua: Pegunungan Bintang Regency, Borme, ca.
<geoCoordinate id="F72C798D3CCD3C5FF29A4D2862384502" direction="south" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
;
<geoCoordinate id="D63620D517A704A6201F5B804556D0D3" direction="east" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
, 1200 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="FD98822B3A4ADEA745F573421665F653" pageId="14" pageNumber="31" type="type material">
<paragraph id="72D28CB6CEDF1BE375371CDA3A73FB01" pageId="14" pageNumber="31">Type material studied.</paragraph>
<paragraph id="C7B11C40CE0940E2B552B09F4333929B" pageId="14" pageNumber="31">
Holotype: male &quot;IRIAN JAYA: Borme ca.
<geoCoordinate id="6D3F4F95268622AAF75C480B0C7B008E" direction="east" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
,
<geoCoordinate id="39B00FEA152E1EF4D4A45C937FFC1B67" direction="south" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
1200 m, 2.9.1993 leg. M. Balke (1)&quot;,
<normalizedToken id="BB25466A3600C3B3C7463458412BE005" originalValue="“HOLOTYPUS”">&quot;HOLOTYPUS&quot;</normalizedToken>
[red], &quot;Copelatus talaki Balke des. 1997&quot; [red] (NHMW). Paratypes: Three females with the same label as the holotype and additionally with red labels &quot;Paratypus Copelatus talaki Balke des. 1997&quot;, one of them with two additional labels &quot;M. Balke 3294&quot; [green] and &quot;M. Balke 6415&quot; [green text] (NHMW).
</paragraph>
</subSubSection>
<subSubSection id="7DB357307250C2A3C68B56BCA374767B" lastPageId="15" lastPageNumber="32" pageId="14" pageNumber="31" type="diagnosis">
<paragraph id="A0B198B84CD91D1BCB822377D2CA86D9" pageId="14" pageNumber="31">Diagnosis.</paragraph>
<paragraph id="8842C13912207321FA128AD0A4E7BBE3" lastPageId="15" lastPageNumber="32" pageId="14" pageNumber="31">
Beetle small (TL-H 3.25-3.4 mm); oblong-oval; reddish brown to brown, with paler head and pronotum, with fine punctation and weakly impressed microreticulation, shiny; pronotum with distinct lateral bead; male antennae simple (Fig. 9); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male
<pageBreakToken id="624CC289C3601ECBA385D52B5537CC9D" pageId="15" pageNumber="32" start="start">protarsomere</pageBreakToken>
5 long and narrow, without concavity, with anterior row of 14 and posterior row of seven relatively short setae; median lobe strongly curved, sharply pointed in lateral view, evenly tapering, with broadly pointed apex in ventral view; paramere with notch on its dorsal side and small, less evidently separated subdistal part with a tuft of dense setae; proximal setae inconspicuous (Fig. 11
<normalizedToken id="5CDB27A9D9A9551451A1A0FF1103FB1D" originalValue="AD">A-D</normalizedToken>
). For complete description, see
<bibRefCitation id="C29D8601A805CA6FD2A5225309783B9E" pageId="15" pageNumber="32">Balke (1998)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D59517C834C1DEACA101707D1C166276" pageId="15" pageNumber="32" type="distribution">
<paragraph id="C35986DD1872BC62BB54BA89F2D9537C" pageId="15" pageNumber="32">Distribution.</paragraph>
<paragraph id="90A20ECD8BD6A79F01F962781EE81719" pageId="15" pageNumber="32">Papua: Pegunungan Bintang Regency (Fig. 12). The species is known only from the type material.</paragraph>
<caption id="0A1B601E411D887E72745D0BC31D0980" pageId="15" pageNumber="32">
<paragraph id="87571C729046E704C713D3A47DB65DDF" pageId="15" pageNumber="32">
Figure 12. Map of New Guinea showing distribution of the species of both groups. The white square with a question mark indicates the male from Aipomek-Tanime area, which most likely belongs to
<taxonomicName id="9255DB59BD58C6DF799B3150524F0C04" lsidName="E. manfredi" pageId="15" pageNumber="32" rank="species" species="manfredi">E. manfredi</taxonomicName>
(Balke, 1998).
</paragraph>
</caption>
</subSubSection>
</treatment>
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