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<document id="A091A1F7C540A2D94C5ED8409BDE6928" ID-CLB-Dataset="147248" ID-DOI="10.11646/zootaxa.5188.5.1" ID-GBIF-Dataset="125c40a8-e8a6-4751-9f63-4a81ac32a805" ID-ISSN="1175-5326" ID-Zenodo-Dep="7099200" ID-ZooBank="46E32E3E-1E31-48D0-99EC-B123789B566E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663742511325" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="AF0087D7FFDEFFE4FF48F9BD1672FE24" docLanguage="en" docName="zootaxa.5188.5.1.pdf" docOrigin="Zootaxa 5188 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geodromicus suensoni Bernhauer 1936" docType="treatment" docVersion="7" lastPageNumber="425" masterDocId="5339FFAFFFC8FFFCFFDFFFD9135BFFAD" masterDocTitle="The plagiatus species group of the genus Geodromicus Redtenbacher, 1857 (Insecta Coleoptera: Staphylinidae: Omaliinae), and additional data on some species from Japan" masterLastPageNumber="429" masterPageNumber="401" pageNumber="423" updateTime="1699382853362" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="121A8CFE9400D86EC379174AB0A48889">The plagiatus species group of the genus Geodromicus Redtenbacher, 1857 (Insecta Coleoptera: Staphylinidae: Omaliinae), and additional data on some species from Japan</mods:title>
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<mods:namePart id="80A6C39F1D548EC05EF3D0D6A4920A59">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="D64A98F1D628B849EBE88D3D00E868E5">2022</mods:date>
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<treatment id="AF0087D7FFDEFFE4FF48F9BD1672FE24" ID-DOI="http://doi.org/10.5281/zenodo.7101136" ID-GBIF-Taxon="200858428" ID-Zenodo-Dep="7101136" LSID="urn:lsid:plazi:treatment:AF0087D7FFDEFFE4FF48F9BD1672FE24" httpUri="http://treatment.plazi.org/id/AF0087D7FFDEFFE4FF48F9BD1672FE24" lastPageId="24" lastPageNumber="425" pageId="22" pageNumber="423">
<subSubSection id="6FB3654AFFDEFFEAFF48F9BD11DAF9D3" box="[151,641,1636,1663]" pageId="22" pageNumber="423" type="nomenclature">
<paragraph id="271636C1FFDEFFEAFF48F9BD11DAF9D3" blockId="22.[151,641,1636,1699]" box="[151,641,1636,1663]" pageId="22" pageNumber="423">
<heading id="7C5E81ADFFDEFFEAFF48F9BD11DAF9D3" bold="true" box="[151,641,1636,1663]" fontSize="11" level="1" pageId="22" pageNumber="423" reason="1">
<taxonomicName id="E0A94D42FFDEFFEAFF48F9BD11DAF9D3" ID-CoL="3FPK9" authority="Bernhauer, 1936" authorityName="Bernhauer" authorityYear="1936" box="[151,641,1636,1663]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="suensoni">
<emphasis id="15DDEAD3FFDEFFEAFF48F9BD11DAF9D3" bold="true" box="[151,641,1636,1663]" pageId="22" pageNumber="423">
<emphasis id="15DDEAD3FFDEFFEAFF48F9BD12F7F9D3" bold="true" box="[151,428,1636,1663]" italics="true" pageId="22" pageNumber="423">Geodromicus suensoni</emphasis>
<bibRefCitation id="43384B30FFDEFFEAFE6CF9BC11DAF9D3" author="Bernhauer, M." box="[435,641,1636,1663]" pageId="22" pageNumber="423" pagination="303 - 325" refId="ref24316" refString="Bernhauer, M. (1936) Neuheiten der palearktischen Staphylinidenfauna II. Pubblicazioni del Museo Entomologico &quot; Pietro Rossi &quot; Duino, 1, 303 - 325." type="journal article" year="1936">Bernhauer, 1936</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="6FB3654AFFDEFFEAFF48F9511203F90E" box="[151,344,1672,1699]" pageId="22" pageNumber="423" type="description">
<paragraph id="271636C1FFDEFFEAFF48F9511203F90E" blockId="22.[151,641,1636,1699]" box="[151,344,1672,1699]" pageId="22" pageNumber="423">
(
<figureCitation id="BF922A44FFDEFFEAFF40F9511273F90E" box="[159,296,1672,1699]" captionStart="FIGURES 41–48" captionStartId="23.[151,264,1902,1926]" captionTargetBox="[167,1425,367,1870]" captionTargetId="figure-84@23.[151,1436,345,1878]" captionTargetPageId="23" captionText="FIGURES 41–48. Aedeagus of Japanese species of Geodromicus (Figs. 41, 43, 45, 47: parameral view, Figs. 42, 44, 46, 48: lateral view): 41–42–G. aokii (Mt. Daisetsuzan), 43–44–G. sprius (holotype), 45–46–G. sinuatus (Kamikochi), 47–48–G. suensoni (Simasima). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7099220" httpUri="https://zenodo.org/record/7099220/files/figure.png" pageId="22" pageNumber="423">Figs. 47–48</figureCitation>
,
<figureCitation id="BF922A44FFDEFFEAFEECF951120BF90E" box="[307,336,1672,1699]" captionStart="FIGURES 49–50" captionStartId="24.[151,264,1321,1345]" captionTargetBox="[151,1436,417,1298]" captionTargetId="figure-113@24.[151,1436,417,1298]" captionTargetPageId="24" captionText="FIGURES 49–50. Distribution of Geodromicus sinuatus (Fig. 49) and G. suensoni (Fig. 50) in Japan. Black circles: specimens studied by author; white circles: previously published records, specimens not seen by author." figureDoi="http://doi.org/10.5281/zenodo.7099224" httpUri="https://zenodo.org/record/7099224/files/figure.png" pageId="22" pageNumber="423">50</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6FB3654AFFDEFFEAFF48F91411D2F88F" pageId="22" pageNumber="423" type="reference_group">
<paragraph id="271636C1FFDEFFEAFF48F9141104F948" blockId="22.[151,717,1741,1826]" box="[151,607,1741,1765]" pageId="22" pageNumber="423">
<treatmentCitationGroup id="07B911EFFFDEFFEAFF48F9141104F948" box="[151,607,1741,1765]" pageId="22" pageNumber="423">
<treatmentCitation id="A60810D0FFDEFFEAFF48F9141104F948" author="Bernhauer, M." box="[151,607,1741,1765]" page="304" pageId="22" pageNumber="423" year="1936">
<taxonomicName id="E0A94D42FFDEFFEAFF48F9141104F948" ID-CoL="3FPK9" authority="Bernhauer, 1936: 304" authorityName="Bernhauer" authorityPageNumber="304" authorityYear="1936" box="[151,607,1741,1765]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="suensoni">
<emphasis id="15DDEAD3FFDEFFEAFF48F9141225F948" box="[151,382,1741,1765]" italics="true" pageId="22" pageNumber="423">Geodromicus suensoni</emphasis>
<bibRefCitation id="43384B30FFDEFFEAFE5BF9141104F948" author="Bernhauer, M." box="[388,607,1741,1765]" pageId="22" pageNumber="423" pagination="303 - 325" refId="ref24316" refString="Bernhauer, M. (1936) Neuheiten der palearktischen Staphylinidenfauna II. Pubblicazioni del Museo Entomologico &quot; Pietro Rossi &quot; Duino, 1, 303 - 325." type="journal article" year="1936">Bernhauer, 1936: 304</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="271636C1FFDEFFEAFF48F9351196F8A9" blockId="22.[151,717,1741,1826]" box="[151,717,1771,1796]" pageId="22" pageNumber="423">
<treatmentCitationGroup id="07B911EFFFDEFFEAFF48F9351196F8A9" box="[151,717,1771,1796]" pageId="22" pageNumber="423">
<taxonomicName id="E0A94D42FFDEFFEAFF48F9351196F8A9" authority=": Watanabe, 1985: 267" authorityName="Watanabe" authorityPageNumber="267" authorityYear="1985" box="[151,717,1771,1796]" class="Insecta" family="Staphylinidae" genus="Psephidonus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="suensoni">
<treatmentCitation id="A60810D0FFDEFFEAFF48F9351106F8AE" author="Watanabe, Y." box="[151,605,1771,1796]" page="267" pageId="22" pageNumber="423" year="1985">
<emphasis id="15DDEAD3FFDEFFEAFF48F9351223F8AE" box="[151,376,1772,1796]" italics="true" pageId="22" pageNumber="423">Psephidonus suensoni</emphasis>
:
<bibRefCitation id="43384B30FFDEFFEAFE5AF9351106F8AE" author="Watanabe, Y." box="[389,605,1771,1796]" pageId="22" pageNumber="423" pagination="264 - 268" refId="ref28592" refString="Watanabe, Y. (1985) Omaliinae. In: Ueno, S. - L., Kurosawa, Y. &amp; Sato, M. (Eds.), The Coleoptera of Japan in Color, 2, pp. 264 - 268, pls. 46 - 47. [Hoikusha, Osaka, in Japanese]" type="journal article" year="1985">Watanabe, 1985: 267</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="A60810D0FFDEFFEAFDB8F9351196F8A9" author="Watanabe, Y." box="[615,717,1772,1796]" page="293" pageId="22" pageNumber="423" year="1990">
<bibRefCitation id="43384B30FFDEFFEAFDB8F9351196F8A9" author="Watanabe, Y." box="[615,717,1772,1796]" pageId="22" pageNumber="423" pagination="59 - 391" refId="ref28714" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">1990: 293</bibRefCitation>
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="271636C1FFDEFFEAFF48F8D311D2F88F" blockId="22.[151,717,1741,1826]" box="[151,649,1802,1826]" pageId="22" pageNumber="423">
<treatmentCitationGroup id="07B911EFFFDEFFEAFF48F8D311D2F88F" box="[151,649,1802,1826]" pageId="22" pageNumber="423">
<treatmentCitation id="A60810D0FFDEFFEAFF48F8D31166F88F" author="Hayashi, Y." box="[151,573,1802,1826]" page="201" pageId="22" pageNumber="423" year="2018">
<taxonomicName id="E0A94D42FFDEFFEAFF48F8D31166F88F" ID-CoL="3FPNV" authority="Hayashi, 2018: 201" authorityName="Hayashi" authorityPageNumber="201" authorityYear="2018" box="[151,573,1802,1826]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="takabai">
<emphasis id="15DDEAD3FFDEFFEAFF48F8D3122AF88F" box="[151,369,1802,1826]" italics="true" pageId="22" pageNumber="423">Geodromicus takabai</emphasis>
<bibRefCitation id="43384B30FFDEFFEAFEA8F8D31166F88F" author="Hayashi, Y." box="[375,573,1802,1826]" pageId="22" pageNumber="423" pagination="201 - 205" refId="ref25687" refString="Hayashi, Y. (2018) Two new species of the genus Geodromicus Redtenbacher (Coleoptera, Staphylinidae, Omalinae [sic] from Japan). Elytra, Tokyo, New Series, 8 (1), 201 - 205." type="journal article" year="2018">Hayashi, 2018: 201</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<emphasis id="15DDEAD3FFDEFFEAFD9CF8D311D2F88F" bold="true" box="[579,649,1802,1826]" pageId="22" pageNumber="423">
<taxonomicNameLabel id="0EEE57A8FFDEFFEAFD9CF8D311D2F88F" box="[579,649,1802,1826]" pageId="22" pageNumber="423">syn. n.</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6FB3654AFFDEFFE4FF48F8881151FF78" lastPageId="24" lastPageNumber="425" pageId="22" pageNumber="423" type="materials_examined">
<paragraph id="271636C1FFDEFFEBFF48F8881045FEEC" blockId="22.[151,1437,1872,2043]" lastBlockId="23.[151,1437,151,321]" lastPageId="23" lastPageNumber="424" pageId="22" pageNumber="423">
<emphasis id="15DDEAD3FFDEFFEAFF48F8881068F8C7" bold="true" box="[151,819,1872,1899]" pageId="22" pageNumber="423">
Type material examined:
<collectingCountry id="5FBE7651FFDEFFEAFE19F889117FF8C7" box="[454,548,1872,1898]" name="Japan" pageId="22" pageNumber="423">JAPAN</collectingCountry>
:
<collectingRegion id="E56DF823FFDEFFEAFDEFF8891074F8C7" box="[560,815,1872,1898]" country="Japan" name="Nagano" pageId="22" pageNumber="423">NAGANO PREFECTURE</collectingRegion>
:
</emphasis>
<typeStatus id="F8128863FFDEFFEAFCE6F88910FAF8C7" box="[825,929,1872,1898]" pageId="22" pageNumber="423" type="holotype">Holotype</typeStatus>
[Left antennomeres 4–11 glued on same card with beetle]
<specimenCount id="31AFFD48FFDEFFEAFEF9F8AC1266F822" box="[294,317,1909,1935]" pageId="22" pageNumber="423" type="female">♀</specimenCount>
: ‘Kamikochi | North[ern]. Alps Japan | 5000 f. Suenson |
<date id="53171001FFDEFFEAFC0EF8AD170EF822" box="[977,1109,1908,1935]" pageId="22" pageNumber="423" value="1934-09-15">15. 9. 1934</date>
’ &lt;handwritten&gt;, ‘Kamikochi 5000 Feet | Northern Alps Japan | E. Suenson |
<date id="53171001FFDEFFEAFD14F8411037F81E" box="[715,876,1944,1971]" pageId="22" pageNumber="423" value="1934-09-15">15 SEP. 1934</date>
’ &lt;printed in blue&gt;, ‘
<taxonomicName id="E0A94D42FFDEFFEAFB84F84117A1F81F" authority="Shp." authorityName="Sharp" authorityYear="1889" box="[1115,1274,1944,1970]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="423" phylum="Arthropoda" rank="species" species="sinuatus">sinuatus Shp.</taxonomicName>
[handwritten] | det. Bernhau[er]’ &lt;printed&gt;, ‘Hs. schmäler | fein p[un]ktiert’ &lt;pink, handwritten&gt;, ‘Suensoni | Brh. Typus | unic.’ &lt;handwritten&gt;, ‘Suensoni | Brnh. Typ. m. | Geodromicus’ &lt;light brown, handwritten&gt;, ‘Chicago NHMus | M.Bernhauer | Collection’ &lt;printed&gt;, ‘
<typeStatus id="F8128863FFDFFFEBFDE1FF4E118CFF1C" box="[574,727,151,177]" pageId="23" pageNumber="424" type="holotype">HOLOTYPE</typeStatus>
| teste D.J.Clarke2014 | GDI Imaging Project’ &lt;violet, printed&gt;, ‘PHOTOGRAPHED | Kelsey Keaton 2014 | Emu Catalog’ &lt;blue, printed&gt;. ‘FMNHINS | 2819543 | FIELD MUSEUM’ &lt;rectangular label, with 2-D barcode, printed&gt;, ‘
<taxonomicName id="E0A94D42FFDFFFEBFC8AFF0610B0FF54" authorityName="Redtenbacher" authorityYear="1857" box="[853,1003,223,249]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="15DDEAD3FFDFFFEBFC8AFF0610B0FF54" box="[853,1003,223,249]" italics="true" pageId="23" pageNumber="424">Geodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="E0A94D42FFDFFFEBFBDBFF061677FF54" authority="Bernhauer, 1936" authorityName="Bernhauer" authorityYear="1936" box="[1028,1324,223,249]" class="Insecta" family="Staphylinidae" genus="Psephidonus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="suensoni">
<emphasis id="15DDEAD3FFDFFFEBFBDBFF06173DFF55" box="[1028,1126,223,248]" italics="true" pageId="23" pageNumber="424">suensoni</emphasis>
<bibRefCitation id="43384B30FFDFFFEBFBAFFF061677FF54" author="Bernhauer, M." box="[1136,1324,223,249]" pageId="23" pageNumber="424" pagination="303 - 325" refId="ref24316" refString="Bernhauer, M. (1936) Neuheiten der palearktischen Staphylinidenfauna II. Pubblicazioni del Museo Entomologico &quot; Pietro Rossi &quot; Duino, 1, 303 - 325." type="journal article" year="1936">Bernhauer, 1936</bibRefCitation>
</taxonomicName>
| Shavrin A.V. det. 2016’ &lt;printed&gt; (
<collectionCode id="41B8AE04FFDFFFEBFE0CFEDA1171FEB0" box="[467,554,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="23" pageNumber="424" type="Museum">FMNH</collectionCode>
). The photographs of the habitus and type labels are available in Arthropod Collections Database of FMNH (last access:
<date id="53171001FFDFFFEBFD50FEFE1049FEEC" box="[655,786,295,321]" pageId="23" pageNumber="424" value="2022-08-23">23.08.2022</date>
).
</paragraph>
<caption id="73D66649FFDFFFEBFF48F8B7111AF863" ID-DOI="http://doi.org/10.5281/zenodo.7099220" ID-Zenodo-Dep="7099220" httpUri="https://zenodo.org/record/7099220/files/figure.png" pageId="23" pageNumber="424" startId="23.[151,264,1902,1926]" targetBox="[167,1425,367,1870]" targetPageId="23">
<paragraph id="271636C1FFDFFFEBFF48F8B7111AF863" blockId="23.[151,1436,1902,1998]" pageId="23" pageNumber="424">
<emphasis id="15DDEAD3FFDFFFEBFF48F8B71203F82B" bold="true" box="[151,344,1902,1926]" pageId="23" pageNumber="424">FIGURES 41–48.</emphasis>
Aedeagus of Japanese species of
<taxonomicName id="E0A94D42FFDFFFEBFD1DF8B71011F82B" authorityName="Redtenbacher" authorityYear="1857" box="[706,842,1902,1926]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="genus">
<emphasis id="15DDEAD3FFDFFFEBFD1DF8B71011F82B" box="[706,842,1902,1926]" italics="true" pageId="23" pageNumber="424">Geodromicus</emphasis>
</taxonomicName>
(Figs. 41, 43, 45, 47: parameral view, Figs. 42, 44, 46, 48: lateral view): 41–42–
<taxonomicName id="E0A94D42FFDFFFEBFE45F84B11F6F807" authority="(Mt. Daisetsuzan)" baseAuthorityName="Mt. Daisetsuzan" box="[410,685,1938,1962]" class="Insecta" family="Staphylinidae" genus="Psephidonus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="aokii">
<emphasis id="15DDEAD3FFDFFFEBFE45F84B12F7F804" box="[410,428,1938,1961]" italics="true" pageId="23" pageNumber="424">G</emphasis>
.
<emphasis id="15DDEAD3FFDFFFEBFE65F84B12B6F807" box="[442,493,1938,1962]" italics="true" pageId="23" pageNumber="424">aokii</emphasis>
(Mt. Daisetsuzan)
</taxonomicName>
, 43–44–
<taxonomicName id="E0A94D42FFDFFFEBFCDBF84B1039F807" authorityName="Watanabe" authorityYear="1990" box="[772,866,1938,1962]" class="Insecta" family="Staphylinidae" genus="Psephidonus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="sprius">
<emphasis id="15DDEAD3FFDFFFEBFCDBF84B104DF804" box="[772,790,1938,1961]" italics="true" pageId="23" pageNumber="424">G</emphasis>
.
<emphasis id="15DDEAD3FFDFFFEBFCFBF84A1039F807" box="[804,866,1939,1962]" italics="true" pageId="23" pageNumber="424">sprius</emphasis>
</taxonomicName>
(holotype), 45–46–
<taxonomicName id="E0A94D42FFDFFFEBFBF2F84B1676F807" authority="(Kamikochi)" baseAuthorityName="Kamikochi" box="[1069,1325,1938,1962]" class="Insecta" family="Staphylinidae" genus="Anthophagus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="15DDEAD3FFDFFFEBFBF2F84B1764F804" box="[1069,1087,1938,1961]" italics="true" pageId="23" pageNumber="424">G</emphasis>
.
<emphasis id="15DDEAD3FFDFFFEBFB92F84A17FAF807" box="[1101,1185,1939,1962]" italics="true" pageId="23" pageNumber="424">sinuatus</emphasis>
(Kamikochi)
</taxonomicName>
, 47–48–
<taxonomicName id="E0A94D42FFDFFFEBFA5BF84B1235F863" authority="(Simasima)" baseAuthorityName="Simasima" class="Insecta" family="Staphylinidae" genus="Psephidonus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="424" phylum="Arthropoda" rank="species" species="suensoni">
<emphasis id="15DDEAD3FFDFFFEBFA5BF84B16CDF804" box="[1412,1430,1938,1961]" italics="true" pageId="23" pageNumber="424">G</emphasis>
.
<emphasis id="15DDEAD3FFDFFFEBFF48F86E13ABF863" box="[151,240,1975,1998]" italics="true" pageId="23" pageNumber="424">suensoni</emphasis>
(Simasima)
</taxonomicName>
. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="271636C1FFD0FFE4FF18FF4F1151FF78" blockId="24.[151,1437,150,393]" pageId="24" pageNumber="425">
<emphasis id="15DDEAD3FFD0FFE4FF18FF4F116FFF1D" bold="true" box="[199,564,150,176]" pageId="24" pageNumber="425">Additional material examined.</emphasis>
<materialsCitation id="97C13C9CFFD0FFE4FDE5FF4F115DFF78" ID-GBIF-Occurrence="3909820306" collectingDate="1966-07-10" collectorName="Y. Watanabe" country="Japan" location="NAGANO PREFECTURE" pageId="24" pageNumber="425" specimenCount="1" specimenCount-male="1" stateProvince="Nagano">
<emphasis id="15DDEAD3FFD0FFE4FDE5FF4F10F3FF1D" bold="true" box="[570,936,150,176]" pageId="24" pageNumber="425">
<collectingCountry id="5FBE7651FFD0FFE4FDE5FF4F11C3FF1D" box="[570,664,150,176]" name="Japan" pageId="24" pageNumber="425">JAPAN</collectingCountry>
:
<collectingRegion id="E56DF823FFD0FFE4FD7BFF4F10FFFF1D" box="[676,932,150,176]" country="Japan" name="Nagano" pageId="24" pageNumber="425">NAGANO PREFECTURE</collectingRegion>
:
</emphasis>
<specimenCount id="31AFFD48FFD0FFE4FC70FF4E1087FF1C" box="[943,988,151,177]" pageId="24" pageNumber="425" type="male">1 ♂</specimenCount>
: ‘Simasima-val. Nagano-pref., Jap.
<date id="53171001FFD0FFE4FAB1FF4E13A2FF78" pageId="24" pageNumber="425" value="1966-07-10">
<collectingDate id="4353E9E9FFD0FFE4FAB1FF4E13A2FF78" pageId="24" pageNumber="425" value="1966-07-10">July 10, 1966</collectingDate>
</date>
<collectorName id="8A5C5317FFD0FFE4FF20FF6212D0FF78" box="[255,395,187,213]" pageId="24" pageNumber="425">Y. Watanabe</collectorName>
leg.’ (cW)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6FB3654AFFD0FFE4FF18FF0717CBFF54" box="[199,1168,222,249]" pageId="24" pageNumber="425" type="distribution">
<paragraph id="271636C1FFD0FFE4FF18FF0717CBFF54" blockId="24.[151,1437,150,393]" box="[199,1168,222,249]" pageId="24" pageNumber="425">
<emphasis id="15DDEAD3FFD0FFE4FF18FF07123AFF55" bold="true" box="[199,353,222,248]" pageId="24" pageNumber="425">Distribution.</emphasis>
The species is known from several localities in Honshu,
<collectingCountry id="5FBE7651FFD0FFE4FC3FFF06177BFF54" box="[992,1056,223,249]" name="Japan" pageId="24" pageNumber="425">Japan</collectingCountry>
(
<figureCitation id="BF922A44FFD0FFE4FBEFFF0617DFFF54" box="[1072,1156,222,249]" captionStart="FIGURES 49–50" captionStartId="24.[151,264,1321,1345]" captionTargetBox="[151,1436,417,1298]" captionTargetId="figure-113@24.[151,1436,417,1298]" captionTargetPageId="24" captionText="FIGURES 49–50. Distribution of Geodromicus sinuatus (Fig. 49) and G. suensoni (Fig. 50) in Japan. Black circles: specimens studied by author; white circles: previously published records, specimens not seen by author." figureDoi="http://doi.org/10.5281/zenodo.7099224" httpUri="https://zenodo.org/record/7099224/files/figure.png" pageId="24" pageNumber="425">Fig. 50</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6FB3654AFFD0FFE4FF18FEDA1672FE24" pageId="24" pageNumber="425" type="discussion">
<paragraph id="271636C1FFD0FFE4FF18FEDA1672FE24" blockId="24.[151,1437,150,393]" pageId="24" pageNumber="425">
<emphasis id="15DDEAD3FFD0FFE4FF18FEDA1260FEB0" bold="true" box="[199,315,259,285]" pageId="24" pageNumber="425">Remarks.</emphasis>
<taxonomicName id="E0A94D42FFD0FFE4FE99FEDA1113FEB1" authorityName="Bernhauer" authorityYear="1936" box="[326,584,259,285]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="suensoni">
<emphasis id="15DDEAD3FFD0FFE4FE99FEDA1113FEB1" box="[326,584,259,285]" italics="true" pageId="24" pageNumber="425">Geodromicus suensoni</emphasis>
</taxonomicName>
was originally described from Kamikochi, central Honshu.
<taxonomicName id="E0A94D42FFD0FFE4FAD9FEDA12FDFEEC" authority="Hayashi, 2018" authorityName="Hayashi" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="takabai">
<emphasis id="15DDEAD3FFD0FFE4FAD9FEDA13B0FEEC" italics="true" pageId="24" pageNumber="425">Geodromicus takabai</emphasis>
<bibRefCitation id="43384B30FFD0FFE4FF24FEFE12FDFEEC" author="Hayashi, Y." box="[251,422,295,321]" pageId="24" pageNumber="425" pagination="201 - 205" refId="ref25687" refString="Hayashi, Y. (2018) Two new species of the genus Geodromicus Redtenbacher (Coleoptera, Staphylinidae, Omalinae [sic] from Japan). Elytra, Tokyo, New Series, 8 (1), 201 - 205." type="journal article" year="2018">Hayashi, 2018</bibRefCitation>
</taxonomicName>
was originally described from
<collectingRegion id="E56DF823FFD0FFE4FCEFFEFE1740FEEC" box="[816,1051,295,321]" country="Japan" name="Ishikawa" pageId="24" pageNumber="425">Ishikawa Prefecture</collectingRegion>
, central Honshu. Based on the morphological features provided in the original description and the shape of the aedeagus (
<bibRefCitation id="43384B30FFD0FFE4FB59FE92167DFEC8" author="Hayashi, Y." box="[1158,1318,331,357]" pageId="24" pageNumber="425" pagination="201 - 205" refId="ref25687" refString="Hayashi, Y. (2018) Two new species of the genus Geodromicus Redtenbacher (Coleoptera, Staphylinidae, Omalinae [sic] from Japan). Elytra, Tokyo, New Series, 8 (1), 201 - 205." type="journal article" year="2018">Hayashi 2018</bibRefCitation>
: fig. 3),
<taxonomicName id="E0A94D42FFD0FFE4FA5EFE9213B0FE24" authorityName="Hayashi" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="takabai">
<emphasis id="15DDEAD3FFD0FFE4FA5EFE9216CEFEC8" box="[1409,1429,331,357]" italics="true" pageId="24" pageNumber="425">G</emphasis>
.
<emphasis id="15DDEAD3FFD0FFE4FF48FEB613B0FE24" box="[151,235,367,393]" italics="true" pageId="24" pageNumber="425">takabai</emphasis>
</taxonomicName>
is conspecific with
<taxonomicName id="E0A94D42FFD0FFE4FE14FEB61114FE25" authorityName="Bernhauer" authorityYear="1936" box="[459,591,367,393]" class="Insecta" family="Staphylinidae" genus="Geodromicus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="425" phylum="Arthropoda" rank="species" species="suensoni">
<emphasis id="15DDEAD3FFD0FFE4FE14FEB61284FE24" box="[459,479,367,393]" italics="true" pageId="24" pageNumber="425">G</emphasis>
.
<emphasis id="15DDEAD3FFD0FFE4FE32FEB61114FE25" box="[493,591,367,392]" italics="true" pageId="24" pageNumber="425">suensoni</emphasis>
</taxonomicName>
, with which it is synonymized here. Aedeagus as in
<figureCitation id="BF922A44FFD0FFE4FB42FEB6167EFE24" box="[1181,1317,367,393]" captionStart="FIGURES 41–48" captionStartId="23.[151,264,1902,1926]" captionTargetBox="[167,1425,367,1870]" captionTargetId="figure-84@23.[151,1436,345,1878]" captionTargetPageId="23" captionText="FIGURES 41–48. Aedeagus of Japanese species of Geodromicus (Figs. 41, 43, 45, 47: parameral view, Figs. 42, 44, 46, 48: lateral view): 41–42–G. aokii (Mt. Daisetsuzan), 43–44–G. sprius (holotype), 45–46–G. sinuatus (Kamikochi), 47–48–G. suensoni (Simasima). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7099220" httpUri="https://zenodo.org/record/7099220/files/figure.png" pageId="24" pageNumber="425">Figs. 47–48</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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