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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.476.8630" ID-PMC="PMC4311692" ID-Pensoft-Pub="1313-2970-476-1" ID-Pensoft-UUID="FC4DC947FF97FF86190BFFD8B82CAB56" ID-PubMed="25685002" ID-Zenodo-Dep="578702" ID-ZooBank="086D71AF8A294F028559C2E0456B5C5B" ModsDocID="1313-2970-476-1" checkinTime="1451244761026" checkinUser="pensoft" docAuthor="Gustafson, Grey T. &amp; Miller, Kelly B." docDate="2015" docId="AEE8990990BE38480CDFB3B9FA8D45AE" docLanguage="en" docName="ZooKeys 476: 1-135" docOrigin="ZooKeys 476" docPubDate="2015-01-23" docSource="http://dx.doi.org/10.3897/zookeys.476.8630" docTitle="Dineutus productus Roberts 1895" docType="treatment" docVersion="5" id="FC4DC947FF97FF86190BFFD8B82CAB56" lastPageNumber="62" masterDocId="FC4DC947FF97FF86190BFFD8B82CAB56" masterDocTitle="The New World whirligig beetles of the genus Dineutus Macleay, 1825 (Coleoptera, Gyrinidae, Gyrininae, Dineutini)" masterLastPageNumber="135" masterPageNumber="1" pageNumber="60" updateTime="1668159852297" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The New World whirligig beetles of the genus Dineutus Macleay, 1825 (Coleoptera, Gyrinidae, Gyrininae, Dineutini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gustafson, Grey T.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131, USA</mods:affiliation>
<mods:nameIdentifier type="email">gtgustafson@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Kelly B.</mods:namePart>
<mods:affiliation>Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-01-23</mods:number>
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<mods:detail type="volume">
<mods:number>476</mods:number>
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<mods:start>1</mods:start>
<mods:end>135</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.476.8630</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.476.8630</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-476-1</mods:identifier>
<mods:identifier type="ZooBank">086D71AF8A294F028559C2E0456B5C5B</mods:identifier>
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<mods:identifier type="Zenodo-Dep">578702</mods:identifier>
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<treatment ID-GBIF-Taxon="152057920" LSID="urn:lsid:plazi:treatment:AEE8990990BE38480CDFB3B9FA8D45AE" httpUri="http://treatment.plazi.org/id/AEE8990990BE38480CDFB3B9FA8D45AE" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60">
<subSubSection pageId="59" pageNumber="60" type="nomenclature">
<paragraph pageId="59" pageNumber="60">
<taxonomicName LSID="AEE89909-90BE-3848-0CDF-B3B9FA8D45AE" authority="Roberts, 1895" authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">Dineutus productus Roberts, 1895</taxonomicName>
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Dineutus productus. A ♀ dorsal habitus B ♀ ventral habitus C ♂ dorsal habitus. D ♂ ventral habitus All scale bars ≈ 2 mm." pageId="59" pageNumber="60">Figures 33</figureCitation>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dineutus productus. A aedeagus dorsal view B aedeagus ventral view C ♂ mesotarsal claws D aedeagus lateral view. Scale bar for C ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">, 34</figureCitation>
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. General distribution map. A Dineutus productus B Dineutus solitarius C Dineutus sublineatus D Dineutus mexicanus, Dineutus truncatus." pageId="59" pageNumber="60">, 54</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="reference_group">
<paragraph pageId="59" pageNumber="60">
<taxonomicName class="Insecta" family="Gyrinidae" genus="Dineutes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dineutes productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">Dineutes productus</taxonomicName>
<bibRefCitation author="Roberts, CH" journalOrPublisher="Transactions of the American Entomological Society" pageId="95" pageNumber="96" pagination="279 - 288" refId="B107" refString="Roberts, CH, 1895. The species of Dineutes of America north of Mexico. Transactions of the American Entomological Society 22: 279 - 288" title="The species of Dineutes of America north of Mexico." volume="22" year="1895">Roberts 1895</bibRefCitation>
: 282,
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus (Cyclinus) productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus" subGenus="Cyclinus">Dineutus (Cyclinus) productus</taxonomicName>
:
<bibRefCitation author="Ochs, G" journalOrPublisher="Entomologische Zeitschrift" pageId="94" pageNumber="95" pagination="129 - 140" refId="B89" refString="Ochs, G, 1926a. Die Dineutini. 2. Tribus der Unterfamilien Enhydrinae Fam. Gyrinidae (Col). A. Aligemeiner Teil. Entomologische Zeitschrift 40: 129 - 140" title="Die Dineutini. 2. Tribus der Unterfamilien Enhydrinae Fam. Gyrinidae (Col). A. Aligemeiner Teil." volume="40" year="1926 a">Ochs 1926a</bibRefCitation>
: 137.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="type locality">
<paragraph pageId="59" pageNumber="60">Type locality.</paragraph>
<paragraph pageId="59" pageNumber="60">USA, Texas.</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="specimens examined">
<paragraph pageId="59" pageNumber="60">Specimens examined.</paragraph>
<paragraph pageId="59" pageNumber="60">17</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="type material examined">
<paragraph pageId="59" pageNumber="60">Type material examined.</paragraph>
<paragraph pageId="59" pageNumber="60">
Syntype (♂ pinned) &quot;Tex./Acc. 4858/Type productus, ♂type # 2, C.H.R./
<taxonomicName class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">Dineutus</taxonomicName>
productus Roberts 1895 Det: L. Cook 2005&quot; AMNH type Catologue 496.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="material examined">
<paragraph pageId="59" pageNumber="60">Material examined.</paragraph>
<paragraph pageId="59" pageNumber="60">
<emphasis bold="true" pageId="59" pageNumber="60">U.S.A.:</emphasis>
<emphasis bold="true" pageId="59" pageNumber="60">Kansas:</emphasis>
Bourbon Co., Ft. Scott, ca. 2 mi NE, Marmaton River, &quot;T25S R25Esec 20 NW 1/4&quot;, 4.viii.1976, leg. S. Hamilton, T. Oldham, SEMC 10549560 (1 ex. KSEM); Elk Co., Elk River at Elk Falls,
<geoCoordinate degrees="37.374416" direction="north" orientation="latitude" precision="1" value="37.374416">37.374416</geoCoordinate>
<geoCoordinate degrees="96.184123" direction="west" orientation="longitude" precision="1" value="-96.18412">-96.184123</geoCoordinate>
, 278m, 22.vi.2014, leg. C. Maier, C. Faris, S. Baca, G.Gustafson, GTG062214B (5 ex. GTGC), Elk River S. of Longton,
<geoCoordinate degrees="37.36960" direction="north" orientation="latitude" precision="1" value="37.3696">37.36960</geoCoordinate>
<geoCoordinate degrees="96.078735" direction="west" orientation="longitude" precision="1" value="-96.078735">-96.078735</geoCoordinate>
, 262m, 22.vi.2014, leg. C. Maier, C. Faris, S. Baca, G.Gustafson, GTG062214A (6 ex. GTGC); Greenwood Co., Lapland, 4mi S Fall River, east branch, 13.ix.1974, leg. D. Huggins, SEMC 1054959 (1 ex. KSEM); Lyon Co., Lyon Co.-Greenwood Co. line, Verdigiris River, &quot;T21S R10E sec 36 SE 1/4&quot;, 16.vii.1976, leg. S. Hamilton, SEMC 1054955, SEMC 1054957 (2 ex. KSEM); Morris Co., Council Grove Lake, outflows and groves, 13.vi.1974, leg. T. Edmonds, SEMC 1054958 (1 ex. KSEM);
<emphasis bold="true" pageId="59" pageNumber="60">Texas:</emphasis>
<normalizedToken originalValue="“Sequin”">&quot;Sequin&quot;</normalizedToken>
, 26.vi.1938, leg. D.W. Craik (2 ex. KSEM).
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="60" type="diagnosis">
<paragraph pageId="59" pageNumber="60">Diagnosis.</paragraph>
<paragraph pageId="59" pageNumber="60">
Male (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Dineutus productus. A ♀ dorsal habitus B ♀ ventral habitus C ♂ dorsal habitus. D ♂ ventral habitus All scale bars ≈ 2 mm." pageId="59" pageNumber="60">33C, D</figureCitation>
): Size: 9.5-9.6 mm. Body form narrowly elongate oval; elytral apices rounded with sutural angle produced to a point, apicolateral sinuation weakly present, irregularities present apically near sutural production, elytral apices strongly deflexed, reticulation of pronotal and elytral discs strongly impressed, producing a bronzy appearance; elytral striae faintly evident, most evident on the subapicomedial portion of the elytral disc; profemora with a very small sub-apicoventral tooth, accompanied by a series of denticles extending proximad, associated with each posterior setigerous puncture of the profemora; protibiae subsinuate; mesotarsal claws (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dineutus productus. A aedeagus dorsal view B aedeagus ventral view C ♂ mesotarsal claws D aedeagus lateral view. Scale bar for C ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">34C</figureCitation>
) long and of similar size, with ventral margin straight; venter darkly colored, usually black to very dark bronzy-brown, with the tibiae and tarsi of mesothoracic and metathoracic legs usually lighter in coloration, as well as apex of abdomen; Aedeagus (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dineutus productus. A aedeagus dorsal view B aedeagus ventral view C ♂ mesotarsal claws D aedeagus lateral view. Scale bar for C ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">34A, B, D</figureCitation>
) median lobe in dorsal view noticeably shorter than parameres, weakly constricted just apicad to middle, acuminate in apical ca. 1/5, apex of acumination flatly rounded, in lateral view median lobe roundly curved dorsally, in dorsal view parameres sinuate laterally after basal 1/3, in apical 1/3 laterally expanded, apically strongly rounded.
</paragraph>
<paragraph pageId="59" pageNumber="60">
Female (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Dineutus productus. A ♀ dorsal habitus B ♀ ventral habitus C ♂ dorsal habitus. D ♂ ventral habitus All scale bars ≈ 2 mm." pageId="59" pageNumber="60">33A, B</figureCitation>
): Size: 9.5-9.9 mm. Body form narrowly elongate oval; elytral apices angled towards very strong sutural production, apicolateral sinuation strongly present and with plica, irregularities present apically near sutural production, elytral apices strongly deflexed, reticulation of pronotal and elytral discs strongly impressed, producing a bronzy appearance; elytral striae faintly evident, most evident on the subapicomedial portion of the elytral disc; profemora without sub-apicoventral tooth; protibiae club shaped, with lateral margin round, weakly expanded apicaly; venter darkly colored, usually black to very dark bronzy-brown, with the tibiae and tarsi of mesothoracic and metathoracic legs usually lighter in coloration, as well as apex of abdomen.
</paragraph>
<caption pageId="59" pageNumber="60" start="Figure 33" startId="F33">
<paragraph pageId="59" pageNumber="60">
<emphasis bold="true" pageId="59" pageNumber="60">Figure 33.</emphasis>
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="59" pageNumber="60">A</emphasis>
♀ dorsal habitus
<emphasis bold="true" pageId="59" pageNumber="60">B</emphasis>
♀ ventral habitus
<emphasis bold="true" pageId="59" pageNumber="60">C</emphasis>
♂ dorsal habitus.
<emphasis bold="true" pageId="59" pageNumber="60">D</emphasis>
♂ ventral habitus All scale bars ≈ 2 mm.
</paragraph>
</caption>
<caption pageId="59" pageNumber="60" start="Figure 34" startId="F34">
<paragraph pageId="59" pageNumber="60">
<emphasis bold="true" pageId="59" pageNumber="60">Figure 34.</emphasis>
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="59" pageNumber="60">A</emphasis>
aedeagus dorsal view
<emphasis bold="true" pageId="59" pageNumber="60">B</emphasis>
aedeagus ventral view
<emphasis bold="true" pageId="59" pageNumber="60">C</emphasis>
♂ mesotarsal claws
<emphasis bold="true" pageId="59" pageNumber="60">D</emphasis>
aedeagus lateral view. Scale bar for
<emphasis bold="true" pageId="59" pageNumber="60">C</emphasis>
≈ 0.10 mm all others ≈ 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="differential diagnosis">
<paragraph pageId="59" pageNumber="60">Differential diagnosis.</paragraph>
<paragraph pageId="59" pageNumber="60">
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
can be distinguished from all other North American
<taxonomicName class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus</emphasis>
</taxonomicName>
in males having the profemora with a small sub-apicoventral tooth, accompanied by a series of denticles that extend proximad, with each denticle being associated with a posterior setigerous puncture of the profemora, elongate mesotarsal claws of equal size, with straight ventral margins (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dineutus productus. A aedeagus dorsal view B aedeagus ventral view C ♂ mesotarsal claws D aedeagus lateral view. Scale bar for C ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">34C</figureCitation>
), and in the form of the aedeagus (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dineutus productus. A aedeagus dorsal view B aedeagus ventral view C ♂ mesotarsal claws D aedeagus lateral view. Scale bar for C ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">34A</figureCitation>
). The species most similar to
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
is
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
. The nominal subspecies of
<taxonomicName authority="serrulatus" authorityName="serrulatus LeConte" authorityYear="1868" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. serrulatus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="serrulatus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus serrulatus</emphasis>
</taxonomicName>
should not be confused with
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
in that it has a restricted range in Florida, and its elytral apices would not easy be confused with those of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
, being regularly rounded, with a strong apicolateral sinuation in both sexes and the dorsum being polished black in appearance.
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
on the other hand tends to be variable in the appearance of the elytra (Fig.
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Dineutus serrulatus analis. A ♀ dorsal habitus B ♀ ventral habitus C ♂ dorsal habitus D ♂ ventral habitus. All scale bars ≈ 2 mm." pageId="59" pageNumber="60">37A, C</figureCitation>
), with some members having the sutural angle produced and the dorsum with strong microreticulation giving the elytra and pronotum a polished bronzed appearance, similar in both respects to
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="59" pageNumber="60">
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
can generally be separated from
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
in having a more elongate narrow body form (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Dineutus productus. A ♀ dorsal habitus B ♀ ventral habitus C ♂ dorsal habitus. D ♂ ventral habitus All scale bars ≈ 2 mm." pageId="59" pageNumber="60">33A, C</figureCitation>
), and being smaller in size.
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
is much more narrowly attenuated anteriorly, compared to
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
, which is broader in body form and larger in size. The ventral coloration of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
is most often black to very dark brownish red, and frequently accompanied by a bronzy metallic sheen, while the venter of
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
is most often red in color, rarely appearing black and often lacking any metallic sheen. When more darkly colored
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
is much more red in hue, compared to when
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
is more lightly colored, having a more brownish hue. A few specimens of
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
examined from southeast Kansas had their venters truly black (KSEM) and for these specimens differences between the male secondary sexual characters as well as differences in the elytra allowed for unambiguous identification. The serration of the elytral apices differs between these two species. In
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
the apices of the elytra only have irregularities present, at most having round bumps that resemble highly reduced serration, while in
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
the serration is present as distinct and fine points.
</paragraph>
<paragraph pageId="59" pageNumber="60">
The males of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
can easily be distinguished from those of
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
in that the profemoral sub-apicoventral tooth is small and accompanied by a series of denticles that extend proximad, with each denticle being associated with a posterior setigerous puncture of the profemora, in
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
there is a profemoral sub-apicoventral tooth as normal, but no accompanying series of denticles. This character appears unique to
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
among all of the North American
<taxonomicName class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus</emphasis>
</taxonomicName>
species. The mesotarsal claws also differ greatly between these two species.
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
males have the mesotarsal claws elongate with their ventral margins straight (Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dineutus productus. A aedeagus dorsal view B aedeagus ventral view C ♂ mesotarsal claws D aedeagus lateral view. Scale bar for C ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">34C</figureCitation>
) and lacking a denticle, while in
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
the mesotarsal claws are small, with their ventral margins curved and possessing a denticle (38F). The aedeagus is by far the most definite way to separate the two species. The aedeagus of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus productus</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dineutus productus. A aedeagus dorsal view B aedeagus ventral view C ♂ mesotarsal claws D aedeagus lateral view. Scale bar for C ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">34A</figureCitation>
) has the median lobe shorter than the parameres, and strongly acuminate apically, with the parameres being sinuate after the basal 1/3 and laterally expanded in the apical 1/3, with the apices rounded. In
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="59" pageNumber="60">Dineutus serrulatus analis</emphasis>
</taxonomicName>
the median lobe (Fig.
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Dineutus serrulatus analis. Liberty Co. Florida specimen aedeagus A dorsal view B ventral view C lateral view; Tennessee specimen aedeagus D dorsal view E ventral view F ♂ mesotarsal claws G aedeagus lateral view; Kansas specimen aedeagus H dorsal view I ventral view J lateral view. Scale bar for F ≈ 0.10 mm all others ≈ 1 mm." pageId="59" pageNumber="60">38A, D, H</figureCitation>
) is as long as the parameres, parallel sided for most its length, having only the apical 1/4 narrowed, with the parameres being parallel sided for most of their length, being only weakly expanded in the apical 1/3 and often obliquely truncate, or obliquely flatly rounded.
</paragraph>
<paragraph lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
Females of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="60" pageNumber="61">
<pageBreakToken pageId="60" pageNumber="61" start="start">Dineutus</pageBreakToken>
productus
</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Dineutus productus. A ♀ dorsal habitus B ♀ ventral habitus C ♂ dorsal habitus. D ♂ ventral habitus All scale bars ≈ 2 mm." pageId="60" pageNumber="61">33A</figureCitation>
) can be distinguished from those of
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="60" pageNumber="61">Dineutus serrulatus analis</emphasis>
</taxonomicName>
in having a much greater apicolateral sinuation of the elytra, the sinuation is so deep that a plica is formed, and the elytral apices have a much steeper angle towards the sutural production than in
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="60" pageNumber="61">Dineutus serrulatus analis</emphasis>
</taxonomicName>
females (Fig.
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Dineutus serrulatus analis. A ♀ dorsal habitus B ♀ ventral habitus C ♂ dorsal habitus D ♂ ventral habitus. All scale bars ≈ 2 mm." pageId="60" pageNumber="61">37A</figureCitation>
), which have a much more shallow apicolateral sinuation and more rounded elytral apices. As mentioned earlier the elytra of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="60" pageNumber="61">Dineutus productus</emphasis>
</taxonomicName>
have irregularities present apically, but not fine serration, as seen in
<taxonomicName authorityName="Regimbart" authorityYear="1883" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus serrulatus subsp. analis" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="serrulatus" subSpecies="analis">
<emphasis italics="true" pageId="60" pageNumber="61">Dineutus serrulatus analis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="61" type="distribution">
<paragraph pageId="60" pageNumber="61">Distribution</paragraph>
<paragraph pageId="60" pageNumber="61">
<emphasis bold="true" pageId="60" pageNumber="61">
(Fig.
<figureCitation captionStart="Figure 54" captionStartId="F54" captionText="Figure 54. General distribution map. A Dineutus productus B Dineutus solitarius C Dineutus sublineatus D Dineutus mexicanus, Dineutus truncatus." pageId="60" pageNumber="61">54A</figureCitation>
).
</emphasis>
Central and south-central United States (
<bibRefCitation author="Roberts, CH" journalOrPublisher="Transactions of the American Entomological Society" pageId="95" pageNumber="96" pagination="279 - 288" refId="B107" refString="Roberts, CH, 1895. The species of Dineutes of America north of Mexico. Transactions of the American Entomological Society 22: 279 - 288" title="The species of Dineutes of America north of Mexico." volume="22" year="1895">Roberts 1895</bibRefCitation>
), into northeastern Mexico (
<bibRefCitation author="Ochs, G" journalOrPublisher="Revista de Entomologia" pageId="95" pageNumber="96" pagination="253 - 300" refId="B96" refString="Ochs, G, 1949. A revision of the Gyrinoidea of Central America (Coleoptera). Revista de Entomologia 20: 253 - 300" title="A revision of the Gyrinoidea of Central America (Coleoptera)." volume="20" year="1949">Ochs 1949</bibRefCitation>
). As this species is so rarely collected, we list here the states and counties where it has been collected and the identification is not in question:
<emphasis bold="true" pageId="60" pageNumber="61">USA:</emphasis>
<emphasis bold="true" pageId="60" pageNumber="61">Kansas:</emphasis>
Bourbon Co., Elk Co, Greenwood Co., Lyon Co., Morris Co. (this study);
<emphasis bold="true" pageId="60" pageNumber="61">Texas:</emphasis>
Bosque Co. [Clifton] (
<bibRefCitation author="Ochs, G" journalOrPublisher="Annals of the Carnegie Museum" pageId="95" pageNumber="96" pagination="123 - 134" refId="B93" refString="Ochs, G, 1929. VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species. Annals of the Carnegie Museum 19: 123 - 134" title="VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species." volume="19" year="1929">Ochs 1929</bibRefCitation>
), Brazos Co. [College Station] (
<bibRefCitation author="Wood, FE" journalOrPublisher="University of Missouri, Columbia, Missouri" pageId="97" pageNumber="98" refId="B130" refString="Wood, FE, 1962. A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere. University of Missouri, Columbia, Missouri" title="A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere" year="1962">Wood 1962</bibRefCitation>
), Comal Co. (
<bibRefCitation author="Wood, FE" journalOrPublisher="University of Missouri, Columbia, Missouri" pageId="97" pageNumber="98" refId="B130" refString="Wood, FE, 1962. A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere. University of Missouri, Columbia, Missouri" title="A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere" year="1962">Wood 1962</bibRefCitation>
), Dallas Co. [Dallas County](
<bibRefCitation author="Ochs, G" journalOrPublisher="Annals of the Carnegie Museum" pageId="95" pageNumber="96" pagination="123 - 134" refId="B93" refString="Ochs, G, 1929. VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species. Annals of the Carnegie Museum 19: 123 - 134" title="VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species." volume="19" year="1929">Ochs 1929</bibRefCitation>
), Dimmit Co. (
<bibRefCitation author="Wood, FE" journalOrPublisher="University of Missouri, Columbia, Missouri" pageId="97" pageNumber="98" refId="B130" refString="Wood, FE, 1962. A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere. University of Missouri, Columbia, Missouri" title="A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere" year="1962">Wood 1962</bibRefCitation>
), Karnes Co. [Kenedy](
<bibRefCitation author="Wood, FE" journalOrPublisher="University of Missouri, Columbia, Missouri" pageId="97" pageNumber="98" refId="B130" refString="Wood, FE, 1962. A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere. University of Missouri, Columbia, Missouri" title="A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere" year="1962">Wood 1962</bibRefCitation>
), Neuces Co. [Corpus Christi](
<bibRefCitation author="Wood, FE" journalOrPublisher="University of Missouri, Columbia, Missouri" pageId="97" pageNumber="98" refId="B130" refString="Wood, FE, 1962. A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere. University of Missouri, Columbia, Missouri" title="A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere" year="1962">Wood 1962</bibRefCitation>
);
<emphasis bold="true" pageId="60" pageNumber="61">Mexico:</emphasis>
Nuevo Leon: Granja Rodriguez on the borders of the Rio Salado alt. 195m (
<bibRefCitation author="Ochs, G" journalOrPublisher="Revista de Entomologia" pageId="95" pageNumber="96" pagination="253 - 300" refId="B96" refString="Ochs, G, 1949. A revision of the Gyrinoidea of Central America (Coleoptera). Revista de Entomologia 20: 253 - 300" title="A revision of the Gyrinoidea of Central America (Coleoptera)." volume="20" year="1949">Ochs 1949</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="60" pageNumber="61" type="habitat">
<paragraph pageId="60" pageNumber="61">Habitat.</paragraph>
<paragraph pageId="60" pageNumber="61">
Lotic species, the Kansas Biological Survey field notes (available at the KSEM) indicate the species was collected from slow moving, cool, mud bottomed streams, that were fairly wide.
<bibRefCitation author="Gustafson, GT" journalOrPublisher="The Coleopterists Bulletin" pageId="92" pageNumber="93" pagination="714 - 718" publicationUrl="10.1649/0010-065X-68.4.714" refId="B37" refString="Gustafson, GT, Maier, CA, Baca, SM, Faris, CK, 2014. Scientific Note: Rediscovery of Lutrochus laticeps Casey, 1893 (Coleoptera: Lutrochidae) and the discovery of Dineutus productus Roberts, 1895 and Dineutus serrulatus analis Regimbart, 1882 (Coleoptera: Gyrinidae) in Kansas, USA, with notes on habitat preference. The Coleopterists Bulletin 68: 714 - 718, DOI: 10.1649/0010-065X-68.4.714" title="Scientific Note: Rediscovery of Lutrochus laticeps Casey, 1893 (Coleoptera: Lutrochidae) and the discovery of Dineutus productus Roberts, 1895 and Dineutus serrulatus analis Regimbart, 1882 (Coleoptera: Gyrinidae) in Kansas, USA, with notes on habitat preference." url="10.1649/0010-065X-68.4.714" volume="68" year="2014">Gustafson et al. (2014)</bibRefCitation>
describe the habitat where
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="60" pageNumber="61">Dineutus productus</emphasis>
</taxonomicName>
was recently collected in detail, accompanied by photos of the habitats. The species has also been taken at light traps placed at water level in the vicinity of rapids, on the borders of the Rio Salado, in Nuevo Leon, Mexico (
<bibRefCitation author="Ochs, G" journalOrPublisher="Revista de Entomologia" pageId="95" pageNumber="96" pagination="253 - 300" refId="B96" refString="Ochs, G, 1949. A revision of the Gyrinoidea of Central America (Coleoptera). Revista de Entomologia 20: 253 - 300" title="A revision of the Gyrinoidea of Central America (Coleoptera)." volume="20" year="1949">Ochs 1949</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="discussion">
<paragraph pageId="60" pageNumber="61">Discussion.</paragraph>
<paragraph pageId="61" pageNumber="62">
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">
<pageBreakToken pageId="61" pageNumber="62" start="start">Dineutus</pageBreakToken>
productus
</emphasis>
</taxonomicName>
appears to be an uncommonly collected species and the reason for this is not exactly clear. Most species found in the United States are uncommon due to a narrow range of endemism (e.g.
<taxonomicName authorityName="Leng" authorityYear="1911" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus robertsi" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="robertsi">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus robertsi</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="LeConte" authorityYear="1878" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus angustus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="angustus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus angustus</emphasis>
</taxonomicName>
), but material examined in this study, as well as records from the literature, suggest the range of this species is not as narrow as the previously mentioned species.
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
was originally described from Texas by
<bibRefCitation author="Roberts, CH" journalOrPublisher="Transactions of the American Entomological Society" pageId="95" pageNumber="96" pagination="279 - 288" refId="B107" refString="Roberts, CH, 1895. The species of Dineutes of America north of Mexico. Transactions of the American Entomological Society 22: 279 - 288" title="The species of Dineutes of America north of Mexico." volume="22" year="1895">Roberts (1895)</bibRefCitation>
, and most records of this species come from this state (
<bibRefCitation author="Regimbart, M" journalOrPublisher="Annales de la Societe entomologique de France" pageId="95" pageNumber="96" pagination="663 - 737" refId="B106" refString="Regimbart, M, 1907. Essai monographique de la famille des Gyrinidae. 3e supplement. Annales de la Societe entomologique de France 76: 663 - 737" title="Essai monographique de la famille des Gyrinidae. 3 e supplement." volume="76" year="1907">
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
1907
</bibRefCitation>
;
<bibRefCitation author="Ochs, G" journalOrPublisher="Annals of the Carnegie Museum" pageId="95" pageNumber="96" pagination="123 - 134" refId="B93" refString="Ochs, G, 1929. VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species. Annals of the Carnegie Museum 19: 123 - 134" title="VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species." volume="19" year="1929">Ochs 1929</bibRefCitation>
;
<bibRefCitation author="Wood, FE" journalOrPublisher="University of Missouri, Columbia, Missouri" pageId="97" pageNumber="98" refId="B130" refString="Wood, FE, 1962. A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere. University of Missouri, Columbia, Missouri" title="A synopsis of the genus Dineutus (Coleoptera: Gyrinidae) in the Western Hemisphere" year="1962">Wood 1962</bibRefCitation>
).
<bibRefCitation author="Young, FN" journalOrPublisher=") Unveristy of Florida Studies Biological Science Series, Gainesville" pageId="97" pageNumber="98" refId="B133" refString="Young, FN, 1954. The Water Beetles of Florida. In: Florida Uo (Ed.) Unveristy of Florida Studies Biological Science Series, Gainesville" title="The Water Beetles of Florida. In: Florida Uo (Ed." year="1954">Young (1954)</bibRefCitation>
claimed to have specimens from the panhandle of Florida, but
<bibRefCitation author="Epler, JH" journalOrPublisher="Bureau of Water Resource Protection, Florida Department of Environmental Protection, Tallahassee" pageId="91" pageNumber="92" refId="B18" refString="Epler, JH, 2010. The Water Beetles of Florida - an identification manual for the families Chrysomelidae, Curculionidae, Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydraenidae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae and Scirtidae." title="The Water Beetles of Florida - an identification manual for the families Chrysomelidae, Curculionidae, Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydraenidae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae and Scirtidae" year="2010">Epler (2010)</bibRefCitation>
re-examined specimens in the FSCA identified by Young as
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
, finding them to belong to other species.
<bibRefCitation author="Hatch, MH" journalOrPublisher="Bulletin of the Brooklyn Entomological Society" pageId="92" pageNumber="93" pagination="101 - 114" refId="B40" refString="Hatch, MH, 1925a. An outline of ecology of Gyrinidae. Bulletin of the Brooklyn Entomological Society 20: 101 - 114" title="An outline of ecology of Gyrinidae." volume="20" year="1925 a">Hatch (1925a)</bibRefCitation>
claimed to have collected the species from the Sangamon River near Decatur, Illinois.
<bibRefCitation author="Ochs, G" journalOrPublisher="Revista de Entomologia" pageId="95" pageNumber="96" pagination="253 - 300" refId="B96" refString="Ochs, G, 1949. A revision of the Gyrinoidea of Central America (Coleoptera). Revista de Entomologia 20: 253 - 300" title="A revision of the Gyrinoidea of Central America (Coleoptera)." volume="20" year="1949">Ochs (1949)</bibRefCitation>
however, questioned the Illinois record as well as
<bibRefCitation author="Regimbart, M" journalOrPublisher="Annales de la Societe entomologique de France" pageId="95" pageNumber="96" pagination="663 - 737" refId="B106" refString="Regimbart, M, 1907. Essai monographique de la famille des Gyrinidae. 3e supplement. Annales de la Societe entomologique de France 76: 663 - 737" title="Essai monographique de la famille des Gyrinidae. 3 e supplement." volume="76" year="1907">
<normalizedToken originalValue="Régimbarts">Regimbart's</normalizedToken>
(1907)
</bibRefCitation>
from the Carolinas, stating all specimens identified as
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
from outside of Texas actually belonged to other species upon his examination. In the same paper
<bibRefCitation author="Ochs, G" journalOrPublisher="Revista de Entomologia" pageId="95" pageNumber="96" pagination="253 - 300" refId="B96" refString="Ochs, G, 1949. A revision of the Gyrinoidea of Central America (Coleoptera). Revista de Entomologia 20: 253 - 300" title="A revision of the Gyrinoidea of Central America (Coleoptera)." volume="20" year="1949">Ochs (1949)</bibRefCitation>
extended the species range southward in to Neuvo Leon, Mexico.
</paragraph>
<paragraph pageId="61" pageNumber="62">
Material examined for this study from Kansas (KSEM, GTGC), was undoubtedly
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
, confirming that the range of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
extends at least that far northward. This adds credence to
<bibRefCitation author="Hatch, MH" journalOrPublisher="Bulletin of the Brooklyn Entomological Society" pageId="92" pageNumber="93" pagination="101 - 114" refId="B40" refString="Hatch, MH, 1925a. An outline of ecology of Gyrinidae. Bulletin of the Brooklyn Entomological Society 20: 101 - 114" title="An outline of ecology of Gyrinidae." volume="20" year="1925 a">
<normalizedToken originalValue="Hatchs">Hatch's</normalizedToken>
(1925a)
</bibRefCitation>
record from Illinois. Furthermore, comparing images of the Sangamon River available online to those provided by
<bibRefCitation author="Gustafson, GT" journalOrPublisher="The Coleopterists Bulletin" pageId="92" pageNumber="93" pagination="714 - 718" publicationUrl="10.1649/0010-065X-68.4.714" refId="B37" refString="Gustafson, GT, Maier, CA, Baca, SM, Faris, CK, 2014. Scientific Note: Rediscovery of Lutrochus laticeps Casey, 1893 (Coleoptera: Lutrochidae) and the discovery of Dineutus productus Roberts, 1895 and Dineutus serrulatus analis Regimbart, 1882 (Coleoptera: Gyrinidae) in Kansas, USA, with notes on habitat preference. The Coleopterists Bulletin 68: 714 - 718, DOI: 10.1649/0010-065X-68.4.714" title="Scientific Note: Rediscovery of Lutrochus laticeps Casey, 1893 (Coleoptera: Lutrochidae) and the discovery of Dineutus productus Roberts, 1895 and Dineutus serrulatus analis Regimbart, 1882 (Coleoptera: Gyrinidae) in Kansas, USA, with notes on habitat preference." url="10.1649/0010-065X-68.4.714" volume="68" year="2014">Gustafson et al. (2014)</bibRefCitation>
, they appear to be the same habitat. The first author was unable to locate
<normalizedToken originalValue="Régimbarts">Regimbart's</normalizedToken>
specimen of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
within the MNHN, in order to confirm
<bibRefCitation author="Regimbart, M" journalOrPublisher="Annales de la Societe entomologique de France" pageId="95" pageNumber="96" pagination="663 - 737" refId="B106" refString="Regimbart, M, 1907. Essai monographique de la famille des Gyrinidae. 3e supplement. Annales de la Societe entomologique de France 76: 663 - 737" title="Essai monographique de la famille des Gyrinidae. 3 e supplement." volume="76" year="1907">
<normalizedToken originalValue="Régimbarts">Regimbart's</normalizedToken>
1907
</bibRefCitation>
record, but
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
was not listed as a species of
<taxonomicName class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus</emphasis>
</taxonomicName>
found in North and South Carolina by
<bibRefCitation author="Sanderson, MW" journalOrPublisher="Aquatic Insects and Oligochaetes of North and South Carolina" pageId="96" pageNumber="97" pagination="10.29 - 10.38" refId="B116" refString="Sanderson, MW, 1982. . Aquatic Insects and Oligochaetes of North and South Carolina Ill: 10.29 - 10.38" volume="Ill" year="1982">Sanderson (1982)</bibRefCitation>
. Given this and the inability to locate the specimen of
<normalizedToken originalValue="Régimbarts">Regimbart's</normalizedToken>
from the Carolinas in his collection at the MNHN, we do not treat
<bibRefCitation author="Regimbart, M" journalOrPublisher="Annales de la Societe entomologique de France" pageId="95" pageNumber="96" pagination="663 - 737" refId="B106" refString="Regimbart, M, 1907. Essai monographique de la famille des Gyrinidae. 3e supplement. Annales de la Societe entomologique de France 76: 663 - 737" title="Essai monographique de la famille des Gyrinidae. 3 e supplement." volume="76" year="1907">
<normalizedToken originalValue="Régimbarts">Regimbart's</normalizedToken>
1907
</bibRefCitation>
record as a part of the species range.
<normalizedToken originalValue="Ochs">Ochs'</normalizedToken>
diagnosis of
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
(1949) matchs our own, therefore we accept his record for the species from Nuevo Leon, Mexico as representing the southern-most extreme for the range. As mentioned above, given the new Kansas records for the species and the habitat photos of the Sangamon River available online, we accept
<bibRefCitation author="Hatch, MH" journalOrPublisher="Bulletin of the Brooklyn Entomological Society" pageId="92" pageNumber="93" pagination="101 - 114" refId="B40" refString="Hatch, MH, 1925a. An outline of ecology of Gyrinidae. Bulletin of the Brooklyn Entomological Society 20: 101 - 114" title="An outline of ecology of Gyrinidae." volume="20" year="1925 a">
<normalizedToken originalValue="Hatchs">Hatch's</normalizedToken>
(1925a)
</bibRefCitation>
record from Illinois, as the northern-most extreme for the
<normalizedToken originalValue="species">species'</normalizedToken>
range.
</paragraph>
<paragraph pageId="61" pageNumber="62">
Most works treating
<taxonomicName authorityName="Roberts" authorityYear="1895" class="Insecta" family="Gyrinidae" genus="Dineutus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dineutus productus" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="productus">
<emphasis italics="true" pageId="61" pageNumber="62">Dineutus productus</emphasis>
</taxonomicName>
only mentioned having examined a few specimens (i.e. small series at most, usually 4 specimens for most studies [
<bibRefCitation author="Roberts, CH" journalOrPublisher="Transactions of the American Entomological Society" pageId="95" pageNumber="96" pagination="279 - 288" refId="B107" refString="Roberts, CH, 1895. The species of Dineutes of America north of Mexico. Transactions of the American Entomological Society 22: 279 - 288" title="The species of Dineutes of America north of Mexico." volume="22" year="1895">Roberts 1895</bibRefCitation>
;
<bibRefCitation author="Regimbart, M" journalOrPublisher="Annales de la Societe entomologique de France" pageId="95" pageNumber="96" pagination="663 - 737" refId="B106" refString="Regimbart, M, 1907. Essai monographique de la famille des Gyrinidae. 3e supplement. Annales de la Societe entomologique de France 76: 663 - 737" title="Essai monographique de la famille des Gyrinidae. 3 e supplement." volume="76" year="1907">
<normalizedToken originalValue="Régimbart">Regimbart</normalizedToken>
1907
</bibRefCitation>
;
<bibRefCitation author="Ochs, G" journalOrPublisher="Annals of the Carnegie Museum" pageId="95" pageNumber="96" pagination="123 - 134" refId="B93" refString="Ochs, G, 1929. VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species. Annals of the Carnegie Museum 19: 123 - 134" title="VII. Notes upon some Gyrinidae in the Carnegie Museum with descriptions of new species." volume="19" year="1929">Ochs 1929</bibRefCitation>
,
<bibRefCitation author="Ochs, G" journalOrPublisher="Revista de Entomologia" pageId="95" pageNumber="96" pagination="253 - 300" refId="B96" refString="Ochs, G, 1949. A revision of the Gyrinoidea of Central America (Coleoptera). Revista de Entomologia 20: 253 - 300" title="A revision of the Gyrinoidea of Central America (Coleoptera)." volume="20" year="1949">1949</bibRefCitation>
;
<bibRefCitation author="Young, FN" journalOrPublisher=") Unveristy of Florida Studies Biological Science Series, Gainesville" pageId="97" pageNumber="98" refId="B133" refString="Young, FN, 1954. The Water Beetles of Florida. In: Florida Uo (Ed.) Unveristy of Florida Studies Biological Science Series, Gainesville" title="The Water Beetles of Florida. In: Florida Uo (Ed." year="1954">Young 1954</bibRefCitation>
]). Recent collecting revealed that the species could be collected, albeit with much difficulty, which may explain this species rarity in collections (
<bibRefCitation author="Gustafson, GT" journalOrPublisher="The Coleopterists Bulletin" pageId="92" pageNumber="93" pagination="714 - 718" publicationUrl="10.1649/0010-065X-68.4.714" refId="B37" refString="Gustafson, GT, Maier, CA, Baca, SM, Faris, CK, 2014. Scientific Note: Rediscovery of Lutrochus laticeps Casey, 1893 (Coleoptera: Lutrochidae) and the discovery of Dineutus productus Roberts, 1895 and Dineutus serrulatus analis Regimbart, 1882 (Coleoptera: Gyrinidae) in Kansas, USA, with notes on habitat preference. The Coleopterists Bulletin 68: 714 - 718, DOI: 10.1649/0010-065X-68.4.714" title="Scientific Note: Rediscovery of Lutrochus laticeps Casey, 1893 (Coleoptera: Lutrochidae) and the discovery of Dineutus productus Roberts, 1895 and Dineutus serrulatus analis Regimbart, 1882 (Coleoptera: Gyrinidae) in Kansas, USA, with notes on habitat preference." url="10.1649/0010-065X-68.4.714" volume="68" year="2014">Gustafson et al. 2014</bibRefCitation>
). Hopefully accurate information on indentifying this species, as well as modern information of habitat and distribution, will help this species become better known.
</paragraph>
</subSubSection>
</treatment>
</document>