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<document id="D8A46F4E6960D7B52C883D6BD8F4C062" ID-DOI="10.37828/em.2024.77.5" ID-ISSN="2336-9744" ID-Zenodo-Dep="14654632" ID-ZooBank="6BAC225E-7706-4046-91AF-DDCFF4D215A3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736952786659" checkinUser="felipe" docAuthor="Volynkin, Anton V. &amp; Černý, Karel" docDate="2024" docId="AE7B87A8FFEEFFD0FF156B5CFCD3F4E8" docLanguage="en" docName="EcolMontenegrina.77.44-68.pdf" docOrigin="Ecologica Montenegrina 77" docSource="https://doi.org/10.37828/em.2024.77.5" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.3:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="3" docTitle="Medioptyx Volynkin &amp; Černý, 2024, gen. n." docType="treatment" docVersion="2" lastPageNumber="63" masterDocId="5242FFD0FFFCFFC3FFA86C26FFD8F72D" masterDocTitle="Taxonomic review of the Thysanoptyx Hampson generic complex (Lepidoptera: Erebidae: Arctiinae: Lithosiini). I. The genus Collitoptyx Volynkin with descriptions of two new genera, two new subgenera and six new species" masterLastPageNumber="68" masterPageNumber="44" pageNumber="62" updateTime="1736955635683" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CFC37CE04349EEA0BDD51DC220F7E540">Taxonomic review of the Thysanoptyx Hampson generic complex (Lepidoptera: Erebidae: Arctiinae: Lithosiini). I. The genus Collitoptyx Volynkin with descriptions of two new genera, two new subgenera and six new species</mods:title>
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<mods:namePart id="E373A027C28F8D526B621DA63F4160B1">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="F8FC32B1D325DCB7210D9CE72C334E83">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia.</mods:affiliation>
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<mods:namePart id="F4607C40BBCA7FC3FC84BBE2FF0F4518">Černý, Karel</mods:namePart>
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<mods:affiliation id="04E4A2CC03B861975A0512C6A7280912">Tiergartenstrasse 27, A- 6020 Innsbruck, Austria. E-mail: natura. cerny @ aon. at; https: // orcid. org / 0000 - 0001 - 7207 - 2255 * Corresponding author. E-mail: monstruncusarctia @ gmail. com; https: // orcid. org / 0000 - 0001 - 9447 - 4925</mods:affiliation>
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<mods:date id="CC34D7666F3657A1FEDA610395C6CD89">2024</mods:date>
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<paragraph id="266D36BEFFEEFFD1FF156B5CFE36F0B9" blockId="18.[189,1191,1914,2005]" box="[189,494,1914,1940]" pageId="18" pageNumber="62">
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Genus
<taxonomicName id="E1D24D3DFFEEFFD1FEBD6B5CFE4CF0B9" ID-CoL="CJMNB" authorityName="Volynkin &amp; Černý" authorityYear="2024" box="[277,404,1914,1940]" class="Insecta" family="Erebidae" genus="Medioptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="14A6EAACFFEEFFD1FEBD6B5CFE4CF0B9" bold="true" box="[277,404,1914,1940]" italics="true" pageId="18" pageNumber="62">Medioptyx</emphasis>
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<taxonomicNameLabel id="0F9557D7FFEEFFD1FE336B5CFE36F0B9" box="[411,494,1914,1940]" pageId="18" pageNumber="62" rank="genus">gen. n.</taxonomicNameLabel>
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https://zoobank.org/
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<subSubSection id="6EC86535FFEEFFD1FF156B9DFC75F0F8" box="[189,941,1978,2005]" pageId="18" pageNumber="62" type="materials_examined">
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<typeStatus id="F969881CFFEEFFD1FF156B9DFF21F0F8" box="[189,249,1979,2005]" pageId="18" pageNumber="62">Type</typeStatus>
species:
<taxonomicName id="E1D24D3DFFEEFFD1FECD6B9CFD9DF0F8" authorityName="Volynkin &amp; Černý" authorityYear="2024" box="[357,581,1978,2005]" class="Insecta" family="Erebidae" genus="Medioptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="62" phylum="Arthropoda" rank="species" species="kishidai" status="sp. nov.">
<emphasis id="14A6EAACFFEEFFD1FECD6B9CFD9DF0F8" box="[357,581,1978,2005]" italics="true" pageId="18" pageNumber="62">Medioptyx kishidai</emphasis>
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<taxonomicNameLabel id="0F9557D7FFEEFFD1FDE56B9CFD57F0F9" box="[589,655,1978,2004]" pageId="18" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
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, by present designation.
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<subSubSection id="6EC86535FFEFFFD0FF156CE2FB18F431" pageId="19" pageNumber="63" type="diagnosis">
<paragraph id="266D36BEFFEFFFD0FF156CE2FB18F431" blockId="19.[189,1399,196,965]" pageId="19" pageNumber="63">
<emphasis id="14A6EAACFFEFFFD0FF156CE2FEEEF7F3" bold="true" box="[189,310,196,222]" pageId="19" pageNumber="63">Diagnosis</emphasis>
. The only known species of the new genus (
<figureCitation id="BEE92A3BFFEFFFD0FCF16CE2FC3DF7F2" box="[857,997,196,223]" captionStart="Figures 2533" captionStartId="9.[189,275,1912,1936]" captionTargetBox="[200,1392,258,1885]" captionTargetId="figure-134@9.[189,1398,257,1893]" captionTargetPageId="9" captionText="Figures 2533. Adults of Medioptyx kishidai sp. n. (25, 26), Thysanoptyx tetragona (27, 28), Caliginoptyx gen. n. spp. (2931), and Thysanoplisna brevimacula (32, 33). Depositories of the specimens: 2530 and 32 in MWM/ZSM (30: ex CKC), 31 in CKC; 33 in NHMUK (©The Trustees of NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.14654644" httpUri="https://zenodo.org/record/14654644/files/figure.png" pageId="19" pageNumber="63">Figs 25, 26</figureCitation>
) has forewing shape, pattern and venation very similar to
<taxonomicName id="E1D24D3DFFEFFFD0FE756CC0FD87F62C" authorityName="Volynkin" authorityYear="2023" box="[477,607,230,257]" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FE756CC0FD87F62C" box="[477,607,230,257]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
</taxonomicName>
(
<figureCitation id="BEE92A3BFFEFFFD0FDC66CC0FD3EF62C" box="[622,742,230,257]" captionStart-0="Figures 18" captionStart-1="Figures 916" captionStart-2="Figures 1724" captionStartId-0="6.[189,275,1912,1936]" captionStartId-1="7.[189,275,1912,1936]" captionStartId-2="8.[189,275,1912,1936]" captionTargetBox-0="[238,1333,259,1885]" captionTargetBox-1="[271,1335,260,1885]" captionTargetBox-2="[217,1373,257,1885]" captionTargetId-0="figure-70@6.[189,1398,257,1893]" captionTargetId-1="figure-102@7.[189,1398,257,1893]" captionTargetId-2="figure-103@8.[189,1398,257,1893]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figures 18. Adults of Collitoptyx s. str. spp. Depositories of the specimens: 1 and 6 in NHMUK (©The Trustees of NHMUK); 25, 7 and 8 in MWM/ZSM (5: ex CKC)." captionText-1="Figures 916. Adults of Collitoptyx s. str. spp. Depositories of the specimens: 912 in CAV; 13 in NHMUK (©The Trustees of NHMUK); 14 in OUMNH; 15and 16 in MWM/ZSM." captionText-2="Figures 1724. Adults of Collitoptyx (Insolitoptyx) meghalaya sp. n. (1720) and Collitoptyx (Confusoptyx) duplex sp. n. (2124). Depositories of the specimens: 17, 18, and 2124 in MWM/ZSM; 19 and 20 in NHMUK (©The Trustees of NHMUK)." figureDoi-0="http://doi.org/10.5281/zenodo.14654634" figureDoi-1="http://doi.org/10.5281/zenodo.14654638" figureDoi-2="http://doi.org/10.5281/zenodo.14654640" httpUri-0="https://zenodo.org/record/14654634/files/figure.png" httpUri-1="https://zenodo.org/record/14654638/files/figure.png" httpUri-2="https://zenodo.org/record/14654640/files/figure.png" pageId="19" pageNumber="63">Figs 124</figureCitation>
) but the significant differences in both male and female genitalia prove its generic status. The male genitalia of
<taxonomicName id="E1D24D3DFFEFFFD0FC966D21FC62F60F" authorityName="Volynkin &amp; Černý" authorityYear="2024" box="[830,954,263,290]" class="Insecta" family="Erebidae" genus="Medioptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="14A6EAACFFEFFFD0FC966D21FC62F60F" box="[830,954,263,290]" italics="true" pageId="19" pageNumber="63">Medioptyx</emphasis>
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<emphasis id="14A6EAACFFEFFFD0FC686D2EFBC9F60F" bold="true" box="[960,1041,264,290]" pageId="19" pageNumber="63">
<taxonomicNameLabel id="0F9557D7FFEFFFD0FC686D2EFBC9F60F" box="[960,1041,264,290]" pageId="19" pageNumber="63" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="BEE92A3BFFEFFFD0FB886D21FB7CF60F" box="[1056,1188,263,290]" captionStart="Figures 4648" captionStartId="15.[189,275,1931,1955]" captionTargetBox="[238,1389,274,1884]" captionTargetId="figure-90@15.[189,1398,253,1889]" captionTargetPageId="15" captionText="Figures 4648. Male genitalia of Medioptyx kishidai (46, 47) and Thysanoptyx tetragona (48). The specimens dissected are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.14654656" httpUri="https://zenodo.org/record/14654656/files/figure.png" pageId="19" pageNumber="63">Figs 46, 47</figureCitation>
) are characterised by the combination of the following features. (1) The conjuga is triangular proximally, rod-like distally, articulated with its counterpart, and both the processus momenti and the tendon are equally well-sclerotised (similar to
<taxonomicName id="E1D24D3DFFEFFFD0FE6E6D4BFD86F6A5" authorityName="Volynkin &amp; Černý" authorityYear="2024" box="[454,606,365,392]" class="Insecta" family="Erebidae" genus="Caliginoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="14A6EAACFFEFFFD0FE6E6D4BFD86F6A5" box="[454,606,365,392]" italics="true" pageId="19" pageNumber="63">Caliginoptyx</emphasis>
</taxonomicName>
<emphasis id="14A6EAACFFEFFFD0FDCD6D4BFD61F6AA" bold="true" box="[613,697,365,391]" pageId="19" pageNumber="63">
<taxonomicNameLabel id="0F9557D7FFEFFFD0FDCD6D4BFD61F6AA" box="[613,697,365,391]" pageId="19" pageNumber="63" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
) whereas in
<taxonomicName id="E1D24D3DFFEFFFD0FCFA6D4BFC0CF6A5" authorityName="Volynkin" authorityYear="2023" box="[850,980,365,392]" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FCFA6D4BFC0CF6A5" box="[850,980,365,392]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
</taxonomicName>
the tendon part performs a skeletal function. (2) The anellus is thick-walled and bears two broad ventro-lateral plates (similar to
<taxonomicName id="E1D24D3DFFEFFFD0FB5D6DA8FE84F6E6" authorityName="Volynkin &amp; Černý" authorityYear="2024" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Insolitoptyx">
<emphasis id="14A6EAACFFEFFFD0FB5D6DA8FAAFF684" box="[1269,1399,398,425]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
(
<emphasis id="14A6EAACFFEFFFD0FF6F6D96FE89F6E6" box="[199,337,432,459]" italics="true" pageId="19" pageNumber="63">Insolitoptyx</emphasis>
)
</taxonomicName>
<emphasis id="14A6EAACFFEFFFD0FECB6D97FE39F6E6" bold="true" box="[355,481,433,459]" pageId="19" pageNumber="63">
<taxonomicNameLabel id="0F9557D7FFEFFFD0FECB6D97FE39F6E6" box="[355,481,433,459]" pageId="19" pageNumber="63" rank="subGenus">subgen. n.</taxonomicNameLabel>
</emphasis>
). (3) The main diverticulum of the vesica bears a sclerotised plate armed with several spike-like cornuti (vs. a single conical cornutus in
<taxonomicName id="E1D24D3DFFEFFFD0FCF66DF4FC39F6C0" authorityName="Volynkin" authorityYear="2023" box="[862,993,466,493]" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FCF66DF4FC39F6C0" box="[862,993,466,493]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
</taxonomicName>
). (4) The sub-ejaculatorial (apical) section of the vesica is proximally enlarged, bears a thorn-like cornutus, and the vesica ejaculatorius originates from its distal part (similar to
<taxonomicName id="E1D24D3DFFEFFFD0FD0D6E33FCE1F51D" authorityName="Hampson" authorityYear="1894" box="[677,825,533,560]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FD0D6E33FCE1F51D" box="[677,825,533,560]" italics="true" pageId="19" pageNumber="63">Thysanoptyx</emphasis>
</taxonomicName>
) whereas the vesica ejaculatorius of
<taxonomicName id="E1D24D3DFFEFFFD0FB5C6E33FAAEF51D" authorityName="Volynkin" authorityYear="2023" box="[1268,1398,533,560]" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FB5C6E33FAAEF51D" box="[1268,1398,533,560]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
</taxonomicName>
originates from the base of the vesica. The female genitalia of
<taxonomicName id="E1D24D3DFFEFFFD0FC0D6E11FBF9F57F" authorityName="Volynkin &amp; Černý" authorityYear="2024" box="[933,1057,567,594]" class="Insecta" family="Erebidae" genus="Medioptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="14A6EAACFFEFFFD0FC0D6E11FBF9F57F" box="[933,1057,567,594]" italics="true" pageId="19" pageNumber="63">Medioptyx</emphasis>
</taxonomicName>
<emphasis id="14A6EAACFFEFFFD0FB816E11FBA5F57C" bold="true" box="[1065,1149,567,593]" pageId="19" pageNumber="63">
<taxonomicNameLabel id="0F9557D7FFEFFFD0FB816E11FBA5F57C" box="[1065,1149,567,593]" pageId="19" pageNumber="63" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="BEE92A3BFFEFFFD0FB256E11FB3BF57F" box="[1165,1251,567,594]" captionStart="Figures 5964" captionStartId="22.[189,275,1845,1869]" captionTargetBox="[189,1391,193,1816]" captionTargetId="figure-105@22.[189,1398,189,1825]" captionTargetPageId="22" captionText="Figures 5964. Female genitalia of the Thysanoptyx generic complex. Depositories of the specimens dissected: 5961 in MWM/ZSM; 62 and 64in NHMUK (©The Trustees of NHMUK); 63 in CKC." figureDoi="http://doi.org/10.5281/zenodo.14654668" httpUri="https://zenodo.org/record/14654668/files/figure.png" pageId="19" pageNumber="63">Fig. 61</figureCitation>
) differ from
<taxonomicName id="E1D24D3DFFEFFFD0FF156E7FFEE7F559" authorityName="Volynkin" authorityYear="2023" box="[189,319,601,628]" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FF156E7FFEE7F559" box="[189,319,601,628]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
</taxonomicName>
(
<figureCitation id="BEE92A3BFFEFFFD0FEFA6E7FFE05F559" box="[338,477,601,628]" captionStart-0="Figures 5258" captionStart-1="Figures 5964" captionStartId-0="21.[189,275,1924,1948]" captionStartId-1="22.[189,275,1845,1869]" captionTargetBox-0="[200,1390,272,1897]" captionTargetBox-1="[189,1391,193,1816]" captionTargetId-0="figure-129@21.[189,1398,269,1905]" captionTargetId-1="figure-105@22.[189,1398,189,1825]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="Figures 5258. Female genitalia of Collitoptyx spp. Depositories of the specimens dissected: 52, 53, 56 and 58 in MWM/ZSM; 54 and 55 in CAV; 57 in OUMNH." captionText-1="Figures 5964. Female genitalia of the Thysanoptyx generic complex. Depositories of the specimens dissected: 5961 in MWM/ZSM; 62 and 64in NHMUK (©The Trustees of NHMUK); 63 in CKC." figureDoi-0="http://doi.org/10.5281/zenodo.14654663" figureDoi-1="http://doi.org/10.5281/zenodo.14654668" httpUri-0="https://zenodo.org/record/14654663/files/figure.png" httpUri-1="https://zenodo.org/record/14654668/files/figure.png" pageId="19" pageNumber="63">Figs 5260</figureCitation>
) in the long, dorso-ventrally flattened and heavily sclerotised ductus bursae (similar to
<taxonomicName id="E1D24D3DFFEFFFD0FEE76E5CFE3BF5B8" authorityName="Hampson" authorityYear="1894" box="[335,483,634,661]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FEE76E5CFE3BF5B8" box="[335,483,634,661]" italics="true" pageId="19" pageNumber="63">Thysanoptyx</emphasis>
</taxonomicName>
), which is short or reduced and entirely or posteriorly membranous in
<taxonomicName id="E1D24D3DFFEFFFD0FF156EBAFEE7F59A" authorityName="Volynkin" authorityYear="2023" box="[189,319,668,695]" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FF156EBAFEE7F59A" box="[189,319,668,695]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
</taxonomicName>
. Compared to
<taxonomicName id="E1D24D3DFFEFFFD0FE436EBAFDA7F59A" authorityName="Hampson" authorityYear="1894" box="[491,639,668,695]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FE436EBAFDA7F59A" box="[491,639,668,695]" italics="true" pageId="19" pageNumber="63">Thysanoptyx</emphasis>
</taxonomicName>
(
<figureCitation id="BEE92A3BFFEFFFD0FD266EBAFD3CF59A" box="[654,740,668,695]" captionStart="Figures 5964" captionStartId="22.[189,275,1845,1869]" captionTargetBox="[189,1391,193,1816]" captionTargetId="figure-105@22.[189,1398,189,1825]" captionTargetPageId="22" captionText="Figures 5964. Female genitalia of the Thysanoptyx generic complex. Depositories of the specimens dissected: 5961 in MWM/ZSM; 62 and 64in NHMUK (©The Trustees of NHMUK); 63 in CKC." figureDoi="http://doi.org/10.5281/zenodo.14654668" httpUri="https://zenodo.org/record/14654668/files/figure.png" pageId="19" pageNumber="63">Fig. 62</figureCitation>
), the female genitalia of the new genus have a gelatinous and rugose posterior section of the corpus bursae with a broad ventral protrusion bearing a posterior sclerotised area, and a membranous and more or less globular anterior section, which is similar to
<taxonomicName id="E1D24D3DFFEFFFD0FF156F27FEB0F431" authorityName="Volynkin" authorityYear="2023" box="[189,360,769,796]" class="Insecta" family="Erebidae" genus="Thysanoplisna" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FF156F27FEB0F431" box="[189,360,769,796]" italics="true" pageId="19" pageNumber="63">Thysanoplisna</emphasis>
</taxonomicName>
but in the latter it also bears a signum, which is absent in the new genus.
</paragraph>
</subSubSection>
<subSubSection id="6EC86535FFEFFFD0FEB46F02FCD3F4E8" pageId="19" pageNumber="63" type="etymology">
<paragraph id="266D36BEFFEFFFD0FEB46F02FEF6F48E" blockId="19.[189,1399,196,965]" pageId="19" pageNumber="63">
<emphasis id="14A6EAACFFEFFFD0FEB46F02FE7CF413" bold="true" box="[284,420,804,830]" pageId="19" pageNumber="63">Etymology</emphasis>
. The genus name is an aggregate of the Latin adjective medius meaning intermediate and the genus- group name
<taxonomicName id="E1D24D3DFFEFFFD0FD016F63FCE5F44D" authorityName="Hampson" authorityYear="1894" box="[681,829,837,864]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FD016F63FCE5F44D" box="[681,829,837,864]" italics="true" pageId="19" pageNumber="63">Thysanoptyx</emphasis>
</taxonomicName>
. The name refers to the genitalia morphology of the new genus, which combines the features characteristic of
<taxonomicName id="E1D24D3DFFEFFFD0FC086F41FBFAF4AF" authorityName="Volynkin" authorityYear="2023" box="[928,1058,871,898]" class="Insecta" family="Erebidae" genus="Collitoptyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FC086F41FBFAF4AF" box="[928,1058,871,898]" italics="true" pageId="19" pageNumber="63">Collitoptyx</emphasis>
</taxonomicName>
and
<taxonomicName id="E1D24D3DFFEFFFD0FBF76F41FB2BF4AF" authorityName="Hampson" authorityYear="1894" box="[1119,1267,871,898]" class="Insecta" family="Erebidae" genus="Thysanoptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="14A6EAACFFEFFFD0FBF76F41FB2BF4AF" box="[1119,1267,871,898]" italics="true" pageId="19" pageNumber="63">Thysanoptyx</emphasis>
</taxonomicName>
. Gender is feminine.
</paragraph>
<paragraph id="266D36BEFFEFFFD0FEB46F8DFCD3F4E8" blockId="19.[189,1399,196,965]" box="[284,779,938,965]" pageId="19" pageNumber="63">
<emphasis id="14A6EAACFFEFFFD0FEB46F8DFE03F4E8" bold="true" box="[284,475,939,965]" pageId="19" pageNumber="63">Species content</emphasis>
. The genus is monotypic.
</paragraph>
</subSubSection>
</treatment>
</document>