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<document id="9A09262025E737A01A7BA1A978263BB5" ID-DOI="10.1649/072.065.0406" ID-ISSN="1938-4394" ID-Zenodo-Dep="10085403" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1622479356237" checkinUser="carolina" docAuthor="Franz, Nico M." docDate="2011" docId="AE6D4D0FFF895C56A364299BFCD8FB16" docLanguage="en" docName="TheColeopBull.65.4.352-362.pdf.imf" docOrigin="The Coleopterists Bulletin 65 (4)" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Melathra Franz, new genus" docType="treatment" docVersion="2" lastPageNumber="359" masterDocId="52543577FF885C51A277283AFFA2FF82" masterDocTitle="Melathra Huyenae Franz, A New Genus And New Species Of Entimine Weevil (Coleoptera: Curculionidae: Entiminae) From Southwestern Hispaniola" masterLastPageNumber="362" masterPageNumber="352" pageNumber="353" updateTime="1699477014241" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="976CF6814E6DD6BDFA600D2DF78301DD">Melathra Huyenae Franz, A New Genus And New Species Of Entimine Weevil (Coleoptera: Curculionidae: Entiminae) From Southwestern Hispaniola</mods:title>
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<mods:name id="50BDBCCFB7045814DF3C17132D1DD6E4" type="personal">
<mods:role id="37CB9E2A2E61BB5785FEB17EECFC0447">
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<mods:namePart id="F4B4E31FE59EFFB181AB335D2BA8DF35">Franz, Nico M.</mods:namePart>
<mods:affiliation id="454C4F1C4BF53A36F78E5C2BAB1B6A99">School of Life Sciences, PO Box 874501, Arizona State University Tempe, AZ 85287, U. S. A.</mods:affiliation>
<mods:nameIdentifier id="C495AC65F6B1C09935576E5D3B668C2F" type="email">nico.franz@asu.edu</mods:nameIdentifier>
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<mods:title id="61F37911F892BAB688948DCD16C8756D">The Coleopterists Bulletin</mods:title>
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<mods:part id="A4D4577F494E4DBE9080FA5EB4C089F7">
<mods:date id="7E5C0C0B814B55CC7E9A6DED69381E4E">2011</mods:date>
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<mods:number id="81FCDFE0CBFF8E7D1E0E2C69F295D5CC">2011-12-01</mods:number>
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<mods:detail id="119D1164BC5676D9CB8A230C9D2A4930" type="volume">
<mods:number id="CA7F2F43C8FE0BBAC508CCF30C9622F9">65</mods:number>
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<mods:classification id="AD2940081E4655D639E660A01E892B4C">journal article</mods:classification>
<mods:identifier id="8873D24777C42FF31B8346F4DB8C3B3B" type="DOI">10.1649/072.065.0406</mods:identifier>
<mods:identifier id="2F908DE78AE2CF78364E8FC422078C60" type="ISSN">1938-4394</mods:identifier>
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<treatment id="AE6D4D0FFF895C56A364299BFCD8FB16" LSID="urn:lsid:plazi:treatment:AE6D4D0FFF895C56A364299BFCD8FB16" httpUri="http://treatment.plazi.org/id/AE6D4D0FFF895C56A364299BFCD8FB16" lastPageId="7" lastPageNumber="359" pageId="1" pageNumber="353">
<subSubSection id="6EDEAF92FF895C50A364299BFD8FFE34" box="[275,557,417,438]" pageId="1" pageNumber="353" type="nomenclature">
<paragraph id="267BFC19FF895C50A364299BFD8FFE34" blockId="1.[275,557,417,465]" box="[275,557,417,438]" pageId="1" pageNumber="353">
<heading id="7D334B75FF895C50A364299BFD8FFE34" box="[275,557,417,438]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="353" reason="2">
<emphasis id="14B0200BFF895C50A364299BFD8FFE34" bold="true" box="[275,557,417,438]" pageId="1" pageNumber="353">
<taxonomicName id="E1C4879AFF895C50A364299BFE14FE34" ID-CoL="8HM6F" authority="Franz" authorityName="Franz" box="[275,438,417,438]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="14B0200BFF895C50A364299BFECCFE34" bold="true" box="[275,366,417,438]" italics="true" pageId="1" pageNumber="353">Melathra</emphasis>
Franz
</taxonomicName>
,
<taxonomicNameLabel id="0F839D70FF895C50A3B5299BFD8FFE34" box="[450,557,417,438]" pageId="1" pageNumber="353" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="6EDEAF92FF895C50A3132986FE79FE53" box="[356,475,443,465]" pageId="1" pageNumber="353" type="description">
<paragraph id="267BFC19FF895C50A3132986FE79FE53" blockId="1.[275,557,417,465]" box="[356,475,443,465]" pageId="1" pageNumber="353">
(
<figureCitation id="BEFFE09CFF895C50A31C2986FE76FE53" box="[363,468,443,465]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStart-4="Figs" captionStartId-0="2.[175,215,1710,1729]" captionStartId-1="3.[196,236,1412,1431]" captionStartId-2="4.[175,215,1279,1298]" captionStartId-3="5.[197,237,1674,1693]" captionStartId-4="6.[175,215,1327,1346]" captionTargetBox-0="[231,1091,191,1687]" captionTargetBox-1="[172,1192,191,1389]" captionTargetBox-2="[155,1169,192,1240]" captionTargetBox-3="[181,1193,191,1618]" captionTargetBox-4="[158,1171,191,1291]" captionTargetId-0="figure-0@2.[231,1092,191,1687]" captionTargetId-1="figure-142@3.[172,1192,191,1389]" captionTargetId-2="figure-192@4.[151,1171,183,1255]" captionTargetId-3="figure-0@5.[173,1193,183,1651]" captionTargetId-4="figure-163@6.[151,1171,191,1303]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="Figs. 1–3. Habitus of Melathra huyenae, female. 1) Dorsal; 2) Lateral; 3) Ventral." captionText-1="Figs. 4–6. Habitus of Melathra huyenae, male. 4) Lateral; (5) Head, dorsal view; (6) Head, lateral view." captionText-2="Figs. 7–9. Mouthparts and wings of Melathra huyenae. 7) Right maxilla, ventral view; 8) Labial prementum, lateral view; 9) Right hind wing. Not illustrated to scale." captionText-3="Figs. 10–12. Female terminalia of Melathra huyenae. 10) Sternum VIII; 11) Coxites and styli; 12) Spermatheca. Not illustrated to scale." captionText-4="Figs. 13–14. Male terminalia of Melathra huyenae. 13) Spiculum gastrale; (14) Tegmen and aedeagus, ventral and lateral views. Not illustrated to scale." figureDoi-0="http://doi.org/10.5281/zenodo.10085405" figureDoi-1="http://doi.org/10.5281/zenodo.10085407" figureDoi-2="http://doi.org/10.5281/zenodo.10085409" figureDoi-3="http://doi.org/10.5281/zenodo.10085411" figureDoi-4="http://doi.org/10.5281/zenodo.10085413" httpUri-0="https://zenodo.org/record/10085405/files/figure.png" httpUri-1="https://zenodo.org/record/10085407/files/figure.png" httpUri-2="https://zenodo.org/record/10085409/files/figure.png" httpUri-3="https://zenodo.org/record/10085411/files/figure.png" httpUri-4="https://zenodo.org/record/10085413/files/figure.png" pageId="1" pageNumber="353">Figs. 1–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6EDEAF92FF895C50A2B229DCFC02FF70" pageId="1" pageNumber="353" type="diagnosis">
<paragraph id="267BFC19FF895C50A2B229DCFC02FF70" blockId="1.[173,660,486,1729]" lastBlockId="1.[707,1194,194,1729]" pageId="1" pageNumber="353">
<emphasis id="14B0200BFF895C50A2B229DCFE8FFE79" bold="true" box="[197,301,486,507]" pageId="1" pageNumber="353">Diagnosis.</emphasis>
<taxonomicName id="E1C4879AFF895C50A34C29DCFE39FE79" box="[315,411,486,507]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A34C29DCFE39FE79" box="[315,411,486,507]" italics="true" pageId="1" pageNumber="353">Melathra</emphasis>
</taxonomicName>
keys to
<taxonomicName id="E1C4879AFF895C50A38829DCFD31FE79" authorityName="Dejean" authorityYear="1834" box="[511,659,486,507]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A38829DCFD31FE79" box="[511,659,486,507]" italics="true" pageId="1" pageNumber="353">Apotomoderes</emphasis>
</taxonomicName>
(“
<taxonomicName id="E1C4879AFF895C50A2C82A3BFE86FD94" box="[191,292,513,534]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="tribe" tribe="Barynotini">Barynotini</taxonomicName>
”) in van Emden (1944), and variously to the naupactine genera assigned to couplets
<quantity id="E13C51FCFF895C50A2DA2A0CFEB9FDC9" box="[173,283,565,587]" metricMagnitude="0" metricUnit="m" metricValue="3.0353" metricValueMax="3.0987999999999998" metricValueMin="2.9718" pageId="1" pageNumber="353" unit="in" value="119.5" valueMax="122.0" valueMin="117.0">117–122 in</quantity>
Anderson (2002;
<emphasis id="14B0200BFF895C50A3A52A0FFE42FDC9" box="[466,480,565,587]" italics="true" pageId="1" pageNumber="353">∼</emphasis>
<taxonomicName id="E1C4879AFF895C50A39D2A0FFD32FDC9" authority="Sahlberg" authorityName="Sahlberg" authorityYear="1823" box="[490,656,565,587]" class="Insecta" family="Curculionidae" genus="Artipus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A39D2A0FFD92FDC8" box="[490,560,565,586]" italics="true" pageId="1" pageNumber="353">Artipus</emphasis>
Sahlberg
</taxonomicName>
,
<taxonomicName id="E1C4879AFF895C50A2DA2A6AFEC2FDE4" authority="Dejean" authorityName="Dejean" authorityYear="1821" box="[173,352,592,614]" class="Insecta" family="Curculionidae" genus="Naupactus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A2DA2A6AFEB1FDE7" box="[173,275,592,613]" italics="true" pageId="1" pageNumber="353">Naupactus</emphasis>
Dejean
</taxonomicName>
) by virtue of the absence of a postocular lobe and vibrissae on the pronotum, the laterally positioned scrobe, the median sulcus on the rostrum, and the laterally positioned eyes. However,
<taxonomicName id="E1C4879AFF895C50A2912A80FEE3FD4D" box="[230,321,698,719]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A2912A80FEE3FD4D" box="[230,321,698,719]" italics="true" pageId="1" pageNumber="353">Melathra</emphasis>
</taxonomicName>
cannot be accommodated within any of the existing North American genera as circumscribed in Anderson (2002), and is indeed more closely related to the predominantly Hispaniolan geonemine genus
<emphasis id="14B0200BFF895C50A3292B1FFE49FCB8" box="[350,491,805,826]" italics="true" pageId="1" pageNumber="353">
<taxonomicName id="E1C4879AFF895C50A3292B1FFE45FCB8" authorityName="Dejean" authorityYear="1834" box="[350,487,805,826]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">Apotomoderes</taxonomicName>
,
</emphasis>
sharing with the latter the following diagnostic features (Franz 2011): labial palps 2-segmented; transition of rostrum to head marked by a circum-capital suture; head with a postocular constriction; eyes laterally positioned on head; profemur with one pointed, cuticular tooth; metatibial apex simple
<emphasis id="14B0200BFF895C50A3F02BFEFE07FC5B" box="[391,421,964,985]" italics="true" pageId="1" pageNumber="353">(i.e</emphasis>
., without an inner flange or an outer bevel); and aedeagus with groups of setae present in the dorsal subapical region. Nevertheless,
<taxonomicName id="E1C4879AFF895C50A3752C2EFEFCFBAB" box="[258,350,1044,1065]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A3752C2EFEFCFBAB" box="[258,350,1044,1065]" italics="true" pageId="1" pageNumber="353">Melathra</emphasis>
</taxonomicName>
is readily distinguished from
<taxonomicName id="E1C4879AFF895C50A2DA2C14FE90FBC1" authorityName="Dejean" authorityYear="1834" box="[173,306,1070,1091]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A2DA2C14FE90FBC1" box="[173,306,1070,1091]" italics="true" pageId="1" pageNumber="353">Apotomoderes</emphasis>
</taxonomicName>
by the following traits (see also Franz 2010a): dorsal half of scrobe passing over eye (not entirely ventrad of eye as in
<taxonomicName id="E1C4879AFF895C50A3CF2C59FD9FFBFA" authorityName="Dejean" authorityYear="1834" box="[440,573,1123,1144]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A3CF2C59FD9FFBFA" box="[440,573,1123,1144]" italics="true" pageId="1" pageNumber="353">Apotomoderes</emphasis>
</taxonomicName>
); eyes in dorsal profile not &quot;tilted&quot; posteriad; pronotum not sexually dimorphic; profemoral tooth small, pointed, not knife-like (as in
<taxonomicName id="E1C4879AFF895C50A3102C89FE4EFB4A" authorityName="Dejean" authorityYear="1834" box="[359,492,1203,1224]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A3102C89FE4EFB4A" box="[359,492,1203,1224]" italics="true" pageId="1" pageNumber="353">Apotomoderes</emphasis>
</taxonomicName>
); protibia without an anteromesally positioned, ridge-like (and often toothed), cuticular projection; elytra with humeri obscure; wings partially “reduced” (not fully developed as in
<taxonomicName id="E1C4879AFF895C50A3682D27FE0EFAB0" authorityName="Dejean" authorityYear="1834" box="[287,428,1309,1330]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A3682D27FE0EFAB0" box="[287,428,1309,1330]" italics="true" pageId="1" pageNumber="353">Apotomoderes</emphasis>
</taxonomicName>
); female sternum VIII elliptical (not triangular); spermatheca with corpus swollen; and median lobe with strongly sclerotized flagellum and paired anterior, narrowly plicate sclerites (absent in
<taxonomicName id="E1C4879AFF895C50A31A2DB2FE55FA1F" authorityName="Dejean" authorityYear="1834" box="[365,503,1416,1437]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A31A2DB2FE55FA1F" box="[365,503,1416,1437]" italics="true" pageId="1" pageNumber="353">Apotomoderes</emphasis>
</taxonomicName>
).
<taxonomicName id="E1C4879AFF895C50A07A2DB2FDC5FA1F" box="[525,615,1416,1437]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A07A2DB2FDC5FA1F" box="[525,615,1416,1437]" italics="true" pageId="1" pageNumber="353">Melathra</emphasis>
</taxonomicName>
furthermore differs from
<taxonomicName id="E1C4879AFF895C50A3FE2D98FDB1FA35" authorityName="Dejean" authorityYear="1834" box="[393,531,1442,1463]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A3FE2D98FDB1FA35" box="[393,531,1442,1463]" italics="true" pageId="1" pageNumber="353">Apotomoderes</emphasis>
</taxonomicName>
by the more uniformly dark brown scale coloration which is interrupted primarily by two anteromesally positioned, rusty brown spots on the elytra; a much less depressed post-epistomal region; a shorter and more foveate (less narrowly linear) median rostral sulcus; and the lack of a row of teeth along the ventral side of the metatibia in females. The aforementioned characteristics jointly serve to separate
<taxonomicName id="E1C4879AFF895C50A04D2E4DFD31F90E" box="[570,659,1655,1676]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="353" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF895C50A04D2E4DFD31F90E" box="[570,659,1655,1676]" italics="true" pageId="1" pageNumber="353">Melathra</emphasis>
</taxonomicName>
also from other as of yet undescribed Caribbean species of entimine weevils that were examined by the author, thereby justifying the erection of this new genus-level taxon.
</paragraph>
</subSubSection>
<subSubSection id="6EDEAF92FF895C56A0AC28CDFCE1FC26" lastPageId="7" lastPageNumber="359" pageId="1" pageNumber="353" type="description">
<paragraph id="267BFC19FF895C56A0AC28CDFD36FBC6" blockId="1.[707,1194,194,1729]" lastBlockId="7.[173,660,194,1729]" lastPageId="7" lastPageNumber="359" pageId="1" pageNumber="353">
<emphasis id="14B0200BFF895C50A0AC28CDFC0FFE8E" bold="true" box="[731,941,247,268]" pageId="1" pageNumber="353">Description. Female</emphasis>
(
<figureCitation id="BEFFE09CFF895C50A1CC28CDFBB4FE8E" box="[955,1046,247,269]" captionStart="Figs" captionStartId="2.[175,215,1710,1729]" captionTargetBox="[231,1091,191,1687]" captionTargetId="figure-0@2.[231,1092,191,1687]" captionTargetPageId="2" captionText="Figs. 1–3. Habitus of Melathra huyenae, female. 1) Dorsal; 2) Lateral; 3) Ventral." figureDoi="http://doi.org/10.5281/zenodo.10085405" httpUri="https://zenodo.org/record/10085405/files/figure.png" pageId="1" pageNumber="353">Figs. 1–3</figureCitation>
). Length
<quantity id="E13C51FCFF895C50A60428CDFC82FEA5" metricMagnitude="-3" metricUnit="m" metricValue="8.085" metricValueMax="9.97" metricValueMin="6.2" pageId="1" pageNumber="353" unit="mm" value="8.085" valueMax="9.97" valueMin="6.2">6.20– 9.97 mm</quantity>
, width
<quantity id="E13C51FCFF895C50A1182928FBA3FEA5" box="[879,1025,273,295]" metricMagnitude="-3" metricUnit="m" metricValue="3.465" metricValueMax="4.17" metricValueMin="2.76" pageId="1" pageNumber="353" unit="mm" value="3.465" valueMax="4.17" valueMin="2.76">2.76–4.17 mm</quantity>
(
<emphasis id="14B0200BFF895C50A664292BFBBDFEA4" box="[1043,1055,273,294]" italics="true" pageId="1" pageNumber="353">n</emphasis>
= 10); shape elongate-pyriform, length/width ratio 2.25–2.50; widest at midregion to posterior 1/3 of elytra. Integument dark (black), legs dark reddish brown. Small, subcircular, subcontiguous, appressed, variously colored scales densely arranged throughout body surface; scales (where not abraised) mostly pale white or grey-silver to tan or darker brown, somewhat translucent, yet characteristically light rusty brown (to orange) and metallic (visible at higher magnification) on lateral sides of pronotum, epipleura, and in anteromesal regions of elytra.
<emphasis id="14B0200BFF895C50A0B42A6BFC9DFDE4" bold="true" box="[707,831,593,614]" pageId="1" pageNumber="353">Mouthparts</emphasis>
: Mandibles (
<figureCitation id="BEFFE09CFF895C50A1B02A6BFBA7FDE4" box="[967,1029,593,614]" captionStart="Figs" captionStartId="2.[175,215,1710,1729]" captionTargetBox="[231,1091,191,1687]" captionTargetId="figure-0@2.[231,1092,191,1687]" captionTargetPageId="2" captionText="Figs. 1–3. Habitus of Melathra huyenae, female. 1) Dorsal; 2) Lateral; 3) Ventral." figureDoi="http://doi.org/10.5281/zenodo.10085405" httpUri="https://zenodo.org/record/10085405/files/figure.png" pageId="1" pageNumber="353">Fig. 3</figureCitation>
) lacking scales, with 5–8 long, several shorter setae, mandibular scar positioned apicolaterally, slightly projected. Maxillae (
<figureCitation id="BEFFE09CFF895C50A15D2A9AFCCBFD37" box="[810,873,672,693]" captionStart="Figs" captionStartId="4.[175,215,1279,1298]" captionTargetBox="[155,1169,192,1240]" captionTargetId="figure-192@4.[151,1171,183,1255]" captionTargetPageId="4" captionText="Figs. 7–9. Mouthparts and wings of Melathra huyenae. 7) Right maxilla, ventral view; 8) Labial prementum, lateral view; 9) Right hind wing. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085409" httpUri="https://zenodo.org/record/10085409/files/figure.png" pageId="1" pageNumber="353">Fig. 7</figureCitation>
) with cardo elongate, basically widely bifurcate, lacking setae; stipes narrowly elongate, with lateral setae; galeo-lacinial complex mesally extending to apex of maxillary palpomere I, apically rounded, covered with short setae throughout, with several lacinial teeth (5–6 narrow and apical, wider and mesal; see Ting 1936), thereafter (along mesal margin) with tuft of long setae; palpiger with transverse row of setae; maxillary palps 3-segmented; I longer than II, with 2 apicolateral setae; II similar in length and wider than III, with multiple circumapical setae; III elongate, with parallel sulci, apically papillate. Labium (
<figureCitation id="BEFFE09CFF895C50A6132BE5FB03FC76" box="[1124,1185,991,1012]" captionStart="Figs" captionStartId="4.[175,215,1279,1298]" captionTargetBox="[155,1169,192,1240]" captionTargetId="figure-192@4.[151,1171,183,1255]" captionTargetPageId="4" captionText="Figs. 7–9. Mouthparts and wings of Melathra huyenae. 7) Right maxilla, ventral view; 8) Labial prementum, lateral view; 9) Right hind wing. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085409" httpUri="https://zenodo.org/record/10085409/files/figure.png" pageId="1" pageNumber="353">Fig. 8</figureCitation>
) with prementum entirely covering maxillary palps; escudate-pentagonal; apical margin medially projected; each apicolateral region with 3–4 long setae; labial palps 2-segmented, not reaching apical margin of prementum; I and II similar in length, subquadrate; I with 1 apicolateral seta; II apically papillate.
<emphasis id="14B0200BFF895C50A1572CA3FCDAFB2C" bold="true" box="[800,888,1177,1198]" pageId="1" pageNumber="353">Rostrum</emphasis>
: Length
<quantity id="E13C51FCFF895C50A1A72CA3FBFFFB2C" box="[976,1117,1176,1198]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" metricValueMax="1.46" metricValueMin="0.96" pageId="1" pageNumber="353" unit="mm" value="1.21" valueMax="1.46" valueMin="0.96">0.96–1.46 mm</quantity>
, rostral/ pronotal length ratio 0.54–0.62, rostral length/width ratio 0.93–1.04. Dorsal outline of rostrum subquadrate; dorsolateral margins anteriorly diverging; apical margin triangularly incised. Nasal plate (
<bibRefCitation id="425581E8FF895C50A0BE2D24FCE7FAB1" author="Vaurie" box="[713,837,1310,1331]" pageId="1" pageNumber="353" pagination="239 - 304" refId="ref5453" refString="Vaurie, P. 1963. A revision of the South American genus Hyphantus (Coleoptera, Curculionidae, Otiorhynchinae). Bulletin of the American Museum of Natural History 125: 239 - 304." type="journal article" year="1963">Vaurie 1963</bibRefCitation>
) weakly defined, slightly concaveinflected, posteriorly not carinate; epistoma forming inversely V-shaped, slightly angulate stripe between nasal plate and basal 3/4 of rostrum. Dorsal surface of rostrum slightly convex, with median, posteriorly narrowing sulcus extending (maximally) to imaginary midpoint between anterior margins of eyes, often shorter. Rostrum in lateral view nearly straight, anteriorly slightly expanded; occipital sutures welldefined, extending to ventral midpoint of rostrum. Ventral surface of rostrum with gular sutures fused (Lyal 1995), anteriorly terminating in large, deep fovea, thereafter hypostomal-labial sutures widely bifurcating, curved, not reaching labial prementum. Antennal insertion near anterior 1/5 of rostrum. Scrobe lateral, strongly curved and posteriorly widened, terminating anteriorly and directed ventrad of eye, though dorsal half passing over eye; anterior 1/2 of ventral margin of scrobe wide, convex, set off from lateral surface of rostrum by deep sulcus. Antennae 11-segmented; scape moderately stout; funicular antennomeres progressing from elongate to equilateral, clavate; club similar in length to funicular antennomeres III–VI; otherwise similar to
<taxonomicName id="E1C4879AFF8B5C52A2BE2EABFEF3F924" authorityName="Dejean" authorityYear="1834" box="[201,337,1681,1702]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF8B5C52A2BE2EABFEF3F924" box="[201,337,1681,1702]" italics="true" pageId="3" pageNumber="355">Apotomoderes</emphasis>
</taxonomicName>
(see Franz 2010a).
<emphasis id="14B0200BFF8B5C52A0692EA8FDF6F925" bold="true" box="[542,596,1682,1703]" pageId="3" pageNumber="355">Head</emphasis>
: Eyes laterally positioned, more or less evenly globular and protruded; separated (in dorsal view) by distance more than anterior-to-posterior length of each eye. Head with apparent postocular constriction extending along entire circumference, rostrum and head in lateral view angulate.
<emphasis id="14B0200BFF8B5C52A18E2E1DFBE0F9BE" bold="true" box="[1017,1090,1575,1596]" pageId="3" pageNumber="355">Thorax</emphasis>
: Pronotum equilateral, length/width ratio 0.92–0.96, pronotal/ elytra length ratio 0.34–0.36, slightly convex and globular; widest near midpoint; surface punctate, punctures homogeneously spaced; median sulcus absent. Anterolateral margins of pronotum slightly projected, postocular lobe and vibrissae absent. Scutellum exposed, semi-circular, covered with scales. Epipleura with mesepisternum triangular; mesepimeron dorsally obliquely narrowed; metepisternum linear, anteriorly abruptly widened; metepimeron entirely covered by elytron. Prosternum longer than mesosternum; procoxal cavities positioned at midpoint, contiguous, prosternal process short, elevated. Mesosternum slightly shorter than metasternum; mesocoxal cavities separated by distance nearly half as long as width of each mesocoxal cavity. Metasternum with median sulcus present as large, transverse fovea positioned anteriorly of posterior margin; metacoxal cavities widely separated; otherwise similar to
<taxonomicName id="E1C4879AFF8C5C55A1122D68FC4AFAE5" authorityName="Dejean" authorityYear="1834" box="[869,1000,1362,1383]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="356" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF8C5C55A1122D68FC4AFAE5" box="[869,1000,1362,1383]" italics="true" pageId="4" pageNumber="356">Apotomoderes</emphasis>
</taxonomicName>
(see Franz 2010a). Metendosternite with stalk wide; hemiducts wide, truncate; dorsal margin of sheath undulate; anterior tendons positioned near midpoint between median keel and base of furcal arms; furcal arms short, slender, diverging at 30–45° in relation to median keel.
<emphasis id="14B0200BFF8C5C55A0952DC8FCB3F985" bold="true" box="[738,785,1522,1543]" pageId="4" pageNumber="356">Legs</emphasis>
: Prothoracic and metathoracic legs each longer than mesothoracic legs. Profemoral/pronotal length ratio 1.09–1.19; profemur with 1 small, pointed (though not knife-like), cuticular tooth inserted at apical 2/5 on anteromesal margin. Protibial/ profemoral length ratio 0.92–1.00; protibia nearly straight, width similar throughout; anteromesal margin with row of 8–10 small, apically obliquely truncate cuticular teeth, each tooth distally with brownish, spiniform seta; protibial apex with anterior margin truncate, setal comb absent; mucro similar in length to tarsal claw. Protarsus with tarsomere I nearly 2X as long as II, elongate, clavate; II and III similar in length, equilateral to transverse, jointly similar in length to V; claws paired, separate, simple. Meso- and metafemora unarmed; mesotibia with anteromesal row of teeth more widely spaced, metatibiae with teeth not well-developed (obscure); metatibial apex simple, mucro absent, lateral margins each with a comb of spiniform setae of increasing length, yet without a bevel or flange (“corbel open”; Thompson 1992), surface surrounding metatarsal condyle glabrous.
<emphasis id="14B0200BFF8E5C57A1652DC8FCF6F985" bold="true" box="[786,852,1522,1543]" pageId="6" pageNumber="358">Elytra</emphasis>
: Length/width ratio 1.77–1.81; widest near midregion; anterior margins jointly slightly wider than posterior margin of pronotum, nearly straight; humeri obscure, rounded; lateral margins subparallel along anterior 3/4, thereafter roundly converging, posteriorly attenuate. Elytra in lateral view plane to slightly convex; posterior declivity apparent though not strongly angulate, nearly straight. Elytra with striae I-IX complete, stria X incomplete and apparent only along marginal anterior 1/4; striae similar in width to intervals; punctures separated by distance similar to width of each puncture; intervals not elevated; with a tuft of aurate, linear, suberect scales at mesal margins of declivity.
<emphasis id="14B0200BFF8F5C56A3792958FEECFEF5" bold="true" box="[270,334,354,375]" pageId="7" pageNumber="359">Wings</emphasis>
(
<figureCitation id="BEFFE09CFF8F5C56A3172958FE3DFEF5" box="[352,415,354,375]" captionStart="Figs" captionStartId="4.[175,215,1279,1298]" captionTargetBox="[155,1169,192,1240]" captionTargetId="figure-192@4.[151,1171,183,1255]" captionTargetPageId="4" captionText="Figs. 7–9. Mouthparts and wings of Melathra huyenae. 7) Right maxilla, ventral view; 8) Labial prementum, lateral view; 9) Right hind wing. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085409" httpUri="https://zenodo.org/record/10085409/files/figure.png" pageId="7" pageNumber="359">Fig. 9</figureCitation>
): Wings present but not fully developed, shorter and more linear in comparison to
<taxonomicName id="E1C4879AFF8F5C56A36D29ACFE00FE29" authorityName="Dejean" authorityYear="1834" box="[282,418,406,427]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF8F5C56A36D29ACFE00FE29" box="[282,418,406,427]" italics="true" pageId="7" pageNumber="359">Apotomoderes</emphasis>
</taxonomicName>
(see Franz 2010a), distally obliquely narrowed, likely not functional, though most alar veins and sclerites well-defined (see
<figureCitation id="BEFFE09CFF8F5C56A2AB29DCFEBBFE79" box="[220,281,486,507]" captionStart="Figs" captionStartId="4.[175,215,1279,1298]" captionTargetBox="[155,1169,192,1240]" captionTargetId="figure-192@4.[151,1171,183,1255]" captionTargetPageId="4" captionText="Figs. 7–9. Mouthparts and wings of Melathra huyenae. 7) Right maxilla, ventral view; 8) Labial prementum, lateral view; 9) Right hind wing. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085409" httpUri="https://zenodo.org/record/10085409/files/figure.png" pageId="7" pageNumber="359">Fig. 9</figureCitation>
).
<emphasis id="14B0200BFF8F5C56A35929DCFE33FE79" bold="true" box="[302,401,486,507]" pageId="7" pageNumber="359">Abdomen</emphasis>
: Venter with segments III and IV jointed, slightly convex and elevated, V-VII separate, the latter with linear and increasingly longer, suberect, aurate scales; VII with several small, shallow, irregularly shaped impressions, lateral margins posteriorly triangularly converging, posterior margin narrowly rounded. Tergum VII lacking strigulate regions. Pygidium (tergum VIII) entirely covered by elytra.
<emphasis id="14B0200BFF8F5C56A3D42A81FDA9FD52" bold="true" box="[419,523,699,720]" pageId="7" pageNumber="359">Terminalia</emphasis>
: Sternum VIII (
<figureCitation id="BEFFE09CFF8F5C56A2C32AEFFEA3FD68" box="[180,257,725,746]" captionStart="Figs" captionStartId="5.[197,237,1674,1693]" captionTargetBox="[181,1193,191,1618]" captionTargetId="figure-0@5.[173,1193,183,1651]" captionTargetPageId="5" captionText="Figs. 10–12. Female terminalia of Melathra huyenae. 10) Sternum VIII; 11) Coxites and styli; 12) Spermatheca. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085411" httpUri="https://zenodo.org/record/10085411/files/figure.png" pageId="7" pageNumber="359">Fig. 10</figureCitation>
) with anterior 3/4 (spiculum ventrale) narrowly stylate; posterior 1/4 (lamina) elliptical, entire (arms jointed), lateral margins continuously rounded, posterior margin slightly emarginate, posterior 1/4 of lamina with increasingly longer setae. Coxites (
<figureCitation id="BEFFE09CFF8F5C56A34D2B60FEDEFCED" box="[314,380,858,879]" captionStart="Figs" captionStartId="5.[197,237,1674,1693]" captionTargetBox="[181,1193,191,1618]" captionTargetId="figure-0@5.[173,1193,183,1651]" captionTargetPageId="5" captionText="Figs. 10–12. Female terminalia of Melathra huyenae. 10) Sternum VIII; 11) Coxites and styli; 12) Spermatheca. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085411" httpUri="https://zenodo.org/record/10085411/files/figure.png" pageId="7" pageNumber="359">Fig. 11</figureCitation>
) similar in length to spiculum ventrale, weakly sclerotized, laterally with sparsely arranged setae; styli more strongly sclerotized, short, ovoid, posteriorly with 2–3 longer setae. Spermatheca (
<figureCitation id="BEFFE09CFF8F5C56A2C32BFEFF5CFC5B" box="[180,254,964,985]" captionStart="Figs" captionStartId="5.[197,237,1674,1693]" captionTargetBox="[181,1193,191,1618]" captionTargetId="figure-0@5.[173,1193,183,1651]" captionTargetPageId="5" captionText="Figs. 10–12. Female terminalia of Melathra huyenae. 10) Sternum VIII; 11) Coxites and styli; 12) Spermatheca. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085411" httpUri="https://zenodo.org/record/10085411/files/figure.png" pageId="7" pageNumber="359">Fig. 12</figureCitation>
) variously C-shaped; collum and ramus not projected and widely separated; corpus large, tumescent; cornu slightly longer than corpus, continuously curved and apically pointing in a direction perpendicular to “axis” of collum, apex acute.
</paragraph>
<caption id="72BBAC91FF8A5C53A2D82E94FC2DF943" ID-DOI="http://doi.org/10.5281/zenodo.10085405" ID-Zenodo-Dep="10085405" box="[175,911,1710,1729]" httpUri="https://zenodo.org/record/10085405/files/figure.png" pageId="2" pageNumber="354" startId="2.[175,215,1710,1729]" targetBox="[231,1091,191,1687]" targetPageId="2">
<paragraph id="267BFC19FF8A5C53A2D82E94FC2DF943" blockId="2.[175,911,1710,1729]" box="[175,911,1710,1729]" pageId="2" pageNumber="354">
<emphasis id="14B0200BFF8A5C53A2D82E94FEA9F943" bold="true" box="[175,267,1710,1729]" pageId="2" pageNumber="354">Figs. 1–3.</emphasis>
Habitus of
<emphasis id="14B0200BFF8A5C53A3F52E94FD81F943" box="[386,547,1710,1729]" italics="true" pageId="2" pageNumber="354">
<taxonomicName id="E1C4879AFF8A5C53A3F52E94FD82F943" box="[386,544,1710,1729]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="354" phylum="Arthropoda" rank="species" species="huyenae">Melathra huyenae</taxonomicName>
,
</emphasis>
female.
<emphasis id="14B0200BFF8A5C53A0182E94FD21F943" bold="true" box="[623,643,1710,1729]" pageId="2" pageNumber="354">1)</emphasis>
Dorsal;
<emphasis id="14B0200BFF8A5C53A0B92E94FD40F943" bold="true" box="[718,738,1710,1729]" pageId="2" pageNumber="354">2)</emphasis>
Lateral;
<emphasis id="14B0200BFF8A5C53A1462E94FCE6F943" bold="true" box="[817,836,1710,1729]" pageId="2" pageNumber="354">3)</emphasis>
Ventral.
</paragraph>
</caption>
<caption id="72BBAC91FF8B5C52A2B32DBEFBCAFA15" ID-DOI="http://doi.org/10.5281/zenodo.10085407" ID-Zenodo-Dep="10085407" box="[196,1128,1412,1431]" httpUri="https://zenodo.org/record/10085407/files/figure.png" pageId="3" pageNumber="355" startId="3.[196,236,1412,1431]" targetBox="[172,1192,191,1389]" targetPageId="3">
<paragraph id="267BFC19FF8B5C52A2B32DBEFBCAFA15" blockId="3.[196,1128,1412,1431]" box="[196,1128,1412,1431]" pageId="3" pageNumber="355">
<emphasis id="14B0200BFF8B5C52A2B32DBEFE82FA15" bold="true" box="[196,288,1412,1431]" pageId="3" pageNumber="355">Figs. 4–6.</emphasis>
Habitus of
<emphasis id="14B0200BFF8B5C52A3E02DBEFD9BFA15" box="[407,569,1412,1431]" italics="true" pageId="3" pageNumber="355">
<taxonomicName id="E1C4879AFF8B5C52A3E02DBEFD97FA15" box="[407,565,1412,1431]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="355" phylum="Arthropoda" rank="species" species="huyenae">Melathra huyenae</taxonomicName>
,
</emphasis>
male.
<emphasis id="14B0200BFF8B5C52A0032DBEFD25FA15" bold="true" box="[628,647,1412,1431]" pageId="3" pageNumber="355">4)</emphasis>
Lateral;
<emphasis id="14B0200BFF8B5C52A0A12DBEFD51FA15" bold="true" box="[726,755,1412,1431]" pageId="3" pageNumber="355">(5)</emphasis>
Head, dorsal view;
<emphasis id="14B0200BFF8B5C52A1D42DBEFC1DFA15" bold="true" box="[931,959,1412,1431]" pageId="3" pageNumber="355">(6)</emphasis>
Head, lateral view.
</paragraph>
</caption>
<caption id="72BBAC91FF8C5C55A2D82CC5FDE9FAAB" ID-DOI="http://doi.org/10.5281/zenodo.10085409" ID-Zenodo-Dep="10085409" httpUri="https://zenodo.org/record/10085409/files/figure.png" pageId="4" pageNumber="356" startId="4.[175,215,1279,1298]" targetBox="[155,1169,192,1240]" targetPageId="4">
<paragraph id="267BFC19FF8C5C55A2D82CC5FDE9FAAB" blockId="4.[151,1172,1278,1321]" pageId="4" pageNumber="356">
<emphasis id="14B0200BFF8C5C55A2D82CC5FEABFA90" bold="true" box="[175,265,1278,1298]" pageId="4" pageNumber="356">Figs. 7–9.</emphasis>
Mouthparts and wings of
<taxonomicName id="E1C4879AFF8C5C55A3812CC4FD2DFA93" box="[502,655,1278,1297]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="356" phylum="Arthropoda" rank="species" species="huyenae">
<emphasis id="14B0200BFF8C5C55A3812CC4FD2DFA93" box="[502,655,1278,1297]" italics="true" pageId="4" pageNumber="356">Melathra huyenae</emphasis>
</taxonomicName>
.
<emphasis id="14B0200BFF8C5C55A0ED2CC5FD0FFA90" bold="true" box="[666,685,1279,1298]" pageId="4" pageNumber="356">7)</emphasis>
Right maxilla, ventral view;
<emphasis id="14B0200BFF8C5C55A1D52CC5FC14FA90" bold="true" box="[930,950,1279,1298]" pageId="4" pageNumber="356">8)</emphasis>
Labial prementum, lateral view;
<emphasis id="14B0200BFF8C5C55A2BA2D2CFF42FAAB" bold="true" box="[205,224,1302,1321]" pageId="4" pageNumber="356">9)</emphasis>
Right hind wing. Not illustrated to scale.
</paragraph>
</caption>
<caption id="72BBAC91FF8D5C54A2B22EB0FED4F937" ID-DOI="http://doi.org/10.5281/zenodo.10085411" ID-Zenodo-Dep="10085411" httpUri="https://zenodo.org/record/10085411/files/figure.png" pageId="5" pageNumber="357" startId="5.[197,237,1674,1693]" targetBox="[181,1193,191,1618]" targetPageId="5">
<paragraph id="267BFC19FF8D5C54A2B22EB0FB08F91F" blockId="5.[173,1194,1674,1717]" box="[197,1194,1674,1693]" pageId="5" pageNumber="357">
<emphasis id="14B0200BFF8D5C54A2B22EB0FE97F91F" bold="true" box="[197,309,1674,1693]" pageId="5" pageNumber="357">Figs. 10–12.</emphasis>
Female terminalia of
<taxonomicName id="E1C4879AFF8D5C54A0732EB0FD02F91F" box="[516,672,1674,1693]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="357" phylum="Arthropoda" rank="species" species="huyenae">
<emphasis id="14B0200BFF8D5C54A0732EB0FD02F91F" box="[516,672,1674,1693]" italics="true" pageId="5" pageNumber="357">Melathra huyenae</emphasis>
</taxonomicName>
.
<emphasis id="14B0200BFF8D5C54A0DA2EB0FD69F91F" bold="true" box="[685,715,1674,1693]" pageId="5" pageNumber="357">10)</emphasis>
Sternum VIII;
<emphasis id="14B0200BFF8D5C54A1262EB0FCCEF91F" bold="true" box="[849,876,1674,1693]" pageId="5" pageNumber="357">11)</emphasis>
Coxites and styli;
<emphasis id="14B0200BFF8D5C54A6652EB0FB92F91F" bold="true" box="[1042,1072,1674,1693]" pageId="5" pageNumber="357">12)</emphasis>
Spermatheca.
</paragraph>
<paragraph id="267BFC19FF8D5C54A2DA2E98FED4F937" blockId="5.[173,1194,1674,1717]" box="[173,374,1698,1717]" pageId="5" pageNumber="357">Not illustrated to scale.</paragraph>
</caption>
<caption id="72BBAC91FF8E5C57A2D82D15FE7BFAD8" ID-DOI="http://doi.org/10.5281/zenodo.10085413" ID-Zenodo-Dep="10085413" httpUri="https://zenodo.org/record/10085413/files/figure.png" pageId="6" pageNumber="358" startId="6.[175,215,1327,1346]" targetBox="[158,1171,191,1291]" targetPageId="6">
<paragraph id="267BFC19FF8E5C57A2D82D15FE7BFAD8" blockId="6.[151,1172,1326,1370]" pageId="6" pageNumber="358">
<emphasis id="14B0200BFF8E5C57A2D82D15FEBCFAC0" bold="true" box="[175,286,1326,1346]" pageId="6" pageNumber="358">Figs. 13–14.</emphasis>
Male terminalia of
<emphasis id="14B0200BFF8E5C57A3A02D14FDDAFAC3" box="[471,632,1326,1345]" italics="true" pageId="6" pageNumber="358">
<taxonomicName id="E1C4879AFF8E5C57A3A02D14FDD6FAC3" box="[471,628,1326,1345]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="358" phylum="Arthropoda" rank="species" species="huyenae">Melathra huyenae</taxonomicName>
.
</emphasis>
<emphasis id="14B0200BFF8E5C57A00B2D15FD38FAC0" bold="true" box="[636,666,1327,1346]" pageId="6" pageNumber="358">13)</emphasis>
Spiculum gastrale;
<emphasis id="14B0200BFF8E5C57A1342D15FCC8FAC0" bold="true" box="[835,874,1327,1346]" pageId="6" pageNumber="358">(14)</emphasis>
Tegmen and aedeagus, ventral and lateral views. Not illustrated to scale.
</paragraph>
</caption>
<paragraph id="267BFC19FF8F5C56A2B22C73FCE1FC26" blockId="7.[173,660,194,1729]" lastBlockId="7.[707,1193,194,1172]" pageId="7" pageNumber="359">
<emphasis id="14B0200BFF8F5C56A2B22C73FF55FBDC" bold="true" box="[197,247,1097,1118]" pageId="7" pageNumber="359">Male</emphasis>
(
<figureCitation id="BEFFE09CFF8F5C56A3762C73FEF5FBDC" box="[257,343,1097,1119]" captionStart="Figs" captionStartId="3.[196,236,1412,1431]" captionTargetBox="[172,1192,191,1389]" captionTargetId="figure-142@3.[172,1192,191,1389]" captionTargetPageId="3" captionText="Figs. 4–6. Habitus of Melathra huyenae, male. 4) Lateral; (5) Head, dorsal view; (6) Head, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10085407" httpUri="https://zenodo.org/record/10085407/files/figure.png" pageId="7" pageNumber="359">Figs. 4–6</figureCitation>
). Overall similar to female,though body shape slightly more slender (not pyriform) and pronotum slightly more globular. Length 6.14– 10.03, width 2.49–4.37 (
<emphasis id="14B0200BFF8F5C56A3C52CA2FE1CFB2F" box="[434,446,1176,1197]" italics="true" pageId="7" pageNumber="359">n</emphasis>
= 10); length/width ratio 2.30–2.47; rostral length
<quantity id="E13C51FCFF8F5C56A3822C8EFD2DFB4B" box="[501,655,1203,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" metricValueMax="1.52" metricValueMin="0.92" pageId="7" pageNumber="359" unit="mm" value="1.22" valueMax="1.52" valueMin="0.92">0.92–1.52 mm</quantity>
; rostral/pronotal length ratio 0.50–0.55, rostral length/width ratio 0.92–0.96; pronotal length/ width ratio 0.91–0.96, pronotal/elytral length ratio 0.36–0.38; profemoral/pronotal length ratio 1.07– 1.22; protibial/profemoral ratio 0.89–0.95; metatibia slightly longer than in female, apically curved, otherwise similar to protibia, row of cuticular teeth and mucro present; elytral length/width ratio 1.73–1.82; elytral declivity rounded, not angulate, less pronounced than in female, mesally positioned tuft of scales absent; venter with segment VII short, posterior margin widely rounded to truncate.
<emphasis id="14B0200BFF8F5C56A2DA2E37FEBAF9A0" bold="true" box="[173,280,1549,1570]" pageId="7" pageNumber="359">Terminalia</emphasis>
: Sternum VIII similar to
<taxonomicName id="E1C4879AFF8F5C56A0792E36FD31F9A3" authorityName="Dejean" authorityYear="1834" box="[526,659,1548,1569]" class="Insecta" family="Curculionidae" genus="Apotomoderes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF8F5C56A0792E36FD31F9A3" box="[526,659,1548,1569]" italics="true" pageId="7" pageNumber="359">Apotomoderes</emphasis>
</taxonomicName>
(see Franz 2010a). Spiculum gastrale (
<figureCitation id="BEFFE09CFF8F5C56A0482E1DFD29F9BE" box="[575,651,1575,1596]" captionStart="Figs" captionStartId="6.[175,215,1327,1346]" captionTargetBox="[158,1171,191,1291]" captionTargetId="figure-163@6.[151,1171,191,1303]" captionTargetPageId="6" captionText="Figs. 13–14. Male terminalia of Melathra huyenae. 13) Spiculum gastrale; (14) Tegmen and aedeagus, ventral and lateral views. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085413" httpUri="https://zenodo.org/record/10085413/files/figure.png" pageId="7" pageNumber="359">Fig. 13</figureCitation>
) longer than median lobe; stylus slightly curved; posteriorly bifurcate, furcal arms elongate-triangular, anteriorly weakly sclerotized, posteriorly diverging and more heavily sclerotized, apices finely acute. Tegmen (
<figureCitation id="BEFFE09CFF8F5C56A3642E96FEC6F943" box="[275,356,1708,1729]" captionStart="Figs" captionStartId="6.[175,215,1327,1346]" captionTargetBox="[158,1171,191,1291]" captionTargetId="figure-163@6.[151,1171,191,1303]" captionTargetPageId="6" captionText="Figs. 13–14. Male terminalia of Melathra huyenae. 13) Spiculum gastrale; (14) Tegmen and aedeagus, ventral and lateral views. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085413" httpUri="https://zenodo.org/record/10085413/files/figure.png" pageId="7" pageNumber="359">Fig. 14</figureCitation>
) more than 1.5X length of median lobe; tegminal apodeme slender, slightly sinuate; basal piece posteriorly Y-shaped. Aedeagus (
<figureCitation id="BEFFE09CFF8F5C56A0BD28CDFCBBFE8E" box="[714,793,247,268]" captionStart="Figs" captionStartId="6.[175,215,1327,1346]" captionTargetBox="[158,1171,191,1291]" captionTargetId="figure-163@6.[151,1171,191,1303]" captionTargetPageId="6" captionText="Figs. 13–14. Male terminalia of Melathra huyenae. 13) Spiculum gastrale; (14) Tegmen and aedeagus, ventral and lateral views. Not illustrated to scale." figureDoi="http://doi.org/10.5281/zenodo.10085413" httpUri="https://zenodo.org/record/10085413/files/figure.png" pageId="7" pageNumber="359">Fig. 14</figureCitation>
) with median lobe (aedeagal pedon) length/width ratio 6.04–6.42; basiventral margin roundly emarginate; lateral margins subparallel, gradually roundly converging along posterior 1/6, apex narrowly rounded. Median lobe in lateral view slightly curved along anterior 3/5, thereafter outline straight to slightly undulate; width similar throughout, except for gradually narrowed posterior 1/6, along which dorsolateral margins (and to a lesser extent the dorsal mesal area) are densely covered with short, fine setae (see also Franz 2010a); apex triangularly narrowed, slightly deflected. Internal sac with variously plicate membranes, though lacking denticulate regions; with well-developed, long and anteriorly and posteriorly strongly sclerotized flagellum (see Wanat 2007; a membranous flagellum was observed in the majority of taxa studied in Franz 2011), anterior end of flagellum with knob-like expansion, midregion not sclerotized, yet posteriorly (along region of ostium) connected to 2 narrowly elongate, subparallel, undulating, plicate sclerites which terminate in plicate “head” region that is acutely narrowed and recurved by 180°; ostium large, elliptical, lateral margins invaginated. Aedeagal apodemes nearly 3/5 as long as median lobe, sclerotized, slender.
</paragraph>
</subSubSection>
<subSubSection id="6EDEAF92FF8F5C56A0AC2B90FBC7FC3D" box="[731,1125,937,959]" pageId="7" pageNumber="359" type="type_taxon">
<paragraph id="267BFC19FF8F5C56A0AC2B90FBC7FC3D" blockId="7.[707,1193,194,1172]" box="[731,1125,937,959]" pageId="7" pageNumber="359">
<emphasis id="14B0200BFF8F5C56A0AC2B90FCC1FC3D" bold="true" box="[731,867,938,959]" pageId="7" pageNumber="359">
<typeStatus id="F97F42BBFF8F5C56A0AC2B90FCA9FC3D" box="[731,779,938,959]" pageId="7" pageNumber="359">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="E1C4879AFF8F5C56A1062B93FBC7FC3D" ID-CoL="3ZDF6" authority="Franz." authorityName="Franz." box="[881,1125,937,959]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="359" phylum="Arthropoda" rank="species" species="huyenae">
<emphasis id="14B0200BFF8F5C56A1062B93FB82FC3C" box="[881,1056,937,958]" italics="true" pageId="7" pageNumber="359">Melathra huyenae</emphasis>
Franz.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="6EDEAF92FF8F5C56A0AC2BFEFCD8FB16" pageId="7" pageNumber="359" type="etymology">
<paragraph id="267BFC19FF8F5C56A0AC2BFEFCD8FB16" blockId="7.[707,1193,194,1172]" pageId="7" pageNumber="359">
<emphasis id="14B0200BFF8F5C56A0AC2BFEFCEFFC5B" bold="true" box="[731,845,964,985]" pageId="7" pageNumber="359">Etymology.</emphasis>
The genus name
<taxonomicName id="E1C4879AFF8F5C56A18F2BFEFBECFC5B" box="[1016,1102,964,985]" class="Insecta" family="Curculionidae" genus="Melathra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="359" phylum="Arthropoda" rank="genus">
<emphasis id="14B0200BFF8F5C56A18F2BFEFBECFC5B" box="[1016,1102,964,985]" italics="true" pageId="7" pageNumber="359">Melathra</emphasis>
</taxonomicName>
is a combination of the Ancient Greek words
<emphasis id="14B0200BFF8F5C56A6332BE5FBD9FC76" box="[1092,1147,991,1012]" italics="true" pageId="7" pageNumber="359">melas</emphasis>
and
<emphasis id="14B0200BFF8F5C56A0B42BC3FD5EFB8C" box="[707,764,1017,1038]" italics="true" pageId="7" pageNumber="359">lathra</emphasis>
(see
<bibRefCitation id="425581E8FF8F5C56A15F2BC0FC39FB8D" author="Brown" box="[808,923,1018,1039]" pageId="7" pageNumber="359" refId="ref4896" refString="Brown, R. W. 1956. Composition of Scientific Words, Revised Edition. Smithsonian Institution Press, Washington, D. C." type="book" year="1956">Brown 1956</bibRefCitation>
). The former signifies “black, dark” and refers to the dark habitus of the weevils, whereas the latter signifies “secretly, stealthy” and refers to the “mysterious character” of the genus, both taxonomically and in terms of life habits. The gender is feminine.
</paragraph>
</subSubSection>
</treatment>
</document>
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