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<document ID-DOI="http://doi.org/10.5281/zenodo.5618973" ID-GBIF-Dataset="53d83d32-58bd-4230-83cd-c0b6ee8fa560" ID-GBIF-Taxon="120621547" ID-Zenodo-Dep="5618973" checkinTime="1462380772451" checkinUser="plazi" docAuthor="Heike Feldhaar, Ulrich Maschwitz &amp; Brigitte Fiala" docDate="2016" docId="AE1287CFFF949C77A647C647FDD90FC9" docLanguage="en" docName="feldhaar_et_al_2016_sociobiology_crematogaster_macaranga_revision.pdf" docOrigin="Sociobiology 63 (1)" docStyle="DocumentStyle{}" docTitle="Crematogaster" docType="treatment" docVersion="10" lastPageId="8" lastPageNumber="659" masterDocId="522BFFB7FF939C7FA562C74AFFDC081F" masterDocTitle="Taxonomic Revision of the Obligate Plant-Ants of the Genus Crematogaster Lund (Hymenoptera: Formicidae: Myrmicinae), Associated with Macaranga Thouars (Euphorbiaceae) on Borneo and the Malay Peninsula" masterLastPageNumber="681" masterPageNumber="651" pageId="7" pageNumber="658" updateTime="1635526472509" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomic Revision of the Obligate Plant-Ants of the Genus Crematogaster Lund (Hymenoptera: Formicidae: Myrmicinae), Associated with Macaranga Thouars (Euphorbiaceae) on Borneo and the Malay Peninsula</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heike Feldhaar</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ulrich Maschwitz</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brigitte Fiala</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Sociobiology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>63</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>651</mods:start>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.13102/sociobiology.v63i1.949</mods:identifier>
<mods:identifier type="GBIF-Dataset">53d83d32-58bd-4230-83cd-c0b6ee8fa560</mods:identifier>
<mods:identifier type="Zenodo-Dep">269814</mods:identifier>
<mods:identifier type="ZooBank">28CD6B09-5459-4EA2-BC4E-5B14322F676D</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5618973" ID-GBIF-Taxon="120621547" ID-Zenodo-Dep="5618973" LSID="urn:lsid:plazi:treatment:AE1287CFFF949C77A647C647FDD90FC9" httpUri="http://treatment.plazi.org/id/AE1287CFFF949C77A647C647FDD90FC9" lastPageId="8" lastPageNumber="659" pageId="7" pageNumber="658">
<subSubSection pageId="7" pageNumber="658" type="nomenclature">
<paragraph blockId="7.[805,1459,269,363]" pageId="7" pageNumber="658">
<emphasis bold="true" box="[805,1176,269,294]" pageId="7" pageNumber="658">
Synonymic list of
<taxonomicName box="[1002,1160,270,294]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1002,1160,270,294]" italics="true" pageId="7" pageNumber="658">Crematogaster</emphasis>
</taxonomicName>
(
</emphasis>
<taxonomicName box="[1175,1450,270,294]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[1175,1450,270,294]" italics="true" pageId="7" pageNumber="658">Crematogaster borneensis</emphasis>
</taxonomicName>
- group
<emphasis bold="true" pageId="7" pageNumber="658">
) species from Borneo and the Malay Peninsula obligately associated with
<taxonomicName box="[1095,1220,339,362]" class="Magnoliopsida" family="Euphorbiaceae" genus="Macaranga" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="658" phylum="Tracheophyta" rank="genus">
<emphasis bold="true" box="[1095,1220,339,362]" italics="true" pageId="7" pageNumber="658">Macaranga</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="658" type="discussion">
<paragraph blockId="7.[805,1460,408,1993]" pageId="7" pageNumber="658">
In this study eight species of
<taxonomicName box="[1300,1459,409,432]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="genus">
<emphasis box="[1300,1459,409,432]" italics="true" pageId="7" pageNumber="658">Crematogaster</emphasis>
</taxonomicName>
(
<taxonomicName box="[814,1097,443,467]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[814,1097,443,467]" italics="true" pageId="7" pageNumber="658">Crematogaster borneensis</emphasis>
</taxonomicName>
-group) are recognized (including five new species), belonging to three species subgroups. Species subgroups were designated based on morphological characters of the queen caste, i.e. queens of the
<taxonomicName box="[1205,1310,547,571]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="decamera">
<emphasis box="[1205,1310,547,571]" italics="true" pageId="7" pageNumber="658">decamera</emphasis>
</taxonomicName>
-subgroup are smaller (WL ranging from 1.7 to 2.3 mm) and have relatively smaller compound eyes (EL&lt;0.5 mm) than queens of the
<taxonomicName box="[805,895,652,675]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="captiosa">
<emphasis box="[805,895,652,675]" italics="true" pageId="7" pageNumber="658">captiosa</emphasis>
</taxonomicName>
-subgroup (WL ranging from 2.2 to 2.9 mm and EL&gt; 0.5 mm) (
<bibRefCitation author="Feldhaar, H. &amp; Fiala, B. &amp; Gadau, J. &amp; Mohamed, M. &amp; Maschwitz, U." box="[932,1169,686,710]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="7" pageNumber="658" pagination="441 - 452" part="27" publicationUrl="http://dx.doi.org/ 10.1016/S1055-" refString="Feldhaar, H., Fiala, B., Gadau, J., Mohamed, M. &amp; Maschwitz, U. (2003 a). Molecular phylogeny of Crematogaster subgenus Decacrema ants (Hymenoptera: Formicidae) and the colonization of Macaranga (Euphorbiaceae) trees. Molecular Phylogenetics and Evolution, 27: 441 - 452. doi: 10.1016 / S 1055 - 7903 (02) 00449 - 9" title="Molecular phylogeny of Crematogaster subgenus Decacrema ants (Hymenoptera: Formicidae) and the colonization of Macaranga (Euphorbiaceae) trees" type="journal article" year="2003" yearSuffix="a">Feldhaar et al., 2003a</bibRefCitation>
;
<bibRefCitation author="Feldhaar, H. &amp; Gadau, J. &amp; Fiala, B." box="[1180,1402,686,710]" editor="M. Glaubrecht" journalOrPublisher="Berlin: Springer" pageId="7" pageNumber="658" pagination="193 - 213" refString="Feldhaar, H., Gadau, J. &amp; Fiala, B. (2010). Speciation in obligately plant-associated Crematogaster ants: Hostdistribution rather than adaption towards specific hosts drives the process. In M. Glaubrecht &amp; H. Schneider (Eds.), Evolution in action - Adaptive radiations and the origins of biodiversity (pp. 193 - 213). Berlin: Springer." title="Speciation in obligately plant-associated Crematogaster ants: Hostdistribution rather than adaption towards specific hosts drives the process" type="book chapter" volumeTitle="Evolution in action" year="2010">Feldhaar et al., 2010</bibRefCitation>
) (see Table S.1 for comparison of morphospecies designations used by our group and synonymized or new species names).
</paragraph>
<paragraph blockId="7.[805,1460,408,1993]" pageId="7" pageNumber="658">
All workers of
<taxonomicName box="[1028,1177,790,813]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="genus">
<emphasis box="[1028,1177,790,813]" italics="true" pageId="7" pageNumber="658">Crematogaster</emphasis>
</taxonomicName>
(
<taxonomicName box="[1189,1450,790,814]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[1189,1450,790,814]" italics="true" pageId="7" pageNumber="658">Crematogaster borneensis</emphasis>
</taxonomicName>
- group) workers including those associated with
<taxonomicName box="[1332,1459,825,848]" class="Magnoliopsida" family="Euphorbiaceae" genus="Macaranga" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="658" phylum="Tracheophyta" rank="genus">
<emphasis box="[1332,1459,825,848]" italics="true" pageId="7" pageNumber="658">Macaranga</emphasis>
</taxonomicName>
have ten antennal segments. The workers of the
<taxonomicName box="[1323,1450,860,883]" class="Magnoliopsida" family="Euphorbiaceae" genus="Macaranga" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="658" phylum="Tracheophyta" rank="genus">
<emphasis box="[1323,1450,860,883]" italics="true" pageId="7" pageNumber="658">Macaranga</emphasis>
</taxonomicName>
- associated
<taxonomicName box="[923,1085,894,917]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="genus">
<emphasis box="[923,1085,894,917]" italics="true" pageId="7" pageNumber="658">Crematogaster</emphasis>
</taxonomicName>
species are all monomorphic. This is in contrast to the other three described species belonging to the former
<taxonomicName box="[968,1092,964,987]" class="Insecta" family="Formicidae" genus="Decacrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="genus">
<emphasis box="[968,1092,964,987]" italics="true" pageId="7" pageNumber="658">Decacrema</emphasis>
</taxonomicName>
from SE-Asia (
<taxonomicName box="[1279,1416,963,987]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="angulosa">
<emphasis box="[1279,1416,963,987]" italics="true" pageId="7" pageNumber="658">C. angulosa</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="biformis">
<emphasis italics="true" pageId="7" pageNumber="658">C. biformis</emphasis>
</taxonomicName>
and
<taxonomicName box="[949,1097,998,1022]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="cephalotes">
<emphasis box="[949,1097,998,1022]" italics="true" pageId="7" pageNumber="658">C. cephalotes</emphasis>
</taxonomicName>
) that all have dimorphic workers (see Supplementary Fig S2.1A to S2.3B). Body size may vary strongly within species although this is dependent on colony size (e.g. TL ranges from ~ 2.1 mm to 3.3 mm in
<taxonomicName box="[847,1123,1137,1160]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="captiosa">
<emphasis box="[847,1123,1137,1160]" italics="true" pageId="7" pageNumber="658">Crematogaster captiosa</emphasis>
</taxonomicName>
workers). Thus, especially in workers relative measures (e.g. SI or RLEG) are more important characters than absolute measurements (e.g. SL or LHT) (see overview of most important distinguishing characters of workers in
<tableCitation box="[1123,1215,1275,1299]" captionStart="Table 1" captionStartId="8.[130,193,131,155]" captionTargetBox="[130,1482,181,1302]" captionTargetPageId="8" captionText="Table 1. Overview of most distinct morphological characters of the Macaranga - associated Crematogaster (borneensis - group) species." httpUri="http://table.plazi.org/id/72C46651FF9B9C77A5E0C7C9FA160884" pageId="7" pageNumber="658" tableUuid="72C46651FF9B9C77A5E0C7C9FA160884">Table 1</tableCitation>
). Workers collected from small colonies are also lighter in colour. In queens shade of body coloration is a useful character, as are absolute measures. If possible, queens should be used for determination as worker characteristics are sometimes less distinctive (see overview of most important characters distinguishing queens in
<tableCitation box="[1077,1157,1483,1507]" captionStart="Table 1" captionStartId="8.[130,193,131,155]" captionTargetBox="[130,1482,181,1302]" captionTargetPageId="8" captionText="Table 1. Overview of most distinct morphological characters of the Macaranga - associated Crematogaster (borneensis - group) species." httpUri="http://table.plazi.org/id/72C46651FF9B9C77A5E0C7C9FA160884" pageId="7" pageNumber="658" tableUuid="72C46651FF9B9C77A5E0C7C9FA160884">Table 1</tableCitation>
).
</paragraph>
<paragraph blockId="7.[805,1460,408,1993]" pageId="7" pageNumber="658">
Most of the diversity of the species is apportioned between the
<taxonomicName box="[944,1049,1552,1576]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="decamera">
<emphasis box="[944,1049,1552,1576]" italics="true" pageId="7" pageNumber="658">decamera</emphasis>
</taxonomicName>
-subgroup and the
<taxonomicName box="[1254,1344,1553,1576]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="captiosa">
<emphasis box="[1254,1344,1553,1576]" italics="true" pageId="7" pageNumber="658">captiosa</emphasis>
</taxonomicName>
-subgroup. The placement of
<taxonomicName box="[1013,1138,1587,1611]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="maryatii">
<emphasis box="[1013,1031,1587,1610]" italics="true" pageId="7" pageNumber="658">C</emphasis>
.
<emphasis box="[1048,1138,1588,1611]" italics="true" pageId="7" pageNumber="658">maryatii</emphasis>
</taxonomicName>
is unclear. In the molecular phylogeny based on mitochondrial DNA this species does not cluster with either group and came out most basal. Based on nuclear DNA (elongation factor 1α) (
<bibRefCitation author="Feldhaar, H. &amp; Gadau, J. &amp; Fiala, B." editor="M. Glaubrecht" journalOrPublisher="Berlin: Springer" pageId="7" pageNumber="658" pagination="193 - 213" refString="Feldhaar, H., Gadau, J. &amp; Fiala, B. (2010). Speciation in obligately plant-associated Crematogaster ants: Hostdistribution rather than adaption towards specific hosts drives the process. In M. Glaubrecht &amp; H. Schneider (Eds.), Evolution in action - Adaptive radiations and the origins of biodiversity (pp. 193 - 213). Berlin: Springer." title="Speciation in obligately plant-associated Crematogaster ants: Hostdistribution rather than adaption towards specific hosts drives the process" type="book chapter" volumeTitle="Evolution in action" year="2010">Feldhaar et al., 2010</bibRefCitation>
) this species is not separated from species of the
<taxonomicName box="[805,895,1761,1784]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="captiosa">
<emphasis box="[805,895,1761,1784]" italics="true" pageId="7" pageNumber="658">captiosa</emphasis>
</taxonomicName>
-group. However, queen morphology and life-history characters resemble that of the
<taxonomicName box="[1153,1258,1795,1819]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="decamera">
<emphasis box="[1153,1258,1795,1819]" italics="true" pageId="7" pageNumber="658">decamera</emphasis>
</taxonomicName>
-group. Small size of queens and workers and early onset of reproduction may be a convergent trait though as this seems to enable usage of less productive and slow growing host plants (
<bibRefCitation author="Feldhaar, H. &amp; Gadau, J. &amp; Fiala, B." editor="M. Glaubrecht" journalOrPublisher="Berlin: Springer" pageId="7" pageNumber="658" pagination="193 - 213" refString="Feldhaar, H., Gadau, J. &amp; Fiala, B. (2010). Speciation in obligately plant-associated Crematogaster ants: Hostdistribution rather than adaption towards specific hosts drives the process. In M. Glaubrecht &amp; H. Schneider (Eds.), Evolution in action - Adaptive radiations and the origins of biodiversity (pp. 193 - 213). Berlin: Springer." title="Speciation in obligately plant-associated Crematogaster ants: Hostdistribution rather than adaption towards specific hosts drives the process" type="book chapter" volumeTitle="Evolution in action" year="2010">Feldhaar et al. 2010</bibRefCitation>
). Therefore, we currently do not place
<taxonomicName box="[1302,1427,1934,1957]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="658" phylum="Arthropoda" rank="species" species="maryatii">
<emphasis box="[1302,1320,1934,1957]" italics="true" pageId="7" pageNumber="658">C</emphasis>
.
<emphasis box="[1337,1427,1934,1957]" italics="true" pageId="7" pageNumber="658">maryatii</emphasis>
</taxonomicName>
in either group.
</paragraph>
</subSubSection>
<caption ID-Table-UUID="72C46651FF9B9C77A5E0C7C9FA160884" box="[130,1482,131,155]" httpUri="http://table.plazi.org/id/72C46651FF9B9C77A5E0C7C9FA160884" pageId="8" pageNumber="659" targetBox="[130,1482,181,1302]" targetIsTable="true" targetPageId="8">
<paragraph blockId="8.[130,1482,131,155]" box="[130,1482,131,155]" pageId="8" pageNumber="659">
<emphasis bold="true" box="[130,210,131,155]" pageId="8" pageNumber="659">Table 1</emphasis>
. Overview of most distinct morphological characters of the
<taxonomicName box="[807,927,132,155]" class="Magnoliopsida" family="Euphorbiaceae" genus="Macaranga" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="659" phylum="Tracheophyta" rank="genus">
<emphasis box="[807,927,132,155]" italics="true" pageId="8" pageNumber="659">Macaranga</emphasis>
</taxonomicName>
-associated
<taxonomicName box="[1044,1197,131,154]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="genus">
<emphasis box="[1044,1197,131,154]" italics="true" pageId="8" pageNumber="659">Crematogaster</emphasis>
</taxonomicName>
(
<taxonomicName box="[1210,1320,131,155]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[1210,1320,131,155]" italics="true" pageId="8" pageNumber="659">borneensis</emphasis>
</taxonomicName>
-group) species.
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="659">
<table box="[130,1482,181,1302]" gridcols="3" gridrows="9" pageId="8" pageNumber="659">
<tr box="[130,1482,181,203]" gridrow="0" pageId="8" pageNumber="659">
<emphasis bold="true" box="[139,1221,181,203]" pageId="8" pageNumber="659">
<th box="[130,289,181,203]" gridcol="0" gridrow="0" pageId="8" pageNumber="659">Species</th>
<th box="[421,976,181,203]" gridcol="1" gridrow="0" pageId="8" pageNumber="659">Worker characters</th>
<th box="[1039,1482,181,203]" gridcol="2" gridrow="0" pageId="8" pageNumber="659">Queen characters</th>
</emphasis>
</tr>
<tr box="[130,1482,222,332]" gridrow="1" pageId="8" pageNumber="659">
<th box="[130,289,222,332]" gridcol="0" gridrow="1" pageId="8" pageNumber="659">
<taxonomicName box="[139,273,266,287]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[139,155,266,287]" italics="true" pageId="8" pageNumber="659">C</emphasis>
.
<emphasis box="[168,273,266,287]" italics="true" pageId="8" pageNumber="659">borneensis</emphasis>
</taxonomicName>
</th>
<td box="[421,976,222,332]" gridcol="1" gridrow="1" pageId="8" pageNumber="659">Propodeal spines present Scape index (SI)&gt; 0.65 RLEG &lt;0.7, rarely above</td>
<td box="[1039,1482,222,332]" gridcol="2" gridrow="1" pageId="8" pageNumber="659">EL&gt; 0.5 mm (to 0.6mm) REL 0.36 - 0.43 OD1&gt; OW Head longer than wide (CI &lt;0.95)</td>
</tr>
<tr box="[130,1482,348,458]" gridrow="2" pageId="8" pageNumber="659">
<th box="[130,289,348,458]" gridcol="0" gridrow="2" pageId="8" pageNumber="659">
<taxonomicName box="[139,279,392,414]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="linsenmairi">
<emphasis box="[139,155,393,414]" italics="true" pageId="8" pageNumber="659">C</emphasis>
.
<emphasis box="[168,279,392,413]" italics="true" pageId="8" pageNumber="659">linsenmairi</emphasis>
</taxonomicName>
</th>
<td box="[421,976,348,458]" gridcol="1" gridrow="2" pageId="8" pageNumber="659">Propodeal spines present Scape index (SI)&gt; 0.65 RLEG&gt; 0.7, rarely below</td>
<td box="[1039,1482,348,458]" gridcol="2" gridrow="2" pageId="8" pageNumber="659">EL rarely &lt;0.65mm REL 0.39 - 0.48 OD1 &lt;OW Scape length&gt;0.75 mm</td>
</tr>
<tr box="[130,1482,474,584]" gridrow="3" pageId="8" pageNumber="659">
<th box="[130,289,474,584]" gridcol="0" gridrow="3" pageId="8" pageNumber="659">
<taxonomicName box="[139,244,518,540]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="hullettii">
<emphasis box="[139,155,519,540]" italics="true" pageId="8" pageNumber="659">C</emphasis>
.
<emphasis box="[168,244,518,539]" italics="true" pageId="8" pageNumber="659">hullettii</emphasis>
</taxonomicName>
</th>
<td box="[421,976,474,584]" gridcol="1" gridrow="3" pageId="8" pageNumber="659">Propodeal spines present Scape index (SI) &lt;0.65 Head as wide as long (CI ~ 1.0) Legs relatively short, RLEG 0.58 to 0.62</td>
<td box="[1039,1482,474,584]" gridcol="2" gridrow="3" pageId="8" pageNumber="659">EL 0.39 0.47 mm (rarely&gt; 0.45) REL 0.32-0.4 OD1&gt; OW ROD between 0.09 - 0.13</td>
</tr>
<tr box="[130,1482,600,710]" gridrow="4" pageId="8" pageNumber="659">
<th box="[130,289,600,710]" gridcol="0" gridrow="4" pageId="8" pageNumber="659">
<taxonomicName box="[139,250,645,666]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="captiosa">
<emphasis box="[139,250,645,666]" italics="true" pageId="8" pageNumber="659">C. captiosa</emphasis>
</taxonomicName>
</th>
<td box="[421,976,600,710]" gridcol="1" gridrow="4" pageId="8" pageNumber="659">Propodeal spines present Scape index (SI) &lt;0.65 RLEG 0.61 to 0.69</td>
<td box="[1039,1482,600,710]" gridcol="2" gridrow="4" pageId="8" pageNumber="659">EL ~ 0.6mm REL 0.38 - 0.48 OD1 &lt;OW Scape length usually &lt;0.75 mm</td>
</tr>
<tr box="[130,1482,726,894]" gridrow="5" pageId="8" pageNumber="659">
<th box="[130,289,726,894]" gridcol="0" gridrow="5" pageId="8" pageNumber="659">
<taxonomicName box="[139,264,799,821]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="decamera">
<emphasis box="[139,264,799,821]" italics="true" pageId="8" pageNumber="659">C. decamera</emphasis>
</taxonomicName>
</th>
<td box="[421,976,726,894]" gridcol="1" gridrow="5" pageId="8" pageNumber="659">Propodeal spines absent Legs relatively long (RLEG 0.71 - 0.85) Head slightly longer than wide (CI 0.94 - 1.02) Postpetiole always wider than petiole (PI &lt;1.0)</td>
<td box="[1039,1482,726,894]" gridcol="2" gridrow="5" pageId="8" pageNumber="659">EL 0.39 - 0.45mm REL 0.29 - 0.35 OD1&gt; OW SL&gt; 0.7 mm CI usually&gt; 1.0 PI &lt;1.0</td>
</tr>
<tr box="[130,1482,911,1050]" gridrow="6" pageId="8" pageNumber="659">
<th box="[130,289,911,1050]" gridcol="0" gridrow="6" pageId="8" pageNumber="659">
<taxonomicName box="[139,250,970,991]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="maryatii">
<emphasis box="[139,250,970,991]" italics="true" pageId="8" pageNumber="659">C. maryatii</emphasis>
</taxonomicName>
</th>
<td box="[421,976,911,1050]" gridcol="1" gridrow="6" pageId="8" pageNumber="659">Very short propodeal spines, sometimes absent Scape index (SI)&gt; 0.65 Head slightly longer than wide (CI 0.94 - 1.02) Petiole as wide or wider than postpetiole, PI&gt; 0.98 RLEG usually &lt;0.7</td>
<td box="[1039,1482,911,1050]" gridcol="2" gridrow="6" pageId="8" pageNumber="659">EL 0.34 - 0.38mm REL 0.28-0.34 OD1&gt; OW PI usually&gt; 1.0 (to 1.12)</td>
</tr>
<tr box="[130,1482,1066,1176]" gridrow="7" pageId="8" pageNumber="659">
<th box="[130,289,1066,1176]" gridcol="0" gridrow="7" pageId="8" pageNumber="659">
<taxonomicName box="[139,289,1110,1131]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="roslihashimi">
<emphasis box="[139,289,1110,1131]" italics="true" pageId="8" pageNumber="659">C. roslihashimi</emphasis>
</taxonomicName>
</th>
<td box="[421,976,1066,1176]" gridcol="1" gridrow="7" pageId="8" pageNumber="659">Propodeal spines absent Scape index (SI)&gt; 0.7 Head distinctly longer than wide (CI: 0.91) Legs realtively long (RLEG 0.71-0.73)</td>
<td box="[1039,1482,1066,1176]" gridcol="2" gridrow="7" pageId="8" pageNumber="659">EL 0.3 0.32; REL &lt;0.28 OD1&gt; OW Elongated head (CI &lt;0.9) PI ≤ 1.0</td>
</tr>
<tr box="[130,1482,1192,1302]" gridrow="8" pageId="8" pageNumber="659">
<th box="[130,289,1192,1302]" gridcol="0" gridrow="8" pageId="8" pageNumber="659">
<taxonomicName box="[139,252,1236,1257]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="claudiae">
<emphasis box="[139,252,1236,1257]" italics="true" pageId="8" pageNumber="659">C. claudiae</emphasis>
</taxonomicName>
</th>
<td box="[421,976,1192,1302]" gridcol="1" gridrow="8" pageId="8" pageNumber="659">Propodeal spines usually absent, when present very short Petiole distinctly wider than postpetiole (PI&gt; 1.15) Legs and scapi short, RLEG &lt;0.69 and SI &lt;0.65</td>
<td box="[1039,1482,1192,1302]" gridcol="2" gridrow="8" pageId="8" pageNumber="659">EL 0.45 - 0.5 mm (REL 0.36-0.39) OD1&gt; OW PI&gt; 1.0 ROD between 0.13 - 0.17</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="8" pageNumber="659" type="synonymic_list">
<paragraph blockId="8.[129,478,1345,1369]" box="[129,478,1345,1369]" pageId="8" pageNumber="659">
<taxonomicName box="[129,389,1345,1369]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="maryatii" status="sp. nov.">
<emphasis bold="true" box="[129,389,1345,1369]" italics="true" pageId="8" pageNumber="659">Crematogaster maryatii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[395,478,1346,1369]" pageId="8" pageNumber="659" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,570,1404,1532]" box="[129,383,1404,1428]" pageId="8" pageNumber="659">
<heading bold="true" box="[129,383,1404,1428]" fontSize="10" level="3" pageId="8" pageNumber="659" reason="6">
<emphasis bold="true" box="[129,383,1404,1428]" pageId="8" pageNumber="659">
<taxonomicName box="[129,267,1404,1428]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="decamera">
<emphasis bold="true" box="[129,267,1404,1428]" italics="true" pageId="8" pageNumber="659">C. decamera</emphasis>
</taxonomicName>
-subgroup
</emphasis>
</heading>
</paragraph>
<paragraph blockId="8.[129,570,1404,1532]" box="[166,570,1438,1463]" pageId="8" pageNumber="659">
<taxonomicName authority="Forel, 1910" authorityName="Forel" authorityYear="1910" box="[166,570,1438,1463]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="decamera">
<emphasis bold="true" box="[166,441,1438,1463]" italics="true" pageId="8" pageNumber="659">Crematogaster decamera</emphasis>
<bibRefCitation author="Forel, A." box="[448,570,1439,1463]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="8" pageNumber="659" pagination="6 - 32" part="54" refString="Forel, A. (1910). Glanures myrmecologiques. Annales de la Societe Entomologique de Belgique, 54: 6 - 32." title="Glanures myrmecologiques" type="journal article" year="1910">Forel, 1910</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="8.[129,570,1404,1532]" box="[166,510,1473,1497]" pageId="8" pageNumber="659">
<taxonomicName box="[166,420,1473,1497]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="hullettii" status="sp. nov.">
<emphasis bold="true" box="[166,420,1473,1497]" italics="true" pageId="8" pageNumber="659">Crematogaster hullettii</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[427,510,1474,1497]" pageId="8" pageNumber="659" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,570,1404,1532]" box="[166,562,1508,1532]" pageId="8" pageNumber="659">
<taxonomicName box="[166,472,1508,1532]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="roslihashimi" status="sp. nov.">
<emphasis bold="true" box="[166,472,1508,1532]" italics="true" pageId="8" pageNumber="659">Crematogaster roslihashimi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[479,562,1509,1532]" pageId="8" pageNumber="659" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[129,367,1567,1591]" pageId="8" pageNumber="659">
<heading bold="true" box="[129,367,1567,1591]" fontSize="10" level="3" pageId="8" pageNumber="659" reason="6">
<emphasis bold="true" box="[129,367,1567,1591]" pageId="8" pageNumber="659">
<taxonomicName box="[129,250,1567,1591]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="captiosa">
<emphasis bold="true" box="[129,250,1567,1591]" italics="true" pageId="8" pageNumber="659">C. captiosa</emphasis>
</taxonomicName>
-subgroup
</emphasis>
</heading>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,593,1601,1625]" pageId="8" pageNumber="659">
<taxonomicName authority="Andre, 1896" authorityName="Andre" authorityYear="1896" box="[166,593,1601,1625]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis bold="true" box="[166,453,1601,1625]" italics="true" pageId="8" pageNumber="659">Crematogaster borneensis</emphasis>
<bibRefCitation author="Andre, E." box="[460,593,1601,1625]" journalOrPublisher="Revue d'Entomologie (Caen)" pageId="8" pageNumber="659" pagination="251 - 265" part="15" refString="Andre, E. (1896). Fourmis nouvelles d'Asie et d'Australie. Revue d'Entomologie (Caen), 15: 251 - 265." title="Fourmis nouvelles d'Asie et d'Australie" type="journal article" year="1896">André, 1896</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,710,1636,1661]" pageId="8" pageNumber="659">
=
<taxonomicName box="[188,335,1636,1660]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[188,335,1636,1660]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
subsp.
<emphasis box="[415,479,1637,1660]" italics="true" pageId="8" pageNumber="659">capax</emphasis>
<bibRefCitation author="Forel, A." box="[486,607,1636,1660]" journalOrPublisher="Revue Suisse de Zoologie" pageId="8" pageNumber="659" pagination="23 - 62" part="19" refString="Forel, A. (1911). Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo. Revue Suisse de Zoologie, 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo" type="journal article" year="1911">Forel, 1911</bibRefCitation>
syn. nov.
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,703,1671,1695]" pageId="8" pageNumber="659">
=
<taxonomicName box="[188,335,1671,1695]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[188,335,1671,1695]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
subsp.
<taxonomicName authority="Forel, 1911" authorityName="Forel" authorityYear="1911" box="[415,600,1671,1695]" class="Magnoliopsida" family="Euphorbiaceae" genus="Macaranga" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="659" phylum="Tracheophyta" rank="species" species="hosei" status="syn. nov.">
<emphasis box="[415,471,1671,1695]" italics="true" pageId="8" pageNumber="659">hosei</emphasis>
<bibRefCitation author="Forel, A." box="[478,600,1671,1695]" journalOrPublisher="Revue Suisse de Zoologie" pageId="8" pageNumber="659" pagination="23 - 62" part="19" refString="Forel, A. (1911). Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo. Revue Suisse de Zoologie, 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo" type="journal article" year="1911">Forel, 1911</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel box="[607,703,1671,1695]" pageId="8" pageNumber="659">syn. nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,756,1705,1730]" pageId="8" pageNumber="659">
=
<taxonomicName box="[188,335,1705,1729]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[188,335,1705,1729]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
subsp.
<emphasis box="[415,525,1705,1729]" italics="true" pageId="8" pageNumber="659">sembilana</emphasis>
<bibRefCitation author="Forel, A." box="[532,653,1705,1730]" journalOrPublisher="Revue Suisse de Zoologie" pageId="8" pageNumber="659" pagination="23 - 62" part="19" refString="Forel, A. (1911). Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo. Revue Suisse de Zoologie, 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo" type="journal article" year="1911">Forel, 1911</bibRefCitation>
syn. nov.
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,783,1740,1765]" pageId="8" pageNumber="659">
=
<taxonomicName box="[186,329,1740,1764]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[186,329,1740,1764]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
<taxonomicName authority="Viehmeyer, 1916" authorityName="Viehmeyer" authorityYear="1916" box="[334,690,1740,1764]" pageId="8" pageNumber="659" rank="variety" status="syn.nov." variety="macarangae">
var.
<emphasis box="[377,508,1741,1764]" italics="true" pageId="8" pageNumber="659">macarangae</emphasis>
<bibRefCitation author="Viehmeyer, H." box="[513,690,1740,1764]" journalOrPublisher="Beobachtet und gesammelt von H. Overbeck. Archiv fur Naturgeschichte, 81 (A" pageId="8" pageNumber="659" pagination="108 - 168" part="1915" refString="Viehmeyer, H. (1916). Ameisen von Singapore. Beobachtet und gesammelt von H. Overbeck. Archiv fur Naturgeschichte, 81 (A 8, 1915): 108 - 168." title="Ameisen von Singapore" type="journal article" year="1916">Viehmeyer, 1916</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel box="[696,783,1741,1765]" pageId="8" pageNumber="659">syn.nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,554,1775,1799]" pageId="8" pageNumber="659">
<taxonomicName authority="Forel, 1910" authorityName="Forel" authorityYear="1910" box="[166,554,1775,1799]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="captiosa">
<emphasis bold="true" box="[166,424,1775,1799]" italics="true" pageId="8" pageNumber="659">Crematogaster captiosa</emphasis>
<bibRefCitation author="Forel, A." box="[431,554,1775,1799]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="8" pageNumber="659" pagination="6 - 32" part="54" refString="Forel, A. (1910). Glanures myrmecologiques. Annales de la Societe Entomologique de Belgique, 54: 6 - 32." title="Glanures myrmecologiques" type="journal article" year="1910">Forel, 1910</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,702,1809,1834]" pageId="8" pageNumber="659">
=
<taxonomicName box="[188,335,1809,1833]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[188,335,1809,1833]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
<taxonomicName authority="Forel, 1911" authorityName="Forel" authorityYear="1911" box="[341,598,1809,1834]" pageId="8" pageNumber="659" rank="variety" status="syn. nov." variety="harpyia">
var.
<emphasis box="[387,470,1809,1833]" italics="true" pageId="8" pageNumber="659">harpyia</emphasis>
<bibRefCitation author="Forel, A." box="[477,598,1809,1834]" journalOrPublisher="Revue Suisse de Zoologie" pageId="8" pageNumber="659" pagination="23 - 62" part="19" refString="Forel, A. (1911). Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo. Revue Suisse de Zoologie, 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo" type="journal article" year="1911">Forel, 1911</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel box="[606,702,1810,1834]" pageId="8" pageNumber="659">syn. nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,694,1844,1869]" pageId="8" pageNumber="659">
=
<taxonomicName box="[188,335,1844,1868]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[188,335,1844,1868]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
<taxonomicName authority="Forel, 1911" authorityName="Forel" authorityYear="1911" box="[341,591,1844,1868]" pageId="8" pageNumber="659" rank="variety" status="syn. nov." variety="insulsa">
var.
<emphasis box="[387,463,1844,1868]" italics="true" pageId="8" pageNumber="659">insulsa</emphasis>
<bibRefCitation author="Forel, A." box="[469,591,1844,1868]" journalOrPublisher="Revue Suisse de Zoologie" pageId="8" pageNumber="659" pagination="23 - 62" part="19" refString="Forel, A. (1911). Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo. Revue Suisse de Zoologie, 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo" type="journal article" year="1911">Forel, 1911</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel box="[598,694,1845,1869]" pageId="8" pageNumber="659">syn. nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,722,1879,1903]" pageId="8" pageNumber="659">
=
<taxonomicName box="[188,335,1879,1903]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[188,335,1879,1903]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
subsp.
<emphasis box="[415,491,1879,1903]" italics="true" pageId="8" pageNumber="659">symbia</emphasis>
<bibRefCitation author="Forel, A." box="[498,619,1879,1903]" journalOrPublisher="Revue Suisse de Zoologie" pageId="8" pageNumber="659" pagination="23 - 62" part="19" refString="Forel, A. (1911). Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo. Revue Suisse de Zoologie, 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo" type="journal article" year="1911">Forel, 1911</bibRefCitation>
syn. nov.
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,716,1913,1938]" pageId="8" pageNumber="659">
=
<taxonomicName box="[188,335,1913,1937]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="borneensis">
<emphasis box="[188,335,1913,1937]" italics="true" pageId="8" pageNumber="659">C. borneensis</emphasis>
</taxonomicName>
subsp.
<emphasis box="[415,485,1914,1937]" italics="true" pageId="8" pageNumber="659">novem</emphasis>
<bibRefCitation author="Forel, A." box="[492,613,1913,1938]" journalOrPublisher="Revue Suisse de Zoologie" pageId="8" pageNumber="659" pagination="23 - 62" part="19" refString="Forel, A. (1911). Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo. Revue Suisse de Zoologie, 19: 23 - 62." title="Fourmis de Borneo, Singapore, Ceylan, etc. recoltees par MM. Haviland, Green, Winkler, Will, Hose, Roepke et Waldo" type="journal article" year="1911">Forel, 1911</bibRefCitation>
syn. nov.
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,550,1948,1972]" pageId="8" pageNumber="659">
<taxonomicName box="[166,460,1948,1972]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="linsenmairi" status="sp. nov.">
<emphasis bold="true" box="[166,460,1948,1972]" italics="true" pageId="8" pageNumber="659">Crematogaster linsenmairi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[467,550,1949,1972]" pageId="8" pageNumber="659" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="8.[129,783,1567,2007]" box="[166,517,1982,2007]" pageId="8" pageNumber="659">
<taxonomicName box="[166,427,1982,2007]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="659" phylum="Arthropoda" rank="species" species="claudiae" status="sp. nov.">
<emphasis bold="true" box="[166,427,1982,2007]" italics="true" pageId="8" pageNumber="659">Crematogaster claudiae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[434,517,1983,2006]" pageId="8" pageNumber="659" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>