310 lines
40 KiB
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310 lines
40 KiB
XML
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<mods:title id="C039516B72190BFB40B537C1180EA85D">A revision of Chrysopidae (Neuroptera) from the late Eocene Florissant Formation Colorado, with description of new species</mods:title>
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<mods:namePart id="B98296B3E50000F194F5D4CCA755578F">Makarkin, Vladimir N.</mods:namePart>
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<mods:affiliation id="671CD020AFD5D5561C6DECBC15345D05">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia</mods:affiliation>
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<mods:namePart id="CD52A756E6636E8CDCFB1599D49924B9">Antell, Gwen S.</mods:namePart>
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<mods:affiliation id="B519FCDD763D98E96C761687A0933A12">Department of Earth Sciences, University of Oxford, Oxford OX 1 3 AN, United Kingdom</mods:affiliation>
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<mods:namePart id="3F27524EE78A1D683E2378A4AA71939F">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="37CAE791884C8675365AB78C4A728FF5">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada sba 48 @ sfu. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:date id="E45298EE460858325DAF61C23680E7C5">2022</mods:date>
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Genus
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<taxonomicName id="E2DF4D1BFFECFFE0E2EBA3D1CC0C75BB" ID-CoL="ZW2" authority="Adams, 1967" authorityName="Adams" authorityYear="1967" box="[238,588,439,466]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
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<emphasis id="17ABEA8AFFECFFE0E2EBA3D1CFE575B8" bold="true" box="[238,421,439,466]" italics="true" pageId="2" pageNumber="303">Archaeochrysa</emphasis>
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<bibRefCitation id="414E4B69FFECFFE0E3AEA3D1CC0C75BB" author="Adams, P. A." box="[427,588,439,465]" pageId="2" pageNumber="303" pagination="215 - 238" refId="ref21987" refString="Adams, P. A. (1967) A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). Bulletin of the Museum of Comparative Zoology, 135, 215 - 238." type="journal article" year="1967">Adams, 1967</bibRefCitation>
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<taxonomicName id="E2DF4D1BFFECFFE0E292A39ACFB5767E" ID-CoL="ZW2" authority="Adams, 1967: 217" authorityName="Adams" authorityPageNumber="217" authorityYear="1967" box="[151,501,507,532]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
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<emphasis id="17ABEA8AFFECFFE0E292A39ACF6F767E" box="[151,303,508,532]" italics="true" pageId="2" pageNumber="303">Archaeochrysa</emphasis>
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<bibRefCitation id="414E4B69FFECFFE0E333A39ACFB5767E" author="Adams, P. A." box="[310,501,507,532]" pageId="2" pageNumber="303" pagination="215 - 238" refId="ref21987" refString="Adams, P. A. (1967) A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). Bulletin of the Museum of Comparative Zoology, 135, 215 - 238." type="journal article" year="1967">Adams, 1967: 217</bibRefCitation>
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, 219, 220, 229; Schlüter 1984: 7;
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<treatmentCitation id="A47E1089FFECFFE0E153A39ACA0B767E" author="Yang, C. K. & Hong, Y. C." box="[854,1099,508,532]" page="20" pageId="2" pageNumber="303" year="1990">
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<bibRefCitation id="414E4B69FFECFFE0E153A39ACA0B767E" author="Yang, C. K. & Hong, Y. C." box="[854,1099,508,532]" pageId="2" pageNumber="303" pagination="15 - 26" refId="ref26487" refString="Yang, C. K. & Hong, Y. C. (1990) Drakochrysa, an Early Cretaceous new genus of Chrysopidae (Insecta: Neuroptera) from Laiyang Basin, Shandong Province. Geoscience, 4 (4), 15 - 26, 1 pl." type="journal article" year="1990">Yang & Hong 1990: 20</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E653A39ACBD87679" author="Willmann, R. & Brooks, S. J." box="[1110,1432,507,532]" page="126" pageId="2" pageNumber="303" year="1991">
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<bibRefCitation id="414E4B69FFECFFE0E653A39ACBD87679" author="Willmann, R. & Brooks, S. J." box="[1110,1432,507,532]" pageId="2" pageNumber="303" pagination="125 - 135" refId="ref26306" refString="Willmann, R. & Brooks, S. J. (1991) Insekten aus der Fur-Formation von Danemark (Moler, ob. Paleozan / unt. Eozan?). 6. Chrysopidae (Neuroptera). Meyniana, 43, 125 - 135." type="journal article" year="1991">Willmann & Brooks 1991: 126</bibRefCitation>
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</treatmentCitation>
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, 128, 130, 131;
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<treatmentCitation id="A47E1089FFECFFE0E365A07CCC377658" author="Adams, P. A. & Penny, N. D." box="[352,631,538,563]" page="39" pageId="2" pageNumber="303" year="1992">
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<bibRefCitation id="414E4B69FFECFFE0E365A07CCC377658" author="Adams, P. A. & Penny, N. D." box="[352,631,538,563]" pageId="2" pageNumber="303" pagination="35 - 41" refId="ref22023" refString="Adams, P. A. & Penny, N. D. (1992 a) Review of the South American genera of Nothochrysinae (Insecta: Neuroptera: Chrysopidae). In: Canard, M., Aspock, H. & Mansell, M. W. (Eds.), Current Research in Neuropterology. Proceedings of the Fourth International Symposium on Neuropterology, 24 - 27 June 1991, Bagneres-de-Luchon, Haute-Garonne, France. Privetely printed, Toulouse, pp. 35 - 41." type="book chapter" year="1992">Adams & Penny 1992a: 39</bibRefCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E084A07CCDE77658" author="Adams, P. A. & Penny, N. D." box="[641,935,538,563]" page="221" pageId="2" pageNumber="303" year="1992">
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<bibRefCitation id="414E4B69FFECFFE0E084A07CCDE77658" author="Adams, P. A. & Penny, N. D." box="[641,935,538,563]" pageId="2" pageNumber="303" pagination="216 - 221" refId="ref22116" refString="Adams, P. A. & Penny, N. D. (1992 b) New genera of Nothochrysinae from South America (Neuroptera: Chrysopidae). Pan-Pacific Entomologist, 68, 216 - 221." type="journal article" year="1992">Adams & Penny 1992b: 221</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E1B7A07CCAC67658" author="Carpenter, F. M." box="[946,1158,538,562]" page="347" pageId="2" pageNumber="303" year="1992">
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<bibRefCitation id="414E4B69FFECFFE0E1B7A07CCAC67658" author="Carpenter, F. M." box="[946,1158,538,562]" pageId="2" pageNumber="303" pagination="1 - 655" refId="ref22887" refString="Carpenter, F. M. (1992) Superclass Hexapoda. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vols. 3 & 4. Geological Society of America, Boulder, Colorado & University of Kansas, Lawrence, Kansas, pp. 1 - 655." type="book chapter" year="1992">Carpenter 1992: 347</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E695A07CCB247658" author="Willmann, R." box="[1168,1380,538,562]" page="242" pageId="2" pageNumber="303" year="1993">
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<bibRefCitation id="414E4B69FFECFFE0E695A07CCB247658" author="Willmann, R." box="[1168,1380,538,562]" pageId="2" pageNumber="303" pagination="239 - 245" refId="ref26238" refString="Willmann, R. (1993) Insekten aus der Fur-Formation von Danemark (Moler, ob. Paleozan / unt. Eozan?). 8. Zwei neue Vertreter der Chrysopidae (Neuroptera). Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 4, 239 - 245. https: // doi. org / 10.1127 / njgpm / 1993 / 1993 / 239" type="journal article" year="1993">Willmann 1993: 242</bibRefCitation>
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, 244;
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<treatmentCitation id="A47E1089FFECFFE0E2C2A05FCF8E763B" author="Penalver, E. & Nel, A. & Martinex-Delclos, X." box="[199,462,569,593]" page="486" pageId="2" pageNumber="303" year="1995">
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<bibRefCitation id="414E4B69FFECFFE0E2C2A05FCF8E763B" author="Penalver, E. & Nel, A. & Martinex-Delclos, X." box="[199,462,569,593]" pageId="2" pageNumber="303" pagination="481 - 487" refId="ref25120" refString="Penalver, E., Nel, A. & Martinex-Delclos, X. (1995) New Nothochrysinae from the spanish Miocene (Neuroptera, Chrysopidae). Bulletin de la Societe Entomologique de France, 100, 481 - 487. https: // doi. org / 10.3406 / bsef. 1995.17194" type="journal article" year="1995">
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Peñalver
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<emphasis id="17ABEA8AFFECFFE0E32DA05CCF16763B" box="[296,342,569,593]" italics="true" pageId="2" pageNumber="303">et al</emphasis>
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. 1995: 486
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E3DCA05FCC83763B" author="Ren, D. & Guo, Z. G." box="[473,707,569,593]" page="471" pageId="2" pageNumber="303" year="1996">
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<bibRefCitation id="414E4B69FFECFFE0E3DCA05FCC83763B" author="Ren, D. & Guo, Z. G." box="[473,707,569,593]" pageId="2" pageNumber="303" pagination="461 - 479" refId="ref25310" refString="Ren, D. & Guo, Z. G. (1996) On the new fossil genera and species of Neuroptera (Insecta) from the Late Jurassic of northeast China. Acta Zootaxonomica Sinica, 21, 461 - 479." type="journal article" year="1996">Ren & Guo 1996: 471</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E0CAA05FCDD4763B" author="Nel, A. & Delclos, X. & Hutin, A." box="[719,916,569,593]" page="67" pageId="2" pageNumber="303" year="2005">
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<bibRefCitation id="414E4B69FFECFFE0E0CAA05FCDD4763B" author="Nel, A. & Delclos, X. & Hutin, A." box="[719,916,569,593]" pageId="2" pageNumber="303" pagination="29 - 68" refId="ref24759" refString="Nel, A., Delclos, X. & Hutin, A. (2005) Mesozoic chrysopid-like Planippenia: a phylogenetic approach (Insecta: Neuroptera). Annales de la Societe Entomologique de France, 41, 29 - 68. https: // doi. org / 10.1080 / 00379271.2005.10697440" type="journal article" year="2005">
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Nel
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<emphasis id="17ABEA8AFFECFFE0E0FEA05CCD69763B" box="[763,809,569,593]" italics="true" pageId="2" pageNumber="303">et al</emphasis>
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. 2005: 67
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</bibRefCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E1A5A05FCABB763B" author="Makarkin, V. N. & Archibald, S. B." box="[928,1275,569,593]" page="134" pageId="2" pageNumber="303" year="2013">
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<bibRefCitation id="414E4B69FFECFFE0E1A5A05FCABB763B" author="Makarkin, V. N. & Archibald, S. B." box="[928,1275,569,593]" pageId="2" pageNumber="303" pagination="123 - 146" refId="ref24341" refString="Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1" type="journal article" year="2013">Makarkin & Archibald 2013: 134</bibRefCitation>
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, 141, 142, 144;
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<treatmentCitation id="A47E1089FFECFFE0E2C2A03ECF98761A" author="Archibald, S. B. & Makarkin, V. N. & Greenwood, D. R. & Gunnell, G. F." box="[199,472,600,624]" page="191" pageId="2" pageNumber="303" year="2014">
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<bibRefCitation id="414E4B69FFECFFE0E2C2A03ECF98761A" author="Archibald, S. B. & Makarkin, V. N. & Greenwood, D. R. & Gunnell, G. F." box="[199,472,600,624]" pageId="2" pageNumber="303" pagination="185 - 191" refId="ref22595" refString="Archibald, S. B., Makarkin, V. N., Greenwood, D. R. & Gunnell, G. F. (2014) The Red Queen and Court Jester in green lacewing evolution: bat predation and global climate change. Palaios, 29, 185 - 191. https: // doi. org / 10.2110 / palo. 2013.089" type="journal article" year="2014">
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Archibald
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<emphasis id="17ABEA8AFFECFFE0E331A03ECF28761A" box="[308,360,600,624]" italics="true" pageId="2" pageNumber="303">et al.</emphasis>
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2014: 191
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;
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<treatmentCitation id="A47E1089FFECFFE0E3E6A03ECCDC761A" author="Makarkin V. N." box="[483,668,600,624]" page="4" pageId="2" pageNumber="303" year="2014">
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<bibRefCitation id="414E4B69FFECFFE0E3E6A03ECCDC761A" author="Makarkin V. N." box="[483,668,600,624]" pageId="2" pageNumber="303" pagination="1 - 7" refId="ref24254" refString="Makarkin V. N. (2014) A new fossil green lacewing (Neuroptera: Chrysopidae) from the Eocene Tadushi Formation, eastern Sikhote-Alin. Far Eastern Entomologist, 272, 1 - 7." type="journal article" year="2014">Makarkin 2014: 4</bibRefCitation>
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;
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<treatmentCitation id="A47E1089FFECFFE0E0A2A03ECA407605" author="Archibald, S. B. & Makarkin, V. N." box="[679,1024,599,624]" page="359" pageId="2" pageNumber="303" year="2015">
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<bibRefCitation id="414E4B69FFECFFE0E0A2A03ECA407605" author="Archibald, S. B. & Makarkin, V. N." box="[679,1024,599,624]" pageId="2" pageNumber="303" pagination="359 - 369" refId="ref22418" refString="Archibald, S. B. & Makarkin, V. N. (2015) A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the Early Eocene of Driftwood Canyon, British Columbia, Canada. Canadian Entomologist, 147, 359 - 369. https: // doi. org / 10.4039 / tce. 2014.53" type="journal article" year="2015">Archibald & Makarkin 2015: 359</bibRefCitation>
|
|
</treatmentCitation>
|
|
, 362, 364;
|
|
<treatmentCitation id="A47E1089FFECFFE0E677A03ECEB076E4" author="Archibald, S. B. & Makarkin, V. N." page="400" pageId="2" pageNumber="303" year="2017">
|
|
<bibRefCitation id="414E4B69FFECFFE0E677A03ECEB076E4" author="Archibald, S. B. & Makarkin, V. N." pageId="2" pageNumber="303" pagination="397 - 400" refId="ref22479" refString="Archibald, S. B. & Makarkin, V. N. (2017) A new fossil green lacewing (Neuroptera: Chrysopidae) from the early Eocene Driftwood Canyon, Canada. Zootaxa, 4324 (2), 397 - 400. https: // doi. org / 10.11646 / zootaxa. 4324.2.13" type="journal article" year="2017">Archibald & Makarkin 2017: 400</bibRefCitation>
|
|
</treatmentCitation>
|
|
;
|
|
<treatmentCitation id="A47E1089FFECFFE0E2FEA010CC4576E4" author="Makarkin, V. N. & Wedmann, S. & Weiterschan, T." box="[251,517,630,654]" page="534" pageId="2" pageNumber="303" year="2018">
|
|
<bibRefCitation id="414E4B69FFECFFE0E2FEA010CC4576E4" author="Makarkin, V. N. & Wedmann, S. & Weiterschan, T." box="[251,517,630,654]" pageId="2" pageNumber="303" pagination="527 - 537" refId="ref24549" refString="Makarkin, V. N., Wedmann, S. & Weiterschan, T. (2018) The first green lacewings from the late Eocene Baltic amber. Acta Palaeontologica Polonica, 63, 527 - 537. https: // doi. org / 10.4202 / app. 00504.2018" type="journal article" year="2018">
|
|
Makarkin
|
|
<emphasis id="17ABEA8AFFECFFE0E366A011CFD076E4" box="[355,400,630,654]" italics="true" pageId="2" pageNumber="303">et al</emphasis>
|
|
. 2018: 534
|
|
</bibRefCitation>
|
|
</treatmentCitation>
|
|
;
|
|
<treatmentCitation id="A47E1089FFECFFE0E00AA010CC8276E4" author="Tauber, C. A." box="[527,706,630,654]" page="14" pageId="2" pageNumber="303" year="2019">
|
|
<bibRefCitation id="414E4B69FFECFFE0E00AA010CC8276E4" author="Tauber, C. A." box="[527,706,630,654]" pageId="2" pageNumber="303" pagination="1 - 18" refId="ref26046" refString="Tauber, C. A. (2019 a) South American Nothochrysinae (Neuroptera, Chrysopidae): I. Description of Nothochrysa ehrenbergi sp. nov. ZooKeys, 866, 1 - 18. [https: // zookeys. pensoft. net / article / 35394 /] https: // doi. org / 10.3897 / zookeys. 866.35394" type="journal article" year="2019">Tauber 2019a: 14</bibRefCitation>
|
|
</treatmentCitation>
|
|
, 15;
|
|
<treatmentCitation id="A47E1089FFECFFE0E0F6A010CDE876E4" author="Tauber, C. A." box="[755,936,630,654]" page="36" pageId="2" pageNumber="303" year="2019">
|
|
<bibRefCitation id="414E4B69FFECFFE0E0F6A010CDE876E4" author="Tauber, C. A." box="[755,936,630,654]" pageId="2" pageNumber="303" pagination="19 - 38" refId="ref26110" refString="Tauber, C. A. (2019 b) South American Nothochrysinae (Neuroptera, Chrysopidae): II. Redescription of Leptochrysa prisca Adams & Penny. ZooKeys, 866, 19 - 38. [https: // zookeys. pensoft. net / article / 35396 /] https: // doi. org / 10.3897 / zookeys. 866.35396" type="journal article" year="2019">Tauber 2019b: 36</bibRefCitation>
|
|
</treatmentCitation>
|
|
.
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6DC56513FFECFFE0E292A0D1CA0A70A3" pageId="2" pageNumber="303" type="type_taxon">
|
|
<paragraph id="25603698FFECFFE0E292A0D1CDBE76BB" blockId="2.[151,1437,695,1226]" box="[151,1022,695,722]" pageId="2" pageNumber="303">
|
|
<emphasis id="17ABEA8AFFECFFE0E292A0D1CF7276BB" bold="true" box="[151,306,695,721]" pageId="2" pageNumber="303">
|
|
<typeStatus id="FA64883AFFECFFE0E292A0D1CE9276BB" box="[151,210,695,721]" pageId="2" pageNumber="303">Type</typeStatus>
|
|
species.
|
|
</emphasis>
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E33CA0D1CCA976B8" ID-CoL="6TKFC" authority="Carpenter, 1935" authorityName="Carpenter" authorityYear="1935" box="[313,745,695,722]" class="Insecta" family="Chrysopidae" genus="Palaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="creedei">
|
|
<emphasis id="17ABEA8AFFECFFE0E33CA0D1CC6B76BB" box="[313,555,695,721]" italics="true" pageId="2" pageNumber="303">Palaeochrysa creedei</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E036A0D1CCA976B8" author="Carpenter, F. M." box="[563,745,695,722]" pageId="2" pageNumber="303" pagination="259 - 271" refId="ref22778" refString="Carpenter, F. M. (1935) Tertiary insects of the family Chrysopidae. Journal of Paleontology, 9, 259 - 271." type="journal article" year="1935">Carpenter, 1935</bibRefCitation>
|
|
</taxonomicName>
|
|
, by original designation.
|
|
</paragraph>
|
|
<paragraph id="25603698FFECFFE0E2C2A0BDCD40707F" blockId="2.[151,1437,695,1226]" pageId="2" pageNumber="303">
|
|
<emphasis id="17ABEA8AFFECFFE0E2C2A0BDCFF9769F" bold="true" box="[199,441,731,757]" pageId="2" pageNumber="303">Emended diagnosis.</emphasis>
|
|
May be distinguished from other genera of
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E1C3A0BDCA37769F" authorityName="Navas" authorityYear="1910" box="[966,1143,731,757]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="subFamily" subFamily="Nothochrysinae">Nothochrysinae</taxonomicName>
|
|
by a combination of the following. Forewing: (1) M forked far distad origin of RP [other genera: proximad to slightly distad]; (2) one crossvein between RA, RP proximad 1r-m present [other genera: absent]; (3)
|
|
<emphasis id="17ABEA8AFFECFFE0E606A142CA5F7757" box="[1027,1055,804,829]" italics="true" pageId="2" pageNumber="303">im</emphasis>
|
|
relatively narrow, elongate [
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E761A142CFF7770B" ID-CoL="3PX3" authority="Schluter, 1982" authorityName="Schluter" authorityYear="1982" class="Insecta" family="Chrysopidae" genus="Cimbrochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E761A142CF48770B" italics="true" pageId="2" pageNumber="303">Cimbrochrysa</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E315A121CFF7770B" author="Schluter, T." box="[272,439,839,865]" pageId="2" pageNumber="303" pagination="257 - 264" refId="ref25389" refString="Schluter, T. (1982) Cimbrochrysa moleriensis n. g. n. sp. und Hypochrysa hercyniensis n. sp., zwei fossile Chrysopidae-Arten (Insecta: Planipennia) aus dem europaischen Tertiar. Neues Jahrbuch fur Geologie und Palaontologie, Monatshefte, 5, 257 - 264. https: // doi. org / 10.1127 / njgpm / 1982 / 1982 / 257" type="journal article" year="1982">Schlüter, 1982</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E3C6A121CD46770B" ID-CoL="3ZRR" authority="Wilmann, 1993" authorityName="Wilmann" authorityYear="1993" box="[451,774,839,865]" class="Insecta" family="Chrysopidae" genus="Danochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E3C6A121CC0C770B" box="[451,588,839,865]" italics="true" pageId="2" pageNumber="303">Danochrysa</emphasis>
|
|
Wilmann, 1993
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E114A121CA0D770B" ID-CoL="62CN7" authority="Makarkin, 2014" authorityName="Makarkin" authorityYear="2014" box="[785,1101,839,865]" class="Insecta" family="Chrysopidae" genus="Asiachrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E114A121CDCD770B" box="[785,909,839,865]" italics="true" pageId="2" pageNumber="303">Asiachrysa</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E193A121CA0D770B" author="Makarkin V. N." box="[918,1101,839,865]" pageId="2" pageNumber="303" pagination="1 - 7" refId="ref24254" refString="Makarkin V. N. (2014) A new fossil green lacewing (Neuroptera: Chrysopidae) from the Eocene Tadushi Formation, eastern Sikhote-Alin. Far Eastern Entomologist, 272, 1 - 7." type="journal article" year="2014">Makarkin, 2014</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E65FA121CF3377EF" ID-CoL="67H3" authority="Makarkin & Archibald, 2013" authorityName="Makarkin & Archibald" authorityYear="2013" class="Insecta" family="Chrysopidae" genus="Okanaganochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E65FA121CB67770B" box="[1114,1319,839,865]" italics="true" pageId="2" pageNumber="303">Okanaganochrysa</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E72AA121CF3377EF" author="Makarkin, V. N. & Archibald, S. B." pageId="2" pageNumber="303" pagination="123 - 146" refId="ref24341" refString="Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1" type="journal article" year="2013">Makarkin & Archibald, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E387A10DCC4C77EF" ID-CoL="5CX8" authorityName="Adams & Penny" authorityYear="1992" box="[386,524,875,901]" class="Insecta" family="Chrysopidae" genus="Leptochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E387A10DCC4C77EF" box="[386,524,875,901]" italics="true" pageId="2" pageNumber="303">Leptochrysa</emphasis>
|
|
</taxonomicName>
|
|
: broadly pentagonal;
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E108A10DCDD477EF" ID-CoL="52NJ" authorityName="Hagen" authorityYear="1966" box="[781,916,875,901]" class="Insecta" family="Chrysopidae" genus="Hypochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E108A10DCDD477EF" box="[781,916,875,901]" italics="true" pageId="2" pageNumber="303">Hypochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E1A3A10DCA0B77EF" ID-CoL="34G9" authorityName="Adams & Penny" authorityYear="1992" box="[934,1099,875,901]" class="Insecta" family="Chrysopidae" genus="Asthenochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E1A3A10DCA0B77EF" box="[934,1099,875,901]" italics="true" pageId="2" pageNumber="303">Asthenochrysa</emphasis>
|
|
</taxonomicName>
|
|
: broadly triangular]; (4) two gradate series in radial space [
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E3F0A1E9CC8277C3" ID-CoL="67H3" authorityName="Makarkin & Archibald" authorityYear="2013" box="[501,706,911,937]" class="Insecta" family="Chrysopidae" genus="Okanaganochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E3F0A1E9CC8277C3" box="[501,706,911,937]" italics="true" pageId="2" pageNumber="303">Okanaganochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E0D6A1E9CD2277C3" ID-CoL="82CV" authorityName="Kimmins" authorityYear="1952" box="[723,866,911,937]" class="Insecta" family="Chrysopidae" genus="Triplochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E0D6A1E9CD2277C3" box="[723,866,911,937]" italics="true" pageId="2" pageNumber="303">Triplochrysa</emphasis>
|
|
</taxonomicName>
|
|
: three;
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E1B8A1E9CA1277C3" ID-CoL="62RB6" authorityName="Esben-Petersen" authorityYear="1917" box="[957,1106,911,937]" class="Insecta" family="Chrysopidae" genus="Dictyochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E1B8A1E9CA1277C3" box="[957,1106,911,937]" italics="true" pageId="2" pageNumber="303">Dictyochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E667A1E9CF1077A7" ID-CoL="NYH" authority="Makarkin & Archibald, 2013" authorityName="Makarkin & Archibald" authorityYear="2013" class="Insecta" family="Chrysopidae" genus="Adamsochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E667A1E9CB4777C3" box="[1122,1287,911,937]" italics="true" pageId="2" pageNumber="303">Adamsochrysa</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E715A1E9CF1077A7" author="Makarkin, V. N. & Archibald, S. B." pageId="2" pageNumber="303" pagination="123 - 146" refId="ref24341" refString="Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1" type="journal article" year="2013">Makarkin & Archibald, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
: four or more]. Hind wing: (5) MA not fused with RP (connecting by crossvein 1r-m) [other genera: fused (except
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E337A1B1CFFC779B" ID-CoL="5CX8" authorityName="Adams & Penny" authorityYear="1992" box="[306,444,983,1009]" class="Insecta" family="Chrysopidae" genus="Leptochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E337A1B1CFFC779B" box="[306,444,983,1009]" italics="true" pageId="2" pageNumber="303">Leptochrysa</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E3CCA1B1CD7B779B" ID-CoL="7N4Z" authority="Wilmann, 1993" authorityName="Wilmann" authorityYear="1993" box="[457,827,982,1009]" class="Insecta" family="Chrysopidae" genus="Stephenbrooksia" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="17ABEA8AFFECFFE0E3CCA1B1CCC3779B" box="[457,643,983,1009]" italics="true" pageId="2" pageNumber="303">Stephenbrooksia</emphasis>
|
|
Wilmann, 1993
|
|
</taxonomicName>
|
|
)]; (6) MP not fused with CuA (connecting by crossvein 2m-cu) [probably all other genera: fused or contacting]
|
|
</paragraph>
|
|
<paragraph id="25603698FFECFFE0E2C2A679CB1470CF" blockId="2.[151,1437,695,1226]" pageId="2" pageNumber="303">
|
|
<emphasis id="17ABEA8AFFECFFE0E2C2A679CFD47053" bold="true" box="[199,404,1055,1081]" pageId="2" pageNumber="303">Species included.</emphasis>
|
|
Six species:
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E02DA679CAD07053" ID-CoL="5VQ5B" authority="Makarkin & Archibald, 2013" authorityName="Makarkin & Archibald" authorityYear="2013" box="[552,1168,1055,1081]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="profracta">
|
|
<emphasis id="17ABEA8AFFECFFE0E02DA679CD027053" box="[552,834,1055,1081]" italics="true" pageId="2" pageNumber="303">Archaeochrysa profracta</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E14CA679CAD07053" author="Makarkin, V. N. & Archibald, S. B." box="[841,1168,1055,1081]" pageId="2" pageNumber="303" pagination="123 - 146" refId="ref24341" refString="Makarkin, V. N. & Archibald, S. B. (2013) A diverse new assemblage of green lacewings (Insecta: Neuroptera: Chrysopidae) from the Early Eocene Okanagan Highlands, western North America. Journal of Paleontology, 87, 123 - 146. https: // doi. org / 10.1666 / 12 - 052 R. 1" type="journal article" year="2013">Makarkin & Archibald, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
(early Eocene of McAbee,
|
|
<collectingRegion id="E71BF87AFFECFFE0E2CEA625CFCE7037" box="[203,398,1091,1117]" country="Canada" name="British Columbia" pageId="2" pageNumber="303">British Columbia</collectingRegion>
|
|
,
|
|
<collectingCountry id="5DC87608FFECFFE0E39CA625CFB27034" box="[409,498,1091,1118]" name="Canada" pageId="2" pageNumber="303">Canada</collectingCountry>
|
|
);
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E001A625CD867034" authority="Archibald & Makarkin, 2015" authorityName="Archibald & Makarkin" authorityYear="2015" box="[516,966,1091,1118]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="sanikwa">
|
|
<emphasis id="17ABEA8AFFECFFE0E001A625CC3E7037" box="[516,638,1091,1117]" italics="true" pageId="2" pageNumber="303">A. sanikwa</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E087A625CD867034" author="Archibald, S. B. & Makarkin, V. N." box="[642,966,1091,1118]" pageId="2" pageNumber="303" pagination="359 - 369" refId="ref22418" refString="Archibald, S. B. & Makarkin, V. N. (2015) A new species of Archaeochrysa Adams (Neuroptera: Chrysopidae) from the Early Eocene of Driftwood Canyon, British Columbia, Canada. Canadian Entomologist, 147, 359 - 369. https: // doi. org / 10.4039 / tce. 2014.53" type="journal article" year="2015">Archibald & Makarkin, 2015</bibRefCitation>
|
|
</taxonomicName>
|
|
(early Eocene of Driftwood Canyon,
|
|
<collectingRegion id="E71BF87AFFECFFE0E762A625CF6F70EB" country="Canada" name="British Columbia" pageId="2" pageNumber="303">British Columbia</collectingRegion>
|
|
,
|
|
<collectingCountry id="5DC87608FFECFFE0E33FA601CFD370E8" box="[314,403,1127,1154]" name="Canada" pageId="2" pageNumber="303">Canada</collectingCountry>
|
|
);
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E3A1A601CC9870EB" ID-CoL="G7N9" authority="Adams, 1967" authorityName="Adams" authorityYear="1967" box="[420,728,1127,1153]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="paranervis">
|
|
<emphasis id="17ABEA8AFFECFFE0E3A1A601CC7C70EB" box="[420,572,1127,1153]" italics="true" pageId="2" pageNumber="303">A. paranervis</emphasis>
|
|
<bibRefCitation id="414E4B69FFECFFE0E044A601CC9870EB" author="Adams, P. A." box="[577,728,1127,1153]" pageId="2" pageNumber="303" pagination="215 - 238" refId="ref21987" refString="Adams, P. A. (1967) A review of the Mesochrysinae and Nothochrysinae (Neuroptera: Chrysopidae). Bulletin of the Museum of Comparative Zoology, 135, 215 - 238." type="journal article" year="1967">Adams, 1967</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E0E7A601CA5070EB" ID-CoL="G7N8" authority="(Cockerell, 1914)" baseAuthorityName="Cockerell" baseAuthorityYear="1914" box="[738,1040,1127,1153]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="fracta">
|
|
<emphasis id="17ABEA8AFFECFFE0E0E7A601CD0370EB" box="[738,835,1127,1153]" italics="true" pageId="2" pageNumber="303">A. fracta</emphasis>
|
|
(
|
|
<bibRefCitation id="414E4B69FFECFFE0E154A601CA4970EB" author="Cockerell, T. D. A." box="[849,1033,1127,1153]" pageId="2" pageNumber="303" pagination="714 - 724" refId="ref23258" refString="Cockerell, T. D. A. (1914) New and little-known insects from the Miocene of Florissant, Colorado. Journal of Geology, 22, 714 - 724. https: // doi. org / 10.1086 / 622186" type="journal article" year="1914">Cockerell, 1914</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E61EA601CAE670EB" authority="Makarkin & Antell & Archibald, 2022" authorityName="Makarkin & Antell & Archibald" authorityYear="2022" box="[1051,1190,1127,1153]" class="Insecta" family="Chrysopidae" genus="Archaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="cockerelli" status="sp. nov.">
|
|
<emphasis id="17ABEA8AFFECFFE0E61EA601CAE670EB" box="[1051,1190,1127,1153]" italics="true" pageId="2" pageNumber="303">A. cockerelli</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="17ABEA8AFFECFFE0E6A8A601CB4670EB" bold="true" box="[1197,1286,1127,1153]" pageId="2" pageNumber="303">
|
|
<taxonomicNameLabel id="0C9857F1FFECFFE0E6A8A601CB4670EB" box="[1197,1286,1127,1153]" pageId="2" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
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|
</emphasis>
|
|
(all from late Eocene of Florissant,
|
|
<collectingRegion id="E71BF87AFFECFFE0E38EA6EDCFB670CF" box="[395,502,1163,1189]" country="United States of America" name="Colorado" pageId="2" pageNumber="303">Colorado</collectingRegion>
|
|
,
|
|
<collectingCountry id="5DC87608FFECFFE0E007A6EDCC7F70CF" box="[514,575,1163,1189]" name="United States of America" pageId="2" pageNumber="303">USA</collectingCountry>
|
|
);
|
|
<taxonomicName id="E2DF4D1BFFECFFE0E057A6EDCDCF70CC" ID-CoL="6TKFC" authority="(Carpenter, 1935)" baseAuthorityName="Carpenter" baseAuthorityYear="1935" box="[594,911,1163,1190]" class="Insecta" family="Chrysopidae" genus="Palaeochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="creedei">
|
|
<emphasis id="17ABEA8AFFECFFE0E057A6EDCC8270CF" box="[594,706,1163,1189]" italics="true" pageId="2" pageNumber="303">A. creedei</emphasis>
|
|
(
|
|
<bibRefCitation id="414E4B69FFECFFE0E0D4A6EDCDC770CC" author="Carpenter, F. M." box="[721,903,1163,1190]" pageId="2" pageNumber="303" pagination="259 - 271" refId="ref22778" refString="Carpenter, F. M. (1935) Tertiary insects of the family Chrysopidae. Journal of Paleontology, 9, 259 - 271." type="journal article" year="1935">Carpenter, 1935</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
(Oligocene of Creede,
|
|
<collectingRegion id="E71BF87AFFECFFE0E690A6EDCB4070CF" box="[1173,1280,1163,1189]" country="United States of America" name="Colorado" pageId="2" pageNumber="303">Colorado</collectingRegion>
|
|
,
|
|
<collectingCountry id="5DC87608FFECFFE0E70EA6EDCB0870CF" box="[1291,1352,1163,1189]" name="United States of America" pageId="2" pageNumber="303">USA</collectingCountry>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="25603698FFECFFE0E2C2A6C9CA0A70A3" blockId="2.[151,1437,695,1226]" box="[199,1098,1199,1226]" pageId="2" pageNumber="303">
|
|
<emphasis id="17ABEA8AFFECFFE0E2C2A6C9CF1270A3" bold="true" box="[199,338,1199,1225]" pageId="2" pageNumber="303">Occurrence</emphasis>
|
|
. Eocene (Ypresian to Priabonian) and Oligocene of North America.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |