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<document id="AEC194A2EB08556455B9B0939F9D323D" ID-DOI="10.5281/zenodo.170840" ID-GBIF-Dataset="c0e87963-d99a-4ac4-ab32-41d41d140df9" ID-ISSN="1175­5326" ID-Zenodo-Dep="170840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459715720803" checkinUser="plazi" docAuthor="Helmy, Tamer, Van, Rob W. M. &amp; Soest" docDate="2005" docId="AD0E87EDFFECC0318C4A9947FB044E87" docLanguage="en" docName="zt00859.pdf" docOrigin="Zootaxa 859" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Amphimedon chloros Ilan et al. 2004" docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="5137FF95FFEFC0378D429C69FF864F51" masterDocTitle="Amphimedon species (Porifera: Niphatidae) from the Gulf of Aqaba, Northern Red Sea: Filling the gaps in the distribution of a common pantropical genus" masterLastPageNumber="18" masterPageNumber="1" pageNumber="4" updateTime="1698207824370" updateUser="plazi">
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<mods:title id="2E7B5AD4472CA301052206DA018305C4">Amphimedon species (Porifera: Niphatidae) from the Gulf of Aqaba, Northern Red Sea: Filling the gaps in the distribution of a common pantropical genus</mods:title>
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<mods:namePart id="650F49C748B9C83FE510EC59188955DB">Helmy, Tamer</mods:namePart>
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<treatment id="AD0E87EDFFECC0318C4A9947FB044E87" ID-DOI="http://doi.org/10.5281/zenodo.6266282" ID-GBIF-Taxon="119334543" ID-Zenodo-Dep="6266282" LSID="urn:lsid:plazi:treatment:AD0E87EDFFECC0318C4A9947FB044E87" httpUri="http://treatment.plazi.org/id/AD0E87EDFFECC0318C4A9947FB044E87" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
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<paragraph id="251836FBFFECC0348C4A9947FCF74A19" blockId="3.[264,881,1326,1352]" box="[264,881,1326,1352]" pageId="3" pageNumber="4">
<heading id="7E508197FFECC0348C4A9947FCF74A19" bold="true" box="[264,881,1326,1352]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<emphasis id="17D3EAE9FFECC0348C4A9947FCF74A19" bold="true" box="[264,881,1326,1352]" pageId="3" pageNumber="4">
<taxonomicName id="E2A74D78FFECC0348C4A9947FD554A19" ID-CoL="D3MV" authority="Ilan et al., 2004" authorityName="Ilan et al." authorityYear="2004" box="[264,723,1326,1352]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="4" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFECC0348C4A9947FD8A4A19" bold="true" box="[264,524,1326,1352]" italics="true" pageId="3" pageNumber="4">Amphimedon chloros</emphasis>
<bibRefCitation id="41364B0AFFECC0348F519947FD554A19" author="Ilan" box="[531,723,1326,1352]" pageId="3" pageNumber="17" refString="Ilan, M., Gugel, J. &amp; Van Soest, R. W. M. (2004) Taxonomy, reproduction and ecology of new and known Red Sea sponges. Sarsia, 89, 1 - 24." type="journal article" year="2004">Ilan et al., 2004</bibRefCitation>
</taxonomicName>
(
<figureCitation id="BD9C2A7EFFECC0348FA09947FCC34A19" box="[738,837,1326,1352]" captionStart="FIGURE 2. A" captionStartId="4.[264,367,1740,1764]" captionTargetBox="[314,1272,313,1713]" captionTargetId="figure@4.[314,1273,304,1713]" captionTargetPageId="4" captionText="FIGURE 2. A. Amphimedon chloros Ilan et al., 2004, habit (scale bar = 1 cm); B. Amphimedon chloros Ilan et al., 2004, structure of the ectosomal skeleton showing the three­dimensional network of spicule tracts (scale bar = 0.5 mm); C F. SEM­images of oxeas of Amphimedon chloros (C), A. dinae sp. nov. (D), A. jalae sp. nov. (E) and A. hamadai sp. nov. (F) (scale bars = 10 µm)." httpUri="https://zenodo.org/record/170842/files/figure.png" pageId="3" pageNumber="4">Figs 2A</figureCitation>
C)
</emphasis>
</heading>
</paragraph>
<paragraph id="251836FBFFECC0348C4A9910FD2E4AA1" blockId="3.[264,1323,1401,1521]" pageId="3" pageNumber="4">
<emphasis id="17D3EAE9FFECC0348C4A9910FE004AC0" bold="true" box="[264,390,1401,1425]" pageId="3" pageNumber="4">Synonymy.</emphasis>
<taxonomicName id="E2A74D78FFECC0348CCD9913FCC84AC1" authority="Keller, 1889: 391" authorityName="Keller" authorityPageNumber="391" authorityYear="1889" box="[399,846,1401,1425]" class="Demospongiae" family="Niphatidae" genus="Dactylochalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="4" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFECC0348CCD9913FD074AC0" box="[399,641,1402,1425]" italics="true" pageId="3" pageNumber="4">Dactylochalina viridis</emphasis>
<bibRefCitation id="41364B0AFFECC0348FC99910FC904AC1" author="Keller" box="[651,790,1401,1424]" pageId="3" pageNumber="17" refString="Keller, C. (1889) Die Spongienfauna des rothen Meeres (I. Halfte). Zeitschrift furwissenschaftliche Zoologie, 48, 311 - 405." type="journal article" year="1889">Keller, 1889</bibRefCitation>
: 391
</taxonomicName>
;?
<taxonomicName id="E2A74D78FFECC0348E2A9913FBEC4AC0" box="[872,1130,1402,1425]" class="Demospongiae" family="Niphatidae" genus="Pachychalina" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="4" phylum="Porifera" rank="species" species="variabilis">
<emphasis id="17D3EAE9FFECC0348E2A9913FBEC4AC0" box="[872,1130,1402,1425]" italics="true" pageId="3" pageNumber="4">Pachychalina variabilis</emphasis>
</taxonomicName>
;
<bibRefCitation id="41364B0AFFECC034893E9910FB734AC1" author="Row" box="[1148,1269,1401,1424]" pageId="3" pageNumber="17" refString="Row, R. W. H. (1911) Reports on the marine biology of the Sudanense Red Sea, from collections made by Cyril Crossland, M. A., B. Sc., F. Z. S. XIX. Report on the Sponges collected by Mr. Cyril Crossland in 1904 - 5. Part II. Non-Calcarea. Journal of the Linnean Society Zoology, 31 (208), 287 - 400." type="journal article" year="1911">Row, 1911</bibRefCitation>
: 321 (not:Dendy, 1890);
<emphasis id="17D3EAE9FFECC0348F4499F3FD6D4AE0" box="[518,747,1434,1457]" italics="true" pageId="3" pageNumber="4">Hemihaliclonaviridis</emphasis>
;
<bibRefCitation id="41364B0AFFECC0348FB799F0FC074AE1" author="Burton" box="[757,897,1433,1456]" pageId="3" pageNumber="17" refString="Burton, M. (1937) Supplement to The Littoral Fauna of Krusadai Island in the Gulf of Manaar. Porifera. Bulletin of the Madras Government Museum (New Series, Natural History Section), 1 (2, 4), 1 - 58, pls I - IX." type="journal article" year="1937">Burton,1937</bibRefCitation>
: 18, pl. Ifig.7 (withadditionalsynonyms, but these appear mostly unsustained);
<taxonomicName id="E2A74D78FFECC0348F9999D3FC304A80" box="[731,950,1466,1489]" class="Demospongiae" family="Callyspongiidae" genus="Callyspongia" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="4" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFECC0348F9999D3FC304A80" box="[731,950,1466,1489]" italics="true" pageId="3" pageNumber="4">Callyspongia viridis</emphasis>
</taxonomicName>
;
<bibRefCitation id="41364B0AFFECC0348E8499D0FBDF4A81" author="Burton" box="[966,1113,1465,1488]" pageId="3" pageNumber="17" refString="Burton, M. (1952) The ' Manihine' expedition to the Gulf of Aqaba 1948 - 1949. Sponges. Bulletin of the British Museum (Natural History) Zoology, 1 (8), 163 - 174." type="journal article" year="1952">Burton, 1952</bibRefCitation>
: 167;
<emphasis id="17D3EAE9FFECC03489DC99D3FE014AA0" italics="true" pageId="3" pageNumber="4">
<taxonomicName id="E2A74D78FFECC03489DC99D3FEFA4AA0" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="4" phylum="Porifera" rank="species" species="chloro">Amphimedon chloro</taxonomicName>
<emphasis id="17D3EAE9FFECC0348C3F99B0FE014AA0" bold="true" box="[381,391,1497,1521]" italics="true" pageId="3" pageNumber="4">s</emphasis>
</emphasis>
<bibRefCitation id="41364B0AFFECC0348CCC99B0FDBE4AA1" author="Ilan" box="[398,568,1497,1520]" pageId="3" pageNumber="17" refString="Ilan, M., Gugel, J. &amp; Van Soest, R. W. M. (2004) Taxonomy, reproduction and ecology of new and known Red Sea sponges. Sarsia, 89, 1 - 24." type="journal article" year="2004">Ilan et al., 2004</bibRefCitation>
: 11, fig. 7.
</paragraph>
</subSubSection>
<subSubSection id="6DBD6570FFECC0348C4A9A48FECC4932" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="251836FBFFECC0348C4A9A48FECC4932" blockId="3.[264,1324,1569,1875]" pageId="3" pageNumber="4">
<emphasis id="17D3EAE9FFECC0348C4A9A48FE79496A" bold="true" box="[264,511,1569,1595]" pageId="3" pageNumber="4">Material examined.</emphasis>
<collectionCode id="43B6AE3EFFECC0348F579A48FDD0496A" box="[533,598,1569,1595]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="3" pageNumber="4">ZMA</collectionCode>
Por. 17340, Shark Observatory,
<quantity id="E25F9B1EFFECC0348EA49A48FB8B496A" box="[998,1037,1569,1595]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="4" unit="m" value="3.0">3m</quantity>
, coll. T. Helmy,
<date id="5119103BFFECC034899F9A48FEC04932" pageId="3" pageNumber="4" value="2000-07-26">26­07­ 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="6DBD6570FFECC0328C7A9A18FEE84DCF" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="description">
<paragraph id="251836FBFFECC0348C7A9A18FE154802" blockId="3.[264,1324,1569,1875]" pageId="3" pageNumber="4">
<emphasis id="17D3EAE9FFECC0348C7A9A18FE5649DA" bold="true" box="[312,464,1649,1675]" pageId="3" pageNumber="4">Description.</emphasis>
Colour: Brown when dried or preserved in alcohol, presumably green in life (not recorded). Shape: Lobate, size
<quantity id="E25F9B1EFFECC0348FAA9AF0FCB549E2" box="[744,819,1689,1715]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="4" unit="cm" value="25.0">25 cm</quantity>
<date id="5119103BFFECC0348E7F9AF0FCF549E2" box="[829,883,1689,1715]" pageId="3" pageNumber="4" value="1910-10">
x
<quantity id="E25F9B1EFFECC0348E149AF0FCF549E2" box="[854,883,1689,1715]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="cm" value="10.0">10</quantity>
</date>
cm, attached to hard substrata, with hollow upright, finger­like projections up to
<quantity id="E25F9B1EFFECC0348E579AA8FCC9498A" box="[789,847,1729,1755]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="4" unit="cm" value="7.0">7 cm</quantity>
high. Oscula are distinct,
<quantity id="E25F9B1EFFECC03489C19AA8FB63498A" box="[1155,1253,1729,1755]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="4" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
wide, and scattered over the surface (
<figureCitation id="BD9C2A7EFFECC0348F309A80FD694852" box="[626,751,1769,1795]" captionStart="FIGURE 2. A" captionStartId="4.[264,367,1740,1764]" captionTargetBox="[314,1272,313,1713]" captionTargetId="figure@4.[314,1273,304,1713]" captionTargetPageId="4" captionText="FIGURE 2. A. Amphimedon chloros Ilan et al., 2004, habit (scale bar = 1 cm); B. Amphimedon chloros Ilan et al., 2004, structure of the ectosomal skeleton showing the three­dimensional network of spicule tracts (scale bar = 0.5 mm); C F. SEM­images of oxeas of Amphimedon chloros (C), A. dinae sp. nov. (D), A. jalae sp. nov. (E) and A. hamadai sp. nov. (F) (scale bars = 10 µm)." httpUri="https://zenodo.org/record/170842/files/figure.png" pageId="3" pageNumber="4">Figure 2A</figureCitation>
). Surface: Optically smooth, slightly bumpy surface. Punctate due to regularly distributed subdermal spaces. Texture: Compressible, soft, easy to tear.
</paragraph>
<caption id="71D86673FFEBC0338C4A9AA5FB254804" httpUri="https://zenodo.org/record/170842/files/figure.png" pageId="4" pageNumber="5" targetBox="[314,1272,313,1713]" targetPageId="4">
<paragraph id="251836FBFFEBC0338C4A9AA5FB254804" blockId="4.[264,1323,1740,1877]" pageId="4" pageNumber="5">
<emphasis id="17D3EAE9FFEBC0338C4A9AA5FE0A49B5" bold="true" box="[264,396,1740,1764]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
A.
<taxonomicName id="E2A74D78FFEBC0338CFA9AA5FCD049B2" authority="Ilan et al., 2004" authorityName="Ilan et al." authorityYear="2004" box="[440,854,1740,1763]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="5" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFEBC0338CFA9AA5FD1B49B2" box="[440,669,1740,1763]" italics="true" pageId="4" pageNumber="5">Amphimedon chloros</emphasis>
Ilan et al., 2004
</taxonomicName>
, habit (scale bar = 1 cm); B.
<taxonomicName id="E2A74D78FFEBC03389DC9AA5FD834859" authority="Ilan et al., 2004" authorityName="Ilan et al." authorityYear="2004" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="5" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFEBC03389DC9AA5FED14858" italics="true" pageId="4" pageNumber="5">Amphimedon chloros</emphasis>
Ilan et al., 2004
</taxonomicName>
, structure of the ectosomal skeleton showing the three­dimensional network of spicule tracts (scale bar = 0.5 mm); CF. SEM­images of oxeas of
<taxonomicName id="E2A74D78FFEBC03389539B7EFB70487F" box="[1041,1270,1815,1838]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="5" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFEBC03389539B7EFB70487F" box="[1041,1270,1815,1838]" italics="true" pageId="4" pageNumber="5">Amphimedon chloros</emphasis>
</taxonomicName>
(C),
<taxonomicName id="E2A74D78FFEBC0338C4A9B57FEDC4804" box="[264,346,1854,1877]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="5" phylum="Porifera" rank="species" species="dinae" status="sp. nov.">
<emphasis id="17D3EAE9FFEBC0338C4A9B57FEDC4804" box="[264,346,1854,1877]" italics="true" pageId="4" pageNumber="5">A.dinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CE05792FFEBC0338C239B57FE294804" box="[353,431,1854,1877]" pageId="4" pageNumber="5" rank="species">sp.nov.</taxonomicNameLabel>
(D),
<taxonomicName id="E2A74D78FFEBC0338C9C9B57FDAD4804" box="[478,555,1854,1877]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="5" phylum="Porifera" rank="species" species="jalae" status="sp. nov.">
<emphasis id="17D3EAE9FFEBC0338C9C9B57FDAD4804" box="[478,555,1854,1877]" italics="true" pageId="4" pageNumber="5">A.jalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CE05792FFEBC0338F709B57FDF84804" box="[562,638,1854,1877]" pageId="4" pageNumber="5" rank="species">sp.nov.</taxonomicNameLabel>
(E) and
<taxonomicName id="E2A74D78FFEBC0338F919B57FCC14804" box="[723,839,1854,1877]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="4" pageNumber="5" phylum="Porifera" rank="species" species="hamadai" status="sp. nov.">
<emphasis id="17D3EAE9FFEBC0338F919B57FCC14804" box="[723,839,1854,1877]" italics="true" pageId="4" pageNumber="5">A.hamadai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CE05792FFEBC0338E0F9B57FC1C4804" box="[845,922,1854,1877]" pageId="4" pageNumber="5" rank="species">sp.nov.</taxonomicNameLabel>
(F) (scale bars = 10 µm).
</paragraph>
</caption>
<paragraph id="251836FBFFEAC0328C7A9D72FC6A4D1F" blockId="5.[264,1323,283,1830]" pageId="5" pageNumber="6">
<emphasis id="17D3EAE9FFEAC0328C7A9D72FE254E64" bold="true" box="[312,419,283,309]" pageId="5" pageNumber="6">Skeleton</emphasis>
. Ectosomal skeleton composed of a three­dimensional reticulation of uni­, pauci­ or plurispicular spicule tracts of 1065 µm diameter forming rounded meshes, 90­ 115­
<quantity id="E25F9B1EFFEAC0328C799D05FE1E4ED7" box="[315,408,364,390]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="6" unit="m" value="130.0">130 m</quantity>
diameter. Spongin is scarce and forms an indistinct thin layer enveloping and connecting the spicules. Choanosomal skeleton an anisotropic arrangement of plurispicular spicule tracts 1080µm diameter, ascending towards and protruding slightly from the surface, causing a micro­conulose surface (
<figureCitation id="BD9C2A7EFFEAC0328E4A9D8DFCE94EAF" box="[776,879,484,510]" captionStart="FIGURE 2. A" captionStartId="4.[264,367,1740,1764]" captionTargetBox="[314,1272,313,1713]" captionTargetId="figure@4.[314,1273,304,1713]" captionTargetPageId="4" captionText="FIGURE 2. A. Amphimedon chloros Ilan et al., 2004, habit (scale bar = 1 cm); B. Amphimedon chloros Ilan et al., 2004, structure of the ectosomal skeleton showing the three­dimensional network of spicule tracts (scale bar = 0.5 mm); C F. SEM­images of oxeas of Amphimedon chloros (C), A. dinae sp. nov. (D), A. jalae sp. nov. (E) and A. hamadai sp. nov. (F) (scale bars = 10 µm)." httpUri="https://zenodo.org/record/170842/files/figure.png" pageId="5" pageNumber="6">Figure 2</figureCitation>
B). Meshes similar in size to those of the ectosome, but more squarish in outline, up to 200 µm in diameter. Spicules: Oxeas (
<figureCitation id="BD9C2A7EFFEAC0328C529E5DFEF34D1F" box="[272,373,564,590]" captionStart="FIGURE 2. A" captionStartId="4.[264,367,1740,1764]" captionTargetBox="[314,1272,313,1713]" captionTargetId="figure@4.[314,1273,304,1713]" captionTargetPageId="4" captionText="FIGURE 2. A. Amphimedon chloros Ilan et al., 2004, habit (scale bar = 1 cm); B. Amphimedon chloros Ilan et al., 2004, structure of the ectosomal skeleton showing the three­dimensional network of spicule tracts (scale bar = 0.5 mm); C F. SEM­images of oxeas of Amphimedon chloros (C), A. dinae sp. nov. (D), A. jalae sp. nov. (E) and A. hamadai sp. nov. (F) (scale bars = 10 µm)." httpUri="https://zenodo.org/record/170842/files/figure.png" pageId="5" pageNumber="6">Figure 2</figureCitation>
C), curved with sharp endings, 36­120­142
<date id="5119103BFFEAC0328E319E5DFC334D1F" box="[883,949,564,590]" pageId="5" pageNumber="6">x 12</date>
µm.
</paragraph>
<paragraph id="251836FBFFEAC0328C7A9E32FEE84DCF" blockId="5.[264,1323,283,1830]" pageId="5" pageNumber="6">
<emphasis id="17D3EAE9FFEAC0328C7A9E32FE274D24" bold="true" box="[312,417,603,629]" pageId="5" pageNumber="6">Ecology.</emphasis>
Found on the reef flat, attached to undersides of rocks or sometimes on silt bottoms.
</paragraph>
</subSubSection>
<subSubSection id="6DBD6570FFEAC0328C7A9EC5FB8D4DBF" pageId="5" pageNumber="6" type="distribution">
<paragraph id="251836FBFFEAC0328C7A9EC5FB8D4DBF" blockId="5.[264,1323,283,1830]" pageId="5" pageNumber="6">
<emphasis id="17D3EAE9FFEAC0328C7A9EC5FE574D97" bold="true" box="[312,465,684,710]" pageId="5" pageNumber="6">Distribution</emphasis>
. Widely distributed in the Gulf of Aqaba (pers. obs. T.H.) and frequently recorded from other areas, such as the coasts of Jeddah and Eilat.
</paragraph>
</subSubSection>
<subSubSection id="6DBD6570FFEAC0328C7A9E95FE104877" pageId="5" pageNumber="6" type="discussion">
<paragraph id="251836FBFFEAC0328C7A9E95FCA5495F" blockId="5.[264,1323,283,1830]" pageId="5" pageNumber="6">
<emphasis id="17D3EAE9FFEAC0328C7A9E95FE2C4C47" bold="true" box="[312,426,764,790]" pageId="5" pageNumber="6">Remarks</emphasis>
. We were able to verify the identity of the collected specimen with
<taxonomicName id="E2A74D78FFEAC03289999E95FDED4C6F" authority="Keller, 1889" authorityName="Keller" authorityYear="1889" class="Demospongiae" family="Niphatidae" genus="Dactylochalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFEAC03289999E95FE4A4C6C" italics="true" pageId="5" pageNumber="6">Dactylochalina viridis</emphasis>
<bibRefCitation id="41364B0AFFEAC0328C949F4DFDED4C6F" author="Keller" box="[470,619,804,830]" pageId="5" pageNumber="17" refString="Keller, C. (1889) Die Spongienfauna des rothen Meeres (I. Halfte). Zeitschrift furwissenschaftliche Zoologie, 48, 311 - 405." type="journal article" year="1889">Keller, 1889</bibRefCitation>
</taxonomicName>
by the examination of a microscopical slide of the Berlin Museum
<typeStatus id="FA1C8859FFEAC0328C349F25FE2F4C37" box="[374,425,844,870]" pageId="5" pageNumber="6">type</typeStatus>
specimen ZMB 7654. The genus
<taxonomicName id="E2A74D78FFEAC0328E769F25FB384C37" authority="Lendenfeld, 1886" authorityName="Lendenfeld" authorityYear="1886" box="[820,1214,844,870]" class="Demospongiae" family="Niphatidae" genus="Dactylochalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="genus">
<emphasis id="17D3EAE9FFEAC0328E769F25FC6F4C34" box="[820,1001,844,869]" italics="true" pageId="5" pageNumber="6">Dactylochalina</emphasis>
<bibRefCitation id="41364B0AFFEAC0328EB29F25FB384C37" author="Lendenfeld" box="[1008,1214,844,870]" pageId="5" pageNumber="17" refString="Lendenfeld, R. Von (1886) [1885] A Monograph of the Australian Sponges. Part V. The Auleninae. Proceedings of the Linnean Society of New South Wales, 10 (3), 282 - 325." type="journal article" year="1886">Lendenfeld, 1886</bibRefCitation>
</taxonomicName>
to which
<bibRefCitation id="41364B0AFFEAC0328C4A9F1DFE214CDF" author="Keller" box="[264,423,884,910]" pageId="5" pageNumber="17" refString="Keller, C. (1889) Die Spongienfauna des rothen Meeres (I. Halfte). Zeitschrift furwissenschaftliche Zoologie, 48, 311 - 405." type="journal article" year="1889">Keller (1889)</bibRefCitation>
assigned this species is a synonym of
<taxonomicName id="E2A74D78FFEAC0328E379F1DFB4D4CDC" box="[885,1227,884,909]" class="Demospongiae" family="Callyspongiidae" genus="Callyspongia" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="genus">
<emphasis id="17D3EAE9FFEAC0328E379F1DFB4D4CDC" box="[885,1227,884,909]" italics="true" pageId="5" pageNumber="6">Callyspongia (Callyspongia)</emphasis>
</taxonomicName>
according to
<bibRefCitation id="41364B0AFFEAC0328C139FF5FCBF4CE7" author="Desqueyroux-Faundez" box="[337,825,924,950]" pageId="5" pageNumber="17" refString="Desqueyroux-Faundez, R. &amp; Valentine, C. (2002) Family Niphatidae Van Soest, 1980. In: Hooper, J. N. A. &amp; Van Soest, R. W. M. (Eds) Systema Porifera: A guide to the Classification of Sponges, pp. 886 - 902. Kluwer Academic / Plenum Publishers, New York, 1705 pp." type="book chapter" year="2002">Desqueyroux­Faúndez &amp; Valentine, 2002</bibRefCitation>
, and thus
<taxonomicName id="E2A74D78FFEAC0328EEF9FF5FB9D4CE4" box="[941,1051,924,949]" class="Demospongiae" family="Niphatidae" genus="Dactylochalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFEAC0328EEF9FF5FB9D4CE4" box="[941,1051,924,949]" italics="true" pageId="5" pageNumber="6">D. viridis</emphasis>
</taxonomicName>
was wrongly assigned.
<bibRefCitation id="41364B0AFFEAC0328C4A9FADFE444C8F" author="Ilan" box="[264,450,964,990]" pageId="5" pageNumber="17" refString="Ilan, M., Gugel, J. &amp; Van Soest, R. W. M. (2004) Taxonomy, reproduction and ecology of new and known Red Sea sponges. Sarsia, 89, 1 - 24." type="journal article" year="2004">Ilan et al., 2004</bibRefCitation>
made it clear that it belongs to the genus
<taxonomicName id="E2A74D78FFEAC0328EF19FADFBC84C8C" box="[947,1102,964,989]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="genus">
<emphasis id="17D3EAE9FFEAC0328EF19FADFBC84C8C" box="[947,1102,964,989]" italics="true" pageId="5" pageNumber="6">Amphimedon</emphasis>
</taxonomicName>
, and consequently they had to give it a new name (
<taxonomicName id="E2A74D78FFEAC0328FC59F85FD794B54" box="[647,767,1004,1029]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFEAC0328FC59F85FD794B54" box="[647,767,1004,1029]" italics="true" pageId="5" pageNumber="6">A. chloros</emphasis>
</taxonomicName>
) because the combination
<taxonomicName id="E2A74D78FFEAC032897F9F85FAAD4B54" box="[1085,1323,1004,1029]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFEAC032897F9F85FAAD4B54" box="[1085,1323,1004,1029]" italics="true" pageId="5" pageNumber="6">Amphimedon viridis</emphasis>
</taxonomicName>
is preoccupied by a clearly different species, the Caribbean
<taxonomicName id="E2A74D78FFEAC0328EB6987DFE494B07" authority="Duchassaing &amp; Michelotti, 1864" authorityName="Duchassaing &amp; Michelotti" authorityYear="1864" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFEAC0328EB6987DFBE34B7C" box="[1012,1125,1044,1069]" italics="true" pageId="5" pageNumber="6">A. viridis</emphasis>
Duchassaing &amp; Michelotti, 1864
</taxonomicName>
.
<bibRefCitation id="41364B0AFFEAC0328C999855FD044B07" author="Burton" box="[475,642,1084,1110]" pageId="5" pageNumber="17" refString="Burton, M. (1937) Supplement to The Littoral Fauna of Krusadai Island in the Gulf of Manaar. Porifera. Bulletin of the Madras Government Museum (New Series, Natural History Section), 1 (2, 4), 1 - 58, pls I - IX." type="journal article" year="1937">Burton (1937)</bibRefCitation>
made Duchassaing &amp; Michelotti's
<taxonomicName id="E2A74D78FFEAC03289609855FB094B04" box="[1058,1167,1084,1109]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFEAC03289609855FB094B04" box="[1058,1167,1084,1109]" italics="true" pageId="5" pageNumber="6">A. viridis</emphasis>
</taxonomicName>
the
<typeStatus id="FA1C8859FFEAC03289819855FB734B07" box="[1219,1269,1084,1110]" pageId="5" pageNumber="6">type</typeStatus>
of a genus
<taxonomicName id="E2A74D78FFEAC0328C17980DFD854B2C" box="[341,515,1124,1149]" class="Demospongiae" family="Niphatidae" genus="Hemihaliclona" higherTaxonomySource="CoL" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="genus">
<emphasis id="17D3EAE9FFEAC0328C17980DFD854B2C" box="[341,515,1124,1149]" italics="true" pageId="5" pageNumber="6">Hemihaliclona</emphasis>
</taxonomicName>
, which falls into synonymy with
<taxonomicName id="E2A74D78FFEAC0328ED4980DFBB74B2C" box="[918,1073,1124,1149]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="genus">
<emphasis id="17D3EAE9FFEAC0328ED4980DFBB74B2C" box="[918,1073,1124,1149]" italics="true" pageId="5" pageNumber="6">Amphimedon</emphasis>
</taxonomicName>
according to
<bibRefCitation id="41364B0AFFEAC0328997980DFD2F4BF7" author="Desqueyroux-Faundez" pageId="5" pageNumber="17" refString="Desqueyroux-Faundez, R. &amp; Valentine, C. (2002) Family Niphatidae Van Soest, 1980. In: Hooper, J. N. A. &amp; Van Soest, R. W. M. (Eds) Systema Porifera: A guide to the Classification of Sponges, pp. 886 - 902. Kluwer Academic / Plenum Publishers, New York, 1705 pp." type="book chapter" year="2002">Desqueyroux­Faúndez &amp; Valentine, 2002</bibRefCitation>
. Burton (l.c.) gave a large number of synonyms for
<taxonomicName id="E2A74D78FFEAC032884C98E5FED54B9C" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="viridis">
<emphasis id="17D3EAE9FFEAC032884C98E5FED54B9C" italics="true" pageId="5" pageNumber="6">D. viridis</emphasis>
</taxonomicName>
including the names of several Caribbean and South Australian species. We cannot here decide whether all these synonyms are proper, but grave doubts are in order in view of the implied cosmopolitan nature of such a widespread species. We did check whether the Red Sea species
<taxonomicName id="E2A74D78FFEAC0328CB99945FCF04A17" authority="Row, 1911" authorityName="Row" authorityYear="1911" box="[507,886,1324,1350]" class="Demospongiae" family="Chalinidae" genus="Reniera" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="tabernacula">
<emphasis id="17D3EAE9FFEAC0328CB99945FD684A14" box="[507,750,1324,1349]" italics="true" pageId="5" pageNumber="6">Reniera tabernacula</emphasis>
<bibRefCitation id="41364B0AFFEAC0328FB79945FCF04A17" author="Row" box="[757,886,1324,1350]" pageId="5" pageNumber="17" refString="Row, R. W. H. (1911) Reports on the marine biology of the Sudanense Red Sea, from collections made by Cyril Crossland, M. A., B. Sc., F. Z. S. XIX. Report on the Sponges collected by Mr. Cyril Crossland in 1904 - 5. Part II. Non-Calcarea. Journal of the Linnean Society Zoology, 31 (208), 287 - 400." type="journal article" year="1911">Row, 1911</bibRefCitation>
</taxonomicName>
, allegedly also a synonym according to Burton, conforms to
<emphasis id="17D3EAE9FFEAC0328F58993DFBB64A3C" box="[538,1072,1364,1389]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="E2A74D78FFEAC0328F58993DFCA74A3C" box="[538,801,1364,1389]" class="Demospongiae" family="Niphatidae" genus="Dactylochalina" higherTaxonomySource="CoL" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="viridis">Dactylochalina viridis</taxonomicName>
/
<taxonomicName id="E2A74D78FFEAC0328E75993DFBB64A3C" box="[823,1072,1364,1389]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="chloros">Amphimedon chloros</taxonomicName>
</emphasis>
by studying slides of the
<typeStatus id="FA1C8859FFEAC0328C779915FEEE4AC7" box="[309,360,1404,1430]" pageId="5" pageNumber="6">type</typeStatus>
of
<taxonomicName id="E2A74D78FFEAC0328CD69915FD0E4AC4" box="[404,648,1404,1429]" class="Demospongiae" family="Chalinidae" genus="Reniera" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="tabernacula">
<emphasis id="17D3EAE9FFEAC0328CD69915FD0E4AC4" box="[404,648,1404,1429]" italics="true" pageId="5" pageNumber="6">Reniera tabernacula</emphasis>
</taxonomicName>
, BMNH 1926.2.11.18.
<taxonomicName id="E2A74D78FFEAC0328EF29915FB254AC4" box="[944,1187,1404,1429]" class="Demospongiae" family="Chalinidae" genus="Reniera" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="tabernacula">
<emphasis id="17D3EAE9FFEAC0328EF29915FB254AC4" box="[944,1187,1404,1429]" italics="true" pageId="5" pageNumber="6">Reniera tabernacula</emphasis>
</taxonomicName>
is a green, flattened and lamellar mass, with a clear
<taxonomicName id="E2A74D78FFEAC0328E4799CDFC714AEC" authority="(Reniera)" baseAuthorityName="Reniera" box="[773,1015,1444,1469]" class="Demospongiae" family="Chalinidae" genus="Haliclona" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="genus">
<emphasis id="17D3EAE9FFEAC0328E4799CDFC714AEC" box="[773,1015,1444,1469]" italics="true" pageId="5" pageNumber="6">Haliclona (Reniera)</emphasis>
</taxonomicName>
<typeStatus id="FA1C8859FFEAC032894399CDFBB54AEF" box="[1025,1075,1444,1470]" pageId="5" pageNumber="6">type</typeStatus>
ladder­like skeleton unlike that of
<taxonomicName id="E2A74D78FFEAC0328CFA99A5FDB54AB4" box="[440,563,1484,1509]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFEAC0328CFA99A5FDB54AB4" box="[440,563,1484,1509]" italics="true" pageId="5" pageNumber="6">A. chloros</emphasis>
</taxonomicName>
, with spicules relatively robust, straight and abruptly pointed, unlike the thinner, curved oxeas of
<taxonomicName id="E2A74D78FFEAC0328FE7999DFC9B495C" box="[677,797,1524,1549]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFEAC0328FE7999DFC9B495C" box="[677,797,1524,1549]" italics="true" pageId="5" pageNumber="6">A. chloros</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="251836FBFFEAC0328C7A9A75FE104877" blockId="5.[264,1323,283,1830]" pageId="5" pageNumber="6">
It is likely that the material named
<taxonomicName id="E2A74D78FFEAC0328FA89A75FB804964" box="[746,1030,1564,1589]" class="Demospongiae" family="Niphatidae" genus="Pachychalina" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="variabilis">
<emphasis id="17D3EAE9FFEAC0328FA89A75FB804964" box="[746,1030,1564,1589]" italics="true" pageId="5" pageNumber="6">Pachychalina variabilis</emphasis>
</taxonomicName>
by
<bibRefCitation id="41364B0AFFEAC032897B9A75FB3D4967" author="Row" box="[1081,1211,1564,1590]" pageId="5" pageNumber="17" refString="Row, R. W. H. (1911) Reports on the marine biology of the Sudanense Red Sea, from collections made by Cyril Crossland, M. A., B. Sc., F. Z. S. XIX. Report on the Sponges collected by Mr. Cyril Crossland in 1904 - 5. Part II. Non-Calcarea. Journal of the Linnean Society Zoology, 31 (208), 287 - 400." type="journal article" year="1911">Row, 1911</bibRefCitation>
is also a specimen of this species, although we did not attempt to verify this. The description of the habit and the skeleton is similar to the one presented above. The original
<taxonomicName id="E2A74D78FFEAC03289CB9A05FD9249FF" authority="Dendy, 1887" authorityName="Dendy" authorityYear="1887" class="Demospongiae" family="Niphatidae" genus="Pachychalina" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="variabilis">
<emphasis id="17D3EAE9FFEAC03289CB9A05FEFE49FC" italics="true" pageId="5" pageNumber="6">Pachychalina variabilis</emphasis>
<bibRefCitation id="41364B0AFFEAC0328C3D9AFDFD9249FF" author="Dendy" box="[383,532,1684,1710]" pageId="5" pageNumber="17" refString="Dendy, A. (1887) Observations on the West-Indian Chalininae, with descriptions of new species. Proceedings Royal Society London, 1887, 503 - 507." type="proceedings paper" year="1887">Dendy, 1887</bibRefCitation>
</taxonomicName>
is Caribbean (
<collectingCountry id="5DB0766BFFEAC0328FFE9AFDFCAD49FF" box="[700,811,1684,1710]" name="Bahamas" pageId="5" pageNumber="6">Bahamas</collectingCountry>
) and was assigned to
<taxonomicName id="E2A74D78FFEAC032896B9AFDFDB04987" authority="Duchassaing, 1850" authorityName="Duchassaing" authorityYear="1850" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="complanata">
<emphasis id="17D3EAE9FFEAC032896B9AFDFEBA4984" italics="true" pageId="5" pageNumber="6">Amphimedon complanata</emphasis>
(Duchassaing, 1850)
</taxonomicName>
by Wiedenmayer (1977) and to
<taxonomicName id="E2A74D78FFEAC0328EF09AD5FB564984" box="[946,1232,1724,1749]" class="Demospongiae" family="Niphatidae" genus="Cribrochalina" kingdom="Animalia" order="Haplosclerida" pageId="5" pageNumber="6" phylum="Porifera" rank="species" species="variabilis">
<emphasis id="17D3EAE9FFEAC0328EF09AD5FB564984" box="[946,1232,1724,1749]" italics="true" pageId="5" pageNumber="6">Cribrochalina variabilis</emphasis>
</taxonomicName>
by
<bibRefCitation id="41364B0AFFEAC03289BF9AD5FE2649AF" pageId="5" pageNumber="17" refString="Van Soest, R. W. M. (1980) Marine sponges from Curacao and other Caribbean localities. Part II. Haplosclerida. Studies on the Fauna of Curacao and other Caribbean Islands, 62 (191), 1 - 173." type="journal article">Van Soest (1980)</bibRefCitation>
; in any case this is a clearly different species not to be confused with the Red Sea species.
</paragraph>
</subSubSection>
<subSubSection id="6DBD6570FFE9C0318C7A9D75FB044E87" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="251836FBFFE9C0318C7A9D75FB044E87" blockId="6.[264,1323,284,470]" pageId="6" pageNumber="7">
<taxonomicName id="E2A74D78FFE9C0318C7A9D75FE344E64" ID-CoL="D3MV" box="[312,434,284,309]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="chloros">
<emphasis id="17D3EAE9FFE9C0318C7A9D75FE344E64" box="[312,434,284,309]" italics="true" pageId="6" pageNumber="7">A. chloros</emphasis>
</taxonomicName>
is not the only green
<taxonomicName id="E2A74D78FFE9C0318F809D75FCDB4E64" ID-CoL="84LCJ" box="[706,861,284,309]" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="6" pageNumber="7" phylum="Porifera" rank="genus">
<emphasis id="17D3EAE9FFE9C0318F809D75FCDB4E64" box="[706,861,284,309]" italics="true" pageId="6" pageNumber="7">Amphimedon</emphasis>
</taxonomicName>
in the Indo­Pacific as there is also
<taxonomicName id="E2A74D78FFE9C03188539D75FDCA4E0F" ID-CoL="D3NW" authority="Fromont (1993)" authorityName="Fromont" authorityYear="1993" class="Demospongiae" family="Niphatidae" genus="Amphimedon" kingdom="Animalia" order="Haplosclerida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="paraviridis">
<emphasis id="17D3EAE9FFE9C03188539D75FE0C4E0C" italics="true" pageId="6" pageNumber="7">A. paraviridis</emphasis>
<bibRefCitation id="41364B0AFFE9C0318CD39D2DFDCA4E0F" author="Fromont" box="[401,588,324,350]" pageId="6" pageNumber="17" refString="Fromont, J. (1993) Descriptions of species of the Haplosclerida (Porifera: Demospongiae) occurring in tropical waters of the Great Barrier Reef. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 10 (1), 7 - 40." type="journal article" year="1993">Fromont (1993)</bibRefCitation>
</taxonomicName>
from North
<collectingCountry id="5DB0766BFFE9C0318F9C9D2DFCCC4E0F" box="[734,842,324,350]" name="Australia" pageId="6" pageNumber="7">Australia</collectingCountry>
and
<collectingCountry id="5DB0766BFFE9C0318EC09D2DFC7E4E0F" box="[898,1016,324,350]" name="Indonesia" pageId="6" pageNumber="7">Indonesia</collectingCountry>
. This is different in shape, thickly encrusting to ramose,
<quantity id="E25F9B1EFFE9C0318F239D05FD354ED7" box="[609,691,364,390]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="7" unit="mm" value="20.0">20 mm</quantity>
in thickness with slightly raised oscules. The skeleton forms a loose reticulation with in addition many interstitial spicules between the fibres, and spicules are longer and thicker: 114204
<date id="5119103BFFE9C0318E599DD5FCC64E87" box="[795,832,444,470]" pageId="6" pageNumber="7">x 4</date>
10.5 µm (
<bibRefCitation id="41364B0AFFE9C0318E829DD5FBF34E87" author="Fromont" box="[960,1141,444,470]" pageId="6" pageNumber="17" refString="Fromont, J. (1993) Descriptions of species of the Haplosclerida (Porifera: Demospongiae) occurring in tropical waters of the Great Barrier Reef. The Beagle, Records of the Northern Territory Museum of Arts and Sciences, 10 (1), 7 - 40." type="journal article" year="1993">Fromont, 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>