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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.191.3070" ID-GBIF-Dataset="e3acf250-1110-4fb0-8736-4433ea8c6a67" ID-PMC="PMC3353492" ID-Pensoft-Pub="1313-2970-191-1" ID-PubMed="22639534" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-191-1" ModsDocOrigin="ZooKeys 191" ModsDocTitle="Australian Assassins, Part II: A review of the new assassin spider genus Zephyrarchaea (Araneae, Archaeidae) from southern Australia" checkinTime="1451249040581" checkinUser="pensoft" docAuthor="Rix, Michael G. &amp; Harvey, Mark S." docDate="2012" docId="AC82BE8BE037A25260BEC7FEBDDD02B2" docLanguage="en" docName="ZooKeys 191: 1-62" docOrigin="ZooKeys 191" docSource="http://dx.doi.org/10.3897/zookeys.191.3070" docTitle="Zephyrarchaea mainae, comb. n." docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="FFC6FFF5FF81B72CFFEFFF8CFF8B347D" masterDocTitle="Australian Assassins, Part II: A review of the new assassin spider genus Zephyrarchaea (Araneae, Archaeidae) from southern Australia" masterLastPageNumber="62" masterPageNumber="1" pageNumber="8" updateTime="1668153862162" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Australian Assassins, Part II: A review of the new assassin spider genus Zephyrarchaea (Araneae, Archaeidae) from southern Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rix, Michael G.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harvey, Mark S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>191</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>62</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.191.3070</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.191.3070</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-191-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152035431" LSID="urn:lsid:plazi:treatment:AC82BE8BE037A25260BEC7FEBDDD02B2" httpUri="http://treatment.plazi.org/id/AC82BE8BE037A25260BEC7FEBDDD02B2" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
<subSubSection pageId="7" pageNumber="8" type="nomenclature">
<paragraph pageId="7" pageNumber="8">
<taxonomicName LSID="http://species-id.net/wiki/Zephyrarchaea_mainae" authority="Platnick, 1991 b" class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea mainae" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mainae">Zephyrarchaea mainae (Platnick, 1991b)</taxonomicName>
<taxonomicNameLabel pageId="7" pageNumber="8">comb. n.</taxonomicNameLabel>
Figs 1
<normalizedToken originalValue="EF5AB">E-F5A-B</normalizedToken>
, 5E6A, 6
<normalizedToken originalValue="CD">C-D</normalizedToken>
, 6F8G9E1020
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="reference_group">
<paragraph pageId="7" pageNumber="8">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Austrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Austrarchaea mainae" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mainae">Austrarchaea mainae</taxonomicName>
Platnick, 1991b: 259, figs 1-7.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="vernacular name">
<paragraph pageId="7" pageNumber="8">Albany Assassin Spider</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="type material">
<paragraph pageId="7" pageNumber="8">Type material.</paragraph>
<paragraph pageId="7" pageNumber="8">Holotype male. Torndirrup National Park (probably near end of Eclipse Island Road), Western Australia, Australia, pitfall trap, 1-6.VI.1983, P. Dyer, J. Lyon (WAM T17683).</paragraph>
<paragraph pageId="7" pageNumber="8">Paratypes. Allotype female, same data as holotype except 3-8.X.1983 (WAM T17684).</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="other material examined">
<paragraph pageId="7" pageNumber="8">Other material examined.</paragraph>
<paragraph lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
AUSTRALIA:
<taxonomicName genus="Western" lsidName="Western (Australia)" pageId="7" pageNumber="8" rank="subGenus" subGenus="Australia">Western Australia</taxonomicName>
:Torndirrup National Park: same data as holotype except 2-9.XI.1983, 1 juvenile (WAM T17682); next to carpark at end of Salmon Hole Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.101944">35°06'07&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.9675">117°58'03&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 14.III.2008, M. Rix, M. Harvey, 1♀, 2 juveniles (WAM T89566DNA: TO-1-J/TO-2-J); same data except 30 April 2008, 1♀ (AMNH); same data, 1♀ (QMB S91206); same data, 1 juvenile (WAM T89567DNA: TO-171-J); next to carpark at base of Isthmus Hill,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.09861">35°05'55&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.967224">117°58'02&quot;E</geoCoordinate>
, sifting elevated leaf litter in coastal
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
and eucalypt grove, 14.III.2008, M. Rix, M. Harvey, 2 juveniles (WAM T89568DNA: TO-3-J/TO-4-J). Bremer Bay: Wellstead Road, S. of Bremer Bay, near Yate Place,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-34.40278">34°24'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="119.378334">119°22'42&quot;E</geoCoordinate>
, sifting elevated leaf litter in
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
grove, 19.VI.2010, M. Rix, J.D. Roberts, 1♂, 1♀ (WAM T118984); same data except 21.IV.2009, M. Rix, 1♂, 1 juvenile (WAM T97463DNA: BB-147-J); same data except 2.V.2008, M. Rix, M. Harvey, J. Newell, 1♀, 1 juvenile (WAM T89563DNA: BB-34-F/BB-35-J). Denmark Region: Gilge Road, 15 km SE. of Denmark,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.054165">35°03'15&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.48028">117°28'49&quot;E</geoCoordinate>
, sifting elevated leaf litter in remnant Karri grove with
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 16.III.2008, M. Rix, M. Harvey, 1♀, 1 juvenile (WAM T89577DNA: GI-14-F/GI-15-J); Mount Hallowell, 7 km SW. of Denmark,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.009445">35°00'34&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.30112">117°18'04&quot;E</geoCoordinate>
, beating grass clumps in Karri forest, 6.XI.2007, M. Moir, D. Jolly, 1 juvenile (WAM T78901); Mount Hallowell, 7 km SW. of Denmark,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.010555">35°00'38&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.299164">117°17'57&quot;E</geoCoordinate>
, beating grass clump in Karri forest, 19.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89590DNA: MH-30-J). Gull Rock National Park: end of Ledge Point Road, near carpark,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.014168">35°00'51&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="118.006386">118°00'23&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 17.III.2008, M. Rix, M. Harvey, 1♀, 1 juvenile (WAM T89578DNA: GR-
<pageBreakToken pageId="8" pageNumber="9" start="start">16</pageBreakToken>
-F/GR-17-J). Mutton Bird Point: end of Mutton Bird Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.048054">35°02'53&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.695">117°41'42&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 19.VI.2010, M. Rix, J.D. &amp; B. Roberts, 1♂, 2♀, 2 juveniles (WAM T118983); end of Mutton Bird Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.04778">35°02'52&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.695">117°41'42&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1♂, 3 juveniles (WAM T89582DNA: MB-21-J); end of Mutton Bird Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.044445">35°02'40&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.69722">117°41'50&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1♀ (WAM T89581DNA: MB-20-F); end of Mutton Bird Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.048332">35°02'54&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.70194">117°42'07&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1♀ (WAM T89584DNA: MB-23-F); end of Mutton Bird Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.045">35°02'42&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.69334">117°41'36&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89579); end of Mutton Bird Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.045277">35°02'43&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.69528">117°41'43&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89580); end of Mutton Bird Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.049442">35°02'58&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.69889">117°41'56&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89583). Porongurup National Park: west of Waddy's Hut,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-34.681667">34°40'54&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.84667">117°50'48&quot;E</geoCoordinate>
, sifting elevated leaf litter in unburnt Karri forest, 306 m, 20.IV.2009, M. Rix, 1♀, 1 juvenile (WAM T97465DNA: PO-166-F/PO-167-J); same data except 6 March 2010, M. Rix, L. Lopardo, 1♀ (WAM T114028DNA: PO-168-F). Torndirrup Peninsula: Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.06472">35°03'53&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.79305">117°47'35&quot;E</geoCoordinate>
, beating vegetation in coastal heathland, 26.IX.2007, M. Harvey, R. Ott, 3 juveniles (WAM T89557); Albany Wind Farm, off Bibbulmun Track boardwalk,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.065834">35°03'57&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.79528">117°47'43&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Magnoliopsida" family="Proteaceae" genus="Banksia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Banksia praemorsa" order="Proteales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="praemorsa">Banksia praemorsa</taxonomicName>
, 12.III.2007, M. Rix, M. Harvey, M. Moir, 1 juvenile (WAM T81097); Albany Wind Farm, NW. of Wind Turbine No. 2,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.061943">35°03'43&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.79611">117°47'46&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89575DNA: WF-12-J); Albany Wind Farm, NW. of Wind Turbine No. 3,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.06222">35°03'44&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.79389">117°47'38&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89576DNA: WF-13-J); Albany Wind Farm, SW. of Wind Turbine No. 5,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.059723">35°03'35&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.789444">117°47'22&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008, M. Rix, M. Harvey, 1♀ (WAM T89574DNA: WF-11-F); along Bibbulmun Track E. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.068054">35°04'05&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.80028">117°48'01&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 27.V.2011, M. Rix, M. Harvey, G. Binford, 1♂ (WAM T118982); along Bibbulmun Track E. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.068054">35°04'05&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.80195">117°48'07&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1♀ (WAM T89585DNA: WF-24-F); along Bibbulmun Track E. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.069443">35°04'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.80723">117°48'26&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89588); along Bibbulmun Track E. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.06889">35°04'08&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.805">117°48'18&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89587DNA: WF-26-J); along Bibbulmun Track E. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.068333">35°04'06&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.802505">117°48'09&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89586DNA: WF-25-J); Cuthbert, W. of Roberts Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.033054">35°01'59&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.79639">117°47'47&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 18.III.2008, M. Rix, M. Harvey, 1♀, 2 juveniles (WAM T89589DNA: WF-28-J/WF-29-J); end of Prescott Vale Road, W. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.049168">35°02'57&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.75833">117°45'30&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008,
<pageBreakToken pageId="9" pageNumber="10" start="start">M</pageBreakToken>
. Rix, M. Harvey, 1♂, 3 juveniles (WAM T89569DNA: WF-5-J/WF-6-J); end of Prescott Vale Road, W. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.052776">35°03'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.76972">117°46'11&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89573DNA: WF-10-J); end of Prescott Vale Road, W. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.052776">35°03'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.76639">117°45'59&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008, M. Rix, M. Harvey, 1♀ (WAM T89572DNA: WF-9-F); end of Prescott Vale Road, W. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.050556">35°03'02&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.757774">117°45'28&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008, M. Rix, M. Harvey, 1♀ (WAM T89570); end of Prescott Vale Road, W. of Albany Wind Farm,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.052776">35°03'10&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.76195">117°45'43&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 15.III.2008, M. Rix, M. Harvey, 1♀ (WAM T89571DNA: WF-8-F). Walpole-Nornalup National Park: Anderson Road,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-34.995277">34°59'43&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="116.87056">116°52'14&quot;E</geoCoordinate>
, sifting elevated leaf litter under curly grass in Marri and
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
forest, 3.V.2008, M. Rix, M. Harvey, 2 juveniles (WAM T89564DNA: WA-36-J/WA-37-J); same data except 24.I.2010, M. Rix, J. Wojcieszek, 1♂, 2♀, 1 juvenile (WAM T114037DNA: WA-162-F). William Bay National Park: near Elephant Rock carpark,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-35.0225">35°01'21&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="117.236664">117°14'12&quot;E</geoCoordinate>
, sifting elevated leaf litter under
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
, 26.VI.2010, M. Rix, 2♀ (WAM T114029DNA: WB-169-F); same data except 29.IV.2008, M. Rix, M. Harvey, 1 juvenile (WAM T89591DNA: WB-31-J); same data except 7.III.2010, M. Rix, L. Lopardo, 2 juveniles (WAM T114030DNA: WB-170-J).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="diagnosis">
<paragraph pageId="9" pageNumber="10">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="10">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea mainae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mainae">Zephyrarchaea mainae</taxonomicName>
can be distinguished from other known congeners except
<taxonomicName class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea janineae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="janineae">Zephyrarchaea janineae</taxonomicName>
sp. n. by the presence of six dorsal, hump-like tubercles on the abdomen (Figs 1
<normalizedToken originalValue="EF">E-F</normalizedToken>
, 6A, 10
<normalizedToken originalValue="AB">A-B</normalizedToken>
); and from
<taxonomicName class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea janineae" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="janineae">Zephyrarchaea janineae</taxonomicName>
sp. n. by the more flattened, rounded apex of tegular sclerite 1 (TS 1) (Figs 10
<normalizedToken originalValue="DF">D-F</normalizedToken>
).
</paragraph>
<paragraph pageId="9" pageNumber="10">This species can also be distinguished from other genotyped taxa (see Fig. 3) by the following two unique nucleotide substitutions for COI and COII (n = 35): A(282), T(1173).</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph pageId="9" pageNumber="10">Description.</paragraph>
<paragraph pageId="9" pageNumber="10">
Holotype male: Total length 3.44; leg I femur 1.81; F1/CL ratio 1.72. Cephalothorax dark reddish-brown; legs tan brown with darker annulations; abdomen mottled grey-brown and beige, with reddish-brown dorsal scute and sclerites (Fig. 10B). Carapace short (CH/CL ratio 1.60); 1.05 long, 1.68 high, 1.03 wide;
<normalizedToken originalValue="neck">'neck'</normalizedToken>
0.60 wide; highest point of pars cephalica (HPC) near posterior third of
<normalizedToken originalValue="head">'head'</normalizedToken>
(ratio of HPC to post-ocular length 0.67), carapace with pronounced concave depression anterior to HPC;
<normalizedToken originalValue="head">'head'</normalizedToken>
not strongly elevated dorsally (post-ocular ratio 0.24) (Fig. 8G). Chelicerae with proximal tuft and additional comb of accessory setae on anterior face of paturon (Figs 5A, 5E, 10C). Abdomen 1.69 long, 1.18 wide; with three pairs of dorsal hump-like tubercles (HT 1-6); dorsal scute fused anteriorly to epigastric sclerites, extending posteriorly to first pair of hump-like tubercles; HT 3-6 each covered by separate dorsal sclerites. Pedipalp fully expanded (see
<bibRefCitation author="Platnick, NI" journalOrPublisher="Bulletin of the British Arachnological Society" pageId="22" pageNumber="23" pagination="259 - 261" title="On Western Australian Austrarchaea (Araneae, Archaeidae)." volume="8" year="1991 b">Platnick 1991b</bibRefCitation>
, figs 4-6). Unexpanded pedipalp (of WAM T89569) (Figs 10
<normalizedToken originalValue="DF">D-F</normalizedToken>
) pyriform, with broad, distally curved embolus supported by conductor sclerites 1-2; tegular sclerite 1 (TS 1) porrect, slightly curved in prolateral view, with flattened, rounded apex; TS 2-3 projecting beyond retro-distal rim of tegulum.
</paragraph>
<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Female (WAM T118983): Total length 3.31; leg I femur 1.97; F1/CL ratio 1.73. Cephalothorax dark reddish-brown; legs tan brown with darker annulations; abdomen mottled grey-brown and beige (Fig. 10A). Carapace short (CH/CL ratio 1.71); 1.14
<pageBreakToken pageId="10" pageNumber="11" start="start">long</pageBreakToken>
, 1.95 high, 1.10 wide;
<normalizedToken originalValue="neck">'neck'</normalizedToken>
0.68 wide; highest point of pars cephalica (HPC) near posterior third of
<normalizedToken originalValue="head">'head'</normalizedToken>
(ratio of HPC to post-ocular length 0.63), carapace with pronounced concave depression anterior to HPC;
<normalizedToken originalValue="head">'head'</normalizedToken>
not strongly elevated dorsally (post-ocular ratio 0.24) (Fig. 9E). Chelicerae without accessory setae on anterior face of paturon. Abdomen 2.00 long, 1.36 wide; with three pairs of dorsal hump-like tubercles (HT 1-6). Internal genitalia (Fig. 10G) with cluster of ≤ 15 sausage-shaped spermathecae either side of gonopore, clusters widely separated along midline of genital plate.
</paragraph>
<paragraph pageId="10" pageNumber="11">
Variation: Males (n = 8): total length 2.82-3.44; carapace length 1.02-1.15; carapace height 1.62-1.80; CH/CL ratio 1.56-1.64. Females (n = 23): total length 2.77-3.82; carapace length 0.94-1.21; carapace height 1.45-1.99; CH/CL ratio 1.55-1.71. Specimens of
<taxonomicName class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea mainae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mainae">Zephyrarchaea mainae</taxonomicName>
from the eastern-most Bremer Bay population have significantly smaller hump-like tubercles on the abdomen compared to populations further west (see Fig. 1F cf. Figs 1E, 10A), and males from this Bremer Bay population also have a more distally paddle-shaped TS 1 morphology similar to
<taxonomicName class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea marki" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="marki">Zephyrarchaea marki</taxonomicName>
sp. n. from Cape Le Grand National Park.
<bibRefCitation author="Rix, MG" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="22" pageNumber="23" pagination="375 - 396" title="Phylogeny and historical biogeography of ancient assassin spiders (Araneae: Archaeidae) in the Australian mesic zone: evidence for Miocene speciation within Tertiary refugia." url="10.1016/j.ympev.2011.10.009" volume="62" year="2012">Rix and Harvey (2012)</bibRefCitation>
showed that the Bremer Bay population of
<taxonomicName class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea mainae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mainae">Zephyrarchaea mainae</taxonomicName>
is genetically distinct (Fig. 3), with no evidence of recent gene flow, indicating possible incipient speciation.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="distribution">
<paragraph pageId="10" pageNumber="11">Distribution and habitat.</paragraph>
<paragraph pageId="10" pageNumber="11">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Zephyrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zephyrarchaea mainae" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mainae">Zephyrarchaea mainae</taxonomicName>
is known from the greater Albany region of southern Western Australia, from Walpole-Nornalup National Park (near Walpole) east to Bremer Bay and north to the Porongurup National Park, with a range centred on the Torndirrup Peninsula south of Albany (Fig. 20). Specimens have been collected by beating and sifting sedges (
<taxonomicName class="Liliopsida" family="Cyperaceae" genus="Lepidosperma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lepidosperma" order="Poales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="genus">Lepidosperma</taxonomicName>
sp.), curly grass (
<taxonomicName class="Liliopsida" family="Restionaceae" genus="Empodisma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Empodisma gracillimum" order="Poales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="gracillimum">Empodisma gracillimum</taxonomicName>
) and low shrubs in dense coastal or near-coastal groves of Peppermint (
<taxonomicName class="Insecta" family="Callidulidae" genus="Agonis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agonis" order="Lepidoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Agonis</taxonomicName>
sp.), with several outlying populations also known from wet Karri (
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus diversicolor" order="Myrtales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="diversicolor">Eucalyptus diversicolor</taxonomicName>
) forest.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="conservation status">
<paragraph pageId="10" pageNumber="11">Conservation status.</paragraph>
<paragraph pageId="10" pageNumber="11">
Thisspecies is listed as threatened under theWestern Australian Wildlife Conservation Act 1950. It is a short-range endemic taxon (
<bibRefCitation author="Harvey, MS" journalOrPublisher="Invertebrate Systematics" pageId="22" pageNumber="23" pagination="555 - 570" title="Short-range endemism among the Australian fauna: some examples from non-marine environments." url="10.1071/IS02009" volume="16" year="2002 b">Harvey 2002b</bibRefCitation>
), with known populations threatened by fire, dieback disease (affecting coastal heathland vegetation), land-clearing and climate change.
</paragraph>
</subSubSection>
</treatment>
</document>