treatments-xml/data/AC/2D/87/AC2D87E6FF8ED828FF3FC67AFD86F3B8.xml
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<document id="82EEDC668D8F8AC89260A4EDAAA6AC77" ID-CLB-Dataset="24959" ID-DOI="10.11646/zootaxa.4695.1.3" ID-GBIF-Dataset="d688d9bc-ea31-4e99-9dcb-e65416752eed" ID-ISSN="1175-5326" ID-Zenodo-Dep="3530113" ID-ZooBank="945B3188-6A62-4B71-8B0F-5FF360A67A75" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573024157255" checkinUser="plazi" docAuthor="Churkin, Sergei V., Kolesnichenko, Kirill A. &amp; Yakovlev, Roman V." docDate="2019" docId="AC2D87E6FF8ED828FF3FC67AFD86F3B8" docLanguage="en" docName="zootaxa.4695.1.3.pdf" docOrigin="Zootaxa 4695 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Plebejus anikini subsp. azhbogdo Churkin &amp; Kolesnichenko &amp; Yakovlev 2019, ssp. n." docType="treatment" docUuid="7155B462-481F-4A1A-BD10-69A5270B3511" docUuidSource="ZooBank" docVersion="7" lastPageNumber="55" masterDocId="5014FF9EFF89D822FFA8C53FFF9CF36F" masterDocTitle="Two new taxa of Plebejus Kluk, 1780 (Lepidoptera, Lycaenidae) from the Gobi desert, Mongolia" masterLastPageNumber="58" masterPageNumber="45" pageNumber="52" updateTime="1698763241038" updateUser="ExternalLinkService">
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<mods:title id="399340F41A551A5FF61D5DF7B3CE4F5E">Two new taxa of Plebejus Kluk, 1780 (Lepidoptera, Lycaenidae) from the Gobi desert, Mongolia</mods:title>
</mods:titleInfo>
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<mods:namePart id="CDEDC90E4BE4E7AA4EDCE8E1FD846E3F">Churkin, Sergei V.</mods:namePart>
</mods:name>
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<mods:namePart id="B60F8729EEA4CE0D726DD5EB59A74E26">Kolesnichenko, Kirill A.</mods:namePart>
</mods:name>
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<mods:roleTerm id="C521B57F6B20CF0AB7ED7FDB2CB25BD0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E1888A2BF9B0B0753B64AEF0BCE07E1C">Yakovlev, Roman V.</mods:namePart>
</mods:name>
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<mods:title id="CDD6A5506791DA70BA8FD2852B77996F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0B95D3BD0245DE3027ED78AC14C19249">
<mods:date id="9B77BAF47CE8EDCCF0855DFE5F973A31">2019</mods:date>
<mods:detail id="0DB520119A2E4E5B6B2268F0D645F9CD" type="pubDate">
<mods:number id="9AFF91BDAD697915BB33DCAD391A2CAD">2019-11-06</mods:number>
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<mods:detail id="A07B7245AB48F865ED88F91E009AA247" type="volume">
<mods:number id="EA48DCE862D016BC3F071E21B656E316">4695</mods:number>
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<mods:identifier id="8D4D496444E08B2DD0301DD75274D7A4" type="CLB-Dataset">24959</mods:identifier>
<mods:identifier id="695B26F5B79AC73FE81C20493E1C3913" type="DOI">10.11646/zootaxa.4695.1.3</mods:identifier>
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</mods:mods>
<treatment id="AC2D87E6FF8ED828FF3FC67AFD86F3B8" ID-GBIF-Taxon="160142631" LSID="urn:lsid:zoobank.org:act:7155B462-481F-4A1A-BD10-69A5270B3511" httpUri="http://treatment.plazi.org/id/AC2D87E6FF8ED828FF3FC67AFD86F3B8" lastPageId="10" lastPageNumber="55" pageId="7" pageNumber="52">
<subSubSection id="6C9E657BFF8ED825FF3FC67AFD7BF0EC" pageId="7" pageNumber="52" type="nomenclature">
<paragraph id="243B36F0FF8ED825FF3FC67AFB9FF030" blockId="7.[151,1027,837,900]" box="[151,1027,837,864]" pageId="7" pageNumber="52">
<heading id="7F73819CFF8ED825FF3FC67AFB9FF030" bold="true" box="[151,1027,837,864]" fontSize="11" level="1" pageId="7" pageNumber="52" reason="1">
<emphasis id="16F0EAE2FF8ED825FF3FC67AFB9FF030" bold="true" box="[151,1027,837,864]" pageId="7" pageNumber="52">
<taxonomicName id="E3844D73FF8ED825FF3FC67AFC34F030" authority="Churkin, Kolesnichenko &amp; Yakovlev" authorityName="Churkin &amp; Kolesnichenko &amp; Yakovlev" authorityYear="2019" box="[151,936,837,864]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="52" phylum="Arthropoda" rank="subSpecies" species="anikini" status="subsp. nov." subSpecies="azhbogdo">
<emphasis id="16F0EAE2FF8ED825FF3FC67AFE4BF00F" bold="true" box="[151,471,837,864]" italics="true" pageId="7" pageNumber="52">Plebejus anikini azhbogdo</emphasis>
Churkin, Kolesnichenko &amp; Yakovlev
</taxonomicName>
,
<taxonomicNameLabel id="0DC35799FF8ED825FC1EC67AFB9FF030" box="[950,1027,837,863]" pageId="7" pageNumber="52" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="243B36F0FF8ED825FF3FC656FD7BF0EC" blockId="7.[151,1027,837,900]" box="[151,743,873,900]" pageId="7" pageNumber="52">
Plate 1 and 2, figs. 13 (
<typeStatus id="FB3F8852FF8ED825FE31C656FD9DF0EB" box="[409,513,873,900]" pageId="7" pageNumber="52" type="holotype">holotype</typeStatus>
), 1418 (
<typeStatus id="FB3F8852FF8ED825FDC1C656FD47F0EC" box="[617,731,873,899]" pageId="7" pageNumber="52" type="paratype">paratypes</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="6C9E657BFF8ED825FF3FC68EFDE7F734" pageId="7" pageNumber="52" type="materials_examined">
<paragraph id="243B36F0FF8ED825FF3FC68EFBF4F080" blockId="7.[151,1437,945,2015]" pageId="7" pageNumber="52">
<materialsCitation id="94EC3CADFF8ED825FF3FC68EFBF4F080" ID-GBIF-Occurrence="2447663072" collectingDate="2013-07-15" collectionCode="ZISP" collectorName="K. Kolesnichenko" country="Mongolia" elevation="1500" latitude="44.575" location="Altai" longLatPrecision="19" longitude="95.038055" municipality="Mongolian Altai" pageId="7" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="Govi-Altay" typeStatus="holotype">
<emphasis id="16F0EAE2FF8ED825FF3FC68EFE91F0A4" bold="true" box="[151,269,945,971]" pageId="7" pageNumber="52">
<typeStatus id="FB3F8852FF8ED825FF3FC68EFE94F0A4" box="[151,264,945,971]" pageId="7" pageNumber="52" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="3282FD79FF8ED825FEBFC68EFEB2F0A4" box="[279,302,945,971]" pageId="7" pageNumber="52" type="male"></specimenCount>
, S
<collectingCountry id="5C937660FF8ED825FEFEC68EFE5AF0A4" box="[342,454,945,971]" name="Mongolia" pageId="7" pageNumber="52">Mongolia</collectingCountry>
,
<collectingMunicipality id="C45FAC8AFF8ED825FE7BC68EFD0DF0A4" box="[467,657,945,971]" pageId="7" pageNumber="52">
Mongolian
<location id="215B602BFF8ED825FDFEC68EFD0DF0A4" LSID="urn:lsid:plazi:treatment:AC2D87E6FF8ED828FF3FC67AFD86F3B8:215B602BFF8ED825FDFEC68EFD0DF0A4" box="[598,657,945,971]" country="Mongolia" latitude="44.575" longLatPrecision="19" longitude="95.038055" municipality="Mongolian Altai" name="Altai" pageId="7" pageNumber="52" stateProvince="Govi-Altay">Altai</location>
</collectingMunicipality>
,
<collectingRegion id="E640F812FF8ED825FD37C68EFCF5F0A4" box="[671,873,945,971]" country="Mongolia" name="Govi-Altay" pageId="7" pageNumber="52">Gobi-Altai aimak</collectingRegion>
,
<locationDeviation id="35892FD2FF8ED825FCDFC68EFBA7F0A4" box="[887,1083,945,971]" location="Altai" pageId="7" pageNumber="52">
<quantity id="E37C9B15FF8ED825FCDFC68EFC5CF0A4" box="[887,960,945,971]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="52" unit="km" value="10.0">10 km</quantity>
ESE Altai
</locationDeviation>
somon, Hadat-Miangan-Bulag spring,
<quantity id="E37C9B15FF8ED825FF43C6EBFEDEF080" box="[235,322,980,1007]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="52" unit="m" value="1500.0">
<elevation id="AFA9D1C3FF8ED825FF43C6EBFEDEF080" box="[235,322,980,1007]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="52" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="503A1030FF8ED825FEE6C6EBFE53F080" box="[334,463,980,1007]" pageId="7" pageNumber="52" value="2013-07-15">
<collectingDate id="407EE9D8FF8ED825FEE6C6EBFE53F080" box="[334,463,980,1007]" pageId="7" pageNumber="52" value="2013-07-15">15.07.2013</collectingDate>
</date>
,
<geoCoordinate id="41B05037FF8ED825FE72C6EAFDFAF080" box="[474,614,981,1007]" degrees="44" direction="north" minutes="34" orientation="latitude" pageId="7" pageNumber="52" precision="15" seconds="30" value="44.575">44°3430”N</geoCoordinate>
,
<geoCoordinate id="41B05037FF8ED825FDD9C6EBFC9BF080" box="[625,775,980,1007]" degrees="095" direction="east" minutes="02" orientation="longitude" pageId="7" pageNumber="52" precision="15" seconds="17" value="95.038055">095°0217”E</geoCoordinate>
,
<collectorName id="89715326FF8ED825FCBBC6EAFC41F080" box="[787,989,981,1007]" pageId="7" pageNumber="52">K. Kolesnichenko</collectorName>
leg. (
<collectionCode id="4295AE35FF8ED825FBB5C6EAFBC0F080" box="[1053,1116,981,1007]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="7" pageNumber="52">ZISP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC6C6FDE7F734" blockId="7.[151,1437,945,2015]" pageId="7" pageNumber="52">
<materialsCitation id="94EC3CADFF8ED825FF6FC6C6FCFEF77C" ID-GBIF-Occurrence="2447663080" box="[199,866,1017,1043]" collectorName="K. Kolesnichenko" pageId="7" pageNumber="52" specimenCount="38" specimenCount-female="18" specimenCount-male="20" typeStatus="paratype">
<emphasis id="16F0EAE2FF8ED825FF6FC6C6FED4F77C" bold="true" box="[199,328,1017,1043]" pageId="7" pageNumber="52">
<typeStatus id="FB3F8852FF8ED825FF6FC6C6FEDFF77C" box="[199,323,1017,1043]" pageId="7" pageNumber="52" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="3282FD79FF8ED825FEF8C6C6FE16F77C" box="[336,394,1017,1043]" pageId="7" pageNumber="52" type="male">20 ♂</specimenCount>
,
<specimenCount id="3282FD79FF8ED825FE3FC6C6FE53F77C" box="[407,463,1017,1043]" pageId="7" pageNumber="52" type="female">18 ♀</specimenCount>
, same data,
<collectorName id="89715326FF8ED825FDCBC6C6FCB2F77C" box="[611,814,1017,1043]" pageId="7" pageNumber="52">K. Kolesnichenko</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="94EC3CADFF8ED825FCC6C6C6FDE7F734" ID-GBIF-Occurrence="2447663079" collectingDate="2017-06-28" collectorName="R. Yakovlev" country="Mongolia" county="Adz-Bogdo Range" elevation="1350" latitude="45.090275" location="Gobi Altai" longLatPrecision="19" longitude="94.80139" municipality="Altai" pageId="7" pageNumber="52" specimenCount="30" specimenCount-female="13" specimenCount-male="17" typeStatus="paratype">
<specimenCount id="3282FD79FF8ED825FCC6C6C6FC35F77C" box="[878,937,1017,1043]" pageId="7" pageNumber="52" type="male">17 ♂</specimenCount>
,
<specimenCount id="3282FD79FF8ED825FC1DC6C6FC72F77C" box="[949,1006,1017,1043]" pageId="7" pageNumber="52" type="female">13 ♀</specimenCount>
, SW
<collectingCountry id="5C937660FF8ED825FB9DC6C6FB39F77C" box="[1077,1189,1017,1043]" name="Mongolia" pageId="7" pageNumber="52">Mongolia</collectingCountry>
,
<location id="215B602BFF8ED825FB18C6C6FABAF77C" LSID="urn:lsid:plazi:treatment:AC2D87E6FF8ED828FF3FC67AFD86F3B8:215B602BFF8ED825FB18C6C6FABAF77C" box="[1200,1318,1017,1043]" country="Mongolia" county="Adz-Bogdo Range" latitude="45.090275" longLatPrecision="19" longitude="94.80139" municipality="Altai" name="Gobi Altai" pageId="7" pageNumber="52">
Gobi
<collectingMunicipality id="C45FAC8AFF8ED825FB46C6C6FABAF77C" box="[1262,1318,1017,1043]" pageId="7" pageNumber="52">Altai</collectingMunicipality>
</location>
aimak,
<locationDeviation id="35892FD2FF8ED825FA28C6C6FEA1F758" location="Bugat" pageId="7" pageNumber="52">
<quantity id="E37C9B15FF8ED825FA28C6C6FF27F758" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="52" unit="km" value="62.0">62 km</quantity>
SSE Bugat
</locationDeviation>
somon, N slopes of
<collectingCounty id="CD5A4E7CFF8ED825FD8CC122FD64F758" box="[548,760,1053,1079]" pageId="7" pageNumber="52">Adz-Bogdo Range</collectingCounty>
, Zoolon-Suuzhijn-Bulag spring,
<geoCoordinate id="41B05037FF8ED825FBC2C123FB69F758" box="[1130,1269,1052,1079]" degrees="45" direction="north" minutes="05" orientation="latitude" pageId="7" pageNumber="52" precision="15" seconds="25" value="45.090275">45°0525”N</geoCoordinate>
,
<geoCoordinate id="41B05037FF8ED825FAA9C123FA04F758" box="[1281,1432,1052,1079]" degrees="094" direction="east" minutes="48" orientation="longitude" pageId="7" pageNumber="52" precision="15" seconds="05" value="94.80139">094°4805”E</geoCoordinate>
,
<quantity id="E37C9B15FF8ED825FF3FC17FFF6FF734" box="[151,243,1088,1115]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="52" unit="m" value="1350.0">
<elevation id="AFA9D1C3FF8ED825FF3FC17FFF6FF734" box="[151,243,1088,1115]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="52" unit="m" value="1350.0">1350 m</elevation>
</quantity>
.a.s.l.,
<date id="503A1030FF8ED825FE9DC17EFE2AF734" box="[309,438,1089,1115]" pageId="7" pageNumber="52" value="2017-06-28">
<collectingDate id="407EE9D8FF8ED825FE9DC17EFE2AF734" box="[309,438,1089,1115]" pageId="7" pageNumber="52" value="2017-06-28">28.06.2017</collectingDate>
</date>
,
<collectorName id="89715326FF8ED825FE69C17EFDD7F734" box="[449,587,1089,1115]" pageId="7" pageNumber="52">R. Yakovlev</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6C9E657BFF8ED82AFF6FC15BFD25F4CA" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52" type="description">
<paragraph id="243B36F0FF8ED825FF6FC15BFEC5F710" blockId="7.[151,1437,945,2015]" box="[199,345,1124,1151]" pageId="7" pageNumber="52">
<emphasis id="16F0EAE2FF8ED825FF6FC15BFECDF710" bold="true" box="[199,337,1124,1151]" pageId="7" pageNumber="52">Description</emphasis>
.
</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC1B6FBB0F7CC" blockId="7.[151,1437,945,2015]" box="[199,1068,1160,1187]" pageId="7" pageNumber="52">
<emphasis id="16F0EAE2FF8ED825FF6FC1B6FE98F7CC" bold="true" box="[199,260,1161,1187]" pageId="7" pageNumber="52">Male</emphasis>
.
<typeStatus id="FB3F8852FF8ED825FEBAC1B6FEE6F7CC" box="[274,378,1161,1187]" pageId="7" pageNumber="52" type="holotype">Holotype</typeStatus>
forewing length
<quantity id="E37C9B15FF8ED825FD92C1B6FD17F7CC" box="[570,651,1161,1187]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="52" unit="mm" value="14.0">14 mm</quantity>
,
<typeStatus id="FB3F8852FF8ED825FD3FC1B6FC9EF7CC" box="[663,770,1161,1187]" pageId="7" pageNumber="52" type="paratype">paratypes</typeStatus>
<quantity id="E37C9B15FF8ED825FCA1C1B6FC15F7CC" box="[777,905,1160,1187]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="7" pageNumber="52" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">14 15 mm</quantity>
in both series.
</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC192FC1CF7A8" blockId="7.[151,1437,945,2015]" box="[199,896,1197,1223]" pageId="7" pageNumber="52">Fringes white with narrow blackish inner part from upperside.</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC1EEFB90F65C" blockId="7.[151,1437,945,2015]" pageId="7" pageNumber="52">Upperside common violet-blue with few scattered white scales in costal part of forewing. Unfresh specimens look dull with obvious dark veins, however, this is merely a result of loss of several scales (see specimen on Pl. 1:13). Black margins thin, darkened end of veins usually not clearly expressed.</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC002FD50F6F0" blockId="7.[151,1437,945,2015]" pageId="7" pageNumber="52">Forewing underside dark gray with brownish hue; submarginal series practically fully reduced (only blackish traces of spots sometimes obvious); postdiscal row almost straight, spots have normal size and each spot, including developed discal spot, surrounded by whitish ring.</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC096FD51F5D0" blockId="7.[151,1437,945,2015]" pageId="7" pageNumber="52">Hindwing underside of same colour as forewing underside, postdiscal area sometimes lighter. Bluish-green basal suffusion narrow but obvious because of dark ground colour. Black spots small, reduced in size, basal series often not complete; postdiscal series complete (8 spots), each spot as well as the discal spot surrounded by whitish ring; cubital spot shifted toward the base. Submarginal spots consist of 3 segments—very thin v-shaped blackish inner segment, reduced and pale yellowish-orange segment, and very small metallic plate. Orange segments usually obvious only in anal and cubital spots; metallic plates distinctively bluish as a rule, their external parts darkened and not shining so that shape of the plates becomes rectangular. Thin white lines or whitish triangles usually developed in area between postdiscal and submarginal series.</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC3F6FC4AF58C" blockId="7.[151,1437,945,2015]" box="[199,982,1736,1763]" pageId="7" pageNumber="52">
<emphasis id="16F0EAE2FF8ED825FF6FC3F6FE82F58C" bold="true" box="[199,286,1737,1763]" pageId="7" pageNumber="52">Female</emphasis>
. FW length
<quantity id="E37C9B15FF8ED825FE02C3F6FDA3F58C" box="[426,575,1736,1763]" metricMagnitude="-2" metricUnit="m" metricValue="1.475" metricValueMax="1.55" metricValueMin="1.4" pageId="7" pageNumber="52" unit="mm" value="14.75" valueMax="15.5" valueMin="14.0">14 15.5 mm</quantity>
in both series, one female 13.0 mm.
</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC3D2FB74F468" blockId="7.[151,1437,945,2015]" box="[199,1256,1773,1799]" pageId="7" pageNumber="52">Fringes whitish, with very thin dark inner part joined with blackish thick spots at end of veins.</paragraph>
<paragraph id="243B36F0FF8ED825FF6FC22EFAF6F4D4" blockId="7.[151,1437,945,2015]" pageId="7" pageNumber="52">Upperside brown with deep violet bases of wings, this colour often widely spreads across disc. In latter case, wings violet (shining or usually dull) and darkened towards margins, often with pale whitish glimpses between veins near wide brownish marginal bands. Rare dark forms mainly brownish with bluish-violet bases of wings. Forewing discal spot developed and often outlined by white ring. Hindwing with full series of black oval submarginal spots between darkened veins, each spot surrounded by pale whitish ring; rarely pale yellowish colour also obvious.</paragraph>
<paragraph id="243B36F0FF8ED82AFF6FC2FAFEB3F394" blockId="7.[151,1437,945,2015]" lastBlockId="8.[151,1436,153,575]" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52">Forewing underside brownish with slightly obvious yellowish segments in submarginal spots. Hindwing un- derside brownish-gray, all main features as in males, submarginal spots more developed with obvious but pale and small yellowish-orange segments, metallic plates usually bigger but blackened externally and fully reduced in upper part of series.</paragraph>
<paragraph id="243B36F0FF81D82AFF6FC43BFE76F2E4" blockId="8.[151,1436,153,575]" pageId="8" pageNumber="53">
<emphasis id="16F0EAE2FF81D82AFF6FC43BFE21F270" bold="true" box="[199,445,260,287]" pageId="8" pageNumber="53">Individual variation.</emphasis>
Not so obvious. Butterflies from Zoolon-Suuzhijn-Bulag are statistically darker and dimmer than specimens collected south of Adz-Bogdo. However, coloured form featured in Plate 1: 16 is numerous in both localities. Females are very variable but fully brownish form is not recorded. Forms with s-curved postdiscal underside row are also absent.
</paragraph>
<paragraph id="243B36F0FF81D82AFF6FC4AAFC6AF150" blockId="8.[151,1436,153,575]" pageId="8" pageNumber="53">
<emphasis id="16F0EAE2FF81D82AFF6FC4AAFEE8F2C0" bold="true" box="[199,372,404,431]" pageId="8" pageNumber="53">Male genitalia</emphasis>
(Pl. 3: 4, Pl. 4: 34, Pl. 5: 710). Labis (parts of the uncus) with long and thin ends, which obviously gracile from lateral view. Valva without developed convexities (ventral convexity, however, slightly expressed), with moderately smoothed outlines. Distal projection (“upper lip”) wide, lower distal projection (“lower lip”) thin and very short, end of lip gradually narrowed. Branches of juxta straight in basal parts, then distinctively curved and strongly moved apart while distal ends again moderately straight.
</paragraph>
<caption id="70FB6678FF81D82AFF3FC371FE00F5C7" ID-DOI="http://doi.org/10.5281/zenodo.3530115" ID-Zenodo-Dep="3530115" httpUri="https://zenodo.org/record/3530115/files/figure.png" pageId="8" pageNumber="53" startId="8.[151,234,1614,1639]" targetBox="[158,1429,596,1557]" targetPageId="8">
<paragraph id="243B36F0FF81D82AFF3FC371FE00F5C7" blockId="8.[151,1437,1614,1704]" pageId="8" pageNumber="53">
<emphasis id="16F0EAE2FF81D82AFF3FC371FE9FF508" bold="true" box="[151,259,1614,1639]" pageId="8" pageNumber="53">PLATE 3.</emphasis>
Male genitalia of
<taxonomicName id="E3844D73FF81D82AFE6AC371FD85F509" authorityName="Kluk" authorityYear="1780" box="[450,537,1614,1638]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="16F0EAE2FF81D82AFE6AC371FD85F509" box="[450,537,1614,1638]" italics="true" pageId="8" pageNumber="53">Plebejus</emphasis>
</taxonomicName>
spp. (lateral view, aedeagus is separated): 1—
<taxonomicName id="E3844D73FF81D82AFC45C370FBFDF509" authority="Churkin &amp; Kolesnichenko &amp; Yakovlev, 2019" authorityName="Churkin &amp; Kolesnichenko &amp; Yakovlev" authorityYear="2019" box="[1005,1121,1614,1638]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="chrisreai" status="sp. nov.">
<emphasis id="16F0EAE2FF81D82AFC45C370FBFDF509" box="[1005,1121,1614,1638]" italics="true" pageId="8" pageNumber="53">P. chrisreai</emphasis>
</taxonomicName>
<emphasis id="16F0EAE2FF81D82AFBCFC370FB03F508" bold="true" box="[1127,1183,1615,1639]" pageId="8" pageNumber="53">
<taxonomicNameLabel id="0DC35799FF81D82AFBCFC370FB03F508" box="[1127,1183,1615,1639]" pageId="8" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratype, Ushig); 2—
<taxonomicName id="E3844D73FF81D82AFA22C370FF77F5EB" authorityName="Yakovlev" authorityYear="2012" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="germani">
<emphasis id="16F0EAE2FF81D82AFA22C370FF77F5EB" italics="true" pageId="8" pageNumber="53">P. germani</emphasis>
</taxonomicName>
(paratype, 30 km NNW of Bulgan); 3—
<taxonomicName id="E3844D73FF81D82AFD24C352FCA5F5EB" authority="anikini" box="[652,825,1644,1668]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="anikini" subSpecies="anikini">
<emphasis id="16F0EAE2FF81D82AFD24C352FCA5F5EB" box="[652,825,1644,1668]" italics="true" pageId="8" pageNumber="53">P. anikini anikini</emphasis>
</taxonomicName>
(paratype, Alag Nuur); 4—
<emphasis id="16F0EAE2FF81D82AFBFDC352FB70F5EB" box="[1109,1260,1644,1668]" italics="true" pageId="8" pageNumber="53">P. a. azhbogdo</emphasis>
<emphasis id="16F0EAE2FF81D82AFB5BC353FAB3F5EB" bold="true" box="[1267,1327,1644,1668]" pageId="8" pageNumber="53">ssp.n.</emphasis>
(paratype, 10 km ESE Altai somon).
</paragraph>
</caption>
<paragraph id="243B36F0FF81D82AFF6FC3E9FC79F432" blockId="8.[151,1437,1750,2029]" pageId="8" pageNumber="53">Aedeagus thin, proximal part with developed ventral inflation but not too widened compared with distal part, latter not thin but with narrowing in the middle and widened near distal end. Aedeagus curved, curving moved distally from joining of proximal and distal parts—thus joining place straight, and distal part further curved downwards. Curving weakly expressed. Proximal part no shorter than distal part.</paragraph>
<paragraph id="243B36F0FF81D82AFF6FC259FD25F4CA" blockId="8.[151,1437,1750,2029]" pageId="8" pageNumber="53">Characters of genitalia in two studied populations very similar, only butterflies from Zoolon-Suuzhijn-Bulag have heavier aedeagus with wider proximal part.</paragraph>
</subSubSection>
<subSubSection id="6C9E657BFF81D82BFF6FC292FE02F4AA" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53" type="diagnosis">
<paragraph id="243B36F0FF81D82AFF6FC292FDABF483" blockId="8.[151,1437,1750,2029]" pageId="8" pageNumber="53">
<emphasis id="16F0EAE2FF81D82AFF6FC292FEDEF4A7" bold="true" box="[199,322,1965,1992]" pageId="8" pageNumber="53">Diagnosis.</emphasis>
The size of both subspecies of
<taxonomicName id="E3844D73FF81D82AFD0BC290FC91F4A7" authorityName="Yakovlev" authorityYear="2012" box="[675,781,1966,1992]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="anikini">
<emphasis id="16F0EAE2FF81D82AFD0BC290FC91F4A7" box="[675,781,1966,1992]" italics="true" pageId="8" pageNumber="53">P. anikini</emphasis>
</taxonomicName>
is nearly the same as in
<taxonomicName id="E3844D73FF81D82AFB89C290FB06F4A7" authorityName="Yakovlev" authorityYear="2012" box="[1057,1178,1967,1992]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="germani">
<emphasis id="16F0EAE2FF81D82AFB89C290FB06F4A7" box="[1057,1178,1967,1992]" italics="true" pageId="8" pageNumber="53">P. germani</emphasis>
</taxonomicName>
, but really large specimens (over
<quantity id="E37C9B15FF81D82AFEB3C2EEFE1CF482" box="[283,384,2001,2029]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="8" pageNumber="53" unit="mm" value="15.5">15.5 mm</quantity>
) are not known.
</paragraph>
<paragraph id="243B36F0FF80D82BFF6FC5A5FF7EF22C" blockId="9.[151,1437,153,323]" pageId="9" pageNumber="54">
<taxonomicName id="E3844D73FF80D82BFF6FC5A5FEACF3DC" authorityName="Yakovlev" authorityYear="2012" box="[199,304,153,179]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="anikini">
<emphasis id="16F0EAE2FF80D82BFF6FC5A5FEACF3DC" box="[199,304,153,179]" italics="true" pageId="9" pageNumber="54">P. anikini</emphasis>
</taxonomicName>
can be easily differentiated from both other Mongolian relatives by a reduced submarginal pattern on the hindwing underside, pale and often unobvious orange lunules and thin dull metallic plates. Compared with
<taxonomicName id="E3844D73FF80D82BFA20C581FF68F394" authorityName="Yakovlev" authorityYear="2012" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="germani">
<emphasis id="16F0EAE2FF80D82BFA20C581FF68F394" italics="true" pageId="9" pageNumber="54">P. germani</emphasis>
</taxonomicName>
,
<taxonomicName id="E3844D73FF80D82BFEAAC5DDFEF7F394" authorityName="Yakovlev" authorityYear="2012" box="[258,363,225,251]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="anikini">
<emphasis id="16F0EAE2FF80D82BFEAAC5DDFEF7F394" box="[258,363,225,251]" italics="true" pageId="9" pageNumber="54">P. anikini</emphasis>
</taxonomicName>
has a brownish (not grayish) underside ground colour, thus, all black spots are distinctively surrounded by the whitish rings, the size of the spots is obviously smaller. The differences with
<taxonomicName id="E3844D73FF80D82BFB06C439FAB2F270" authorityName="Churkin &amp; Kolesnichenko &amp; Yakovlev" authorityYear="2019" box="[1198,1326,261,287]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="chrisreai">
<emphasis id="16F0EAE2FF80D82BFB06C439FAB2F270" box="[1198,1326,261,287]" italics="true" pageId="9" pageNumber="54">P. chrisreai</emphasis>
</taxonomicName>
are listed above.
</paragraph>
<caption id="70FB6678FF80D82BFF3FC14FFBDDF7C3" ID-DOI="http://doi.org/10.5281/zenodo.3530117" ID-Zenodo-Dep="3530117" httpUri="https://zenodo.org/record/3530117/files/figure.png" pageId="9" pageNumber="54" startId="9.[151,234,1136,1161]" targetBox="[162,1430,345,1103]" targetPageId="9">
<paragraph id="243B36F0FF80D82BFF3FC14FFBDDF7C3" blockId="9.[151,1437,1136,1197]" pageId="9" pageNumber="54">
<emphasis id="16F0EAE2FF80D82BFF3FC14FFE9FF7E6" bold="true" box="[151,259,1136,1161]" pageId="9" pageNumber="54">PLATE 4.</emphasis>
Valva of
<taxonomicName id="E3844D73FF80D82BFECCC14FFE27F7E7" authorityName="Kluk" authorityYear="1780" box="[356,443,1136,1160]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="16F0EAE2FF80D82BFECCC14FFE27F7E7" box="[356,443,1136,1160]" italics="true" pageId="9" pageNumber="54">Plebejus</emphasis>
</taxonomicName>
spp. (lateral view; 1, 2, 3—with joined juxta): 1—
<taxonomicName id="E3844D73FF80D82BFC15C14EFBADF7E7" authority="Churkin &amp; Kolesnichenko &amp; Yakovlev, 2019" authorityName="Churkin &amp; Kolesnichenko &amp; Yakovlev" authorityYear="2019" box="[957,1073,1136,1160]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="chrisreai" status="sp. nov.">
<emphasis id="16F0EAE2FF80D82BFC15C14EFBADF7E7" box="[957,1073,1136,1160]" italics="true" pageId="9" pageNumber="54">P. chrisreai</emphasis>
</taxonomicName>
<emphasis id="16F0EAE2FF80D82BFB9EC14FFBF3F7E7" bold="true" box="[1078,1135,1136,1160]" pageId="9" pageNumber="54">
<taxonomicNameLabel id="0DC35799FF80D82BFB9EC14FFBF3F7E7" box="[1078,1135,1136,1160]" pageId="9" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratype, Ushig); 2—
<taxonomicName id="E3844D73FF80D82BFAF2C14EFF56F7C3" authorityName="Yakovlev" authorityYear="2012" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="germani">
<emphasis id="16F0EAE2FF80D82BFAF2C14EFF56F7C3" italics="true" pageId="9" pageNumber="54">P. germani</emphasis>
</taxonomicName>
(35 km SW Uench); 3, 4—
<emphasis id="16F0EAE2FF80D82BFE4AC1AAFDEBF7C3" box="[482,631,1172,1196]" italics="true" pageId="9" pageNumber="54">P. a. azhbogdo</emphasis>
<emphasis id="16F0EAE2FF80D82BFDD5C1ABFD5CF7C3" bold="true" box="[637,704,1172,1196]" pageId="9" pageNumber="54">ssp. n.</emphasis>
(paratypes, 10 km ESE Altai somon).
</paragraph>
</caption>
<paragraph id="243B36F0FF80D82BFF6FC1C1FD8DF6EB" blockId="9.[151,1437,1278,1989]" pageId="9" pageNumber="54">
The new subspecies differs from the nominotypical
<taxonomicName id="E3844D73FF80D82BFCB7C1C0FC17F676" authorityName="Yakovlev" authorityYear="2012" box="[799,907,1279,1305]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="anikini">
<emphasis id="16F0EAE2FF80D82BFCB7C1C0FC17F676" box="[799,907,1279,1305]" italics="true" pageId="9" pageNumber="54">P. anikini</emphasis>
</taxonomicName>
by the following distinctions: females always have developed violet-blue colour on upperside, submarginal pattern on the hindwing underside is strongly reduced so that the metallic plates often lose the rounded shape and half (at least) of the submarginal spots have neither an orange colour nor metallic scales.
</paragraph>
<paragraph id="243B36F0FF80D82BFF6FC0B1FB6FF57A" blockId="9.[151,1437,1278,1989]" pageId="9" pageNumber="54">
The new taxon belongs to
<taxonomicName id="E3844D73FF80D82BFE43C0B0FDCFF6C6" authorityName="Yakovlev" authorityYear="2012" box="[491,595,1423,1449]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="anikini">
<emphasis id="16F0EAE2FF80D82BFE43C0B0FDCFF6C6" box="[491,595,1423,1449]" italics="true" pageId="9" pageNumber="54">P. anikini</emphasis>
</taxonomicName>
because it has the same combination of specific features of the male genita-lia—labis are thin and long, the valva has no visible sharp convexities at the dorsal and ventral surfaces (contrary to
<taxonomicName id="E3844D73FF80D82BFF3FC0E8FE93F69F" authorityName="Yakovlev" authorityYear="2012" box="[151,271,1495,1520]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="germani">
<emphasis id="16F0EAE2FF80D82BFF3FC0E8FE93F69F" box="[151,271,1495,1520]" italics="true" pageId="9" pageNumber="54">P. germani</emphasis>
</taxonomicName>
), the lower lip is very narrow and shortened (contrary to both other species), the curving of the aedeagus is distally moved from the place of joining of the proximal and distal parts (contrary to
<taxonomicName id="E3844D73FF80D82BFBC1C0C4FB7EF57B" authorityName="Yakovlev" authorityYear="2012" box="[1129,1250,1531,1556]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="germani">
<emphasis id="16F0EAE2FF80D82BFBC1C0C4FB7EF57B" box="[1129,1250,1531,1556]" italics="true" pageId="9" pageNumber="54">P. germani</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="243B36F0FF80D82BFF6FC320FD0DF5CB" blockId="9.[151,1437,1278,1989]" pageId="9" pageNumber="54">
Some characters of the genitalia look even more similar to that of
<taxonomicName id="E3844D73FF80D82BFC1AC320FBAEF556" authorityName="Churkin &amp; Kolesnichenko &amp; Yakovlev" authorityYear="2019" box="[946,1074,1567,1593]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="chrisreai">
<emphasis id="16F0EAE2FF80D82BFC1AC320FBAEF556" box="[946,1074,1567,1593]" italics="true" pageId="9" pageNumber="54">P. chrisreai</emphasis>
</taxonomicName>
, except for the shape of the lips (the ratio of the distal and proximal parts of the aedeagus is also different). The branches of juxta are not straight but not gradually convex—it looks absolutely specific but the juxta of
<taxonomicName id="E3844D73FF80D82BFC2EC358FC6CF5EE" authorityName="Yakovlev" authorityYear="2012" box="[902,1008,1639,1665]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="anikini">
<emphasis id="16F0EAE2FF80D82BFC2EC358FC6CF5EE" box="[902,1008,1639,1665]" italics="true" pageId="9" pageNumber="54">P. anikini</emphasis>
</taxonomicName>
is not studied because the genitalia of the available male were not cut into sclerites.
</paragraph>
<paragraph id="243B36F0FF80D82BFF6FC391FAE0F437" blockId="9.[151,1437,1278,1989]" pageId="9" pageNumber="54">
Compared with the nominate taxon, the aedeagus of
<emphasis id="16F0EAE2FF80D82BFC8EC390FC53F5A6" box="[806,975,1711,1737]" italics="true" pageId="9" pageNumber="54">P. a. azhbogdo</emphasis>
is more curved (however, this character seems variable), the lower lip has a different shape (in
<emphasis id="16F0EAE2FF80D82BFD57C3ECFC19F582" box="[767,901,1747,1773]" italics="true" pageId="9" pageNumber="54">
P. a.
<taxonomicName id="E3844D73FF80D82BFC9FC3ECFC19F582" authorityName="Yakovlev" authorityYear="2012" box="[823,901,1747,1773]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="anikini">anikini</taxonomicName>
</emphasis>
the lower lip is very short and very narrow distally, while in the new taxon it is only gradually narrowed), the cutout between the lips is no more developed than in
<taxonomicName id="E3844D73FF80D82BFF3FC224FE93F45B" authorityName="Yakovlev" authorityYear="2012" box="[151,271,1819,1844]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="germani">
<emphasis id="16F0EAE2FF80D82BFF3FC224FE93F45B" box="[151,271,1819,1844]" italics="true" pageId="9" pageNumber="54">P. germani</emphasis>
</taxonomicName>
or
<taxonomicName id="E3844D73FF80D82BFE9BC224FE38F45B" box="[307,420,1819,1844]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="crisreai">
<emphasis id="16F0EAE2FF80D82BFE9BC224FE38F45B" box="[307,420,1819,1844]" italics="true" pageId="9" pageNumber="54">P. crisreai</emphasis>
</taxonomicName>
, while in
<taxonomicName id="E3844D73FF80D82BFDBAC224FDE7F45A" authorityName="Yakovlev" authorityYear="2012" box="[530,635,1819,1845]" class="Insecta" family="Lycaenidae" genus="Plebejus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="54" phylum="Arthropoda" rank="species" species="anikini">
<emphasis id="16F0EAE2FF80D82BFDBAC224FDE7F45A" box="[530,635,1819,1845]" italics="true" pageId="9" pageNumber="54">P. anikini</emphasis>
</taxonomicName>
this cutout is spectacularly expressed. The taxonomic value of the latter feature is unknown. Theoretically, if all these distinctions are confirmed, the status of the taxon can be changed.
</paragraph>
<paragraph id="243B36F0FF80D82BFF6FC25DFE02F4AA" blockId="9.[151,1437,1278,1989]" pageId="9" pageNumber="54">
<emphasis id="16F0EAE2FF80D82BFF6FC25DFEDFF412" bold="true" box="[199,323,1890,1917]" pageId="9" pageNumber="54">Bionomics</emphasis>
(Pl. 6: 2). The true foodplant is unknown. Both series were collected on flowers and grass near the spring, the species of bush surrounding the oasis is unknown. The situation seems very similar to that of nominate subspecies (see above).
</paragraph>
</subSubSection>
<subSubSection id="6C9E657BFF83D828FF6FC5A7FD86F3B8" pageId="10" pageNumber="55" type="distribution">
<paragraph id="243B36F0FF83D828FF6FC5A7FD86F3B8" blockId="10.[199,1350,152,215]" pageId="10" pageNumber="55">
<emphasis id="16F0EAE2FF83D828FF6FC5A7FEC5F3DC" bold="true" box="[199,345,152,179]" pageId="10" pageNumber="55">Distribution</emphasis>
(Pl. 7). The subspecies inhabits foothills of the Azh-Bogdo massive in Transaltaian Gobi.
<emphasis id="16F0EAE2FF83D828FF6FC582FED4F3B8" bold="true" box="[199,328,189,215]" pageId="10" pageNumber="55">Etymology</emphasis>
. Toponimic name.
</paragraph>
</subSubSection>
</treatment>
</document>