128 lines
16 KiB
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128 lines
16 KiB
XML
<document id="456B3E3E29C3D9327B16D7C1B9C841A3" ID-DOI="10.11646/zootaxa.5448.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11231272" ID-ZooBank="08716B9F-13DF-4661-B2F6-BE8AE565DE97" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1716278333334" checkinUser="plazi" docAuthor="Samathi, Adun" docDate="2024" docId="AC0D87DA932C3A7236F0BACCDBD92527" docLanguage="en" docName="zootaxa.5448.1.4.pdf" docOrigin="Zootaxa 5448 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kinnareemimus khonkaenensis Buffetaut, Suteethorn, and Tong 2009" docType="treatment" docVersion="4" lastPageNumber="72" masterDocId="5034FFA293283A773667BB33D92E243A" masterDocTitle="Phylogenetic position of Kinnareemimus khonkaenensis (Dinosauria: Theropoda: Ornithomimosauria) from the Lower Cretaceous of Thailand" masterLastPageNumber="84" masterPageNumber="67" pageNumber="71" updateTime="1716398006620" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A1F09FD1651235BE9F4D8D8828A4B013">Phylogenetic position of Kinnareemimus khonkaenensis (Dinosauria: Theropoda: Ornithomimosauria) from the Lower Cretaceous of Thailand</mods:title>
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<mods:namePart id="98248A04D78F7BB9199F81B1CAA5F0F8">Samathi, Adun</mods:namePart>
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<mods:affiliation id="EE4C9451E0FFF0237D60C29E6BA22806">Biodiversity and Conservation Research Unit, Walai Rukhavej Botanical Research Institute, Mahasarakham University, Maha Sarakham, Thailand. & Dinosaur Research Unit, Mahasarakham University, Maha Sarakham, Thailand.</mods:affiliation>
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<mods:title id="9B72374D6F34A5246C2A86B745011F58">Zootaxa</mods:title>
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<mods:date id="913F81FEAA45D8A978651DF546D66395">2024</mods:date>
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<mods:number id="E5C22B5E7C8471EF7509E8CC36ED6AF3">2024-05-09</mods:number>
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<mods:identifier id="5A282A4C622D7949E8836394708A54C1" type="DOI">10.11646/zootaxa.5448.1.4</mods:identifier>
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<treatment id="AC0D87DA932C3A7236F0BACCDBD92527" ID-DOI="http://doi.org/10.5281/zenodo.11245414" ID-Zenodo-Dep="11245414" LSID="urn:lsid:plazi:treatment:AC0D87DA932C3A7236F0BACCDBD92527" httpUri="http://treatment.plazi.org/id/AC0D87DA932C3A7236F0BACCDBD92527" lastPageId="5" lastPageNumber="72" pageId="4" pageNumber="71">
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<subSubSection id="6CBE6547932C3A7336F0BACCDAD42623" box="[151,1018,511,538]" pageId="4" pageNumber="71" type="nomenclature">
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<paragraph id="241B36CC932C3A7336F0BACCDAD42623" blockId="4.[151,1018,511,538]" box="[151,1018,511,538]" pageId="4" pageNumber="71">
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<heading id="7F5381A0932C3A7336F0BACCDAD42623" bold="true" box="[151,1018,511,538]" fontSize="11" level="1" pageId="4" pageNumber="71" reason="1">
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<taxonomicName id="E3A44D4F932C3A7336F0BACCDAD42623" authority="Buffetaut, Suteethorn, and Tong, 2009" authorityName="Buffetaut, Suteethorn, and Tong" authorityYear="2009" box="[151,1018,511,538]" class="Reptilia" family="Ornithomimidae" genus="Kinnareemimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="71" phylum="Chordata" rank="species" species="khonkaenensis">
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<emphasis id="16D0EADE932C3A7336F0BACCDAD42623" bold="true" box="[151,1018,511,538]" pageId="4" pageNumber="71">
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<emphasis id="16D0EADE932C3A7336F0BACCDB3B2620" bold="true" box="[151,533,511,538]" italics="true" pageId="4" pageNumber="71">Kinnareemimus khonkaenensis</emphasis>
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<bibRefCitation id="40354B3D932C3A73347ABACCDAD42623" author="Buffetaut, E. & Suteethorn, V. & Tong, H." box="[541,1018,511,537]" pageId="4" pageNumber="71" pagination="229 - 243" refId="ref7825" refString="Buffetaut, E., Suteethorn, V. & Tong, H. (2009) An early ' ostrich dinosaur' (Theropoda: Ornithomimosauria) from the Early Cretaceous Sao Khua Formation of NE Thailand. In: Buffetaut, E., Cuny, G., Le Loeuff, J. & Suteethorn, V. (Eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315. Geological Society, London, pp. 229 - 243. https: // doi. org / 10.1144 / SP 315.16" type="book chapter" year="2009">Buffetaut, Suteethorn, and Tong, 2009</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6CBE6547932C3A7336F0B974DA5123CA" pageId="4" pageNumber="71" type="materials_examined">
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<paragraph id="241B36CC932C3A7336F0B974D9C726F7" blockId="4.[151,1437,583,717]" pageId="4" pageNumber="71">
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<emphasis id="16D0EADE932C3A7336F0B974D9D1265B" bold="true" box="[151,255,583,609]" pageId="4" pageNumber="71">Material</emphasis>
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: The
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<typeStatus id="FB1F886E932C3A733726B974D88D265B" box="[321,419,583,609]" pageId="4" pageNumber="71" type="holotype">holotype</typeStatus>
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SM-PW5A-100 consists of an incomplete left third metatarsal with the distal end and part of the shaft. The referred materials consist of metatarsals, pedal phalanges, pedal ungual phalanx, tibiae, fibula, pubis fragment, and dorsal and caudal vertebrae (
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<figureCitation id="BC9F2A49932C3A7334A4B9BCDA282693" box="[707,774,655,681]" captionStart="FIGURE 3" captionStartId="3.[151,250,1121,1145]" captionTargetBox="[162,1425,192,1088]" captionTargetId="figure-16@3.[151,1436,181,1097]" captionTargetPageId="3" captionText="FIGURE 3. Left tibia of Kinnareemimus in anterior (A, B), posterior (C), medial (D), lateral (E, F), proximal (G, H), and distal (I, J) views. Left fibula of Kinnareemimus in proximal (K) and lateral (L) views. Abbreviations: cc, cnemial crest; fc, fibular crest; ig, intercondylar groove; it, incisura tibialis; lc, lateral condyle; lm, lateral malleolus; mc, medial condyle; mm, medial malleolus. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.11231282" httpUri="https://zenodo.org/record/11231282/files/figure.png" pageId="4" pageNumber="71">Fig. 3</figureCitation>
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,
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<figureCitation id="BC9F2A49932C3A733573B9BCDA0B2693" box="[788,805,655,681]" captionStart="FIGURE 4" captionStartId="4.[151,250,1683,1707]" captionTargetBox="[187,1401,751,1650]" captionTargetId="figure-111@4.[177,1411,741,1660]" captionTargetPageId="4" captionText="FIGURE 4. Left metatarsals of Kinnareemimus in proximal (A, C, G, N), anterior (B, D, E, I, J), posterior (H, L), and distal (F, K, M) views. Incomplete left metatarsal III SM-PW5A-100 (E-H), mid shaft of the left metatarsal III SM-PW5A-103 (I, L, M), and distal portion of the left metatarsal III SM-PW5A-107 (J, K, N). Cross section of metatarsal III SM-PW5A-100 (G), SM-PW5A-103 (M), and SM-PW5A-107 (N). Abbreviation: bif: bifurcation of posterior ridge; elp: extensor ligament pit; nap: nonarticular platform; prid: posterior ridge; arrowhead = medial expansion. Scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.11231286" httpUri="https://zenodo.org/record/11231286/files/figure.png" pageId="4" pageNumber="71">4</figureCitation>
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). For the description of the material, see
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<bibRefCitation id="40354B3D932C3A733294B9BCD9CB26F7" author="Buffetaut, E. & Suteethorn, V. & Tong, H." pageId="4" pageNumber="71" pagination="229 - 243" refId="ref7825" refString="Buffetaut, E., Suteethorn, V. & Tong, H. (2009) An early ' ostrich dinosaur' (Theropoda: Ornithomimosauria) from the Early Cretaceous Sao Khua Formation of NE Thailand. In: Buffetaut, E., Cuny, G., Le Loeuff, J. & Suteethorn, V. (Eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315. Geological Society, London, pp. 229 - 243. https: // doi. org / 10.1144 / SP 315.16" type="book chapter" year="2009">
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Buffetaut
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<emphasis id="16D0EADE932C3A733303B9A3DCBB2693" box="[1380,1429,655,681]" italics="true" pageId="4" pageNumber="71">et al</emphasis>
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. (2009)
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</bibRefCitation>
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.
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</paragraph>
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<caption id="70DB6644932C3A7336F0BDA0DD652301" ID-DOI="http://doi.org/10.5281/zenodo.11231286" ID-Zenodo-Dep="11231286" httpUri="https://zenodo.org/record/11231286/files/figure.png" pageId="4" pageNumber="71" startId="4.[151,250,1683,1707]" targetBox="[187,1401,751,1650]" targetPageId="4" targetType="figure">
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<paragraph id="241B36CC932C3A7336F0BDA0DD652301" blockId="4.[151,1437,1683,1851]" pageId="4" pageNumber="71">
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<emphasis id="16D0EADE932C3A7336F0BDA0D83A2291" bold="true" box="[151,276,1683,1707]" pageId="4" pageNumber="71">FIGURE 4.</emphasis>
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Left metatarsals of
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<taxonomicName id="E3A44D4F932C3A733784BDA7DBAC2291" box="[483,642,1684,1707]" class="Reptilia" genus="Kinnareemimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="4" pageNumber="71" phylum="Chordata" rank="genus">
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<emphasis id="16D0EADE932C3A733784BDA7DBAC2291" box="[483,642,1684,1707]" italics="true" pageId="4" pageNumber="71">Kinnareemimus</emphasis>
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</taxonomicName>
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in proximal (A, C, G, N), anterior (B, D, E, I, J), posterior (H, L), and distal (F, K, M) views. Incomplete left metatarsal III SM-PW5A-100 (E-H), mid shaft of the left metatarsal III SM-PW5A-103 (I, L, M), and distal portion of the left metatarsal III SM-PW5A-107 (J, K, N). Cross section of metatarsal III SM-PW5A-100 (G), SM-PW5A-103 (M), and SM-PW5A-107 (N). Abbreviation: bif: bifurcation of posterior ridge; elp: extensor ligament pit; nap: nonarticular platform; prid: posterior ridge; arrowhead = medial expansion. Scale bar = 5 cm.
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</paragraph>
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</caption>
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<paragraph id="241B36CC932C3A7336A0BC5AD82C2392" blockId="4.[151,1436,1897,2032]" pageId="4" pageNumber="71">
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<emphasis id="16D0EADE932C3A7336A0BC5AD80723B9" bold="true" box="[199,297,1897,1923]" pageId="4" pageNumber="71">Locality</emphasis>
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: Phu Wiang 5 Locality, so-called “Sum Ya Ka”, Phu Wiang Mountain,
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<collectingRegion id="E660F82E932C3A7332FBBC59DCB623BE" box="[1180,1432,1898,1924]" country="Thailand" name="Khon Kaen" pageId="4" pageNumber="71">Khon Kaen Province</collectingRegion>
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,
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<collectingCountry id="5CB3765C932C3A7336F0BCBDD9D02392" box="[151,254,1934,1960]" name="Thailand" pageId="4" pageNumber="71">Thailand</collectingCountry>
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.
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</paragraph>
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<paragraph id="241B36CC932C3A7336A0BC82DA5123CA" blockId="4.[151,1436,1897,2032]" pageId="4" pageNumber="71">
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<emphasis id="16D0EADE932C3A7336A0BC82D80723F1" bold="true" box="[199,297,1969,1995]" pageId="4" pageNumber="71">Horizon</emphasis>
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: Sao Khua Formation, Early Cretaceous, ~late Barremian (see
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<bibRefCitation id="40354B3D932C3A733598BC81DDDE23F6" author="Buffetaut, E. & Suteethorn, V. & Tong, H." box="[1023,1264,1970,1996]" pageId="4" pageNumber="71" pagination="229 - 243" refId="ref7825" refString="Buffetaut, E., Suteethorn, V. & Tong, H. (2009) An early ' ostrich dinosaur' (Theropoda: Ornithomimosauria) from the Early Cretaceous Sao Khua Formation of NE Thailand. In: Buffetaut, E., Cuny, G., Le Loeuff, J. & Suteethorn, V. (Eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315. Geological Society, London, pp. 229 - 243. https: // doi. org / 10.1144 / SP 315.16" type="book chapter" year="2009">
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Buffetaut
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<emphasis id="16D0EADE932C3A733215BC80DD8B23F6" box="[1138,1189,1970,1996]" italics="true" pageId="4" pageNumber="71">et al</emphasis>
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. 2009
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</bibRefCitation>
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;
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<bibRefCitation id="40354B3D932C3A73329ABC81D9F123CA" author="Samathi, A. & Chanthasit, P. & Sander, P. M." pageId="4" pageNumber="71" pagination="201 - 215" refId="ref11292" refString="Samathi, A., Chanthasit, P. & Sander, P. M. (2019 b) A review of theropod dinosaurs from the Late Jurassic to mid-Cretaceous of Southeast Asia. Annales de Paleontologie, 105 (3), 201 - 215. https: // doi. org / 10.1016 / j. annpal. 2019.03.003." type="journal article" year="2019">
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Samathi
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<emphasis id="16D0EADE932C3A733305BC80DCBB23F6" box="[1378,1429,1970,1996]" italics="true" pageId="4" pageNumber="71">et al</emphasis>
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. 2019b
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</bibRefCitation>
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) or late Valanginian–early Hauterivian (
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<bibRefCitation id="40354B3D932C3A7334C7BCE5DA5D23CA" author="Tucker, R. T. & Hyland, E. G. & Gates, T. A. & King, M. R. & Roberts, E. M. & Foley, E. K. & Berndt, D. & Hanta, R. & Khansubha, S. & Aswasereelert, W. & Zanno, L. E." box="[672,883,2006,2032]" pageId="4" pageNumber="71" pagination="111107" refId="ref11891" refString="Tucker, R. T., Hyland, E. G., Gates, T. A., King, M. R., Roberts, E. M., Foley, E. K., Berndt, D., Hanta, R., Khansubha, S., Aswasereelert, W. & Zanno, L. E. (2022) Age, depositional history, and paleoclimatic setting of Early Cretaceous dinosaur assemblages from the Sao Khua Formation (Khorat Group), Thailand. Palaeogeography, Palaeoclimatology, Palaeoecology, 601, 111107. https: // doi. org / 10.1016 / j. palaeo. 2022.111107" type="journal article" year="2022">
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Tucker
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<emphasis id="16D0EADE932C3A733490BCE4DA0623CA" box="[759,808,2006,2032]" italics="true" pageId="4" pageNumber="71">et al</emphasis>
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. 2022
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="6CBE6547932D3A7236A0BBA5DBD92527" pageId="5" pageNumber="72" type="diagnosis">
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<paragraph id="241B36CC932D3A7236A0BBA5DBD92527" blockId="5.[151,1437,150,285]" pageId="5" pageNumber="72">
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<emphasis id="16D0EADE932D3A7236A0BBA5D814248A" bold="true" box="[199,314,150,176]" pageId="5" pageNumber="72">Diagnosis</emphasis>
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: An ornithomimosaur with a combination of the following characters: pedal unguals ventrally flattened, with flexor fossa (as in other ornithomimosaurs), subarctometatarsalian condition of the metatarsals (as in basal ornithomimosaurs), and a triangular cross-section of the metatarsal III closer to the distal end (as in derived ornithomimids) (modified from
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<bibRefCitation id="40354B3D932D3A723798BA30DBC52527" author="Buffetaut, E. & Suteethorn, V. & Tong, H." box="[511,747,259,285]" pageId="5" pageNumber="72" pagination="229 - 243" refId="ref7825" refString="Buffetaut, E., Suteethorn, V. & Tong, H. (2009) An early ' ostrich dinosaur' (Theropoda: Ornithomimosauria) from the Early Cretaceous Sao Khua Formation of NE Thailand. In: Buffetaut, E., Cuny, G., Le Loeuff, J. & Suteethorn, V. (Eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315. Geological Society, London, pp. 229 - 243. https: // doi. org / 10.1144 / SP 315.16" type="book chapter" year="2009">
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Buffetaut
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<emphasis id="16D0EADE932D3A723408BA30DB8F2527" box="[623,673,259,285]" italics="true" pageId="5" pageNumber="72">et al</emphasis>
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||
. 2009
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</bibRefCitation>
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||
).
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</paragraph>
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||
</subSubSection>
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||
</treatment>
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||
</document> |