252 lines
32 KiB
XML
252 lines
32 KiB
XML
<document id="A5B25952C33AA61E69D5BA22B2F7E1D1" ID-DOI="10.11646/zootaxa.5514.6.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13914946" ID-ZooBank="5060EA2A-C13C-49DC-90F4-806DB55DC2FE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728582282270" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="AB068796BC4CFFD9FF152FF501849F35" docLanguage="en" docName="zootaxa.5514.6.7.pdf" docOrigin="Zootaxa 5514 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.6.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphlosotiba zingiberiphila Shavrin, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="595" masterDocId="573FFFEEBC49FFDFFF82296805389844" masterDocTitle="New species and records of Paraphloeostiba Steel, 1960 and Xanthonomus Bernhauer, 1926 from the eastern Palaearctic, Oriental and Papuan regions (Coleoptera: Staphylinidae: Omaliinae)" masterLastPageNumber="597" masterPageNumber="589" pageNumber="594" updateTime="1728583085933" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FA46F99FE3C8B845BD9592D8E61CEC72">New species and records of Paraphloeostiba Steel, 1960 and Xanthonomus Bernhauer, 1926 from the eastern Palaearctic, Oriental and Papuan regions (Coleoptera: Staphylinidae: Omaliinae)</mods:title>
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<mods:namePart id="0FEE23C41BCC925011A178177583CDB2">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="31660000EA295E2C7265F01B49F716DB">text</mods:typeOfResource>
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<mods:title id="39D65DA4B048EAF992984382EE2DFFB6">Zootaxa</mods:title>
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<mods:part id="D889B686796ECEF19BD68316BF240362">
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<mods:date id="7C451CBB704C5A891E7C17AAC2FE0BA3">2024</mods:date>
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<mods:number id="A24F90DEB6FF9BBB8FE69143D9AF6E6A">2024-10-02</mods:number>
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<mods:number id="801493BBB2589F60074F3299994AE895">5514</mods:number>
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<mods:number id="AB41B3115078D8AAD41A9A9410388CE6">6</mods:number>
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<mods:start id="3E09B91CF276678A940A5614A1EE15E3">589</mods:start>
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<mods:url id="D04918B6700C68CD5166A90AB6CBB83E">http://dx.doi.org/10.11646/zootaxa.5514.6.7</mods:url>
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<mods:classification id="9F6D324405FD277B1CB21CC794CB9ECC">journal article</mods:classification>
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<mods:identifier id="B5820DE42B5E44AD82917CEC818B48BF" type="DOI">10.11646/zootaxa.5514.6.7</mods:identifier>
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<mods:identifier id="BF4B0630C5B8910DAF201F3A910B5DEC" type="ISSN">1175-5326</mods:identifier>
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<treatment id="AB068796BC4CFFD9FF152FF501849F35" LSID="urn:lsid:plazi:treatment:AB068796BC4CFFD9FF152FF501849F35" httpUri="http://treatment.plazi.org/id/AB068796BC4CFFD9FF152FF501849F35" lastPageId="6" lastPageNumber="595" pageId="5" pageNumber="594">
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<subSubSection id="6BB5650BBC4CFFDAFF152FF5075B9EFC" box="[151,611,1693,1720]" pageId="5" pageNumber="594" type="nomenclature">
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<paragraph id="23103680BC4CFFDAFF152FF5075B9EFC" blockId="5.[151,611,1693,1756]" box="[151,611,1693,1720]" pageId="5" pageNumber="594">
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<heading id="785881ECBC4CFFDAFF152FF5075B9EFC" bold="true" box="[151,611,1693,1720]" fontSize="11" level="1" pageId="5" pageNumber="594" reason="1">
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<emphasis id="11DBEA92BC4CFFDAFF152FF5075B9EFC" bold="true" box="[151,611,1693,1720]" pageId="5" pageNumber="594">
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<taxonomicName id="E4AF4D03BC4CFFDAFF152FF504C79EFC" authorityName="Shavrin" authorityYear="2024" box="[151,511,1693,1720]" class="Insecta" family="Staphylinidae" genus="Paraphlosotiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="zingiberiphila" status="sp. nov.">
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<emphasis id="11DBEA92BC4CFFDAFF152FF504C79EFC" bold="true" box="[151,511,1693,1720]" italics="true" pageId="5" pageNumber="594">Paraphlosotiba zingiberiphila</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0AE857E9BC4CFFDAFD842FF6075B9EFC" box="[518,611,1694,1720]" pageId="5" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6BB5650BBC4CFFDAFF152FAA04219E98" box="[151,281,1729,1756]" pageId="5" pageNumber="594" type="description">
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<paragraph id="23103680BC4CFFDAFF152FAA04219E98" blockId="5.[151,611,1693,1756]" box="[151,281,1729,1756]" pageId="5" pageNumber="594">
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(
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<figureCitation id="BB942A05BC4CFFDAFF1D2FAA04299E98" box="[159,273,1729,1756]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="5" pageNumber="594">Figs 9–15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="6BB5650BBC4CFFDAFF152E6204749F9C" pageId="5" pageNumber="594" type="materials_examined">
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<paragraph id="23103680BC4CFFDAFF152E6201A59F28" blockId="5.[151,1437,1801,2008]" pageId="5" pageNumber="594">
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<emphasis id="11DBEA92BC4CFFDAFF152E62047F9F67" bold="true" box="[151,327,1801,1828]" pageId="5" pageNumber="594">Type material.</emphasis>
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<typeStatus id="FC148822BC4CFFDAFECD2E62048F9F60" box="[335,439,1802,1828]" pageId="5" pageNumber="594" type="holotype">Holotype</typeStatus>
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<specimenCount id="35A9FD09BC4CFFDAFE422E6204ED9F60" box="[448,469,1802,1828]" count="1" pageId="5" pageNumber="594" type="male">♂</specimenCount>
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(dissected): ‘
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<collectingRegion id="E16BF862BC4CFFDAFDEE2E6207EC9F60" box="[620,724,1802,1828]" country="Malaysia" name="Sabah" pageId="5" pageNumber="594">SABAH</collectingRegion>
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:
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<quantity id="E4579B65BC4CFFDAFD622E6106269F60" box="[736,798,1801,1828]" metricMagnitude="1" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="594" unit="ft" value="250.0">250ft</quantity>
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, [underlined by yellow] | 30mls. Sandakan | - Keningau Rd, |
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<date id="57111040BC4CFFDAFF502E46047C9F0C" box="[210,324,1838,1864]" pageId="5" pageNumber="594" value="1977-10-19">19.x.1977</date>
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’ <printed>, ‘ginger | flowers’ <printed>, ‘M.E. Bacchus | B.M. 1978-48’ <printed>, ‘
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<typeStatus id="FC148822BC4CFFDAFB752E4600A89F0C" box="[1271,1424,1838,1864]" pageId="5" pageNumber="594" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="E4AF4D03BC4CFFDAFF152E3A046A9F28" authorityName="Steel" authorityYear="1960" box="[151,338,1874,1900]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
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<emphasis id="11DBEA92BC4CFFDAFF152E3A046A9F28" box="[151,338,1874,1900]" italics="true" pageId="5" pageNumber="594">Paraphloeostiba</emphasis>
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</taxonomicName>
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<taxonomicName id="E4AF4D03BC4CFFDAFEE72E3A073A9F28" authorityName="Shavrin" authorityYear="2024" box="[357,514,1874,1900]" class="Insecta" family="Staphylinidae" genus="Paraphlosotiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="zingiberiphila" status="sp. nov.">
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<emphasis id="11DBEA92BC4CFFDAFEE72E3A073A9F28" box="[357,514,1874,1900]" italics="true" pageId="5" pageNumber="594">zingiberiphila</emphasis>
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</taxonomicName>
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<emphasis id="11DBEA92BC4CFFDAFD882E3A075B9F28" bold="true" box="[522,611,1874,1900]" pageId="5" pageNumber="594">
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<taxonomicNameLabel id="0AE857E9BC4CFFDAFD882E3A075B9F28" box="[522,611,1874,1900]" pageId="5" pageNumber="594" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2024’ <red, printed> (
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<collectionCode id="45BEAE45BC4CFFDAFBB42E3A01A89F28" box="[1078,1168,1874,1900]" pageId="5" pageNumber="594">BMNH</collectionCode>
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).
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</paragraph>
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<paragraph id="23103680BC4CFFDAFF452E1E04749F9C" blockId="5.[151,1437,1801,2008]" pageId="5" pageNumber="594">
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<typeStatus id="FC148822BC4CFFDAFF452E1E04009FD4" box="[199,312,1910,1936]" pageId="5" pageNumber="594" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="35A9FD09BC4CFFDAFEC72E1E04BF9FD4" box="[325,391,1910,1936]" count="3" pageId="5" pageNumber="594" type="female">3 ♀♀</specimenCount>
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(
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<specimenCount id="35A9FD09BC4CFFDAFE1A2E1E070D9FD4" box="[408,565,1910,1936]" count="1" pageId="5" pageNumber="594" type="generic">one specimen</specimenCount>
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dissected;
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<specimenCount id="35A9FD09BC4CFFDAFD342E1E066A9FD4" box="[694,850,1910,1936]" count="1" pageId="5" pageNumber="594" type="generic">one specimen</specimenCount>
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without left antennomeres 8–11): same data as the holotype, with additional red printed label: ‘
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<typeStatus id="FC148822BC4CFFDAFDFF2EF206359FF0" box="[637,781,1946,1972]" pageId="5" pageNumber="594" type="paratype">PARATYPE</typeStatus>
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|
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<taxonomicName id="E4AF4D03BC4CFFDAFCA22EF206E39FF0" authorityName="Steel" authorityYear="1960" box="[800,987,1946,1972]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
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<emphasis id="11DBEA92BC4CFFDAFCA22EF206E39FF0" box="[800,987,1946,1972]" italics="true" pageId="5" pageNumber="594">Paraphloeostiba</emphasis>
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</taxonomicName>
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<taxonomicName id="E4AF4D03BC4CFFDAFC6F2EF201B29FF0" authorityName="Shavrin" authorityYear="2024" box="[1005,1162,1946,1972]" class="Insecta" family="Staphylinidae" genus="Paraphlosotiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="zingiberiphila" status="sp. nov.">
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<emphasis id="11DBEA92BC4CFFDAFC6F2EF201B29FF0" box="[1005,1162,1946,1972]" italics="true" pageId="5" pageNumber="594">zingiberiphila</emphasis>
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</taxonomicName>
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<emphasis id="11DBEA92BC4CFFDAFB122EF201F09FF0" bold="true" box="[1168,1224,1946,1972]" pageId="5" pageNumber="594">
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<taxonomicNameLabel id="0AE857E9BC4CFFDAFB122EF201F09FF0" box="[1168,1224,1946,1972]" pageId="5" pageNumber="594" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2024’ (
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<collectionCode id="45BEAE45BC4CFFDAFF672ED604079F9C" box="[229,319,1982,2008]" pageId="5" pageNumber="594">BMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="6BB5650BBC4FFFD9FF4529FE06E09EF9" pageId="6" pageNumber="595" type="description">
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<paragraph id="23103680BC4FFFD9FF4529FE004F98BD" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
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<emphasis id="11DBEA92BC4FFFD9FF4529FE046198F4" bold="true" box="[199,345,150,176]" pageId="6" pageNumber="595">Description.</emphasis>
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Measurements (n=4): HW: 0.29–0.33; HL: 0.16–0.17; OL: 0.08–0.10; AL (averaged): 0.41; PL: 0.22–0.30; PWmax: 0.36–0.42; PWmin: 0.31–0.37; ESL: 0.30–0.35; EW: 0.43–0.45; MTbL (
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<typeStatus id="FC148822BC4FFFD9FB2329D300329891" box="[1185,1290,187,213]" pageId="6" pageNumber="595" type="holotype">holotype</typeStatus>
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): 0.22; MTrL (
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<typeStatus id="FC148822BC4FFFD9FF1D29B7043098BD" box="[159,264,223,249]" pageId="6" pageNumber="595" type="holotype">holotype</typeStatus>
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): 0.10 (MTrL 1–4: 0.03; MTrL 5: 0.07); AW: 0.45–0.48; AedL: 0.37; BL: 1.42–1.60 (
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<typeStatus id="FC148822BC4FFFD9FB4129B7001398BD" box="[1219,1323,223,249]" pageId="6" pageNumber="595" type="holotype">holotype</typeStatus>
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: 1.47).
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</paragraph>
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<paragraph id="23103680BC4FFFD9FF45286B04DE99B1" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">Body reddish-brown, with slightly paler elytra and intersegmental membranes of abdomen; antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellowish. Head with sparse and fine punctation, slightly denser in middle; punctation of pronotum indistinct, sparse and fine; punctation of elytra moderately dense, large and deep, finer around scutellum, finer and sparser along suture; abdomen with indistinct, fine and sparse punctation. Head with fine transverse microreticulation; pronotum with dense and relatively coarse isodiametric microsculpture; scutellum with dense isodiametric meshes; elytra with dense sculpture, transverse around scutellum and isodiametric in middle and apical portions.</paragraph>
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<paragraph id="23103680BC4FFFD9FF45289707FE9AB5" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">Head with slightly elevated infraorbital portions, 1.8–1.9 times as broad as long; postocular carina indistinct, acute. Ocelli located about level of posterior margins of eyes; distance between ocelli 1.3–1.5 times as long as distance between ocellus and posterior margin of eyes. Antenna reaching basal part of pronotum when reclined, with distinctly transverse antennomeres 6–10; antennomere 3 about as long as and about twice narrower than antennomere 2, 4 small, twice shorter than 3, 5 about as wide as long, short, but slightly longer and broader than 4, 6 distinctly longer and broader than 5, 7 indistinctly broader than 6, 8–10 slightly longer and distinctly broader than 7, apical antennomere about twice longer than 10.</paragraph>
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<paragraph id="23103680BC4FFFD9FF452B93043E9B19" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">Pronotum 1.4–1.6 times as broad as long, 1.3–1.6 times as broad as head, widest in middle, more narrowed posteriad than anteriad; middle portion with two wide and shallow depressions; lateral margins indistinctly crenulate.</paragraph>
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<paragraph id="23103680BC4FFFD9FF452A0F018F9BC5" blockId="6.[151,1437,150,1905]" box="[199,1207,871,897]" pageId="6" pageNumber="595">Elytra slightly convex, 1.2–1.4 times as broad as long, 1.1–1.3 times as long as pronotum.</paragraph>
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<paragraph id="23103680BC4FFFD9FF452AE307BF9BE1" blockId="6.[151,1437,150,1905]" box="[199,647,907,933]" pageId="6" pageNumber="595">Abdomen distinctly broader than elytra.</paragraph>
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<paragraph id="23103680BC4FFFD9FF452AC701399C39" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
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Male. Posterior margin of abdominal tergite VIII truncate (
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<figureCitation id="BB942A05BC4FFFD9FCF12AC706F09B8D" box="[883,968,943,969]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="6" pageNumber="595">Fig. 11</figureCitation>
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||
). Posterior margin of abdominal sternite VIII widely concave (
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<figureCitation id="BB942A05BC4FFFD9FE172ABB04D39BA9" box="[405,491,979,1005]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="6" pageNumber="595">Fig. 12</figureCitation>
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). Aedeagus with wide basal part, gradually narrowed toward small rounded apex; parameres moderately narrow, with slightly broadened apical portions, not reaching apex of median lobe, with two short apical and three preapical setae, and several additional short setae along inner margin of each paramere; internal sac moderately wide and long, with three elongate teeth in about middle and narrow elongate sclerotized structure in basal portion (
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<figureCitation id="BB942A05BC4FFFD9FE3C2D0B073A9C39" box="[446,514,1123,1149]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="6" pageNumber="595">Fig. 9</figureCitation>
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). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="BB942A05BC4FFFD9FC292D0B06C59C39" box="[939,1021,1123,1149]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="6" pageNumber="595">Fig. 10</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="23103680BC4FFFD9FF452DEF01689C81" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
|
||
Female. Posterior margin of abdominal tergite VIII straight (
|
||
<figureCitation id="BB942A05BC4FFFD9FCF92DEF06E89CE5" box="[891,976,1159,1185]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="6" pageNumber="595">Fig. 13</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII rounded (
|
||
<figureCitation id="BB942A05BC4FFFD9FEBB2DC304B59C81" box="[313,397,1195,1221]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="6" pageNumber="595">Fig. 14</figureCitation>
|
||
). Accessory sclerite not recognized. Spermatheca as in
|
||
<figureCitation id="BB942A05BC4FFFD9FC7B2DC301749C81" box="[1017,1100,1194,1221]" captionStart="FIGURES 9–15" captionStartId="7.[151,264,1533,1557]" captionTargetBox="[159,1428,189,1501]" captionTargetId="figure-16@7.[151,1436,181,1509]" captionTargetPageId="7" captionText="FIGURES 9–15. Paraphloeostiba zingiberiphila: 9—aedeagus, parameral view, 10—aedeagus, lateral view, 11—male abdominal tergite VIII, 12—male abdominal sternite VIII, 13—female abdominal tergite VIII, 14—female abdominal sternite VIII, 15—spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13914952" httpUri="https://zenodo.org/record/13914952/files/figure.png" pageId="6" pageNumber="595">Fig. 15</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="23103680BC4FFFD9FF452DA601619DD9" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
|
||
<emphasis id="11DBEA92BC4FFFD9FF452DA604979CAD" bold="true" box="[199,431,1230,1257]" pageId="6" pageNumber="595">Comparative notes.</emphasis>
|
||
Regarding the shape of the small body, shortened elytra and similar shape of the aedeagus,
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FF152D9B046C9D49" authorityName="Shavrin" authorityYear="2024" box="[151,340,1267,1293]" class="Insecta" family="Staphylinidae" genus="Paraphlosotiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="zingiberiphila" status="sp. nov.">
|
||
<emphasis id="11DBEA92BC4FFFD9FF152D9B05909D48" box="[151,168,1267,1292]" italics="true" pageId="6" pageNumber="595">P</emphasis>
|
||
.
|
||
<emphasis id="11DBEA92BC4FFFD9FF352D9B046C9D49" box="[183,340,1267,1293]" italics="true" pageId="6" pageNumber="595">zingiberiphila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="11DBEA92BC4FFFD9FEDE2D9B048E9D49" bold="true" box="[348,438,1267,1293]" pageId="6" pageNumber="595">
|
||
<taxonomicNameLabel id="0AE857E9BC4FFFD9FEDE2D9B048E9D49" box="[348,438,1267,1293]" pageId="6" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is more similar to
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FD092D9B06289D49" authorityName="Shavrin" authorityYear="2024" box="[651,784,1267,1293]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="brevicula" status="sp. nov.">
|
||
<emphasis id="11DBEA92BC4FFFD9FD092D9B06289D49" box="[651,784,1267,1293]" italics="true" pageId="6" pageNumber="595">P. brevicula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="11DBEA92BC4FFFD9FC9A2D9B064A9D49" bold="true" box="[792,882,1267,1293]" pageId="6" pageNumber="595">
|
||
<taxonomicNameLabel id="0AE857E9BC4FFFD9FC9A2D9B064A9D49" box="[792,882,1267,1293]" pageId="6" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(see above), from which can be distinguished by the finer punctation of the head and the pronotum, narrower median lobe and different structure of the internal sac. Besides that, based on the presence of two wide longitudinal impressions, similar shape of the aedeagus and the presence of setation on inner margin of each paramere,
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FD7B2C37068C9D3D" authorityName="Shavrin" authorityYear="2024" box="[761,948,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphlosotiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="zingiberiphila" status="sp. nov.">
|
||
<emphasis id="11DBEA92BC4FFFD9FD7B2C3706329D3C" box="[761,778,1375,1400]" italics="true" pageId="6" pageNumber="595">P</emphasis>
|
||
.
|
||
<emphasis id="11DBEA92BC4FFFD9FC952C37068C9D3D" box="[791,948,1375,1401]" italics="true" pageId="6" pageNumber="595">zingiberiphila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="11DBEA92BC4FFFD9FC382C37012A9D3D" bold="true" box="[954,1042,1375,1401]" pageId="6" pageNumber="595">
|
||
<taxonomicNameLabel id="0AE857E9BC4FFFD9FC382C37012A9D3D" box="[954,1042,1375,1401]" pageId="6" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FB1D2C3700099D3D" authorityName="Shavrin" authorityYear="2020" box="[1183,1329,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="brancuccii">
|
||
<emphasis id="11DBEA92BC4FFFD9FB1D2C3700099D3D" box="[1183,1329,1375,1401]" italics="true" pageId="6" pageNumber="595">P. brancuccii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FABC2C3700A49D3C" authorityName="Shavrin" authorityYear="2024" box="[1342,1436,1375,1400]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="iriana">
|
||
<emphasis id="11DBEA92BC4FFFD9FABC2C3700A49D3C" box="[1342,1436,1375,1400]" italics="true" pageId="6" pageNumber="595">P. iriana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FF452CEB04689DD9" authorityName="Shavrin" authorityYear="2024" box="[199,336,1411,1437]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="okapensis">
|
||
<emphasis id="11DBEA92BC4FFFD9FF452CEB04689DD9" box="[199,336,1411,1437]" italics="true" pageId="6" pageNumber="595">P. okapensis</emphasis>
|
||
</taxonomicName>
|
||
. It can be distinguished from these species by the following features:
|
||
</paragraph>
|
||
<paragraph id="23103680BC4FFFD9FF452CCF04469DA1" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
|
||
from
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FE862CCF04AF9D85" authorityName="Shavrin" authorityYear="2020" box="[260,407,1447,1473]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="brancuccii">
|
||
<emphasis id="11DBEA92BC4FFFD9FE862CCF04AF9D85" box="[260,407,1447,1473]" italics="true" pageId="6" pageNumber="595">P. brancuccii</emphasis>
|
||
</taxonomicName>
|
||
by the slightly darker body, finer microsculpture of the forebody, slightly broader elytra and narrower parameres;
|
||
</paragraph>
|
||
<paragraph id="23103680BC4FFFD9FF452C87066C9E69" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
|
||
from
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FE872C87045D9E4C" authorityName="Shavrin" authorityYear="2024" box="[261,357,1519,1544]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="iriana">
|
||
<emphasis id="11DBEA92BC4FFFD9FE872C87045D9E4C" box="[261,357,1519,1544]" italics="true" pageId="6" pageNumber="595">P. iriana</emphasis>
|
||
</taxonomicName>
|
||
by the paler body, sparser punctation of the head and the pronotum, broader median lobe of the shorter aedeagus and broader apical portions of the parameres;
|
||
</paragraph>
|
||
<paragraph id="23103680BC4FFFD9FF452F5F06C09E31" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
|
||
from
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FE842F5F04A89E15" authorityName="Shavrin" authorityYear="2024" box="[262,400,1591,1617]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="okapensis">
|
||
<emphasis id="11DBEA92BC4FFFD9FE842F5F04A89E15" box="[262,400,1591,1617]" italics="true" pageId="6" pageNumber="595">P. okapensis</emphasis>
|
||
</taxonomicName>
|
||
by the darker body, sparser punctation of the head and the pronotum, broader median lobe, longer parameres with broader apical portion, and shorter parameral setation.
|
||
</paragraph>
|
||
<paragraph id="23103680BC4FFFD9FF452F1706E09EF9" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">From all these species it can be distinguished by finer punctation of the head and the pronotum, the shorter elytra and details of the external and internal morphology of the aedeagus.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BB5650BBC4FFFD9FF452FAE043E9F41" pageId="6" pageNumber="595" type="distribution">
|
||
<paragraph id="23103680BC4FFFD9FF452FAE043E9F41" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
|
||
<emphasis id="11DBEA92BC4FFFD9FF452FAE04599EA4" bold="true" box="[199,353,1734,1760]" pageId="6" pageNumber="595">Distribution.</emphasis>
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FEED2FAF07EE9EA5" authorityName="Shavrin" authorityYear="2024" box="[367,726,1735,1761]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="595" phylum="Arthropoda" rank="species" species="zingiberiphila" status="sp. nov.">
|
||
<emphasis id="11DBEA92BC4FFFD9FEED2FAF07EE9EA5" box="[367,726,1735,1761]" italics="true" pageId="6" pageNumber="595">Paraphloeostiba zingiberiphila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="11DBEA92BC4FFFD9FD672FAF067F9EA5" bold="true" box="[741,839,1735,1761]" pageId="6" pageNumber="595">
|
||
<taxonomicNameLabel id="0AE857E9BC4FFFD9FD672FAF067F9EA5" box="[741,839,1735,1761]" pageId="6" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="FC148822BC4FFFD9FB082FAF01829EA5" box="[1162,1210,1735,1761]" pageId="6" pageNumber="595">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="E16BF862BC4FFFD9FAD32FAF00A09EA5" box="[1361,1432,1735,1761]" country="Malaysia" name="Sabah" pageId="6" pageNumber="595">Sabah</collectingRegion>
|
||
,
|
||
<collectingCountry id="5BB87610BC4FFFD9FF152F8304399F41" box="[151,257,1771,1797]" name="Malaysia" pageId="6" pageNumber="595">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BB5650BBC4FFFD9FF452E6701849F35" pageId="6" pageNumber="595" type="etymology">
|
||
<paragraph id="23103680BC4FFFD9FF452E6704E59F09" blockId="6.[151,1437,150,1905]" pageId="6" pageNumber="595">
|
||
<emphasis id="11DBEA92BC4FFFD9FF452E6704769F6D" bold="true" box="[199,334,1807,1833]" pageId="6" pageNumber="595">Etymology.</emphasis>
|
||
The name of the new species is derived from the fact that the beetles were collected from ginger flowers (
|
||
<taxonomicName id="E4AF4D03BC4FFFD9FF792E5B04F59F09" box="[251,461,1843,1869]" class="Liliopsida" family="Zingiberaceae" genus="Zingiber" kingdom="Plantae" order="Zingiberales" pageId="6" pageNumber="595" phylum="Tracheophyta" rank="species" species="officinale">
|
||
<emphasis id="11DBEA92BC4FFFD9FF792E5B04F59F09" box="[251,461,1843,1869]" italics="true" pageId="6" pageNumber="595">Zingiber officinale</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="23103680BC4FFFD9FF452E3E01849F35" blockId="6.[151,1437,150,1905]" box="[199,1212,1878,1905]" pageId="6" pageNumber="595">
|
||
<emphasis id="11DBEA92BC4FFFD9FF452E3E04739F34" bold="true" box="[199,331,1878,1904]" pageId="6" pageNumber="595">Bionomics.</emphasis>
|
||
Specimens were collected at low elevation about
|
||
<quantity id="E4579B65BC4FFFD9FCF92E3F068C9F35" box="[891,948,1879,1905]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="6" pageNumber="595" unit="m" value="76.0">76 m</quantity>
|
||
a.s.l. in ginger flowers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |