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<document id="7E9A4944BB7A5CFF6AE05E3203C678C9" ID-DOI="10.5281/zenodo.13229438" ID-ISSN="2345-7600" ID-Zenodo-Dep="13229438" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722894242153" checkinUser="felipe" docAuthor="Li, Xinzheng &amp; Komai, Tomoyuki" docDate="2003" docId="AA7E8793FFA8F6416C11F266FC23F98A" docLanguage="en" docName="RafflesBZool.51.2.257-275.pdf" docOrigin="Raffles Bulletin of Zoology 51 (2)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Heterocarpus A. Milne Edwards 1881" docType="treatment" docVersion="2" lastPageNumber="259" masterDocId="5647FFEBFFAAF6436F6CF656FF96FFE2" masterDocTitle="Pandaloid Shrimps From The Northern South China Sea, With Description Of A New Species Of Plesionika (Crustacea: Decapoda: Caridea)" masterLastPageNumber="275" masterPageNumber="257" pageNumber="259" updateTime="1722902804154" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="70F882876674AAB36D4320A4E8321F21">Pandaloid Shrimps From The Northern South China Sea, With Description Of A New Species Of Plesionika (Crustacea: Decapoda: Caridea)</mods:title>
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<treatment id="AA7E8793FFA8F6416C11F266FC23F98A" ID-DOI="http://doi.org/10.5281/zenodo.13229410" ID-Zenodo-Dep="13229410" LSID="urn:lsid:plazi:treatment:AA7E8793FFA8F6416C11F266FC23F98A" httpUri="http://treatment.plazi.org/id/AA7E8793FFA8F6416C11F266FC23F98A" lastPageNumber="259" pageId="2" pageNumber="259">
<subSubSection id="6ACD650EFFA8F6416C11F266FA1EFBAA" box="[893,1416,1072,1097]" pageId="2" pageNumber="259" type="nomenclature">
<paragraph id="22683685FFA8F6416C11F266FA1EFBAA" blockId="2.[893,1416,1072,1097]" box="[893,1416,1072,1097]" pageId="2" pageNumber="259">
<heading id="792081E9FFA8F6416C11F266FA1EFBAA" bold="true" box="[893,1416,1072,1097]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="259" reason="2">
<emphasis id="10A3EA97FFA8F6416C11F266FA1EFBAA" bold="true" box="[893,1416,1072,1097]" pageId="2" pageNumber="259">
Genus
<taxonomicName id="E5D74D06FFA8F6416CA3F267FA1EFBAA" ID-CoL="9CKZ4" authority="A. Milne Edwards, 1881" authorityName="A. Milne Edwards" authorityYear="1881" box="[975,1416,1072,1097]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="10A3EA97FFA8F6416CA3F267FBF0FBAB" bold="true" box="[975,1126,1073,1097]" italics="true" pageId="2" pageNumber="259">Heterocarpus</emphasis>
A.
<bibRefCitation id="46464B74FFA8F6416BFDF266FA1EFBAA" author="Milne Edwards, A" box="[1169,1416,1072,1096]" pageId="2" pageNumber="259" pagination="1 - 16" refId="ref19747" refString="Milne Edwards, A., 1881. Description de quelques Crustaces Macroures provenant des grandes profondeurs de la mer des Antilles. Annales des Sciences Naturelles, Zoologie, (6) 11 (4): 1 - 16." type="journal article" year="1881">Milne Edwards, 1881</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6ACD650EFFA8F6416C2FF227FC23F98A" pageId="2" pageNumber="259" type="discussion">
<paragraph id="22683685FFA8F6416C2FF227FC23F98A" blockId="2.[835,1474,1137,1640]" pageId="2" pageNumber="259">
<emphasis id="10A3EA97FFA8F6416C2FF227FC57FB6B" bold="true" box="[835,961,1137,1161]" italics="true" pageId="2" pageNumber="259">Remarks. –</emphasis>
Liu &amp; Zhong (1994) listed the following eight species of
<taxonomicName id="E5D74D06FFA8F6416CDBF2C7FBDDFB4A" authorityName="A. Milne Edwards" authorityYear="1881" box="[951,1099,1169,1192]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="10A3EA97FFA8F6416CDBF2C7FBDDFB4A" box="[951,1099,1169,1192]" italics="true" pageId="2" pageNumber="259">Heterocarpus</emphasis>
</taxonomicName>
from Chinese waters:
<taxonomicName id="E5D74D06FFA8F6416A28F2C7FC2CFB2A" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="alphonsi">
<emphasis id="10A3EA97FFA8F6416A28F2C7FA54FB4A" box="[1348,1474,1169,1192]" italics="true" pageId="2" pageNumber="259">H. alphonsi</emphasis>
<bibRefCitation id="46464B74FFA8F6416C2FF2E7FC2CFB2A" author="Bate, C" box="[835,954,1201,1224]" pageId="2" pageNumber="259" pagination="1 - 942" refId="ref17171" refString="Bate, C. S., 1888. Report of the Crustacea Macrura collected by the Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876. 24: 1 - 942, pls 1 - 157." type="journal article" year="1888">Bate, 1888</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E5D74D06FFA8F6416CA8F2E7FB2EFB2A" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[964,1208,1201,1224]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="10A3EA97FFA8F6416CA8F2E7FBADFB2A" box="[964,1083,1201,1224]" italics="true" pageId="2" pageNumber="259">H. dorsalis</emphasis>
<bibRefCitation id="46464B74FFA8F6416B2DF2E7FB2EFB2A" author="Bate, C" box="[1089,1208,1201,1224]" pageId="2" pageNumber="259" pagination="1 - 942" refId="ref17171" refString="Bate, C. S., 1888. Report of the Crustacea Macrura collected by the Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876. 24: 1 - 942, pls 1 - 157." type="journal article" year="1888">Bate, 1888</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E5D74D06FFA8F6416BAEF2E7FA28FB2A" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[1218,1470,1201,1224]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="10A3EA97FFA8F6416BAEF2E7FAD7FB2A" box="[1218,1345,1201,1224]" italics="true" pageId="2" pageNumber="259">H. gibbosus</emphasis>
<bibRefCitation id="46464B74FFA8F6416A2BF2E7FA28FB2A" author="Bate, C" box="[1351,1470,1201,1224]" pageId="2" pageNumber="259" pagination="1 - 942" refId="ref17171" refString="Bate, C. S., 1888. Report of the Crustacea Macrura collected by the Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876. 24: 1 - 942, pls 1 - 157." type="journal article" year="1888">Bate, 1888</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E5D74D06FFA8F6416C2FF287FBD9FB0A" authority="Bate, 1888" authorityName="Bate" authorityYear="1888" box="[835,1103,1233,1256]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="laevigatus">
<emphasis id="10A3EA97FFA8F6416C2FF287FC45FB0A" box="[835,979,1233,1256]" italics="true" pageId="2" pageNumber="259">H. laevigatus</emphasis>
<bibRefCitation id="46464B74FFA8F6416CB6F287FBD9FB0A" author="Bate, C" box="[986,1103,1233,1256]" pageId="2" pageNumber="259" pagination="1 - 942" refId="ref17171" refString="Bate, C. S., 1888. Report of the Crustacea Macrura collected by the Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876. 24: 1 - 942, pls 1 - 157." type="journal article" year="1888">Bate, 1888</bibRefCitation>
</taxonomicName>
(as
<emphasis id="10A3EA97FFA8F6416B10F287FB49FB0A" box="[1148,1247,1233,1256]" italics="true" pageId="2" pageNumber="259">lavigatus</emphasis>
[sic]);
<taxonomicName id="E5D74D06FFA8F6416A42F287FC73FAEA" authority="De Man, 1917" authorityName="De Man" authorityYear="1917" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="parvispina">
<emphasis id="10A3EA97FFA8F6416A42F287FA54FB0A" box="[1326,1474,1233,1256]" italics="true" pageId="2" pageNumber="259">H. parvispina</emphasis>
<bibRefCitation id="46464B74FFA8F6416C2FF2A7FC73FAEA" author="De Man, J" box="[835,997,1265,1288]" pageId="2" pageNumber="259" pagination="279 - 284" refId="ref18539" refString="De Man, J. G., 1917. Diagnoses of New Species of Macrurous Decapod Crustacea from the Siboga-Expedition. Zoologische Mededeelingen, 3 (4): 279 - 284." type="journal article" year="1917">De Man, 1917</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E5D74D06FFA8F6416C9EF2A7FA85FAEA" authority="De Man, 1917" authorityName="De Man" authorityYear="1917" box="[1010,1299,1265,1288]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="10A3EA97FFA8F6416C9EF2A7FBFFFAEA" box="[1010,1129,1265,1288]" italics="true" pageId="2" pageNumber="259">H. sibogae</emphasis>
<bibRefCitation id="46464B74FFA8F6416B1DF2A7FA85FAEA" author="De Man, J" box="[1137,1299,1265,1288]" pageId="2" pageNumber="259" pagination="279 - 284" refId="ref18539" refString="De Man, J. G., 1917. Diagnoses of New Species of Macrurous Decapod Crustacea from the Siboga-Expedition. Zoologische Mededeelingen, 3 (4): 279 - 284." type="journal article" year="1917">De Man, 1917</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="E5D74D06FFA8F6416A4CF2A7FBE4FACA" authority="Alcock &amp; Anderson, 1894" authorityName="Alcock &amp; Anderson" authorityYear="1894" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="tricarinatus">
<emphasis id="10A3EA97FFA8F6416A4CF2A7FA54FAEA" box="[1312,1474,1265,1288]" italics="true" pageId="2" pageNumber="259">H. tricarinatus</emphasis>
Alcock &amp; Anderson, 1894
</taxonomicName>
; and
<taxonomicName id="E5D74D06FFA8F6416BDFF347FCEAFAAA" authority="Alcock, 1901" authorityName="Alcock" authorityYear="1901" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="woodmasoni">
<emphasis id="10A3EA97FFA8F6416BDFF347FAF4FACA" box="[1203,1378,1297,1320]" italics="true" pageId="2" pageNumber="259">H. woodmasoni</emphasis>
Alcock, 1901
</taxonomicName>
.
<bibRefCitation id="46464B74FFA8F6416CF8F367FBD4FAAA" author="Crosnier, A" box="[916,1090,1329,1352]" pageId="2" pageNumber="259" pagination="57 - 103" refId="ref18259" refString="Crosnier, A., 1988. Sur les Heterocarpus (Crustacea, Decapoda, Pandalidae) du sud-ouest de l'ocean Indien Remarques sur d'autres especes ouest-pacifiques du genre et description de quatre taxa nouveaux. Bulletin du Museum national d'Histoire Naturelle, Paris 4 e ser., 10, section A, no. 1: 57 - 103, pls. 1 - 4." type="journal article" year="1988">Crosnier (1988)</bibRefCitation>
concluded that
<taxonomicName id="E5D74D06FFA8F6416B94F367FAEFFAAA" authorityName="Bate" authorityYear="1888" box="[1272,1401,1329,1352]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="alphonsi">
<emphasis id="10A3EA97FFA8F6416B94F367FAEFFAAA" box="[1272,1401,1329,1352]" italics="true" pageId="2" pageNumber="259">H. alphonsi</emphasis>
</taxonomicName>
was a junior subjective synonym of
<taxonomicName id="E5D74D06FFA8F6416BE4F307FA97FA8A" authorityName="Bate" authorityYear="1888" box="[1160,1281,1361,1384]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="10A3EA97FFA8F6416BE4F307FA97FA8A" box="[1160,1281,1361,1384]" italics="true" pageId="2" pageNumber="259">H. dorsalis</emphasis>
</taxonomicName>
.
<bibRefCitation id="46464B74FFA8F6416A7AF307FA54FA8A" author="Crosnier, A" box="[1302,1474,1361,1384]" pageId="2" pageNumber="259" pagination="57 - 103" refId="ref18259" refString="Crosnier, A., 1988. Sur les Heterocarpus (Crustacea, Decapoda, Pandalidae) du sud-ouest de l'ocean Indien Remarques sur d'autres especes ouest-pacifiques du genre et description de quatre taxa nouveaux. Bulletin du Museum national d'Histoire Naturelle, Paris 4 e ser., 10, section A, no. 1: 57 - 103, pls. 1 - 4." type="journal article" year="1988">Crosnier (1988)</bibRefCitation>
also showed that two species were confounded under the name
<taxonomicName id="E5D74D06FFA8F6416CE8F3C7FC6CFA4A" authorityName="De Man" authorityYear="1917" box="[900,1018,1425,1448]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="10A3EA97FFA8F6416CE8F3C7FC6CFA4A" box="[900,1018,1425,1448]" italics="true" pageId="2" pageNumber="259">H. sibogae</emphasis>
</taxonomicName>
,
<taxonomicName id="E5D74D06FFA8F6416B6BF3C7FBEBFA4A" authorityName="De Man" authorityYear="1917" box="[1031,1149,1425,1448]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="10A3EA97FFA8F6416B6BF3C7FBEBFA4A" box="[1031,1149,1425,1448]" italics="true" pageId="2" pageNumber="259">H. sibogae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B9057ECFFA8F6416BEFF3C7FB30FA4A" box="[1155,1190,1425,1448]" pageId="2" pageNumber="259" sensu="stricto">
<emphasis id="10A3EA97FFA8F6416BEFF3C7FB09FA4A" box="[1155,1183,1425,1448]" italics="true" pageId="2" pageNumber="259">s.s</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="E5D74D06FFA8F6416BB7F3C7FCEDFA2A" authority="Crosnier, 1988" authorityName="Crosnier" authorityYear="1988" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="10A3EA97FFA8F6416BB7F3C7FAC1FA4A" box="[1243,1367,1425,1448]" italics="true" pageId="2" pageNumber="259">H. hayashii</emphasis>
<bibRefCitation id="46464B74FFA8F6416A33F3C7FCEDFA2A" author="Crosnier, A" pageId="2" pageNumber="259" pagination="57 - 103" refId="ref18259" refString="Crosnier, A., 1988. Sur les Heterocarpus (Crustacea, Decapoda, Pandalidae) du sud-ouest de l'ocean Indien Remarques sur d'autres especes ouest-pacifiques du genre et description de quatre taxa nouveaux. Bulletin du Museum national d'Histoire Naturelle, Paris 4 e ser., 10, section A, no. 1: 57 - 103, pls. 1 - 4." type="journal article" year="1988">Crosnier, 1988</bibRefCitation>
</taxonomicName>
. The two species often occur in the same catch of deepwater trawl (
<bibRefCitation id="46464B74FFA8F6416CB8F387FBEFFA0A" author="Hayashi, K" box="[980,1145,1489,1512]" pageId="2" pageNumber="259" pagination="1 - 176" refId="ref18924" refString="Hayashi, K. - I., 1986. Penaeoidea and Caridea. In: Baba, K., K. I. Hayashi &amp; M. Toriyama (eds.), Decapod Crustaceans from Continental Shelf and Slope around Japan. Japan Fisheries Resource Conservation Association, Tokyo, Pp. 1 - 336, pls 1 - 176." type="book chapter" year="1986">Hayashi, 1986</bibRefCitation>
; Chan &amp; Yu, 1987; personal observation). Therefore, it is possible that the Chinese records of
<taxonomicName id="E5D74D06FFA8F6416CD4F047FBB8F9CA" authorityName="De Man" authorityYear="1917" box="[952,1070,1553,1576]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="10A3EA97FFA8F6416CD4F047FBB8F9CA" box="[952,1070,1553,1576]" italics="true" pageId="2" pageNumber="259">H. sibogae</emphasis>
</taxonomicName>
contain the two species. During this study,
<taxonomicName id="E5D74D06FFA8F6416CFEF067FB83F9AA" authorityName="Crosnier" authorityYear="1988" box="[914,1045,1585,1608]" class="Malacostraca" family="Pandalidae" genus="Heterocarpus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="hayashii">
<emphasis id="10A3EA97FFA8F6416CFEF067FB83F9AA" box="[914,1045,1585,1608]" italics="true" pageId="2" pageNumber="259">H. hayashii</emphasis>
</taxonomicName>
has been identified from the present collection.
</paragraph>
</subSubSection>
</treatment>
</document>
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