244 lines
22 KiB
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244 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.898.47118" ID-GBIF-Dataset="6a9aaa1b-b903-4472-865d-7edce422a2dc" ID-PMC="PMC6917629" ID-Pensoft-Pub="1313-2970-898-1" ID-Pensoft-UUID="278CAE686DE65920AF5B0FE8BA1BBDC1" ID-PubMed="31866732" ID-ZooBank="7B407C809E644F5995C2B3229CF78C6B" ModsDocID="1313-2970-898-1" checkinTime="1575997091264" checkinUser="pensoft" docAuthor="Godunko, Roman J., Neumann, Christian & Staniczek, Arnold H." docDate="2019" docId="AA629EF8A7B55C928F180E561D8DBDAA" docLanguage="en" docName="ZooKeys 898: 1-26" docOrigin="ZooKeys 898" docSource="http://dx.doi.org/10.3897/zookeys.898.47118" docTitle="Siphloplecton landolti Godunko, Neumann & Staniczek, 2019, sp. nov." docType="treatment" docUuid="F5338779-9D55-48B5-9542-7357704A8A64" docUuidSource="ZooBank" docVersion="5" id="278CAE686DE65920AF5B0FE8BA1BBDC1" lastPageNumber="1" masterDocId="278CAE686DE65920AF5B0FE8BA1BBDC1" masterDocTitle="Revision of fossil Metretopodidae (Insecta, Ephemeroptera) in Baltic amber - Part 4: Description of two new species of Siphloplecton Clemens, 1915, with notes on the new S. jaegeri species group and with key to fossil male adults of Siphloplecton" masterLastPageNumber="26" masterPageNumber="1" pageNumber="1" updateTime="1668168222060" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of fossil Metretopodidae (Insecta, Ephemeroptera) in Baltic amber - Part 4: Description of two new species of Siphloplecton Clemens, 1915, with notes on the new S. jaegeri species group and with key to fossil male adults of Siphloplecton</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Godunko, Roman J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neumann, Christian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Staniczek, Arnold H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>898</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>26</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.898.47118</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.898.47118</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-898-1</mods:identifier>
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<mods:identifier type="ZooBank">7B407C809E644F5995C2B3229CF78C6B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">278CAE686DE65920AF5B0FE8BA1BBDC1</mods:identifier>
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<treatment ID-GBIF-Taxon="160689326" LSID="urn:lsid:zoobank.org:act:F5338779-9D55-48B5-9542-7357704A8A64" httpUri="http://treatment.plazi.org/id/AA629EF8A7B55C928F180E561D8DBDAA" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="AA629EF8-A7B5-5C92-8F18-0E561D8DBDAA" class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton landolti" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="landolti">Siphloplecton landolti</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (photographs) A general dorsal view B general ventral view C cubital field of right forewing D right hind wing in dorsal view E head and thorax in ventral view F genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure5">Figures 5</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (line drawings) A head and thorax in dorsal view B head and thorax in ventral view." figureDoi="10.3897/zookeys.898.47118.figure6" httpUri="https://binary.pensoft.net/fig/363721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure6">6</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (line drawing): genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure7" httpUri="https://binary.pensoft.net/fig/363722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure7">7</figureCitation>
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<tableCitation captionStartId="T1" httpUri="http://table.plazi.org/id/23F53EE7FB0FC4D865035A6D181A4855" pageId="0" pageNumber="1" tableUuid="23F53EE7FB0FC4D865035A6D181A4855">; Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Male imago in Baltic amber (Eocene), SMNS
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<normalizedToken originalValue="BB–">BB-</normalizedToken>
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2377. Well preserved specimen in clear amber, well visible in dorsoventral aspect. Each of both forewings bent upwards approximately at half length (left forewing partly twisted); left hind wing twisted; genitalia visible in ventral and, partly, lateral view.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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For measurements see Table
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<tableCitation captionStartId="T1" httpUri="http://table.plazi.org/id/23F53EE7FB0FC4D865035A6D181A4855" pageId="0" pageNumber="1" tableUuid="23F53EE7FB0FC4D865035A6D181A4855">1</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Colouration pale, dorsal side slightly darker than ventral side. Brownish diffuse maculation on thorax and abdomen preserved. Ocelli well preserved. Eyes large, medially contiguous; brownish maculation on eye surface partly preserved; antennae longer than head (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (photographs) A general dorsal view B general ventral view C cubital field of right forewing D right hind wing in dorsal view E head and thorax in ventral view F genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure5">
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Figs 5
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, E
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</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (line drawings) A head and thorax in dorsal view B head and thorax in ventral view." figureDoi="10.3897/zookeys.898.47118.figure6" httpUri="https://binary.pensoft.net/fig/363721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure6">
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6
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton landolti" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="landolti">
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<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton landolti</emphasis>
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</taxonomicName>
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sp. nov., SMNS,
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<normalizedToken originalValue="BB–">BB-</normalizedToken>
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2377, holotype, male imago (photographs)
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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general dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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general ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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cubital field of right forewing
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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right hind wing in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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head and thorax in ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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genitalia in ventral view.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.898.47118.figure6" httpUri="https://binary.pensoft.net/fig/363721" pageId="0" pageNumber="1" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 6.</emphasis>
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton landolti" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="landolti">
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<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton landolti</emphasis>
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</taxonomicName>
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sp. nov., SMNS,
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<normalizedToken originalValue="BB–">BB-</normalizedToken>
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2377, holotype, male imago (line drawings)
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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head and thorax in dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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head and thorax in ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
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with traces of brownish pigmentation on dorsal and ventral sides;
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<normalizedToken originalValue="“Verlumung”">"Verlumung"</normalizedToken>
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mainly on lateral sides near wing bases. Lateroparapsidal suture elongate, typical for
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton</emphasis>
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</taxonomicName>
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; no conspecific pigmentation around lateroparapsidal suture; mesonotal suture medially bulged (
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (line drawings) A head and thorax in dorsal view B head and thorax in ventral view." figureDoi="10.3897/zookeys.898.47118.figure6" httpUri="https://binary.pensoft.net/fig/363721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure6">Fig. 6A</figureCitation>
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); furcasternal protuberances of mesothorax contiguous (
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (line drawings) A head and thorax in dorsal view B head and thorax in ventral view." figureDoi="10.3897/zookeys.898.47118.figure6" httpUri="https://binary.pensoft.net/fig/363721" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure6">Fig. 6B</figureCitation>
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); lateral aspect of thorax hardly visible.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wings</emphasis>
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translucent, hyaline, not pigmented, venation well visible; pterostigma translucent, without pigmentation, with simple veins. Cubital field of forewing well visible on right wing only (twisted on left wing): one distinct pair of intercalary veins (iCu2 and iCu3) situated close to CuP and connected with CuP and CuA; one long intercalary vein (iCu1) connected with CuA and also with iCu2 by crossveins; additional short intercalaries present, ending at hind margin of wing (one of them between iCu1 and iCu2, basally joining both veins) (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (photographs) A general dorsal view B general ventral view C cubital field of right forewing D right hind wing in dorsal view E head and thorax in ventral view F genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure5">Fig. 5C</figureCitation>
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). Hind wings with triads RS, MA and MP, 0.31
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<normalizedToken originalValue="×">x</normalizedToken>
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forewing length; costal process bluntly pointed (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (photographs) A general dorsal view B general ventral view C cubital field of right forewing D right hind wing in dorsal view E head and thorax in ventral view F genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure5">Fig. 5D</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
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damaged; both right fore- and hind legs lost; tarsus of left foreleg only partly preserved (for details see Table
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<tableCitation captionStartId="T1" httpUri="http://table.plazi.org/id/23F53EE7FB0FC4D865035A6D181A4855" pageId="0" pageNumber="1" tableUuid="23F53EE7FB0FC4D865035A6D181A4855">1</tableCitation>
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). Tibia of left foreleg paler than femur and tarsus. Left foretibia with strong, sharply pointed setae along outer margin. Tibia of left middle and hind legs black; preserved part of first tarsomere of left middle leg black. Tibia of middle and hind legs with trace of tibiopatellar suture; first tarsomere of middle and hind legs fused with tibia. Tarsal claws lost (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (photographs) A general dorsal view B general ventral view C cubital field of right forewing D right hind wing in dorsal view E head and thorax in ventral view F genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure5">
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Fig. 5
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdominal</emphasis>
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segments completely preserved, sterna visibly paler than terga; genitalia dark.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Styliger</emphasis>
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plate angulate, mediocaudally deeply incised; these incisions with pronounced, broad, triangular projections; medial projection not large, blunt apically (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (photographs) A general dorsal view B general ventral view C cubital field of right forewing D right hind wing in dorsal view E head and thorax in ventral view F genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure5">Figs 5F</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (line drawing): genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure7" httpUri="https://binary.pensoft.net/fig/363722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure7">7</figureCitation>
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). Basal segment of forceps basally relatively wide; forceps 4-segmented, segment 4 approximately 3.30 times longer than wide; length ratio of segment 3 to segment 4 approximately 1:1. Penis lobes distinctly elongated, well separated from each other apically, ellipsoidal; medial sclerite rounded apically; lateral and medial sclerite markedly separated on outer side with inconspicuous incision between sclerites; features of ventral surface of penis lobes not visible (
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (photographs) A general dorsal view B general ventral view C cubital field of right forewing D right hind wing in dorsal view E head and thorax in ventral view F genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure5" httpUri="https://binary.pensoft.net/fig/363720" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure5">Figs 5F</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Siphloplecton landolti sp. nov., SMNS, BB- 2377, holotype, male imago (line drawing): genitalia in ventral view." figureDoi="10.3897/zookeys.898.47118.figure7" httpUri="https://binary.pensoft.net/fig/363722" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.898.47118.figure7">7</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.898.47118.figure7" httpUri="https://binary.pensoft.net/fig/363722" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton landolti" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="landolti">
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<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton landolti</emphasis>
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</taxonomicName>
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sp. nov., SMNS,
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<normalizedToken originalValue="BB–">BB-</normalizedToken>
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2377, holotype, male imago (line drawing): genitalia in ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paracercus</emphasis>
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vestigial; cerci partly lost.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="comments">
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<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The new species can be placed within the
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<taxonomicName genus="Pseudoneuroptera" lsidName="Pseudoneuroptera jaegeri" pageId="0" pageNumber="1" rank="species" species="jaegeri">
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<emphasis italics="true" pageId="0" pageNumber="1">jaegeri</emphasis>
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</taxonomicName>
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species group based on the characteristic shape of the penis lobes, the arrangement of cubital intercalaries of the forewing and the sharply pointed setae along the outer margin of the foretibia.
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<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton landolti" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="landolti">
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||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton landolti</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be separated from the closely related
|
||
<taxonomicName lsidName="S. jaegeri" pageId="0" pageNumber="1" rank="species" species="jaegeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. jaegeri</emphasis>
|
||
</taxonomicName>
|
||
by (1) shape of styliger with relatively small and apically rounded medial projection; (2) relatively wide base of basal forceps segment compared to adjoining apical part of plate; (3) proportions of last forceps segments; (4) ellipsoidal shape of penis lobes with medial sclerite rounded at tip.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The body measurements of
|
||
<taxonomicName lsidName="S. landolti" pageId="0" pageNumber="1" rank="species" species="landolti">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. landolti</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are comparable to other representatives of the
|
||
<taxonomicName lsidName="S. jaegeri" pageId="0" pageNumber="1" rank="species" species="jaegeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. jaegeri</emphasis>
|
||
</taxonomicName>
|
||
species group, but the ratio of hind/forewing length is lower in comparison with
|
||
<taxonomicName lsidName="S. jaegeri" pageId="0" pageNumber="1" rank="species" species="jaegeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. jaegeri</emphasis>
|
||
</taxonomicName>
|
||
itself.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Following our tradition of naming new fossil species of
|
||
<taxonomicName class="Insecta" family="Metretopodidae" genus="Siphloplecton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siphloplecton" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siphloplecton</emphasis>
|
||
</taxonomicName>
|
||
after Swiss ephemeropterists, this species is named after Peter Landolt, Lausanne, to honour his contributions to the knowledge of Swiss mayflies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |