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<document id="80E083EC59F28AA3630DFEFB938CDDC7" ID-DOI="10.5281/zenodo.194387" ID-GBIF-Dataset="512294f5-a88a-4d26-b7bf-d56ba038f5bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="194387" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460203440707" checkinUser="plazi" docAuthor="Cartwright, David I." docDate="2010" docId="AA5980201868FF919DE83CFBFDAD1E38" docLanguage="en" docName="zt02415p021.pdf" docOrigin="Zootaxa 2415" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Wellsomina Cartwright, 2010, gen. nov." docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="5660F858186DFF979D7F3A7CFFCF1A34" masterDocTitle="Descriptions of 2 new genera and 13 new species of caddisflies from Australia (Trichoptera: Ecnomidae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="6" updateTime="1698236701404" updateUser="plazi">
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<mods:title id="D71917618633CD63B42DB5D0AD78E318">Descriptions of 2 new genera and 13 new species of caddisflies from Australia (Trichoptera: Ecnomidae)</mods:title>
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<mods:namePart id="4302FEF35388A55F4AB3CC92DCAD8FF8">Cartwright, David I.</mods:namePart>
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<mods:date id="D67BBF47F09DCCDE9E6D0EFED7D61282">2010</mods:date>
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<treatment id="AA5980201868FF919DE83CFBFDAD1E38" ID-DOI="http://doi.org/10.5281/zenodo.6202138" ID-GBIF-Taxon="124957045" ID-Zenodo-Dep="6202138" LSID="urn:lsid:plazi:treatment:AA5980201868FF919DE83CFBFDAD1E38" httpUri="http://treatment.plazi.org/id/AA5980201868FF919DE83CFBFDAD1E38" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="6AEA62BD1868FF929DE83CFBFE3F1C95" box="[151,496,1671,1697]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="224F31361868FF929DE83CFBFE3F1C95" blockId="5.[151,496,1671,1697]" box="[151,496,1671,1697]" pageId="5" pageNumber="6">
<heading id="7907865A1868FF929DE83CFBFE3F1C95" bold="true" box="[151,496,1671,1697]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="1084ED241868FF929DE83CFBFE3F1C95" bold="true" box="[151,496,1671,1697]" pageId="5" pageNumber="6">
Genus
<taxonomicName id="E5F04AB51868FF929D913CFBFEB61C95" box="[238,377,1671,1697]" class="Insecta" family="Ecnomidae" genus="Wellsomina" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1084ED241868FF929D913CFBFEB61C95" bold="true" box="[238,377,1671,1697]" italics="true" pageId="5" pageNumber="6">Wellsomina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB7505F1868FF929CFD3CFBFE3F1C95" box="[386,496,1671,1697]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6AEA62BD1868FF929DE83CAEFB391CDD" box="[151,1270,1746,1770]" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="224F31361868FF929DE83CAEFB391CDD" blockId="5.[151,1270,1746,1770]" box="[151,1270,1746,1770]" pageId="5" pageNumber="6">
<typeStatus id="FD4B8F941868FF929DE83CAEFF031CDD" box="[151,204,1746,1769]" pageId="5" pageNumber="6">Type</typeStatus>
species.
<taxonomicName id="E5F04AB51868FF929C533CAEFE3B1CDD" box="[300,500,1746,1769]" class="Insecta" family="Ecnomidae" genus="Wellsomina" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="stuarti" status="sp. nov.">
<emphasis id="1084ED241868FF929C533CAEFE3B1CDD" box="[300,500,1746,1769]" italics="true" pageId="5" pageNumber="6">Wellsomina stuarti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BB7505F1868FF929C843CAEFD9E1CDE" box="[507,593,1746,1770]" pageId="5" pageNumber="6" rank="species">
<emphasis id="1084ED241868FF929C843CAEFD9E1CDE" bold="true" box="[507,593,1746,1770]" pageId="5" pageNumber="6">sp. nov.</emphasis>
</taxonomicNameLabel>
by original designation, from the Northern Territory,
<collectingCountry id="5AE771A61868FF9299F23CAEFB3D1CDD" box="[1165,1266,1746,1769]" name="Australia" pageId="5" pageNumber="6">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6AEA62BD1868FF929DE83D6BFD891DD8" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="224F31361868FF929DE83D6BFD891DD8" blockId="5.[151,1437,1815,2028]" pageId="5" pageNumber="6">
<emphasis id="1084ED241868FF929DE83D6BFED01D05" bold="true" box="[151,287,1815,1841]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
Forewings each with forks 2, 3, 4 and 5, both forks 2 and 3 relatively short and with long footstalks, and hind wings each with humeral lobe and forks 2, 3 and 5 present, fork 2 relatively short with long footstalk and fork 3 shorter than footstalk. In male genitalia, superior appendages relatively short, complex; inferior appendages fused basally, usually with pair of processes distally. In female genitalia, abdominal sternite VIII basally has a single slender or robust mesal process; segment X is relatively long, usually forming a slender ovipositor.
</paragraph>
</subSubSection>
<subSubSection id="6AEA62BD186BFF919DBB3AEBFBFC18D6" pageId="6" pageNumber="7" type="description">
<paragraph id="224F3136186BFF919DBB3AEBFBFC18D6" blockId="6.[151,1437,151,1036]" pageId="6" pageNumber="7">
<emphasis id="1084ED24186BFF919DBB3AEBFE931A85" bold="true" box="[196,348,151,177]" pageId="6" pageNumber="7">Description.</emphasis>
Smallish adults, head dorsally with occipital warts aligned transversely; scutellum with oval scutal wart (
<figureCitation id="BACB2DB3186BFF919C543AC1FEBA1AE3" box="[299,373,189,215]" captionStart="FIGURES 7 11" captionStartId="5.[151,272,1532,1556]" captionTargetBox="[302,1282,203,1499]" captionTargetId="figure@5.[287,1299,173,1517]" captionTargetPageId="5" captionText="FIGURES 7 11. Wellsomina spp. 7 8: Wellsomina stuarti sp. nov. 7: Forewing and hind wing. 8: Head and thorax, dorsal view. 9 11: Wellsomina baru sp. nov. male genitalia in left lateral, ventral and dorsal views." httpUri="https://zenodo.org/record/194388/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
). Length of forewing: male
<quantity id="E5089CD3186BFF919FB53AC1FC921AE3" box="[714,861,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="6" pageNumber="7" unit="mm" value="2.75" valueMax="3.1" valueMin="2.4">2.43.1 mm</quantity>
, female
<quantity id="E5089CD3186BFF919EBB3AC1FB971AE3" box="[964,1112,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="6" pageNumber="7" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.63.1 mm</quantity>
. Wing venation: forewings relatively narrow, each with length about 3.34.0 times width, R1 divided distally, forks 2, 3, 4 and 5 present, fork 2 relatively short, fork 2 nygma absent, fork 2 footstalk relatively very long, length between 1.43.6 times length of cross-vein
<emphasis id="1084ED24186BFF919CA73B51FDCD1B72" box="[472,514,301,327]" italics="true" pageId="6" pageNumber="7">r -m</emphasis>
, length fork 2 ranges from 1.32.5 times length of fork 3, fork 3 very short, length fork 3 about 0.51.2 times length of footstalk, footstalk fork 3 very long ranges from 2.67.0 times length of cross-vein
<emphasis id="1084ED24186BFF919CE43B04FE7F1BA5" box="[411,432,376,401]" italics="true" pageId="6" pageNumber="7">m</emphasis>
, cross-veins
<emphasis id="1084ED24186BFF919F283B04FD4C1BA5" box="[599,643,376,402]" italics="true" pageId="6" pageNumber="7">r -m</emphasis>
and
<emphasis id="1084ED24186BFF919FBD3B04FD181BA5" box="[706,727,376,401]" italics="true" pageId="6" pageNumber="7">m</emphasis>
usually contiguous or nearly oppose at footstalk fork 3, discoidal cell closed, length about 3 times width; hind wings relatively narrow, each with length between 3.8 4.7 times width, forks 2, 3 and 5 present, all relatively short, fork 2 footstalk relatively very long, length range 2.05.0 times length of cross-vein
<emphasis id="1084ED24186BFF919F483B94FDAE1835" box="[567,609,488,514]" italics="true" pageId="6" pageNumber="7">r -m</emphasis>
, humeral lobe developed, discoidal cell narrow, length about 3 times width, 2 anal veins (
<figureCitation id="BACB2DB3186BFF919CF23871FE171813" box="[397,472,525,551]" captionStart="FIGURES 7 11" captionStartId="5.[151,272,1532,1556]" captionTargetBox="[302,1282,203,1499]" captionTargetId="figure@5.[287,1299,173,1517]" captionTargetPageId="5" captionText="FIGURES 7 11. Wellsomina spp. 7 8: Wellsomina stuarti sp. nov. 7: Forewing and hind wing. 8: Head and thorax, dorsal view. 9 11: Wellsomina baru sp. nov. male genitalia in left lateral, ventral and dorsal views." httpUri="https://zenodo.org/record/194388/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
). Male genitalia with superior appendages relatively short, complex with robust spiny setae apically; phallus generally robust, tube- like; inferior appendages depressed, elongate, fused basally and typically with pair of processes distally (
<figureCitation id="BACB2DB3186BFF919E653824FC561846" box="[794,921,600,626]" captionStart="FIGURES 7 11" captionStartId="5.[151,272,1532,1556]" captionTargetBox="[302,1282,203,1499]" captionTargetId="figure@5.[287,1299,173,1517]" captionTargetPageId="5" captionText="FIGURES 7 11. Wellsomina spp. 7 8: Wellsomina stuarti sp. nov. 7: Forewing and hind wing. 8: Head and thorax, dorsal view. 9 11: Wellsomina baru sp. nov. male genitalia in left lateral, ventral and dorsal views." httpUri="https://zenodo.org/record/194388/files/figure.png" pageId="6" pageNumber="7">Figs 9, 10</figureCitation>
,
<figureCitation id="BACB2DB3186BFF919ED93824FC3A1846" box="[934,1013,600,626]" captionStart="FIGURES 12 20" captionStartId="8.[151,271,1974,1998]" captionTargetBox="[256,1318,589,1933]" captionTargetId="figure@8.[246,1341,551,1960]" captionTargetPageId="8" captionText="FIGURES 12 20. Wellsomina spp. Male genitalia in left lateral, ventral and dorsal views. 12 14: Wellsomina tam sp. nov. 15 17: Wellsomina bucera sp. nov. 18 20: Wellsomina naumanni sp. nov." httpUri="https://zenodo.org/record/194389/files/figure.png" pageId="6" pageNumber="7">12, 13</figureCitation>
). Female genitalia with abdominal sternite VIII relatively broad, basally with slender or robust mesal process, with small groups of long hairs distolaterally, segment IX relatively short, broad-based, distally tapered slightly, segment X relatively long, typically forming slender ovipositor tipped by pair of small cerci (
<figureCitation id="BACB2DB3186BFF919EE238B4FBE918D6" box="[925,1062,712,738]" captionStart="FIGURES 45 53" captionStartId="18.[151,273,1708,1732]" captionTargetBox="[300,1275,336,1659]" captionTargetId="figure@18.[282,1305,306,1686]" captionTargetPageId="18" captionText="FIGURES 45 53. Wellsomina spp. Female genitalia in ventral view. 45: Wellsomina baru sp. nov. 46: Wellsomina ramosa sp. nov. 47: Wellsomina saeta sp. nov. 48: Wellsomina furca sp. nov. 49: Wellsomina gumurr sp. nov. 50: Wellsomina stuarti sp. nov. 51: Wellsomina jalgoodoo sp. nov. 52: Wellsomina ampulla sp. nov. 53: Wellsomina mattina sp. nov." httpUri="https://zenodo.org/record/194393/files/figure.png" pageId="6" pageNumber="7">Figs 4553</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6AEA62BD186BFF919DBB3891FCA61918" pageId="6" pageNumber="7" type="etymology">
<paragraph id="224F3136186BFF919DBB3891FCA61918" blockId="6.[151,1437,151,1036]" pageId="6" pageNumber="7">
<emphasis id="1084ED24186BFF919DBB3891FE961933" bold="true" box="[196,345,749,775]" pageId="6" pageNumber="7">Etymology.</emphasis>
<emphasis id="1084ED24186BFF919C1C3891FE6B1932" box="[355,420,749,774]" italics="true" pageId="6" pageNumber="7">Wells</emphasis>
- named after Dr Alice Wells, for her contribution to the study of
<taxonomicName id="E5F04AB5186BFF9199953891FAB31933" box="[1258,1404,749,775]" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" rank="order">Trichoptera</taxonomicName>
in
<collectingCountry id="5AE771A6186BFF919DE8396EFEC91918" box="[151,262,786,812]" name="Australia" pageId="6" pageNumber="7">Australia</collectingCountry>
, combined with part of the generic name
<taxonomicName id="E5F04AB5186BFF919F94396EFCAD191F" box="[747,866,786,811]" class="Insecta" family="Ecnomidae" genus="Ecnomina" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="1084ED24186BFF919F94396EFCAD191F" box="[747,866,786,811]" italics="true" pageId="6" pageNumber="7">Ecnomina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="6AEA62BD186BFF919DBB394BFDAD1E38" pageId="6" pageNumber="7" type="discussion">
<paragraph id="224F3136186BFF919DBB394BFDAD1E38" blockId="6.[151,1437,151,1036]" pageId="6" pageNumber="7">
<emphasis id="1084ED24186BFF919DBB394BFE8F1965" bold="true" box="[196,320,823,849]" pageId="6" pageNumber="7">Remarks.</emphasis>
<taxonomicName id="E5F04AB5186BFF919C363944FE1A1965" box="[329,469,824,849]" class="Insecta" family="Ecnomidae" genus="Wellsomina" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1084ED24186BFF919C363944FE1A1965" box="[329,469,824,849]" italics="true" pageId="6" pageNumber="7">Wellsomina</emphasis>
</taxonomicName>
is a widespread northern Australian endemic genus most similar to the species
<taxonomicName id="E5F04AB5186BFF919DE83921FEA11942" box="[151,366,861,886]" class="Insecta" family="Ecnomidae" genus="Ecnomina" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kavinia">
<emphasis id="1084ED24186BFF919DE83921FEA11942" box="[151,366,861,886]" italics="true" pageId="6" pageNumber="7">Ecnomina kavinia</emphasis>
</taxonomicName>
from
<collectingCountry id="5AE771A6186BFF919CC63921FDBE1943" box="[441,625,861,887]" name="New Caledonia" pageId="6" pageNumber="7">New Caledonia</collectingCountry>
(
<bibRefCitation id="46614CC7186BFF919FFE3921FC631943" author="Ward" box="[641,940,861,887]" pageId="6" pageNumber="20" refString="Ward, J. B. &amp; Schefter, P. W. (2000) A new genus and twenty new species of New Caledonian Ecnomidae (Trichoptera). Records of the Canterbury Museum, 14, 55 - 87." type="journal article" year="2000">Ward and Schefter, 2000</bibRefCitation>
) which also has relatively short forewing forks 2 and 3 (
<bibRefCitation id="46614CC7186BFF919C3439FEFDB219A8" author="Ward" box="[331,637,898,924]" pageId="6" pageNumber="20" refString="Ward, J. B. &amp; Schefter, P. W. (2000) A new genus and twenty new species of New Caledonian Ecnomidae (Trichoptera). Records of the Canterbury Museum, 14, 55 - 87." type="journal article" year="2000">Ward and Schefter 2000</bibRefCitation>
).
<taxonomicName id="E5F04AB5186BFF919FED39FEFCD219AF" box="[658,797,898,923]" class="Insecta" family="Ecnomidae" genus="Wellsomina" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="1084ED24186BFF919FED39FEFCD219AF" box="[658,797,898,923]" italics="true" pageId="6" pageNumber="7">Wellsomina</emphasis>
</taxonomicName>
species differ from
<taxonomicName id="E5F04AB5186BFF91996539FEFB5819AF" box="[1050,1175,898,923]" class="Insecta" family="Ecnomidae" genus="Ecnomina" kingdom="Animalia" order="Trichoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kavinia">
<emphasis id="1084ED24186BFF91996539FEFB5819AF" box="[1050,1175,898,923]" italics="true" pageId="6" pageNumber="7">E. kavinia</emphasis>
</taxonomicName>
in several hind wing characters including absence of fork
<quantity id="E5089CD3186BFF919F3639D4FDB919F6" box="[585,630,936,962]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="7" unit="in" value="3.0">3 in</quantity>
each wing, the presence of a humeral lobe and the relative length of fork 2. The genus comprises 12 species from across northern
<collectingCountry id="5AE771A6186BFF919E1439B1FC1419D3" box="[875,987,973,999]" name="Australia" pageId="6" pageNumber="7">Australia</collectingCountry>
: northern Western
<collectingCountry id="5AE771A6186BFF9199C739B1FAE819D3" box="[1208,1319,973,999]" name="Australia" pageId="6" pageNumber="7">Australia</collectingCountry>
, Northern Territory and northeastern Queensland.
</paragraph>
</subSubSection>
</treatment>
</document>