treatments-xml/data/AA/55/87/AA5587BF5C2DFFA0FF22FCD0FC166374.xml

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<mods:title id="1BD09566389619D2883157DF9BC8BAE7">Two new genera and five new species of Tullbergiidae (Collembola) from the southern Appalachian Mountains of North America, with redescription of Tullbergia clavata Mills</mods:title>
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<mods:date id="91BCE0B95412A0603A90C06F41990FB8">2016</mods:date>
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<paragraph id="224336A95C2DFFA4FF22FCD0FD14627D" blockId="3.[151,759,794,853]" box="[151,759,794,820]" pageId="3" pageNumber="454">
<heading id="790B81C55C2DFFA4FF22FCD0FD14627D" bold="true" box="[151,759,794,820]" fontSize="11" level="1" pageId="3" pageNumber="454" reason="1">
<emphasis id="1088EABB5C2DFFA4FF22FCD0FD14627D" bold="true" box="[151,759,794,820]" pageId="3" pageNumber="454">
<taxonomicName id="E5FC4D2A5C2DFFA4FF22FCD0FDF9627D" authority="Mills, 1934" authorityName="Mills" authorityYear="1934" box="[151,538,794,820]" class="Entognatha" family="Tullbergiidae" genus="Ameritulla" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="clavata" status="comb. nov.">
<emphasis id="1088EABB5C2DFFA4FF22FCD0FE95627D" bold="true" box="[151,374,794,820]" italics="true" pageId="3" pageNumber="454">Ameritulla clavata</emphasis>
(
<bibRefCitation id="466D4B585C2DFFA4FE33FCD0FDF2627D" author="Mills" box="[390,529,794,820]" pageId="3" pageNumber="454" refString="Mills, H. B. (1934) A monograph of the Collembola of Iowa. Collegiate Press, Ames, Iowa, 143 pp." type="book" year="1934">Mills, 1934</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C2DFFA4FD94FCD0FD14627D" box="[545,759,794,820]" pageId="3" pageNumber="454">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="224336A95C2DFFA4FF22FCF6FEAA621D" blockId="3.[151,759,794,853]" box="[151,329,828,853]" pageId="3" pageNumber="454">
<figureCitation id="BAC72A2C5C2DFFA4FF22FCF6FF3F621C" box="[151,220,828,853]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="3" pageNumber="454">Figs 1</figureCitation>
, 
<figureCitation id="BAC72A2C5C2DFFA4FF5FFCF7FF19621C" box="[234,250,829,853]" captionStart="FIGURE 2" captionStartId="5.[151,250,1761,1783]" captionTargetBox="[153,1433,199,1739]" captionTargetId="figure@5.[153,1433,193,1740]" captionTargetPageId="5" captionText="FIGURE 2. Ameritulla clavata n. comb. A) Postantennal organ, lectotype. B) Anterior cephalic pseudocellus, Sequatchie Co., Tennessee, specimen. C) Ventral chaetotaxy of Abd. VI, Haywood Co., North Carolina, specimen. Scales: A, B, 10 µm; C, 50 µm." httpUri="https://zenodo.org/record/256861/files/figure.png" pageId="3" pageNumber="454">2</figureCitation>
, 3A, B
</paragraph>
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<subSubSection id="6AE665225C2DFFA4FF22FC4EFCB762F0" pageId="3" pageNumber="454" type="reference_group">
<paragraph id="224336A95C2DFFA4FF22FC4EFCB762F0" blockId="3.[151,852,900,953]" pageId="3" pageNumber="454">
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<taxonomicName id="E5FC4D2A5C2DFFA4FF22FC4EFE0D62D3" authority="Mills, 1934:38" authorityName="Mills" authorityPageNumber="38" authorityYear="1934" box="[151,494,900,922]" class="Entognatha" family="Tullbergiidae" genus="Tullbergia" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="1088EABB5C2DFFA4FF22FC4EFEB262D3" box="[151,337,900,922]" italics="true" pageId="3" pageNumber="454">Tullbergia clavata</emphasis>
<treatmentCitation id="A35D10B85C2DFFA4FEE3FC4EFE0D62D3" author="Mills" box="[342,494,900,922]" page="38" pageId="3" pageNumber="454" year="1934">
<bibRefCitation id="466D4B585C2DFFA4FEE3FC4EFE3362D3" author="Mills" box="[342,464,900,922]" pageId="3" pageNumber="454" refString="Mills, H. B. (1934) A monograph of the Collembola of Iowa. Collegiate Press, Ames, Iowa, 143 pp." type="book" year="1934">Mills, 1934</bibRefCitation>
:38
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</taxonomicName>
; 
<treatmentCitation id="A35D10B85C2DFFA4FE4CFC4EFD0F62D3" author="Waltz" box="[505,748,900,922]" page="115" pageId="3" pageNumber="454" year="1995">
<bibRefCitation id="466D4B585C2DFFA4FE4CFC4EFD2162D3" author="Waltz" box="[505,706,900,922]" pageId="3" pageNumber="454" refString="Waltz, R. D. &amp; Hart, J. W. (1995) First report of males of Tullbergia clavata (Collembola: Onychiuridae). Entomological News, 106 (3), 115 - 116." type="journal article" year="1995">Waltz &amp; Hart, 1995</bibRefCitation>
:115
</treatmentCitation>
(males). 
<taxonomicName id="E5FC4D2A5C2DFFA4FF22FC69FDD562F0" authority="Bagnall, 1947:444" authorityName="Bagnall" authorityPageNumber="444" authorityYear="1947" box="[151,566,931,953]" class="Collembola" family="Tullbergiidae" genus="Mesaphorura" higherTaxonomySource="CoL" kingdom="Animalia" order="Poduromorpha" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="1088EABB5C2DFFA4FF22FC69FE9262F0" box="[151,369,931,953]" italics="true" pageId="3" pageNumber="454">Mesaphorura clavata</emphasis>
<treatmentCitation id="A35D10B85C2DFFA4FECDFC69FDD562F0" author="Bagnall" box="[376,566,931,953]" page="444" pageId="3" pageNumber="454" year="1947">
<bibRefCitation id="466D4B585C2DFFA4FECDFC69FDEB62F0" author="Bagnall" box="[376,520,931,953]" pageId="3" pageNumber="454" refString="Bagnall, R. S. (1947) Contributions towards a knowledge of Tullbergiidae (Collembola-Onychiuroidea), I - III. Annals and Magazine of Natural History, 11, 435 - 444. http: // dx. doi. org / 10.1080 / 00222934708654653" type="journal article" year="1947">Bagnall, 1947</bibRefCitation>
:444
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="A35D10B85C2DFFA4FDF5FC6EFCB262F0" author="Dunger" box="[576,849,931,953]" page="49" pageId="3" pageNumber="454" year="2011">
<bibRefCitation id="466D4B585C2DFFA4FDF5FC6EFCD262F0" author="Dunger" box="[576,817,931,953]" pageId="3" pageNumber="454" refString="Dunger, W. &amp; Schlitt, B. (2011) Synopses on Palearctic Collembola. Tullbergiidae. Soil Organisms, 83, 1 - 168." type="journal article" year="2011">Dunger &amp; Schlitt, 2011</bibRefCitation>
:49
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="6AE665225C2DFFA4FF22FC2CFD7064C2" pageId="3" pageNumber="454" type="materials_examined">
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<materialsCitation id="92943CF45C2DFFA4FF22FC2CFD4E656A" ID-GBIF-Occurrence="1306554495" collectingDate="1932-04-02" collectionCode="INHS" collectorName="H. B. Mills" location="Ames" pageId="3" pageNumber="454" specimenCount="1" stateProvince="Iowa" typeStatus="lectotype">
<emphasis id="1088EABB5C2DFFA4FF22FC2CFE6162B6" bold="true" box="[151,386,998,1023]" pageId="3" pageNumber="454">Material examined.</emphasis>
Four cotypes examined: cotype female designated as 
<typeStatus id="FD47880B5C2DFFA4FC44FC2CFBB962B6" box="[1009,1114,998,1023]" pageId="3" pageNumber="454" type="lectotype">lectotype</typeStatus>
, 
<location id="272360725C2DFFA4FBDDFC2DFB4F62B6" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FBDDFC2DFB4F62B6" box="[1128,1196,999,1023]" name="Ames" pageId="3" pageNumber="454" stateProvince="Iowa">Ames</location>
, 
<collectingRegion id="E038F84B5C2DFFA4FB0FFC2DFB1662B7" box="[1210,1269,999,1022]" country="United States of America" name="Iowa" pageId="3" pageNumber="454">Iowa</collectingRegion>
, 
<collectingDate id="4606E9815C2DFFA4FAB7FC2DFA7B62B6" box="[1282,1432,998,1023]" pageId="3" pageNumber="454" value="1932-04-02">
<date id="564210695C2DFFA4FAB7FC2DFA7B62B6" box="[1282,1432,998,1023]" pageId="3" pageNumber="454" value="1932-04-02">2 April 1932</date>
</collectingDate>
, 
<collectionCode id="44EDAE6C5C2DFFA4FF22FBC0FF3B656B" box="[151,216,1034,1058]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="3" pageNumber="454">INHS</collectionCode>
slide number 529.347, 
<collectorName id="8F09537F5C2DFFA4FE58FBC1FD88656A" box="[493,619,1034,1059]" pageId="3" pageNumber="454">H.B. Mills</collectorName>
, coll.
</materialsCitation>
; 
<materialsCitation id="92943CF45C2DFFA4FD09FBC0FCE4650E" ID-GBIF-Occurrence="1306554479" collectingDate="1932-04-03" collectorName="H. B. Mills" location="Ames" pageId="3" pageNumber="454" specimenCount="2" stateProvince="Iowa" typeStatus="paralectotype">
<typeStatus id="FD47880B5C2DFFA4FD09FBC0FCB0656A" box="[700,851,1034,1059]" pageId="3" pageNumber="454" type="paralectotype">paralectotype</typeStatus>
female and 
<typeStatus id="FD47880B5C2DFFA4FC53FBC0FB9E656A" box="[998,1149,1034,1059]" pageId="3" pageNumber="454" type="paralectotype">paralectotype</typeStatus>
juvenile, 
<location id="272360725C2DFFA4FB46FBC0FAD4656B" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FB46FBC0FAD4656B" box="[1267,1335,1034,1058]" name="Ames" pageId="3" pageNumber="454" stateProvince="Iowa">Ames</location>
, 
<collectingRegion id="E038F84B5C2DFFA4FAF0FBC1FA9C656B" box="[1349,1407,1035,1058]" country="United States of America" name="Iowa" pageId="3" pageNumber="454">Iowa</collectingRegion>
, 
<collectingDate id="4606E9815C2DFFA4FA3BFBC0FEF5650E" pageId="3" pageNumber="454" value="1932-04-03">
<date id="564210695C2DFFA4FA3BFBC0FEF5650E" pageId="3" pageNumber="454" value="1932-04-03">3 April 1932</date>
</collectingDate>
, slides 529.345, 529.346, 
<collectorName id="8F09537F5C2DFFA4FDF2FBE5FD26650E" box="[583,709,1070,1095]" pageId="3" pageNumber="454">H.B. Mills</collectorName>
, coll.
</materialsCitation>
; 
<materialsCitation id="92943CF45C2DFFA4FCA3FBE4FD7064C2" ID-GBIF-Occurrence="1306554480" collectingDate="1931-11-05" collectingDateMax="2011" collectingDateMin="1931-11-05" collectionCode="GRSM, WMM, AHSLC" collectorName="H. M. Harris &amp; E. C. Bernard &amp; P. L. Jennings &amp; W. M. Merrit &amp; Tow String Road &amp; A. Mayor &amp; Newfound Gap &amp; R. T. Allen &amp; S. M. Allen &amp; H. S. Allen &amp; Purchase Knob" county="Sequatchie County" latitude="35.490818" location="University" longLatPrecision="1" longitude="-83.80165" municipality="Traer" pageId="3" pageNumber="454" specimenCode="AHSLC07024" specimenCount="1" stateProvince="Indiana" typeStatus="paralectotype">
<typeStatus id="FD47880B5C2DFFA4FCA3FBE4FC4E650E" box="[790,941,1070,1095]" pageId="3" pageNumber="454" type="paralectotype">paralectotype</typeStatus>
female, 
<collectingMunicipality id="C227ACD35C2DFFA4FBA1FBE5FBB2650F" box="[1044,1105,1071,1094]" pageId="3" pageNumber="454">Traer</collectingMunicipality>
, 
<collectingRegion id="E038F84B5C2DFFA4FBD5FBE4FB54650E" box="[1120,1207,1070,1095]" country="United States of America" name="Indiana" pageId="3" pageNumber="454">Indiana</collectingRegion>
, 
<collectingDate id="4606E9815C2DFFA4FB70FBE4FA74650E" box="[1221,1431,1070,1095]" pageId="3" pageNumber="454" value="1931-11-05">
<date id="564210695C2DFFA4FB70FBE4FA74650E" box="[1221,1431,1070,1095]" pageId="3" pageNumber="454" value="1931-11-05">5 November 1931</date>
</collectingDate>
, slide 529.348, 
<collectorName id="8F09537F5C2DFFA4FEF0FB99FE366522" box="[325,469,1106,1131]" pageId="3" pageNumber="454">H.M. Harris</collectorName>
, coll. Additional specimens examined, all in 
<location id="272360725C2DFFA4FC5BFB98FB876522" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FC5BFB98FB876522" box="[1006,1124,1106,1131]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="University" pageId="3" pageNumber="454" stateProvince="Indiana">University</location>
of 
<collectingRegion id="E038F84B5C2DFFA4FB26FB99FAE56523" box="[1171,1286,1107,1130]" country="United States of America" name="Tennessee" pageId="3" pageNumber="454">Tennessee</collectingRegion>
Entomology Collection: one female, 
<collectingRegion id="E038F84B5C2DFFA4FE11FBBDFDF965C7" box="[420,538,1143,1166]" country="United States of America" name="Tennessee" pageId="3" pageNumber="454">Tennessee</collectingRegion>
, 
<collectingCounty id="CB224E255C2DFFA4FD90FBBCFD1965C6" box="[549,762,1142,1167]" pageId="3" pageNumber="454">Sequatchie County</collectingCounty>
, 
<location id="272360725C2DFFA4FCB0FBBCFC9365C6" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FCB0FBBCFC9365C6" box="[773,880,1142,1167]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Whitwell" pageId="3" pageNumber="454" stateProvince="Indiana">Whitwell</location>
, 
<location id="272360725C2DFFA4FCCEFBBCFBCF65C7" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FCCEFBBCFBCF65C7" box="[891,1068,1142,1167]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Kimmons Farm" pageId="3" pageNumber="454" stateProvince="Indiana">Kimmons Farm</location>
“ 
<location id="272360725C2DFFA4FB88FBBCFB3F65C6" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FB88FBBCFB3F65C6" box="[1085,1244,1142,1167]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Moonshadow" pageId="3" pageNumber="454" stateProvince="Indiana">Moonshadow</location>
”, to right of cave mouth, 
<collectingDate id="4606E9815C2DFFA4FF5BFB50FE6865FB" box="[238,395,1178,1203]" pageId="3" pageNumber="454" value="1997-05-15">
<date id="564210695C2DFFA4FF5BFB50FE6865FB" box="[238,395,1178,1203]" pageId="3" pageNumber="454" value="1997-05-15">15 May 1997</date>
</collectingDate>
, sample 1997-37, 
<collectorName id="8F09537F5C2DFFA4FDEBFB50FD1165FA" box="[606,754,1178,1203]" pageId="3" pageNumber="454">E.C. Bernard</collectorName>
&amp; 
<collectorName id="8F09537F5C2DFFA4FCAFFB51FC5765FA" box="[794,948,1178,1203]" pageId="3" pageNumber="454">P.L. Jennings</collectorName>
, colls.; one female, 
<collectingRegion id="E038F84B5C2DFFA4FB2EFB50FAAA65FA" box="[1179,1353,1178,1203]" country="United States of America" name="North Carolina" pageId="3" pageNumber="454">North Carolina</collectingRegion>
, 
<location id="272360725C2DFFA4FAE3FB50FF08659E" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FAE3FB50FF08659E" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Swain County" pageId="3" pageNumber="454" stateProvince="Indiana">Swain County</location>
, 
<collectionCode id="44EDAE6C5C2DFFA4FF40FB75FEA4659E" box="[245,327,1215,1239]" country="0" httpUri="http://grbio.org/cool/d016-ijfq" name="Great Smoky Mountains National Park" pageId="3" pageNumber="454">GRSM</collectionCode>
, 
<location id="272360725C2DFFA4FEE7FB75FDE7659E" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FEE7FB75FDE7659E" box="[338,516,1214,1239]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Lost Cove Trail" pageId="3" pageNumber="454" stateProvince="Indiana">Lost Cove Trail</location>
, leaf litter, 
<geoCoordinate id="47C8506E5C2DFFA4FD34FB74FCF7659E" box="[641,788,1214,1239]" direction="north" orientation="latitude" pageId="3" pageNumber="454" precision="1" value="35.490818">35°29.449’N</geoCoordinate>
<geoCoordinate id="47C8506E5C2DFFA4FCAFFB75FC51659E" box="[794,946,1215,1239]" direction="west" orientation="longitude" pageId="3" pageNumber="454" precision="1" value="-83.80165">83°48.099’W</geoCoordinate>
, 
<collectingDate id="4606E9815C2DFFA4FC08FB75FBA4659E" box="[957,1095,1215,1239]" pageId="3" pageNumber="454" value="2004-05-08">
<date id="564210695C2DFFA4FC08FB75FBA4659E" box="[957,1095,1215,1239]" pageId="3" pageNumber="454" value="2004-05-08">8 May 2004</date>
</collectingDate>
, sample 
<collectingDate id="4606E9815C2DFFA4FB12FB75FAE2659E" box="[1191,1281,1215,1239]" pageId="3" pageNumber="454" value="2004" valueMax="2011">
<date id="564210695C2DFFA4FB12FB75FAE2659E" box="[1191,1281,1215,1239]" pageId="3" pageNumber="454" value="2004" valueMax="2011">2004-11</date>
</collectingDate>
<collectionCode id="44EDAE6C5C2DFFA4FAB4FB75FAB1659E" box="[1281,1362,1215,1239]" country="0" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35218" name="Witte Memorial Museum" pageId="3" pageNumber="454">WMM</collectionCode>
, 
<collectorName id="8F09537F5C2DFFA4FAE8FB75FF0165B2" pageId="3" pageNumber="454">W.M. Merrit</collectorName>
, coll.; two females, 
<collectingRegion id="E038F84B5C2DFFA4FE55FB28FD7165B2" box="[480,658,1250,1275]" country="United States of America" name="North Carolina" pageId="3" pageNumber="454">North Carolina</collectingRegion>
, 
<location id="272360725C2DFFA4FD10FB28FCAD65B3" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FD10FB28FCAD65B3" box="[677,846,1250,1275]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Swain County" pageId="3" pageNumber="454" stateProvince="Indiana">Swain County</location>
, 
<collectionCode id="44EDAE6C5C2DFFA4FCD5FB28FC5165B3" box="[864,946,1250,1274]" country="0" httpUri="http://grbio.org/cool/d016-ijfq" name="Great Smoky Mountains National Park" pageId="3" pageNumber="454">GRSM</collectionCode>
, culvert along 
<collectorName id="8F09537F5C2DFFA4FBD8FB29FADE65B2" box="[1133,1341,1250,1275]" pageId="3" pageNumber="454">Tow String Road</collectorName>
, among liverworts, 
<date id="564210695C2DFFA4FEA3FACDFE366456" box="[278,469,1286,1311]" pageId="3" pageNumber="454" value="2004-10-27">
<collectingDate id="4606E9815C2DFFA4FEA3FACDFE366456" box="[278,469,1286,1311]" pageId="3" pageNumber="454" value="2004-10-27">27 October 2004</collectingDate>
</date>
, 
<collectorName id="8F09537F5C2DFFA4FE55FACDFD906456" box="[480,627,1286,1311]" pageId="3" pageNumber="454">E.C. Bernard</collectorName>
&amp; 
<collectorName id="8F09537F5C2DFFA4FD23FACDFCE66456" box="[662,773,1287,1311]" pageId="3" pageNumber="454">A. Mayor</collectorName>
, colls.; one female, 
<collectingRegion id="E038F84B5C2DFFA4FC51FACCFB6C6456" box="[996,1167,1286,1311]" country="United States of America" name="North Carolina" pageId="3" pageNumber="454">North Carolina</collectingRegion>
, 
<location id="272360725C2DFFA4FB2FFACCFAD86456" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FB2FFACCFAD86456" box="[1178,1339,1286,1311]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Swain County" pageId="3" pageNumber="454" stateProvince="Indiana">Swain County</location>
, 
<collectionCode id="44EDAE6C5C2DFFA4FAF3FACDFA7B6456" box="[1350,1432,1287,1311]" country="0" httpUri="http://grbio.org/cool/d016-ijfq" name="Great Smoky Mountains National Park" pageId="3" pageNumber="454">GRSM</collectionCode>
, 
<collectorName id="8F09537F5C2DFFA4FF22FAE0FEA6640B" box="[151,325,1322,1347]" pageId="3" pageNumber="454">Newfound Gap</collectorName>
, 
<collectingDate id="4606E9815C2DFFA4FEE5FAE0FE04640A" box="[336,487,1322,1347]" pageId="3" pageNumber="454" value="2005-06-22">
<date id="564210695C2DFFA4FEE5FAE0FE04640A" box="[336,487,1322,1347]" pageId="3" pageNumber="454" value="2005-06-22">22 June 2005</date>
</collectingDate>
, sample RTA-05-028, 
<collectorName id="8F09537F5C2DFFA4FD54FAE1FCBB640A" box="[737,856,1322,1347]" pageId="3" pageNumber="454">R.T. Allen</collectorName>
, 
<collectorName id="8F09537F5C2DFFA4FCD6FAE0FC3D640A" box="[867,990,1322,1347]" pageId="3" pageNumber="454">S.M. Allen</collectorName>
&amp; 
<collectorName id="8F09537F5C2DFFA4FBB4FAE0FB99640A" box="[1025,1146,1322,1347]" pageId="3" pageNumber="454">H.S. Allen</collectorName>
, colls.; six females, 
<collectingRegion id="E038F84B5C2DFFA4FAEEFAE0FF19642E" country="United States of America" name="North Carolina" pageId="3" pageNumber="454">North Carolina</collectingRegion>
, 
<location id="272360725C2DFFA4FEBFFA84FE35642E" LSID="urn:lsid:plazi:treatment:AA5587BF5C2DFFA0FF22FCD0FC166374:272360725C2DFFA4FEBFFA84FE35642E" box="[266,470,1358,1383]" county="Sequatchie County" latitude="35.490818" longLatPrecision="1" longitude="-83.80165" municipality="Traer" name="Haywood County" pageId="3" pageNumber="454" stateProvince="Indiana">Haywood County</location>
, 
<collectionCode id="44EDAE6C5C2DFFA4FE53FA85FDDB642E" box="[486,568,1359,1383]" country="0" httpUri="http://grbio.org/cool/d016-ijfq" name="Great Smoky Mountains National Park" pageId="3" pageNumber="454">GRSM</collectionCode>
, 
<collectorName id="8F09537F5C2DFFA4FDF2FA84FD1A642E" box="[583,761,1358,1383]" pageId="3" pageNumber="454">Purchase Knob</collectorName>
, leaf litter in heavy rhododendron thicket, 
<collectingDate id="4606E9815C2DFFA4FB42FA85FA7B642E" box="[1271,1432,1359,1383]" pageId="3" pageNumber="454" value="2007-05-24">
<date id="564210695C2DFFA4FB42FA85FA7B642E" box="[1271,1432,1359,1383]" pageId="3" pageNumber="454" value="2007-05-24">24 May 2007</date>
</collectingDate>
, sample 
<specimenCode id="725A9ED25C2DFFA4FF58FAB8FE7064C3" box="[237,403,1394,1418]" collectionCode="AHSLC" pageId="3" pageNumber="454">
<collectionCode id="44EDAE6C5C2DFFA4FF58FAB8FEA864C3" box="[237,331,1394,1418]" pageId="3" pageNumber="454">AHSLC</collectionCode>
07024
</specimenCode>
EB, 
<collectorName id="8F09537F5C2DFFA4FE74FAB8FDB564C2" box="[449,598,1394,1419]" pageId="3" pageNumber="454">E.C. Bernard</collectorName>
, coll.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6AE665225C2DFFA0FF72FA5CFCBD6055" lastPageId="7" lastPageNumber="458" pageId="3" pageNumber="454" type="description">
<paragraph id="224336A95C2DFFA4FF72FA5CFC8E6752" blockId="3.[151,1437,998,2031]" pageId="3" pageNumber="454">
<emphasis id="1088EABB5C2DFFA4FF72FA5CFE9764E6" bold="true" box="[199,372,1430,1455]" pageId="3" pageNumber="454">Redescription.</emphasis>
White. Length of three adults labeled as cotypes 921‒1137 µm, length of eight additional adults 895‒1204 µm (mean of 
<specimenCount id="34FAFD205C2DFFA4FDAFFA70FD59649A" box="[538,698,1466,1491]" pageId="3" pageNumber="454" type="generic">11 specimens</specimenCount>
= 1014 µm). Length ratio of Ant. I:II:III:IV approximately 1:1.1:1.6:2.2. Pseudocellus shape 
<typeStatus id="FD47880B5C2DFFA4FDA1FA15FDA864BE" box="[532,587,1503,1527]" pageId="3" pageNumber="454">Type</typeStatus>
III (crescentic) (
<figureCitation id="BAC72A2C5C2DFFA4FCB2FA14FCA864BE" box="[775,843,1502,1527]" captionStart="FIGURE 2" captionStartId="5.[151,250,1761,1783]" captionTargetBox="[153,1433,199,1739]" captionTargetId="figure@5.[153,1433,193,1740]" captionTargetPageId="5" captionText="FIGURE 2. Ameritulla clavata n. comb. A) Postantennal organ, lectotype. B) Anterior cephalic pseudocellus, Sequatchie Co., Tennessee, specimen. C) Ventral chaetotaxy of Abd. VI, Haywood Co., North Carolina, specimen. Scales: A, B, 10 µm; C, 50 µm." httpUri="https://zenodo.org/record/256861/files/figure.png" pageId="3" pageNumber="454">Fig. 2</figureCitation>
B), formula 11/010/01011. Pseudopores absent. Anal spines stout, slightly curved, about 1.5× longer than hind unguis.
</paragraph>
<paragraph id="224336A95C2DFFA4FF72F9ECFD3E6786" blockId="3.[151,1437,998,2031]" pageId="3" pageNumber="454">Granules on dorsum of head of small to medium size (diameter 1.4‒2.3 µm, mean = 2.0 µm), evenly distributed, appearing round or broadly oval. Diameter of Th. III granules 1.3‒2.0 µm, mean = 1.6 µm. Distinct mid-dorsal longitudinal rows of granules not present. Posterior region of all segments except pronotum and Abd. VI with 8‒12 transverse rows of tightly packed granules (Figs 3A, B). Granules of Abd. VI not in distinct rows, diameter 1.4‒2.1 µm (mean = 1.6 µm) (Figs 3A, B).</paragraph>
<paragraph id="224336A95C2DFFA4FF72F910FF2666A6" blockId="3.[151,1437,998,2031]" pageId="3" pageNumber="454">
Labial palpus (
<figureCitation id="BAC72A2C5C2DFFA4FEC1F910FE5667BB" box="[372,437,1754,1779]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="3" pageNumber="454">Fig. 1</figureCitation>
H) with papillae A‒E, papilla spine A stout, rounded at tip; B‒E pointed. Eight guard setae present: a1 and b1 minute, papilliform, papilla B with one additional guard seta, papilla D with two guards, papilla E with three. Maxillary palpus with one sublobal hair, spine of papilla slightly longer than basal seta (
<figureCitation id="BAC72A2C5C2DFFA4FADAF8E8FF406616" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="3" pageNumber="454">
Fig. 
<date id="564210695C2DFFA4FF22F88DFF406616" box="[151,163,1863,1887]" pageId="3" pageNumber="454">1</date>
</figureCitation>
I). Mandible with four blunt apical teeth (
<figureCitation id="BAC72A2C5C2DFFA4FDC1F88CFD576616" box="[628,692,1862,1887]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="3" pageNumber="454">Fig. 1</figureCitation>
J). Prelabral setae two, stout, curved; labrum with 11 setae in three rows (5-4-2), setae of distal pair and middle setae of middle row stronger and longer than other labral setae (
<figureCitation id="BAC72A2C5C2DFFA4FADAF8A0FF4166EE" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="3" pageNumber="454">Fig. 1</figureCitation>
E). Postantennal organ elongated, in deep trough, with two rows of oval vesicles, each with 18‒22 vesicles (
<figureCitation id="BAC72A2C5C2DFFA4FADAF844FF456683" captionStart="FIGURE 2" captionStartId="5.[151,250,1761,1783]" captionTargetBox="[153,1433,199,1739]" captionTargetId="figure@5.[153,1433,193,1740]" captionTargetPageId="5" captionText="FIGURE 2. Ameritulla clavata n. comb. A) Postantennal organ, lectotype. B) Anterior cephalic pseudocellus, Sequatchie Co., Tennessee, specimen. C) Ventral chaetotaxy of Abd. VI, Haywood Co., North Carolina, specimen. Scales: A, B, 10 µm; C, 50 µm." httpUri="https://zenodo.org/record/256861/files/figure.png" pageId="3" pageNumber="454">Fig. 2</figureCitation>
A). Trough of PAO closed at medial end by one or more granules, not extending to fovea between setae sd1 and sd3.
</paragraph>
<caption id="768366215C2AFFA3FF22F8CBFABE66DB" httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="4" pageNumber="455" targetBox="[168,1421,206,1762]" targetPageId="4">
<paragraph id="224336A95C2AFFA3FF22F8CBFABE66DB" blockId="4.[151,1436,1793,1938]" pageId="4" pageNumber="455">
<emphasis id="1088EABB5C2AFFA3FF22F8CBFEF4665F" bold="true" box="[151,279,1793,1815]" pageId="4" pageNumber="455">FIGURE 1.</emphasis>
<taxonomicName id="E5FC4D2A5C2AFFA3FE94F8CBFE01665E" box="[289,482,1793,1815]" class="Entognatha" family="Tullbergiidae" genus="Ameritulla" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="clavata" status="comb. nov.">
<emphasis id="1088EABB5C2AFFA3FE94F8CBFE01665E" box="[289,482,1793,1815]" italics="true" pageId="4" pageNumber="455">Ameritulla clavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C2AFFA3FE59F8C8FDA8665E" box="[492,587,1793,1815]" pageId="4" pageNumber="455">n. comb.</taxonomicNameLabel>
, lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II‒V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm.
</paragraph>
</caption>
<caption id="768366215C2BFFA2FF22F92BFF5C667D" httpUri="https://zenodo.org/record/256861/files/figure.png" pageId="5" pageNumber="456" targetBox="[153,1433,199,1739]" targetPageId="5">
<paragraph id="224336A95C2BFFA2FF22F92BFF5C667D" blockId="5.[151,1435,1761,1844]" pageId="5" pageNumber="456">
<emphasis id="1088EABB5C2BFFA2FF22F92BFEF067BF" bold="true" box="[151,275,1761,1783]" pageId="5" pageNumber="456">FIGURE 2.</emphasis>
<taxonomicName id="E5FC4D2A5C2BFFA2FEACF92BFE3467BE" box="[281,471,1761,1783]" class="Entognatha" family="Tullbergiidae" genus="Ameritulla" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="clavata" status="comb. nov.">
<emphasis id="1088EABB5C2BFFA2FEACF92BFE3467BE" box="[281,471,1761,1783]" italics="true" pageId="5" pageNumber="456">Ameritulla clavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C2BFFA2FE68F928FDD567BE" box="[477,566,1761,1783]" pageId="5" pageNumber="456">n. comb.</taxonomicNameLabel>
A) Postantennal organ, lectotype. B) Anterior cephalic pseudocellus, Sequatchie Co., Tennessee, specimen. C) Ventral chaetotaxy of Abd. VI, Haywood Co., North Carolina, specimen. Scales: A, B, 10 µm; C, 50 µm.
</paragraph>
</caption>
<paragraph id="224336A95C2BFFA1FF72F897FCA86055" blockId="5.[151,1437,1884,2017]" lastBlockId="6.[151,1436,151,392]" lastPageId="6" lastPageNumber="457" pageId="5" pageNumber="456">
Ant. I with 7 setae, Ant. II with 11 setae. Sense organ of Ant. III (
<figureCitation id="BAC72A2C5C2BFFA2FC10F896FC07663C" box="[933,996,1884,1909]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="5" pageNumber="456">Fig. 1</figureCitation>
F) with two exposed, converging sense clubs and two minute, weakly bilobed or trilobed sensory rods partially covered by prominent cuticular lobe; three proximal guard setae. Ventral sense club stout; all Ant. III setae pointed (
<figureCitation id="BAC72A2C5C2BFFA2FC7CF86EFBEB66F4" box="[969,1032,1956,1981]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="5" pageNumber="456">Fig. 1</figureCitation>
G). Dorsal side of Ant. IV with five arcuate sensilla; microsensillum rod-like, in shallow groove; subapical organite clavate, extending from cup-like depression; apical bulb spherical (
<figureCitation id="BAC72A2C5C28FFA1FDABFF5DFD8361F9" box="[542,608,151,176]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="6" pageNumber="457">Fig. 1</figureCitation>
F). Most longer dorsal and lateral setae of Ant. IV thin but sensilliform with blunt tips, only one pointed seta mid-dorsally. Ventrally, Ant. IV with several sensilliform setae of various lengths; ventral sensory field not defined, short setae a mix of ordinary and thin sensilliform setae with finely rounded apices surrounding larger sensilliform seta (
<figureCitation id="BAC72A2C5C28FFA1FD51FECEFCC76055" box="[740,804,260,285]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="6" pageNumber="457">Fig. 1</figureCitation>
G).
</paragraph>
<paragraph id="224336A95C28FFA1FF72FEEDFCDD60C1" blockId="6.[151,1436,151,392]" pageId="6" pageNumber="457">
All leg setae pointed. Trochanter with 4 setae, femur with 10 setae. Fore and middle tibiotarsi with 15 setae, A and B setal whorls with 7 and 8 setae, respectively; hind tibiotarsus with 14 setae. Unguis stout, without teeth or tunica-like expansion; unguiculus pointed, minute (
<figureCitation id="BAC72A2C5C28FFA1FD62FEA5FCF560C1" box="[727,790,367,392]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="6" pageNumber="457">Fig. 1</figureCitation>
K).
</paragraph>
<paragraph id="224336A95C28FFA1FF22FA03FDC36754" pageId="6" pageNumber="457">
<table id="50FCC4095C280058FF22FA03FA7F6754" box="[151,1436,1481,1565]" gridcols="3" gridrows="3" pageId="6" pageNumber="457">
<tr id="9CCC34EB5C280058FF22FA03FA7F64A9" box="[151,1436,1481,1504]" gridrow="0" pageId="6" pageNumber="457">
<th id="DF1D5D975C280058FF22FA03FDD464A9" box="[151,567,1481,1504]" gridcol="0" gridrow="0" pageId="6" pageNumber="457">
<emphasis id="1088EABB5C28FFA1FF22FA03FEF76497" bold="true" box="[151,276,1481,1503]" pageId="6" pageNumber="457">FIGURE 3.</emphasis>
A, B) 
<taxonomicName id="E5FC4D2A5C28FFA1FEE8FA03FDFE6496" box="[349,541,1481,1503]" class="Entognatha" family="Tullbergiidae" genus="Ameritulla" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="clavata" status="n.">
<emphasis id="1088EABB5C28FFA1FEE8FA03FDFE6496" box="[349,541,1481,1503]" italics="true" pageId="6" pageNumber="457">Ameritulla clavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C28FFA1FD91FA00FDD46496" box="[548,567,1482,1503]" pageId="6" pageNumber="457">n.</taxonomicNameLabel>
</th>
<th id="DF1D5D975C280058FD8BFA03FC6364A9" box="[574,896,1481,1504]" gridcol="1" gridrow="0" pageId="6" pageNumber="457">comb. A) Abdominal segments</th>
<th id="DF1D5D975C280058FC32FA03FA7F64A9" box="[903,1436,1481,1504]" gridcol="2" gridrow="0" pageId="6" pageNumber="457">IV‒VI, dorsal view. B) Abdominal segments V‒VI,</th>
</tr>
<tr id="9CCC34EB5C280058FF22FA22FA7F64B7" box="[151,1436,1512,1534]" gridrow="1" pageId="6" pageNumber="457">
<th id="DF1D5D975C280058FF22FA22FDD464B7" box="[151,567,1512,1534]" gridcol="0" gridrow="1" pageId="6" pageNumber="457">
lectotype, lateral view. C) 
<taxonomicName id="E5FC4D2A5C28FFA1FE19FA22FDD564B7" box="[428,566,1512,1534]" class="Collembola" family="Tullbergiidae" genus="Mesaphorura" higherTaxonomySource="CoL" kingdom="Animalia" order="Poduromorpha" pageId="6" pageNumber="457" phylum="Arthropoda" rank="genus">
<emphasis id="1088EABB5C28FFA1FE19FA22FDD564B7" box="[428,566,1512,1534]" italics="true" pageId="6" pageNumber="457">Mesaphorura</emphasis>
</taxonomicName>
</th>
<td id="DF1D5D975C280058FD8BFA22FC6364B7" box="[574,896,1512,1534]" gridcol="1" gridrow="1" pageId="6" pageNumber="457">
<taxonomicName id="E5FC4D2A5C28FFA1FD8BFA23FD5364B7" authority="Abd." authorityName="Abd." box="[574,688,1512,1534]" class="Collembola" family="Tullbergiidae" genus="Mesaphorura" higherTaxonomySource="CoL" kingdom="Animalia" order="Poduromorpha" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="yosii">
<emphasis id="1088EABB5C28FFA1FD8BFA23FD8D64B7" box="[574,622,1513,1534]" italics="true" pageId="6" pageNumber="457">yosii</emphasis>
, Abd.
</taxonomicName>
VI and posterior of
</td>
<td id="DF1D5D975C280058FC32FA22FA7F64B7" box="[903,1436,1512,1534]" gridcol="2" gridrow="1" pageId="6" pageNumber="457">Abd. V, dorsal view, showing prominent crescentic</td>
</tr>
<tr id="9CCC34EB5C280058FF22F9CDFA7F6754" box="[151,1436,1543,1565]" gridrow="2" pageId="6" pageNumber="457" rowspan-1="1" rowspan-2="1">
<th id="DF1D5D975C280058FF22F9CDFDD46754" box="[151,567,1543,1565]" gridcol="0" gridrow="2" pageId="6" pageNumber="457">ridges. Scales: A, B, 50 µm; C, 20 µm.</th>
</tr>
</table>
</paragraph>
<paragraph id="224336A95C28FFA1FF72F983FBC66617" blockId="6.[151,1437,1609,2030]" pageId="6" pageNumber="457">
Dorsally, most cephalic setae of similar length, sd5, oc2, g3 and p5 longer than others (
<figureCitation id="BAC72A2C5C28FFA1FB74F983FAE7672B" box="[1217,1284,1609,1634]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="6" pageNumber="457">Fig. 1</figureCitation>
A). Six setae between posterior cephalic pseudocelli, seta p1 about 1.5× the length of p2. Two medial setae present anterior to a0. Seta 
<date id="564210695C28FFA1FF48F958FEFA67E3" box="[253,281,1682,1706]" pageId="6" pageNumber="457">v2</date>
present, 
<date id="564210695C28FFA1FE30F958FE4267E3" box="[389,417,1682,1706]" pageId="6" pageNumber="457">v1</date>
absent. Pronotum with 8 setae. Mesonotum and metanotum (
<figureCitation id="BAC72A2C5C28FFA1FBD9F95BFB4C67E3" box="[1132,1199,1681,1706]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="6" pageNumber="457">Fig. 1</figureCitation>
B) each with m-row consisting of m1, m4 and m5; setae a2, a4 and p5 macrosetae; p3 longer than p4 but shorter than p5; p1 and p2 lengths equal; microsensillum in shallow pit near long sensilliform seta. On Abd. I‒V seta p3 distinctly sensilliform. Abd. I‒III lacking m-setae, setae a2 and p5 longer than other setae. On Abd. IV setae a2, a4 and p5 long; small m2-seta present. Abd. V lacking seta p1; p2 long, displaced far forward near a2; a4 and p5 macrosetae. Abd. VI with two unpaired medial setae (m0, p0), without swellings or tubercles.
</paragraph>
<paragraph id="224336A95C28FFA1FF72F8A3FBF166EF" blockId="6.[151,1437,1609,2030]" pageId="6" pageNumber="457">
Pleural regions of Abd. I‒V with 1, 2, 2, 6, 1 setae, respectively; sensilliform setae present on Abd. II‒IV, anteriorly inserted on Abd. II and III, posteriorly inserted on Abd. IV (
<figureCitation id="BAC72A2C5C28FFA1FC18F847FC0F66EF" box="[941,1004,1933,1958]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="6" pageNumber="457">Fig. 1</figureCitation>
B).
</paragraph>
<paragraph id="224336A95C28FFA0FF72F87BFCBD6055" blockId="6.[151,1437,1609,2030]" lastBlockId="7.[151,1437,151,573]" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457">
Ventrally, head with three pairs of medial setae. Mesosternum and metasternum each with two setae. Ventral tube with 4+4 setae on valves and 2+2 setae on base. Ventral abdominal setae typically with some asymmetry, especially on Abd. IV (
<figureCitation id="BAC72A2C5C29FFA0FE15FF5DFE0361F9" box="[416,480,151,176]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="7" pageNumber="458">Fig. 1</figureCitation>
C). Ventral setae p2 and p5 on Abd. II, p5 on Abd. III and IV, and posterior setae of Abd. V sensilliform. Remnant tenacular and furcal regions absent on Abd. IV and V. Sternum of Abd. V with six setae surrounding the genital aperture and two microsetae on the anterior lip; Abd. VI sternum with full complement of setae (a1‒5, m2‒5, p1‒5, va1‒3) (
<figureCitation id="BAC72A2C5C29FFA0FD71FECEFCE46055" box="[708,775,260,285]" captionStart="FIGURE 1" captionStartId="4.[151,250,1793,1815]" captionTargetBox="[168,1421,206,1762]" captionTargetId="figure@4.[151,1436,193,1772]" captionTargetPageId="4" captionText="FIGURE 1. Ameritulla clavata n. comb., lectotype female unless otherwise noted. A) Head, dorsal view. B) Thorax and abdomen, dorsal chaetotaxy. C) Abdominal segments II ‒ V, ventral chaetotaxy, specimen from Great Smoky Mountains National Park (GRSM). D) Venter of Abd. VI, specimen from GRSM. E) Labrum. F) Third and fourth antennal segments, dorsal view. G) Fourth antennal segment, ventral view. H) Labial palpus. I) Outer lobe of maxilla. J) Apex of mandible. K) Middle femur and tibiotarsus, ventral view. Scales: A, 50 µm; B, C, 50 µm; D, H, I, 20 µm; E, 20 µm; F, G, H, K, 20 µm." httpUri="https://zenodo.org/record/256860/files/figure.png" pageId="7" pageNumber="458">Figs 1</figureCitation>
D, 2C).
</paragraph>
</subSubSection>
<subSubSection id="6AE665225C29FFA0FF72FEEDFC166374" pageId="7" pageNumber="458" type="discussion">
<paragraph id="224336A95C29FFA0FF72FEEDFDAC60BD" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="458">
<emphasis id="1088EABB5C29FFA0FF72FEEDFED86009" bold="true" box="[199,315,295,320]" pageId="7" pageNumber="458">Remarks.</emphasis>
<taxonomicName id="E5FC4D2A5C29FFA0FEF7FEEDFDF76009" box="[322,532,295,320]" class="Entognatha" family="Tullbergiidae" genus="Ameritulla" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="clavata" status="comb. nov.">
<emphasis id="1088EABB5C29FFA0FEF7FEEDFDF76009" box="[322,532,295,320]" italics="true" pageId="7" pageNumber="458">Ameritulla clavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C29FFA0FDA9FEEDFD616009" box="[540,642,295,320]" pageId="7" pageNumber="458">
<emphasis id="1088EABB5C29FFA0FDA9FEEDFD616009" bold="true" box="[540,642,295,320]" pageId="7" pageNumber="458">n. comb.</emphasis>
</taxonomicNameLabel>
is easily recognized as the only large North American tullbergiid with pseudocelli absent from both Th. I and Th. III. The only other tullbergiids with this arrangement are the smaller 
<taxonomicName id="E5FC4D2A5C29FFA0FF22FEA5FE9460C1" box="[151,375,367,392]" class="Entognatha" family="Tullbergiidae" genus="Mixturatulla" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="ozwini" status="gen. nov.">
<emphasis id="1088EABB5C29FFA0FF22FEA5FE9460C1" box="[151,375,367,392]" italics="true" pageId="7" pageNumber="458">Mixturatulla ozwini</emphasis>
</taxonomicName>
<emphasis id="1088EABB5C29FFA0FE37FEA5FDC560C1" bold="true" box="[386,550,367,392]" pageId="7" pageNumber="458">
<taxonomicNameLabel id="0BBB57C05C29FFA0FE37FEA5FE3760C1" box="[386,468,367,392]" pageId="7" pageNumber="458" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
and 
<taxonomicName id="E5FC4D2A5C29FFA0FDD6FEA5FC4E60C1" box="[611,941,367,392]" class="Entognatha" family="Tullbergiidae" genus="Psammophorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="miniclavata" status="sp. nov.">
<emphasis id="1088EABB5C29FFA0FDD6FEA5FC4E60C1" box="[611,941,367,392]" italics="true" pageId="7" pageNumber="458">Psammophorura miniclavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C29FFA0FC02FEA5FC1A60C1" box="[951,1017,367,392]" pageId="7" pageNumber="458" rank="species">
<emphasis id="1088EABB5C29FFA0FC02FEA5FC1A60C1" bold="true" box="[951,1017,367,392]" pageId="7" pageNumber="458">n. sp.</emphasis>
</taxonomicNameLabel>
Oddly, 
<taxonomicName id="E5FC4D2A5C29FFA0FBEDFEBBFB1860C1" box="[1112,1275,367,392]" class="Entognatha" family="Tullbergiidae" genus="Psammophorura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="miniclavata" status="sp. nov.">
<emphasis id="1088EABB5C29FFA0FBEDFEBBFB1860C1" box="[1112,1275,367,392]" italics="true" pageId="7" pageNumber="458">P. miniclavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C29FFA0FAB0FEA5FAA460C1" box="[1285,1351,367,392]" pageId="7" pageNumber="458" rank="species">
<emphasis id="1088EABB5C29FFA0FAB0FEA5FAA460C1" bold="true" box="[1285,1351,367,392]" pageId="7" pageNumber="458">n. sp.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="E5FC4D2A5C29FFA0FA31FEBAFF0960E4" class="Entognatha" family="Tullbergiidae" genus="Ameritulla" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="clavata" status="comb. nov.">
<emphasis id="1088EABB5C29FFA0FA31FEBAFF0960E4" italics="true" pageId="7" pageNumber="458">A. clavata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C29FFA0FF47FE5EFEBA60E4" box="[242,345,404,429]" pageId="7" pageNumber="458">
<emphasis id="1088EABB5C29FFA0FF47FE5EFEBA60E4" bold="true" box="[242,345,404,429]" pageId="7" pageNumber="458">n. comb.</emphasis>
</taxonomicNameLabel>
have the same pseudocellar formula (11/010/01011). The constant presence of seta m2 on Abd. IV is a distinctive character and may be unique in this family. 
<taxonomicName id="E5FC4D2A5C29FFA0FCD1FE7DFBDD6099" box="[868,1086,439,464]" class="Entognatha" family="Tullbergiidae" genus="Ameritulla" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="obscura" status="sp. nov.">
<emphasis id="1088EABB5C29FFA0FCD1FE7DFBDD6099" box="[868,1086,439,464]" italics="true" pageId="7" pageNumber="458">Ameritulla obscura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0BBB57C05C29FFA0FBF2FE7DFB6B6099" box="[1095,1160,439,464]" pageId="7" pageNumber="458" rank="species">
<emphasis id="1088EABB5C29FFA0FBF2FE7DFB6B6099" bold="true" box="[1095,1160,439,464]" pageId="7" pageNumber="458">n. sp.</emphasis>
</taxonomicNameLabel>
(below) has one pair of pseudocelli on Th. II‒III and Abd. I‒V.
</paragraph>
<paragraph id="224336A95C29FFA0FF72FE35FC166374" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="458">
<bibRefCitation id="466D4B585C29FFA0FF72FE35FE576351" author="Waltz" box="[199,436,511,536]" pageId="7" pageNumber="458" refString="Waltz, R. D. &amp; Hart, J. W. (1995) First report of males of Tullbergia clavata (Collembola: Onychiuridae). Entomological News, 106 (3), 115 - 116." type="journal article" year="1995">Waltz &amp; Hart (1995)</bibRefCitation>
reported on the presence of males in two 
<collectingRegion id="E038F84B5C29FFA0FC2DFE35FC0F6351" box="[920,1004,511,536]" country="United States of America" name="Indiana" pageId="7" pageNumber="458">Indiana</collectingRegion>
locations and minor variability in the lengths of some setae, but ascribed setal differences to intraspecies variation.
</paragraph>
</subSubSection>
</treatment>
</document>