treatments-xml/data/A9/68/87/A96887E8FFD2FF84FF43FD52FDFA0707.xml

<document id="D639FCF68D80565166CCA1523B76B1D5" ID-CLB-Dataset="36851" ID-DOI="10.11646/zootaxa.3799.1.1" ID-GBIF-Dataset="d9f29df5-52ff-451f-a367-f9fdb29b180f" ID-ISSN="1175-5326" ID-Zenodo-Dep="286090" ID-ZooBank="56DD05E1-C61C-4D37-9454-396840EB67C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460422790352" checkinUser="plazi" docAuthor="Gibbs, David" docDate="2014" docId="A96887E8FFD2FF84FF43FD52FDFA0707" docLanguage="en" docName="zt03799p085.pdf" docOrigin="Zootaxa 3799 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Usia Latreille 1802, sensu stricto" docType="key" docVersion="5" lastPageNumber="21" masterDocId="5551FF90FFD5FF90FFD4FFCAFFF60616" masterDocTitle="A world revision of the bee fly tribe Usiini (Diptera, Bombyliidae) Part 2: Usia sensu stricto" masterLastPageNumber="85" masterPageNumber="1" pageNumber="8" updateTime="1698330265731" updateUser="ExternalLinkService">
<mods:mods id="D05D22516BA3BD0B08AF93BC821324D8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3E0DD89252E01171A9E50A8EFEEF50EC">
<mods:title id="1F16557E5DCF28E0273A010698305190">A world revision of the bee fly tribe Usiini (Diptera, Bombyliidae) Part 2: Usia sensu stricto</mods:title>
</mods:titleInfo>
<mods:name id="D5CA12178361D8B4D5010C8E567BCE0D" type="personal">
<mods:role id="7BB272B7AF940FE22EC2728EC5775C5C">
<mods:roleTerm id="7F180DB30AB5E87A66043D9E51982162">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A76834CA76E72D25E42E57786E3B110B">Gibbs, David</mods:namePart>
</mods:name>
<mods:typeOfResource id="6FFFE5589A1CCDD31017E31BD942F51F">text</mods:typeOfResource>
<mods:relatedItem id="4F229B0BAF3EE00E588F2FEC16209F5F" type="host">
<mods:titleInfo id="859DB55D8BA1CB40F56FC94176643ACA">
<mods:title id="C8370289EA013AF851D0B7586A2E5959">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D304A6F299A8218ED73147F3B7DD0C98">
<mods:date id="1FFBA8709BA73FD24E81890D198AE5FB">2014</mods:date>
<mods:detail id="A9F79C826AA4714B92616E5C9A905E3C" type="pubDate">
<mods:number id="75C97E40CD49C2FEFB1A88C57126257B">2014-05-22</mods:number>
</mods:detail>
<mods:detail id="C06899646AD94965B685AAB6AFA3C192" type="volume">
<mods:number id="89AA7A21D0C92FDB0A654F6C41CAE509">3799</mods:number>
</mods:detail>
<mods:detail id="114C403797998B3B0512102F00B3B85C" type="issue">
<mods:number id="28BB5095B541FF3B507EDBC05E6EB0DD">1</mods:number>
</mods:detail>
<mods:extent id="EEFFA47990356E8B8F3113496AAFA3BD" unit="page">
<mods:start id="5A0935C47373B024AA80BF41843B3470">1</mods:start>
<mods:end id="0803826E08DD70E5B67871C96DCC70F5">85</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E922A369B9DACF60391209C54CA21D42">journal article</mods:classification>
<mods:identifier id="85E3738CD9A6E545B57A3FC995791C70" type="CLB-Dataset">36851</mods:identifier>
<mods:identifier id="1CF5801AB8E1E8E7C2715934A4F64688" type="DOI">10.11646/zootaxa.3799.1.1</mods:identifier>
<mods:identifier id="23DBACFA053A82753EF01FC2566DB934" type="GBIF-Dataset">d9f29df5-52ff-451f-a367-f9fdb29b180f</mods:identifier>
<mods:identifier id="98F457B009C337E164AE7027BCA90266" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="97A0140D86FA20D53F83F951F6875F5A" type="Zenodo-Dep">286090</mods:identifier>
<mods:identifier id="B015A9014D7813DF15ED704E78E394BC" type="ZooBank">56DD05E1-C61C-4D37-9454-396840EB67C0</mods:identifier>
</mods:mods>
<treatment id="A96887E8FFD2FF84FF43FD52FDFA0707" ID-DOI="http://doi.org/10.5281/zenodo.6135474" ID-GBIF-Taxon="194203641" ID-Zenodo-Dep="6135474" LSID="urn:lsid:plazi:treatment:A96887E8FFD2FF84FF43FD52FDFA0707" httpUri="http://treatment.plazi.org/id/A96887E8FFD2FF84FF43FD52FDFA0707" lastPageId="20" lastPageNumber="21" pageId="7" pageNumber="8">
<subSubSection id="69DB6575FFD2FF97FF43FD52FDBE04A7" box="[151,584,663,690]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="217E36FEFFD2FF97FF43FD52FDBE04A7" blockId="7.[151,584,663,690]" box="[151,584,663,690]" pageId="7" pageNumber="8">
<heading id="7A368192FFD2FF97FF43FD52FDBE04A7" bold="true" box="[151,584,663,690]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="13B5EAECFFD2FF97FF43FD52FDBE04A7" bold="true" box="[151,584,663,690]" pageId="7" pageNumber="8">
Key to Species of 
<emphasis id="13B5EAECFFD2FF97FEA7FD5DFDBE04A7" bold="true" box="[371,584,663,689]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="E6C14D7DFFD2FF97FEA7FD5DFE5C04A7" ID-CoL="63VJK" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[371,426,663,689]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="stricto">Usia</taxonomicName>
<taxonomicNameLabel id="08865797FFD2FF97FE65FD5DFDBE04A7" box="[433,584,663,689]" pageId="7" pageNumber="8" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFD2FF97FF43FD15FE7D05E5" pageId="7" pageNumber="8" type="discussion">
<paragraph id="217E36FEFFD2FF97FF43FD15FE7D05E5" blockId="7.[151,1437,734,1047]" pageId="7" pageNumber="8">As with most parasitoid insects, Usiini can be exceptionally variable, some specimens of the same species being several times the size of the smallest examples. Such large discrepancies in size can often be accompanied by differences in proportions, width of frons, density of dusting, hairiness, surface sculpture and many other minor variations. While every effort has been made to avoid the more variable characters, there will always be specimens that will not run cleanly through a key or may even run to the wrong species. If in any doubt, it is essential to dissect and examine the male and/or female genitalia. Even this precaution might not be adequate for some females. Such specimens collected outside of an expected range and/or without associated males should be treated with extreme caution.</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFD2FF97FF43FC35FC450201" box="[151,947,1022,1047]" pageId="7" pageNumber="8" type="multiple">
<paragraph id="217E36FEFFD2FF97FF43FC35FC450201" blockId="7.[151,1437,734,1047]" box="[151,947,1022,1047]" pageId="7" pageNumber="8">
[♂ 
<taxonomicName id="E6C14D7DFFD2FF97FF68FC35FEC30201" box="[188,309,1022,1047]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="calva">
<emphasis id="13B5EAECFFD2FF97FF68FC35FEC30201" box="[188,309,1022,1047]" italics="true" pageId="7" pageNumber="8">Usia calva</emphasis>
</taxonomicName>
and ♀ 
<taxonomicName id="E6C14D7DFFD2FF97FE53FC35FDE50200" box="[391,531,1023,1046]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="13B5EAECFFD2FF97FE53FC35FDE50200" box="[391,531,1023,1046]" italics="true" pageId="7" pageNumber="8">U. incognita</emphasis>
</taxonomicName>
unknown so not included in the key]
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFD2FF83FF43FB8CFA6D002D" lastPageId="19" lastPageNumber="20" pageId="7" pageNumber="8" type="key">
<keyStep id="9A352E5BFFD2FF97FF43FB8CFA6D026F" pageId="7" pageNumber="8">
<paragraph id="217E36FEFFD2FF97FF43FB8CFA6D024D" blockId="7.[151,1436,1094,1321]" box="[151,1435,1094,1115]" pageId="7" pageNumber="8">
<keyLead id="9A3095CBFFD2FF97FF43FB8CFA6D024D" box="[151,1435,1094,1115]" pageId="7" pageNumber="8">1. Males............................................................................................... 2</keyLead>
</paragraph>
<paragraph id="217E36FEFFD2FF97FF43FBAEFA6D026F" blockId="7.[151,1436,1094,1321]" box="[151,1435,1124,1145]" pageId="7" pageNumber="8">
<keyLead id="9A3095CBFFD2FF97FF43FBAEFA6D026F" box="[151,1435,1124,1145]" pageId="7" pageNumber="8">- Females............................................................................................ 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFD2FF97FF43FB4BFA6A02C7" pageId="7" pageNumber="8">
<paragraph id="217E36FEFFD2FF97FF43FB4BFA6D02A5" blockId="7.[151,1436,1094,1321]" pageId="7" pageNumber="8">
<keyLead id="9A3095CBFFD2FF97FF43FB4BFA6D02A5" pageId="7" pageNumber="8">2. Occiput densely grey dusted obscuring black ground colour, except in many species, small triangular area behind ocellar tubercle shining black...................................................................................... 3</keyLead>
</paragraph>
<paragraph id="217E36FEFFD2FF97FF43FB76FA6A02C7" blockId="7.[151,1436,1094,1321]" box="[151,1436,1212,1233]" pageId="7" pageNumber="8">
<keyLead id="9A3095CBFFD2FF97FF43FB76FA6A02C7" box="[151,1436,1212,1233]" pageId="7" pageNumber="8">- Occiput shining black, at least immediately behind eyes, any grey dusting present thin not obscuring ground colour....... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFD2FF97FF43FB13FA6A033F" pageId="7" pageNumber="8">
<paragraph id="217E36FEFFD2FF97FF43FB13FA6D031D" blockId="7.[151,1436,1094,1321]" pageId="7" pageNumber="8">
<keyLead id="9A3095CBFFD2FF97FF43FB13FA6D031D" pageId="7" pageNumber="8">
3. Frons laterally with outstanding hairs almost as long as those on ocellar tubercle and extending about half way to front of frons (
<figureCitation id="B9FA2A7BFFD2FF97FF33FB3CFEC1031D" box="[231,311,1270,1291]" captionStart="FIGURE 1" captionStartId="7.[151,250,2010,2032]" captionTargetId="figure@7.[450,1129,1370,1980]" captionTargetPageId="7" captionText="FIGURE 1. ♂ U. bicolor frons." figureDoi="http://doi.org/10.5281/zenodo.6376797" httpUri="https://zenodo.org/record/6376797/files/figure.png" pageId="7" pageNumber="8">Figure 1</figureCitation>
)............................................................................................ 4
</keyLead>
</paragraph>
<paragraph id="217E36FEFFD2FF97FF43FADEFA6A033F" blockId="7.[151,1436,1094,1321]" box="[151,1436,1300,1321]" pageId="7" pageNumber="8">
<keyLead id="9A3095CBFFD2FF97FF43FADEFA6A033F" box="[151,1436,1300,1321]" pageId="7" pageNumber="8">- Frons lacking long hairs forward of anterior ocellus, any hairs in this position short and procumbent.................... 5</keyLead>
</paragraph>
</keyStep>
<caption id="75BE6676FFD2FF97FF43F810FE2A01E6" ID-DOI="http://doi.org/10.5281/zenodo.6376797" ID-Zenodo-Dep="6376797" box="[151,476,2010,2032]" httpUri="https://zenodo.org/record/6376797/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,2010,2032]" targetBox="[457,1122,1377,1977]" targetPageId="7">
<paragraph id="217E36FEFFD2FF97FF43F810FE2A01E6" blockId="7.[151,476,2010,2032]" box="[151,476,2010,2032]" pageId="7" pageNumber="8">
<emphasis id="13B5EAECFFD2FF97FF43F810FEE501E6" bold="true" box="[151,275,2010,2032]" pageId="7" pageNumber="8">FIGURE 1.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFD2FF97FEE6F811FE6C01E6" ID-CoL="8FL5Y" box="[306,410,2010,2032]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="13B5EAECFFD2FF97FEE6F811FE6C01E6" box="[306,410,2010,2032]" italics="true" pageId="7" pageNumber="8">U. bicolor</emphasis>
</taxonomicName>
frons.
</paragraph>
</caption>
<keyStep id="9A352E5BFFDDFF98FF43FF5DFA6A0729" pageId="8" pageNumber="9">
<paragraph id="217E36FEFFDDFF98FF43FF5DFA6A06F1" blockId="8.[151,1436,151,320]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FF5DFA6A06F1" pageId="8" pageNumber="9">
4. A shorter haired, neater species, almost all hairs on mesonotum of same length, none longer than hairs on ocellar tubercle. Tibia covered with short, mostly adpressed setae. Epandrium more obviously swollen basally, sides concave apically, apices of gonocoxite convergent, tips touching or almost so (Plate III).................................... ♂ 
<taxonomicName id="E6C14D7DFFDDFF98FB20FF18FA6A06F1" ID-CoL="8FL5Y" authority="Macquart" authorityName="Macquart" box="[1268,1436,210,232]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="13B5EAECFFDDFF98FB20FF18FACF06FE" bold="true" box="[1268,1337,210,232]" italics="true" pageId="8" pageNumber="9">bicolor</emphasis>
Macquart
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDDFF98FF43FF25FA6A0729" blockId="8.[151,1436,151,320]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FF25FA6A0729" pageId="8" pageNumber="9">
- Vestiture longer and with uneven, shaggy appearance. Mesonotum with both short hairs and much longer ones which are often as long as those on ocellar tubercle (
<figureCitation id="B9FA2A7BFFDDFF98FDEEFEC7FD780734" box="[570,654,269,290]" captionStart="FIGURE 2" captionStartId="8.[151,250,1045,1067]" captionTargetId="figure@8.[358,1228,369,1024]" captionTargetPageId="8" captionText="FIGURE 2. ♂ U. crinipes head and mesonotum." figureDoi="http://doi.org/10.5281/zenodo.6376799" httpUri="https://zenodo.org/record/6376799/files/figure.png" pageId="8" pageNumber="9">Figure 2</figureCitation>
). Tibia usually with outstanding hairs. Epandrium more rectangular, sides straight apically, apices of gonocoxite parallel, tips distinctly separated (Plate VI)........................... ♂ 
<taxonomicName id="E6C14D7DFFDDFF98FAD7FEE0FA6A0729" ID-CoL="8FL63" authority="Becker" authorityName="Becker" box="[1283,1436,298,320]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="crinipes">
<emphasis id="13B5EAECFFDDFF98FAD7FEE0FAA70756" bold="true" box="[1283,1361,298,320]" italics="true" pageId="8" pageNumber="9">crinipes</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="75BE6676FFDDFF98FF43FBDFFD71023D" ID-DOI="http://doi.org/10.5281/zenodo.6376799" ID-Zenodo-Dep="6376799" box="[151,647,1045,1067]" httpUri="https://zenodo.org/record/6376799/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1045,1067]" targetBox="[358,1228,369,1023]" targetPageId="8">
<paragraph id="217E36FEFFDDFF98FF43FBDFFD71023D" blockId="8.[151,647,1045,1067]" box="[151,647,1045,1067]" pageId="8" pageNumber="9">
<emphasis id="13B5EAECFFDDFF98FF43FBDFFEE4023D" bold="true" box="[151,274,1045,1067]" pageId="8" pageNumber="9">FIGURE 2.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDDFF98FEE6FBDCFE55023D" ID-CoL="8FL63" box="[306,419,1046,1067]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="crinipes">
<emphasis id="13B5EAECFFDDFF98FEE6FBDCFE55023D" box="[306,419,1046,1067]" italics="true" pageId="8" pageNumber="9">U. crinipes</emphasis>
</taxonomicName>
head and mesonotum.
</paragraph>
</caption>
<keyStep id="9A352E5BFFDDFF98FF43FBA4FA6D02A8" pageId="8" pageNumber="9">
<paragraph id="217E36FEFFDDFF98FF43FBA4FA6D02B7" blockId="8.[151,1436,1134,1774]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FBA4FA6D02B7" pageId="8" pageNumber="9">
5. Femora and tibia entirely yellow; tergites with conspicuous yellow apical margins. Epandrium and gonocoxites distinctive..................................................................................♂ 
<taxonomicName id="E6C14D7DFFDDFF98FB78FB41FA6D02B7" ID-CoL="8FL6T" authority="Paramonov" authorityName="Paramonov" box="[1196,1435,1163,1185]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="13B5EAECFFDDFF98FB78FB41FADE02B7" bold="true" box="[1196,1320,1163,1185]" italics="true" pageId="8" pageNumber="9">transcaspica</emphasis>
Paramonov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDDFF98FF43FB63FA6D02A8" blockId="8.[151,1436,1134,1774]" box="[151,1435,1193,1214]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FB63FA6D02A8" box="[151,1435,1193,1214]" pageId="8" pageNumber="9">- Legs and abdomen entirely shining blackish or aeneous....................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDDFF98FF43FB0CFA6D02EF" pageId="8" pageNumber="9">
<paragraph id="217E36FEFFDDFF98FF43FB0CFA6A02CD" blockId="8.[151,1436,1134,1774]" box="[151,1436,1222,1243]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FB0CFA6A02CD" box="[151,1436,1222,1243]" pageId="8" pageNumber="9">6. Gonocoxite either side with characteristic blunt-ended ‘horn’ which can usually be seen in pinned specimens............. 7</keyLead>
</paragraph>
<paragraph id="217E36FEFFDDFF98FF43FB2EFA6D02EF" blockId="8.[151,1436,1134,1774]" box="[151,1435,1252,1273]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FB2EFA6D02EF" box="[151,1435,1252,1273]" pageId="8" pageNumber="9">- Gonocoxite lacking such ‘horns’..........................................................................8</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDDFF98FF43FACBFAF603D0" pageId="8" pageNumber="9">
<paragraph id="217E36FEFFDDFF98FF43FACBFA6D0347" blockId="8.[151,1436,1134,1774]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FACBFA6D0347" pageId="8" pageNumber="9">
7. A very short-haired species, disc of mesonotum almost naked, usual vestiture reduced to tiny blackish spicules marginally, absent or practically invisible on disc. Oral margin very narrow, narrower than apical diameter of palps. Anepisternum shining at least along anterior and dorsal margin. Gonostylus T-shaped............................... ♂ 
<taxonomicName id="E6C14D7DFFDDFF98FB13FAF1FA6D0347" ID-CoL="8FL5S" authority="Paramonov" authorityName="Paramonov" box="[1223,1435,1339,1361]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aeneoides">
<emphasis id="13B5EAECFFDDFF98FB13FAF1FADE0347" bold="true" box="[1223,1320,1339,1361]" italics="true" pageId="8" pageNumber="9">aeneoides</emphasis>
Paramonov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDDFF98FF43FA93FAF603D0" blockId="8.[151,1436,1134,1774]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FA93FAF603D0" pageId="8" pageNumber="9">
- Mesonotum with covering of short pale yellowish-white hairs, most clearly seen on notopleuron. Oral margin wider, equal to apical diameter of palps. Anepisternum entirely or mostly dusted, no more than narrow anterodorsal shining strip. Gonostylus T-shaped with extra tooth externally....................................................... ♂ 
<taxonomicName id="E6C14D7DFFDDFF98FB3DFA59FA6D03BF" ID-CoL="8FL6Q" authority="Paramonov" authorityName="Paramonov" box="[1257,1435,1427,1449]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="13B5EAECFFDDFF98FB3DFA59FADF03BF" bold="true" box="[1257,1321,1427,1449]" italics="true" pageId="8" pageNumber="9">similis</emphasis>
Paramonov
</taxonomicName>
[
<emphasis id="13B5EAECFFDDFF98FF3DFA7BFB0403D0" box="[233,1266,1456,1478]" italics="true" pageId="8" pageNumber="9">From here on external features will often be inadequate to arrive at a reliable name, always check genitalia</emphasis>
.]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDDFF98FF43FA04FA6A0060" pageId="8" pageNumber="9">
<paragraph id="217E36FEFFDDFF98FF43FA04FA6D0008" blockId="8.[151,1436,1134,1774]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43FA04FA6D0008" pageId="8" pageNumber="9">
8. Wing sub-basally yellowish contrasting with dark infuscation from humeral crossvein to middle of wing (
<figureCitation id="B9FA2A7BFFDDFF98FB01FA04FAC603F5" box="[1237,1328,1486,1507]" captionStart="FIGURE 3" captionStartId="9.[151,250,869,891]" captionTargetId="figure@9.[358,1228,193,848]" captionTargetPageId="9" captionText="FIGURE 3. ♂ U. aenea wing of long-haired form." figureDoi="http://doi.org/10.5281/zenodo.6376801" httpUri="https://zenodo.org/record/6376801/files/figure.png" pageId="8" pageNumber="9">Figures 3</figureCitation>
&amp; 
<figureCitation id="B9FA2A7BFFDDFF98FA9BFA04FAAA03F5" box="[1359,1372,1486,1507]" captionStart="FIGURE 4" captionStartId="9.[151,250,1769,1791]" captionTargetId="figure@9.[358,1228,1092,1747]" captionTargetPageId="9" captionText="FIGURE 4. ♂ U. aenea wing of short-haired form." figureDoi="http://doi.org/10.5281/zenodo.6376803" httpUri="https://zenodo.org/record/6376803/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
) (very variable and occasional specimens lack this feature so will only be determinable by dissection) and grey dust spots on frons clearly separated...................................................................................... 9
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDDFF98FF43F9ECFA6A0060" blockId="8.[151,1436,1134,1774]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43F9ECFA6A0060" pageId="8" pageNumber="9">
- Wing either clear or if infuscated then without contrasting yellowish area basal to humeral crossvein. If with yellow base and vague infuscation beyond humeral crossvein (
<taxonomicName id="E6C14D7DFFDDFF98FD77F98EFCDF004F" ID-CoL="8FL82" box="[675,809,1603,1625]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="13B5EAECFFDDFF98FD77F98EFCDF004F" box="[675,809,1603,1625]" italics="true" pageId="8" pageNumber="9">U. greatheadi</emphasis>
</taxonomicName>
) then dust spots on frons connected anterior to anterior ocellus, 
<figureCitation id="B9FA2A7BFFDDFF98FF35F9ABFA6A0060" box="[225,1436,1633,1654]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="10.[151,250,1764,1786]" captionStartId-1="11.[151,250,869,891]" captionStartId-2="11.[151,250,1736,1758]" captionStartId-3="12.[151,250,869,891]" captionStartId-4="12.[151,250,1940,1962]" captionTargetId-0="figure@10.[358,1228,1087,1742]" captionTargetId-1="figure@11.[358,1228,193,848]" captionTargetId-2="figure@11.[358,1228,1060,1715]" captionTargetId-3="figure@12.[358,1228,193,848]" captionTargetId-4="figure@12.[358,1228,1263,1918]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="12" captionText-0="FIGURE 6. ♂ U. greatheadi head and mesonotum dorsal." captionText-1="FIGURE 7. ♂ U. unicolor head and mesonotum dorsal." captionText-2="FIGURE 8. ♂ U. lata head and mesonotum dorsal." captionText-3="FIGURE 9. ♂ U. angustifrons head and mesonotum dorsal." captionText-4="FIGURE 10. ♂ U. anatoliensis head and mesonotum dorsal." figureDoi-0="http://doi.org/10.5281/zenodo.6376808" figureDoi-1="http://doi.org/10.5281/zenodo.6376810" figureDoi-2="http://doi.org/10.5281/zenodo.6376812" figureDoi-3="http://doi.org/10.5281/zenodo.6376814" figureDoi-4="http://doi.org/10.5281/zenodo.6376816" httpUri-0="https://zenodo.org/record/6376808/files/figure.png" httpUri-1="https://zenodo.org/record/6376810/files/figure.png" httpUri-2="https://zenodo.org/record/6376812/files/figure.png" httpUri-3="https://zenodo.org/record/6376814/files/figure.png" httpUri-4="https://zenodo.org/record/6376816/files/figure.png" pageId="8" pageNumber="9">Figure 6............................................................................................10</figureCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDDFF99FF43F9B4FA6D0204" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<paragraph id="217E36FEFFDDFF98FF43F9B4FA6D00F8" blockId="8.[151,1436,1134,1774]" pageId="8" pageNumber="9">
<keyLead id="9A3095CBFFDDFF98FF43F9B4FA6D00F8" pageId="8" pageNumber="9">
9. Longer haired species, longest ocellar hairs longer than narrowest width of frons, third antennal segment about 
<date id="557F103EFFDDFF98FACDF9B4FAC70085" box="[1305,1329,1662,1683]" pageId="8" pageNumber="9">2x</date>
scape plus pedicel, isolated spots of infuscation over origin of R4, r-m and m-cu crossveins, 1st basal cell clear subapically, 2nd basal cell largely clear (
<figureCitation id="B9FA2A7BFFDDFF98FEB3F971FE4A00C6" box="[359,444,1723,1744]" captionStart="FIGURE 3" captionStartId="9.[151,250,869,891]" captionTargetId="figure@9.[358,1228,193,848]" captionTargetPageId="9" captionText="FIGURE 3. ♂ U. aenea wing of long-haired form." figureDoi="http://doi.org/10.5281/zenodo.6376801" httpUri="https://zenodo.org/record/6376801/files/figure.png" pageId="8" pageNumber="9">Figure 3</figureCitation>
). Epandrium shorter, about as long as wide, curving tip of aedeagus viewed laterally almost parallel. 
<collectingCountry id="59D6766EFFDDFF98FF35F913FEEE00F8" box="[225,280,1753,1774]" name="Spain" pageId="8" pageNumber="9">Spain</collectingCountry>
.....................................................................♂ 
<taxonomicName id="E6C14D7DFFDDFF98FBB9F912FB0500F8" ID-CoL="8FL5Q" authority="Rossi" authorityName="Rossi" box="[1133,1267,1752,1774]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="13B5EAECFFDDFF98FBB9F912FB5E00F8" bold="true" box="[1133,1192,1752,1774]" italics="true" pageId="8" pageNumber="9">aenea</emphasis>
(Rossi)
</taxonomicName>
long-haired form
</keyLead>
</paragraph>
<caption id="75BE6676FFDCFF99FF43FCAFFD6A056D" ID-DOI="http://doi.org/10.5281/zenodo.6376801" ID-Zenodo-Dep="6376801" box="[151,668,869,891]" httpUri="https://zenodo.org/record/6376801/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,869,891]" targetBox="[358,1228,193,847]" targetPageId="9">
<paragraph id="217E36FEFFDCFF99FF43FCAFFD6A056D" blockId="9.[151,668,869,891]" box="[151,668,869,891]" pageId="9" pageNumber="10">
<emphasis id="13B5EAECFFDCFF99FF43FCAFFEE4056D" bold="true" box="[151,274,869,891]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDCFF99FEE6FCACFE7B056D" ID-CoL="8FL5Q" box="[306,397,870,891]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="13B5EAECFFDCFF99FEE6FCACFE7B056D" box="[306,397,870,891]" italics="true" pageId="9" pageNumber="10">U. aenea</emphasis>
</taxonomicName>
wing of long-haired form.
</paragraph>
</caption>
<paragraph id="217E36FEFFDCFF99FF43FC6BFA6D0204" blockId="9.[151,1436,929,1043]" pageId="9" pageNumber="10">
<keyLead id="9A3095CBFFDCFF99FF43FC6BFA6D0204" pageId="9" pageNumber="10">
- Shorter haired species, longest ocellar hairs shorter than narrowest width of frons, third antennal segment about 1.5x scape plus pedicel, infuscation of wing filling 1st and 2nd basal cell so without isolated darker spots over origin of R4, r-m and m-cu crossveins (
<figureCitation id="B9FA2A7BFFDCFF99FEF6FC2AFE8E05E3" box="[290,376,992,1013]" captionStart="FIGURE 4" captionStartId="9.[151,250,1769,1791]" captionTargetId="figure@9.[358,1228,1092,1747]" captionTargetPageId="9" captionText="FIGURE 4. ♂ U. aenea wing of short-haired form." figureDoi="http://doi.org/10.5281/zenodo.6376803" httpUri="https://zenodo.org/record/6376803/files/figure.png" pageId="9" pageNumber="10">Figure 4</figureCitation>
). Epandrium longer, about as 1.5 times longer than wide, curving tip of aedeagus capped with a triangular, membranous structure. 
<collectingCountry id="59D6766EFFDCFF99FE16FC37FE1B0204" box="[450,493,1021,1042]" name="Italy" pageId="9" pageNumber="10">Italy</collectingCountry>
................................................. ..♂ 
<taxonomicName id="E6C14D7DFFDCFF99FBB3FC37FB1B0204" ID-CoL="8FL5Q" authority="Rossi" authorityName="Rossi" box="[1127,1261,1021,1043]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="13B5EAECFFDCFF99FBB3FC37FB540205" bold="true" box="[1127,1186,1021,1043]" italics="true" pageId="9" pageNumber="10">aenea</emphasis>
(Rossi)
</taxonomicName>
short-haired form
</keyLead>
</paragraph>
</keyStep>
<caption id="75BE6676FFDCFF99FF43F923FD5700E9" ID-DOI="http://doi.org/10.5281/zenodo.6376803" ID-Zenodo-Dep="6376803" box="[151,673,1769,1791]" httpUri="https://zenodo.org/record/6376803/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1769,1791]" targetBox="[358,1228,1092,1746]" targetPageId="9">
<paragraph id="217E36FEFFDCFF99FF43F923FD5700E9" blockId="9.[151,673,1769,1791]" box="[151,673,1769,1791]" pageId="9" pageNumber="10">
<emphasis id="13B5EAECFFDCFF99FF43F923FEE400E9" bold="true" box="[151,274,1769,1791]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDCFF99FEE6F920FE7800E9" ID-CoL="8FL5Q" box="[306,398,1770,1791]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="13B5EAECFFDCFF99FEE6F920FE7800E9" box="[306,398,1770,1791]" italics="true" pageId="9" pageNumber="10">U. aenea</emphasis>
</taxonomicName>
wing of short-haired form.
</paragraph>
</caption>
<keyStep id="9A352E5BFFDCFF99FF43F8EEFA6A0141" pageId="9" pageNumber="10">
<paragraph id="217E36FEFFDCFF99FF43F8EEFA6D012F" blockId="9.[151,1436,1828,1879]" box="[151,1435,1828,1849]" pageId="9" pageNumber="10">
<keyLead id="9A3095CBFFDCFF99FF43F8EEFA6D012F" box="[151,1435,1828,1849]" pageId="9" pageNumber="10">
10. Anal lobe very narrow, linear, clearly narrower then anal cell (
<figureCitation id="B9FA2A7BFFDCFF99FCE1F8EEFC71012F" box="[821,903,1828,1849]" captionStart="FIGURE 5" captionStartId="10.[151,250,869,891]" captionTargetId="figure@10.[358,1228,193,848]" captionTargetPageId="10" captionText="FIGURE 5. ♂ U. unicolor wing." figureDoi="http://doi.org/10.5281/zenodo.6376806" httpUri="https://zenodo.org/record/6376806/files/figure.png" pageId="9" pageNumber="10">Figure 5</figureCitation>
).........................................11
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDCFF99FF43F888FA6A0141" blockId="9.[151,1436,1828,1879]" box="[151,1436,1858,1879]" pageId="9" pageNumber="10">
<keyLead id="9A3095CBFFDCFF99FF43F888FA6A0141" box="[151,1436,1858,1879]" pageId="9" pageNumber="10">- Anal lobe broad with a convex margin, as broad as or broader than anal cell......................................12</keyLead>
</paragraph>
</keyStep>
<caption id="75BE6676FFDFFF9AFF43FCAFFE11056D" ID-DOI="http://doi.org/10.5281/zenodo.6376806" ID-Zenodo-Dep="6376806" box="[151,487,869,891]" httpUri="https://zenodo.org/record/6376806/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,869,891]" targetBox="[358,1227,193,847]" targetPageId="10">
<paragraph id="217E36FEFFDFFF9AFF43FCAFFE11056D" blockId="10.[151,487,869,891]" box="[151,487,869,891]" pageId="10" pageNumber="11">
<emphasis id="13B5EAECFFDFFF9AFF43FCAFFEE4056D" bold="true" box="[151,274,869,891]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDFFF9AFEE6FCACFE51056D" ID-CoL="8FL6V" box="[306,423,869,891]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="13B5EAECFFDFFF9AFEE6FCACFE51056D" box="[306,423,869,891]" italics="true" pageId="10" pageNumber="11">U. unicolor</emphasis>
</taxonomicName>
wing.
</paragraph>
</caption>
<keyStep id="9A352E5BFFDFFF9AFF43FC6BFA6D01BD" pageId="10" pageNumber="11">
<paragraph id="217E36FEFFDFFF9AFF43FC6BFA6A0218" blockId="10.[151,1436,929,1038]" pageId="10" pageNumber="11">
<keyLead id="9A3095CBFFDFFF9AFF43FC6BFA6A0218" pageId="10" pageNumber="11">
11. Frons narrow, lateral dust spots separated anteriorly, coalescing across narrow part. Mesonotal vestiture relatively dense, neat and short, hairs just forward of middle of disc proclinate, shorter than those on upper occiput. Anterior slope of mesonotum undusted except on post pronotal lobe. Wing faintly infuscated fading into yellower base (
<figureCitation id="B9FA2A7BFFDFFF9AFB86FC16FB5E05E7" box="[1106,1192,988,1009]" captionStart="FIGURE 6" captionStartId="10.[151,250,1764,1786]" captionTargetId="figure@10.[358,1228,1087,1742]" captionTargetPageId="10" captionText="FIGURE 6. ♂ U. greatheadi head and mesonotum dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376808" httpUri="https://zenodo.org/record/6376808/files/figure.png" pageId="10" pageNumber="11">Figure 6</figureCitation>
). Shape of gonostyli and apec of epandrium characteristic....................................................♂ 
<taxonomicName id="E6C14D7DFFDFFF9AFB75FC32FABF0218" ID-CoL="8FL82" authority="Gibbs" authorityName="Gibbs" box="[1185,1353,1016,1038]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="greatheadi" status="sp. nov.">
<emphasis id="13B5EAECFFDFFF9AFB75FC32FAFC0218" bold="true" box="[1185,1290,1016,1038]" italics="true" pageId="10" pageNumber="11">greatheadi</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFDFFF9AFA9BFC32FA6A0218" box="[1359,1436,1016,1038]" pageId="10" pageNumber="11" rank="species">
<emphasis id="13B5EAECFFDFFF9AFA9BFC32FA6A0218" bold="true" box="[1359,1436,1016,1038]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<caption id="75BE6676FFDFFF9AFF43F92EFD1100EC" ID-DOI="http://doi.org/10.5281/zenodo.6376808" ID-Zenodo-Dep="6376808" box="[151,743,1764,1786]" httpUri="https://zenodo.org/record/6376808/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1764,1786]" targetBox="[358,1228,1087,1741]" targetPageId="10">
<paragraph id="217E36FEFFDFFF9AFF43F92EFD1100EC" blockId="10.[151,743,1764,1786]" box="[151,743,1764,1786]" pageId="10" pageNumber="11">
<emphasis id="13B5EAECFFDFFF9AFF43F92EFEE400EC" bold="true" box="[151,274,1764,1786]" pageId="10" pageNumber="11">FIGURE 6.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDFFF9AFEE6F92FFE4800EC" ID-CoL="8FL82" box="[306,446,1764,1786]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="13B5EAECFFDFFF9AFEE6F92FFE4800EC" box="[306,446,1764,1786]" italics="true" pageId="10" pageNumber="11">U. greatheadi</emphasis>
</taxonomicName>
head and mesonotum dorsal.
</paragraph>
</caption>
<paragraph id="217E36FEFFDFFF9AFF43F8F4FA6D01BD" blockId="10.[151,1436,1854,1964]" pageId="10" pageNumber="11">
<keyLead id="9A3095CBFFDFFF9AFF43F8F4FA6D01BD" pageId="10" pageNumber="11">
- Frons broader, lateral dust spots broader and coalescing immediately above antennae. Mesonotal vestiture much longer more sparsely haired, hairs just forward of middle of disc erect to slightly reclinate, longer than those on upper occiput. Anterior slope of mesonotum conspicuously dusted between paramedian lines and post pronotal lobes, undusted centrally (
<figureCitation id="B9FA2A7BFFDFFF9AFAEEF8B3FA660198" box="[1338,1424,1913,1934]" captionStart="FIGURE 7" captionStartId="11.[151,250,869,891]" captionTargetId="figure@11.[358,1228,193,848]" captionTargetPageId="11" captionText="FIGURE 7. ♂ U. unicolor head and mesonotum dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376810" httpUri="https://zenodo.org/record/6376810/files/figure.png" pageId="10" pageNumber="11">Figure 7</figureCitation>
). Wing clear. Genitalia characteristic...........................................................♂ 
<taxonomicName id="E6C14D7DFFDFFF9AFADFF85CFA6D01BD" ID-CoL="8FL6V" authority="Loew" authorityName="Loew" box="[1291,1435,1942,1964]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="13B5EAECFFDFFF9AFADFF85CFAA901BA" bold="true" box="[1291,1375,1942,1964]" italics="true" pageId="10" pageNumber="11">unicolor</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="75BE6676FFDEFF9BFF43FCAFFD26056D" ID-DOI="http://doi.org/10.5281/zenodo.6376810" ID-Zenodo-Dep="6376810" box="[151,720,869,891]" httpUri="https://zenodo.org/record/6376810/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,869,891]" targetBox="[358,1227,193,846]" targetPageId="11">
<paragraph id="217E36FEFFDEFF9BFF43FCAFFD26056D" blockId="11.[151,720,869,891]" box="[151,720,869,891]" pageId="11" pageNumber="12">
<emphasis id="13B5EAECFFDEFF9BFF43FCAFFEE4056D" bold="true" box="[151,274,869,891]" pageId="11" pageNumber="12">FIGURE 7.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDEFF9BFEE6FCACFE51056D" ID-CoL="8FL6V" box="[306,423,869,891]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="13B5EAECFFDEFF9BFEE6FCACFE51056D" box="[306,423,869,891]" italics="true" pageId="11" pageNumber="12">U. unicolor</emphasis>
</taxonomicName>
head and mesonotum dorsal.
</paragraph>
</caption>
<keyStep id="9A352E5BFFDEFF9BFF43FC68FA6D0121" pageId="11" pageNumber="12">
<paragraph id="217E36FEFFDEFF9BFF43FC68FA6D05E4" blockId="11.[151,1436,930,1010]" pageId="11" pageNumber="12">
<keyLead id="9A3095CBFFDEFF9BFF43FC68FA6D05E4" pageId="11" pageNumber="12">
12. Hairs around margin of scutellum shorter than length of scutellum; hairs on disc of mesonotum shorter than width of frons (
<figureCitation id="B9FA2A7BFFDEFF9BFF33FC75FECD05C2" box="[231,315,959,980]" captionStart="FIGURE 8" captionStartId="11.[151,250,1736,1758]" captionTargetId="figure@11.[358,1228,1060,1715]" captionTargetPageId="11" captionText="FIGURE 8. ♂ U. lata head and mesonotum dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376812" httpUri="https://zenodo.org/record/6376812/files/figure.png" pageId="11" pageNumber="12">Figure 8</figureCitation>
); wings more or less infuscated, most intense basally along leading edge; frons between grey dust spots smooth and shining. Genitalia relatively small, characteristic....................................................♂ 
<taxonomicName id="E6C14D7DFFDEFF9BFAEEFC16FA6D05E4" ID-CoL="8FL6G" authority="Loew" authorityName="Loew" box="[1338,1435,988,1010]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="13B5EAECFFDEFF9BFAEEFC16FAA905E4" bold="true" box="[1338,1375,988,1010]" italics="true" pageId="11" pageNumber="12">lata</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<caption id="75BE6676FFDEFF9BFF43F902FD5400C8" ID-DOI="http://doi.org/10.5281/zenodo.6376812" ID-Zenodo-Dep="6376812" box="[151,674,1736,1758]" httpUri="https://zenodo.org/record/6376812/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1736,1758]" targetBox="[358,1227,1060,1714]" targetPageId="11">
<paragraph id="217E36FEFFDEFF9BFF43F902FD5400C8" blockId="11.[151,674,1736,1758]" box="[151,674,1736,1758]" pageId="11" pageNumber="12">
<emphasis id="13B5EAECFFDEFF9BFF43F902FEE400C8" bold="true" box="[151,274,1736,1758]" pageId="11" pageNumber="12">FIGURE 8.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDEFF9BFEE6F903FE8F00C8" ID-CoL="8FL6G" box="[306,377,1736,1758]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="13B5EAECFFDEFF9BFEE6F903FE8F00C8" box="[306,377,1736,1758]" italics="true" pageId="11" pageNumber="12">U. lata</emphasis>
</taxonomicName>
head and mesonotum dorsal.
</paragraph>
</caption>
<paragraph id="217E36FEFFDEFF9BFF43F8CFFA6D0121" blockId="11.[151,1436,1797,2024]" pageId="11" pageNumber="12">
<keyLead id="9A3095CBFFDEFF9BFF43F8CFFA6D0121" pageId="11" pageNumber="12">- Hairs around margin of scutellum longer than length of scutellum; hairs on disc of mesonotum longer than width of frons; wings clear, no more than yellowish; frons between grey dust spots wrinkled or matt. Genitalia relatively larger..........13</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDEFF9BFF43F8F5FA6D01BA" pageId="11" pageNumber="12">
<paragraph id="217E36FEFFDEFF9BFF43F8F5FA6A0164" blockId="11.[151,1436,1797,2024]" pageId="11" pageNumber="12">
<keyLead id="9A3095CBFFDEFF9BFF43F8F5FA6A0164" pageId="11" pageNumber="12">
13. Larger species relatively densely covered in rather shaggy, yellowish hairs. Cuticle of mesonotum shining black. Genitalia characteristic.............................................................................♂ 
<taxonomicName id="E6C14D7DFFDEFF9BFAC2F896FA6A0164" ID-CoL="8FQ4W" authority="Strobl" authorityName="Strobl" box="[1302,1436,1884,1906]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pubera">
<emphasis id="13B5EAECFFDEFF9BFAC2F896FAAD0164" bold="true" box="[1302,1371,1884,1906]" italics="true" pageId="11" pageNumber="12">pubera</emphasis>
Strobl
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDEFF9BFF43F8B0FA6D01BA" blockId="11.[151,1436,1797,2024]" pageId="11" pageNumber="12">
<keyLead id="9A3095CBFFDEFF9BFF43F8B0FA6D01BA" pageId="11" pageNumber="12">
- Smaller species less densely covered in rather fine, whitish hairs. Cuticle of mesonotum shagreened, matt. Genitalia characteristic..............................................................................♂ 
<taxonomicName id="E6C14D7DFFDEFF9BFB18F85DFA6D01BA" ID-CoL="8FL6D" authority="Paramonov" authorityName="Paramonov" box="[1228,1435,1943,1965]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="13B5EAECFFDEFF9BFB18F85DFADF01BB" bold="true" box="[1228,1321,1943,1965]" italics="true" pageId="11" pageNumber="12">incognita</emphasis>
Paramonov
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDEFF9CFF43F87FFA6A05A0" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<paragraph id="217E36FEFFDEFF9BFF43F87FFA6A01F1" blockId="11.[151,1436,1797,2024]" pageId="11" pageNumber="12">
<keyLead id="9A3095CBFFDEFF9BFF43F87FFA6A01F1" pageId="11" pageNumber="12">
14. Frons at narrowest point no wider than diameter of anterior ocellus; anterior part of frons entirely grey dusted obscuring ground colour (
<figureCitation id="B9FA2A7BFFDEFF9BFEA0F818FE3101F1" box="[372,455,2002,2023]" captionStart="FIGURE 9" captionStartId="12.[151,250,869,891]" captionTargetId="figure@12.[358,1228,193,848]" captionTargetPageId="12" captionText="FIGURE 9. ♂ U. angustifrons head and mesonotum dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376814" httpUri="https://zenodo.org/record/6376814/files/figure.png" pageId="11" pageNumber="12">Figure 9</figureCitation>
)............................................................. ..♂ 
<taxonomicName id="E6C14D7DFFDEFF9BFB00F818FA6A01F1" ID-CoL="8FL5V" authority="Becker" authorityName="Becker" box="[1236,1436,2002,2024]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFDEFF9BFB00F818FAA701FE" bold="true" box="[1236,1361,2002,2024]" italics="true" pageId="11" pageNumber="12">angustifrons</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
<caption id="75BE6676FFD9FF9CFF43FCAFFD0E056D" ID-DOI="http://doi.org/10.5281/zenodo.6376814" ID-Zenodo-Dep="6376814" box="[151,760,869,891]" httpUri="https://zenodo.org/record/6376814/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,869,891]" targetBox="[358,1227,193,847]" targetPageId="12">
<paragraph id="217E36FEFFD9FF9CFF43FCAFFD0E056D" blockId="12.[151,760,869,891]" box="[151,760,869,891]" pageId="12" pageNumber="13">
<emphasis id="13B5EAECFFD9FF9CFF43FCAFFEE4056D" bold="true" box="[151,274,869,891]" pageId="12" pageNumber="13">FIGURE 9.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFD9FF9CFEE6FCACFE39056D" ID-CoL="8FL5V" box="[306,463,869,891]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFD9FF9CFEE6FCACFE39056D" box="[306,463,869,891]" italics="true" pageId="12" pageNumber="13">U. angustifrons</emphasis>
</taxonomicName>
head and mesonotum dorsal.
</paragraph>
</caption>
<paragraph id="217E36FEFFD9FF9CFF43FC6BFA6A05A0" blockId="12.[151,1436,929,1214]" box="[151,1436,929,950]" pageId="12" pageNumber="13">
<keyLead id="9A3095CBFFD9FF9CFF43FC6BFA6A05A0" box="[151,1436,929,950]" pageId="12" pageNumber="13">- Frons more widely separated; at least small triangular area above antennal bases undusted, ground colour clearly visible...15</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFD9FF9CFF43FC74FA6D05E7" pageId="12" pageNumber="13">
<paragraph id="217E36FEFFD9FF9CFF43FC74FA6D05C5" blockId="12.[151,1436,929,1214]" box="[151,1435,958,980]" pageId="12" pageNumber="13">
<keyLead id="9A3095CBFFD9FF9CFF43FC74FA6D05C5" box="[151,1435,958,980]" pageId="12" pageNumber="13">
15. Lower half of frons entirely yellow in ground colour (see 
<figureCitation id="B9FA2A7BFFD9FF9CFCDEFC74FC9105C5" box="[778,871,958,979]" captionStart="FIGURE 15" captionStartId="15.[151,250,1617,1639]" captionTargetId="figure@15.[358,1228,941,1596]" captionTargetPageId="15" captionText="FIGURE 15. ♀ U. notata frons dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376827" httpUri="https://zenodo.org/record/6376827/files/figure.png" pageId="12" pageNumber="13">Figure 15</figureCitation>
)..................................♂ 
<taxonomicName id="E6C14D7DFFD9FF9CFAF5FC74FA6D05C5" ID-CoL="8FL6L" authority="Loew" authorityName="Loew" box="[1313,1435,958,980]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="13B5EAECFFD9FF9CFAF5FC74FAA905C2" bold="true" box="[1313,1375,958,980]" italics="true" pageId="12" pageNumber="13">notata</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFD9FF9CFF43FC16FA6D05E7" blockId="12.[151,1436,929,1214]" box="[151,1435,988,1009]" pageId="12" pageNumber="13">
<keyLead id="9A3095CBFFD9FF9CFF43FC16FA6D05E7" box="[151,1435,988,1009]" pageId="12" pageNumber="13">- Lower half of frons black, concolorous with vertex, at most small yellow spots laterally below dust spots............... 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFD9FF9CFF43FC33FA6A0295" pageId="12" pageNumber="13">
<paragraph id="217E36FEFFD9FF9CFF43FC33FA6A023D" blockId="12.[151,1436,929,1214]" pageId="12" pageNumber="13">
<keyLead id="9A3095CBFFD9FF9CFF43FC33FA6A023D" pageId="12" pageNumber="13">16. Acrosticals_proclinate16.. Most acrostichals proclinate; if a small shining black species from Europe with a few hairs nearest head erect to reclinate then katepisternum dusted and dust spots on frons confluent in front of anterior ocellus...........17</keyLead>
</paragraph>
<paragraph id="217E36FEFFD9FF9CFF43FBFEFA6A0295" blockId="12.[151,1436,929,1214]" pageId="12" pageNumber="13">
<keyLead id="9A3095CBFFD9FF9CFF43FBFEFA6A0295" pageId="12" pageNumber="13">- At least anterior acrostichals reclinate or too short to judge; if a small shining black species from the Middle East with both proclinate and reclinate acrostichals or anterior reclinate acrostichals missing then katepisternum subshining and undusted on ventral half and dust spots on frons well separated...........................................................19</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFD9FF9DFF43FB46FA6A06F1" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<paragraph id="217E36FEFFD9FF9CFF43FB46FA6A02A8" blockId="12.[151,1436,929,1214]" pageId="12" pageNumber="13">
<keyLead id="9A3095CBFFD9FF9CFF43FB46FA6A02A8" pageId="12" pageNumber="13">
17. Hairs_on17. Hairs on disc of mesonotum short, adpressed, orientated inward or forward (
<figureCitation id="B9FA2A7BFFD9FF9CFBE7FB46FB6502B7" box="[1075,1171,1164,1185]" captionStart="FIGURE 10" captionStartId="12.[151,250,1940,1962]" captionTargetId="figure@12.[358,1228,1263,1918]" captionTargetPageId="12" captionText="FIGURE 10. ♂ U. anatoliensis head and mesonotum dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376816" httpUri="https://zenodo.org/record/6376816/files/figure.png" pageId="12" pageNumber="13">Figure 10</figureCitation>
); frons wider, separated by............................................................................. ♂ 
<taxonomicName id="E6C14D7DFFD9FF9CFB47FB62FABF02A8" ID-CoL="8FL72" authority="Gibbs" authorityName="Gibbs" box="[1171,1353,1192,1214]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anatoliensis" status="sp. nov.">
<emphasis id="13B5EAECFFD9FF9CFB47FB62FAFC02A8" bold="true" box="[1171,1290,1192,1214]" italics="true" pageId="12" pageNumber="13">anatoliensis</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFD9FF9CFA9BFB62FA6A02A8" box="[1359,1436,1192,1214]" pageId="12" pageNumber="13" rank="species">
<emphasis id="13B5EAECFFD9FF9CFA9BFB62FA6A02A8" bold="true" box="[1359,1436,1192,1214]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<caption id="75BE6676FFD9FF9CFF43F85EFCF701BC" ID-DOI="http://doi.org/10.5281/zenodo.6376816" ID-Zenodo-Dep="6376816" box="[151,769,1940,1962]" httpUri="https://zenodo.org/record/6376816/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1940,1962]" targetBox="[358,1227,1263,1917]" targetPageId="12">
<paragraph id="217E36FEFFD9FF9CFF43F85EFCF701BC" blockId="12.[151,769,1940,1962]" box="[151,769,1940,1962]" pageId="12" pageNumber="13">
<emphasis id="13B5EAECFFD9FF9CFF43F85EFEE901BC" bold="true" box="[151,287,1940,1962]" pageId="12" pageNumber="13">FIGURE 10.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFD9FF9CFEEBF85FFE2F01BC" ID-CoL="8FL72" box="[319,473,1940,1962]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anatoliensis">
<emphasis id="13B5EAECFFD9FF9CFEEBF85FFE2F01BC" box="[319,473,1940,1962]" italics="true" pageId="12" pageNumber="13">U. anatoliensis</emphasis>
</taxonomicName>
head and mesonotum dorsal.
</paragraph>
</caption>
<paragraph id="217E36FEFFD8FF9DFF43FF5DFA6A06F1" blockId="13.[151,1436,151,349]" pageId="13" pageNumber="14">
<keyLead id="9A3095CBFFD8FF9DFF43FF5DFA6A06F1" pageId="13" pageNumber="14">- Hairs on disc of mesonotum longer, erect, not so clearly orientated inward or forward; frons narrower, separated by about distance across lateral ocelli, grey dust spots on frons contiguous; anal lobe narrow, linear or only slightly convex, narrower then anal cell............................................................................................18</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFD8FF9DFF43FF25FA6A02A9" pageId="13" pageNumber="14">
<paragraph id="217E36FEFFD8FF9DFF43FF25FA6D074A" blockId="13.[151,1436,151,349]" pageId="13" pageNumber="14">
<keyLead id="9A3095CBFFD8FF9DFF43FF25FA6D074A" pageId="13" pageNumber="14">
18. Wing_uniformly18. Wing uniformly brown infuscated, except for yellower base, veins beyond humeral crossvein brown; anal lobe very narrow, linear (
<figureCitation id="B9FA2A7BFFD8FF9DFE0AFEC7FDB60734" box="[478,576,269,290]" captionStart="FIGURE 11" captionStartId="13.[151,250,1074,1096]" captionTargetId="figure@13.[358,1228,398,1053]" captionTargetPageId="13" captionText="FIGURE 11. ♂ U. manca wing." figureDoi="http://doi.org/10.5281/zenodo.6376818" httpUri="https://zenodo.org/record/6376818/files/figure.png" pageId="13" pageNumber="14">Figure 11</figureCitation>
); eyes separated by about distance across lateral ocelli; longer-haired species, anterior acrostichals longer then bare paramedian lines; epandrium with median unsclerotized area not exceeding mid point................................................................................................. ♂ 
<taxonomicName id="E6C14D7DFFD8FF9DFAC8FE8DFA6D074A" ID-CoL="8FL6H" authority="Loew" authorityName="Loew" box="[1308,1435,327,349]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="manca">
<emphasis id="13B5EAECFFD8FF9DFAC8FE8DFAA8074B" bold="true" box="[1308,1374,327,349]" italics="true" pageId="13" pageNumber="14">manca</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<caption id="75BE6676FFD8FF9DFF43FBF8FE16025E" ID-DOI="http://doi.org/10.5281/zenodo.6376818" ID-Zenodo-Dep="6376818" box="[151,480,1074,1096]" httpUri="https://zenodo.org/record/6376818/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1074,1096]" targetBox="[358,1227,398,1052]" targetPageId="13">
<paragraph id="217E36FEFFD8FF9DFF43FBF8FE16025E" blockId="13.[151,480,1074,1096]" box="[151,480,1074,1096]" pageId="13" pageNumber="14">
<emphasis id="13B5EAECFFD8FF9DFF43FBF8FEE8025E" bold="true" box="[151,286,1074,1096]" pageId="13" pageNumber="14">FIGURE 11.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFD8FF9DFEE9FBF9FE56025E" ID-CoL="8FL6H" box="[317,416,1075,1096]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="manca">
<emphasis id="13B5EAECFFD8FF9DFEE9FBF9FE56025E" box="[317,416,1075,1096]" italics="true" pageId="13" pageNumber="14">U. manca</emphasis>
</taxonomicName>
wing.
</paragraph>
</caption>
<subSection id="484E2F15FFD8FF9DFF43FBA5FA6A02A9" pageId="13" pageNumber="14" type="multiple">
<paragraph id="217E36FEFFD8FF9DFF43FBA5FA6A02A9" blockId="13.[151,1436,1135,1216]" pageId="13" pageNumber="14">
<keyLead id="9A3095CBFFD8FF9DFF43FBA5FA6A02A9" pageId="13" pageNumber="14">
i- Wing yellowish, veins beyond humeral crossvein yellow-brown. Anal lobe narrower than anal cell but clearly with convex hind margin (
<figureCitation id="B9FA2A7BFFD8FF9DFEB5FB47FE3402B4" box="[353,450,1165,1186]" captionStart="FIGURE 12" captionStartId="13.[151,250,1941,1963]" captionTargetId="figure@13.[358,1228,1265,1920]" captionTargetPageId="13" captionText="FIGURE 12. ♂ U. putilla wing." figureDoi="http://doi.org/10.5281/zenodo.6376821" httpUri="https://zenodo.org/record/6376821/files/figure.png" pageId="13" pageNumber="14">Figure 12</figureCitation>
). Eyes separated by less than distance across lateral ocelli. Shorter-haired species, anterior acrostichals shorter than bare paramedian lines. Epandrium with median unsclerotized area exceeding mid point....... ♂ 
<taxonomicName id="E6C14D7DFFD8FF9DFAC6FB60FA6A02A9" ID-CoL="8FL6P" authority="Becker" authorityName="Becker" box="[1298,1436,1194,1216]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFD8FF9DFAC6FB60FAA402D6" bold="true" box="[1298,1362,1194,1216]" italics="true" pageId="13" pageNumber="14">putilla</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
</subSection>
</keyStep>
<caption id="75BE6676FFD8FF9DFF43F85FFE1601BD" ID-DOI="http://doi.org/10.5281/zenodo.6376821" ID-Zenodo-Dep="6376821" box="[151,480,1941,1963]" httpUri="https://zenodo.org/record/6376821/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1941,1963]" targetBox="[358,1228,1265,1919]" targetPageId="13">
<paragraph id="217E36FEFFD8FF9DFF43F85FFE1601BD" blockId="13.[151,480,1941,1963]" box="[151,480,1941,1963]" pageId="13" pageNumber="14">
<emphasis id="13B5EAECFFD8FF9DFF43F85FFEE901BD" bold="true" box="[151,287,1941,1963]" pageId="13" pageNumber="14">FIGURE 12.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFD8FF9DFEEBF85CFE5601BD" ID-CoL="8FL6P" box="[319,416,1941,1963]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFD8FF9DFEEBF85CFE5601BD" box="[319,416,1941,1963]" italics="true" pageId="13" pageNumber="14">U. putilla</emphasis>
</taxonomicName>
wing.
</paragraph>
</caption>
<keyStep id="9A352E5BFFDBFF9EFF43FF5DFA6D06F1" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43FF5DFA6A06DC" blockId="14.[151,1436,151,437]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FF5DFA6A06DC" pageId="14" pageNumber="15">19. Marginal_hairs...................................................................................... 19. Marginal hairs of scutellum shorter than half mid-length of scutellum...........................................20</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43FF18FA6D06F1" blockId="14.[151,1436,151,437]" box="[151,1435,210,231]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FF18FA6D06F1" box="[151,1435,210,231]" pageId="14" pageNumber="15">- Marginal hairs of scutellum longer than length of scutellum or almost so......................................... 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF43FF25FA6D074A" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43FF25FBC20729" blockId="14.[151,1436,151,437]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FF25FBC20729" pageId="14" pageNumber="15">
20. Occiput_entirely20. Occiput entirely shining black. Genitalia characteristic (Plate II). 
<collectingCountry id="59D6766EFFDBFF9EFBCDFF25FB700712" box="[1049,1158,239,260]" name="Turkey" pageId="14" pageNumber="15">Asia Minor</collectingCountry>
......................................................................................................♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFB7EFEC6FABF0734" ID-CoL="8FL74" authority="Gibbs" authorityName="Gibbs" box="[1194,1353,268,290]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="annetteae" status="sp. nov.">
<emphasis id="13B5EAECFFDBFF9EFB7EFEC6FAFC0734" bold="true" box="[1194,1290,268,290]" italics="true" pageId="14" pageNumber="15">annetteae</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFDBFF9EFA9BFEC6FA6D0734" box="[1359,1435,268,290]" pageId="14" pageNumber="15" rank="species">
<emphasis id="13B5EAECFFDBFF9EFA9BFEC6FA6D0734" bold="true" box="[1359,1435,268,290]" pageId="14" pageNumber="15">sp. nov.</emphasis>
</taxonomicNameLabel>
[
<taxonomicName id="E6C14D7DFFDBFF9EFF3DFEE0FE810729" ID-CoL="8FL5Z" authority="Loew" authorityName="Loew" box="[233,375,297,319]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="calva">
<emphasis id="13B5EAECFFDBFF9EFF3DFEE0FECC0729" box="[233,314,297,319]" italics="true" pageId="14" pageNumber="15">U. calva</emphasis>
Loew
</taxonomicName>
, with very short mesonotal hairs, will probably run here but ♂ unknown]
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43FE8DFA6D074A" blockId="14.[151,1436,151,437]" box="[151,1435,327,348]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FE8DFA6D074A" box="[151,1435,327,348]" pageId="14" pageNumber="15">- Occiput not entirely shining black, at least some thin dusting below around mouth cavity. Africa...................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF49FEAFFA6D02EC" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF49FEAFFA6A07A3" blockId="14.[151,1436,151,437]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF49FEAFFA6A07A3" pageId="14" pageNumber="15">
21. Epandrium with, truncate, slightly apically expanded, apicolateral elongations that are longer than their apical width, usually quite visible in pinned specimens (
<figureCitation id="B9FA2A7BFFDBFF9EFDF9FE48FD7B0781" box="[557,653,386,407]" captionStart="FIGURE 13" captionStartId="14.[151,250,1162,1184]" captionTargetId="figure@14.[358,1228,486,1141]" captionTargetPageId="14" captionText="FIGURE 13. ♂ U. cornigera gonocoxite lateral view." figureDoi="http://doi.org/10.5281/zenodo.6376823" httpUri="https://zenodo.org/record/6376823/files/figure.png" pageId="14" pageNumber="15">Figure 13</figureCitation>
). Gonostyli with long, curved process on inner face basally................................................................................................♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFB7EFE55FABF07A2" ID-CoL="8FL7C" authority="Gibbs" authorityName="Gibbs" box="[1194,1353,415,437]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cornigera" status="sp. nov.">
<emphasis id="13B5EAECFFDBFF9EFB7EFE55FAFC07A3" bold="true" box="[1194,1290,415,437]" italics="true" pageId="14" pageNumber="15">cornigera</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFDBFF9EFA9BFE55FA6A07A3" box="[1359,1436,415,437]" pageId="14" pageNumber="15" rank="species">
<emphasis id="13B5EAECFFDBFF9EFA9BFE55FA6A07A3" bold="true" box="[1359,1436,415,437]" pageId="14" pageNumber="15">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<caption id="75BE6676FFDBFF9EFF43FB40FD4D02B6" ID-DOI="http://doi.org/10.5281/zenodo.6376823" ID-Zenodo-Dep="6376823" box="[151,699,1162,1184]" httpUri="https://zenodo.org/record/6376823/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1162,1184]" targetBox="[358,1228,486,1140]" targetPageId="14">
<paragraph id="217E36FEFFDBFF9EFF43FB40FD4D02B6" blockId="14.[151,699,1162,1184]" box="[151,699,1162,1184]" pageId="14" pageNumber="15">
<emphasis id="13B5EAECFFDBFF9EFF43FB40FEE902B6" bold="true" box="[151,287,1162,1184]" pageId="14" pageNumber="15">FIGURE 13.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFEEBFB41FE3402B6" ID-CoL="8FL7C" box="[319,450,1163,1184]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cornigera">
<emphasis id="13B5EAECFFDBFF9EFEEBFB41FE3402B6" box="[319,450,1163,1184]" italics="true" pageId="14" pageNumber="15">U. cornigera</emphasis>
</taxonomicName>
gonocoxite lateral view.
</paragraph>
</caption>
<paragraph id="217E36FEFFDBFF9EFF43FB0DFA6D02EC" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FB0DFA6D02EC" pageId="14" pageNumber="15">- Any apicolateral extensions of epandrium not or hardly longer than apical width, gonostyli lacking long, curved basal process...................................................................................................22</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF43FAC8FA6D03BC" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43FAC8FA6A0344" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FAC8FA6A0344" pageId="14" pageNumber="15">
22. Very_short-haired22. Very short-haired species, hairs on ocellar tubercle and mesonotum shorter than distance between lateral ocelli. Epandrium much longer than broad, protuberant below tip of abdomen, usually enclosing gonocoxite and aedeagus. Genitalia characteristic.....................................................................♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFAC6FAF6FA6A0344" ID-CoL="8FL5P" authority="Becker" authorityName="Becker" box="[1298,1436,1340,1362]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="accola">
<emphasis id="13B5EAECFFDBFF9EFAC6FAF6FAA40344" bold="true" box="[1298,1362,1340,1362]" italics="true" pageId="14" pageNumber="15">accola</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43FA90FA6D03BC" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FA90FA6D03BC" pageId="14" pageNumber="15">
- Longer-haired species, hairs on ocellar tubercle usually and those on mesonotum longer than distance between lateral ocelli. Epandrium much broader than long, almost as wide as abdomen at that point, hooked tip of aedeagus protruding beyond apex of epandrium and gonocoxite, genitalia characteristic..........................................♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFB26FA5EFA6D03BC" ID-CoL="8FL69" authority="Fabricius" authorityName="Fabricius" box="[1266,1435,1428,1450]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="florea">
<emphasis id="13B5EAECFFDBFF9EFB26FA5EFADB03BC" bold="true" box="[1266,1325,1428,1450]" italics="true" pageId="14" pageNumber="15">florea</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF43FA78FA6D0009" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43FA78FA6A03F2" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FA78FA6A03F2" pageId="14" pageNumber="15">
23. Genitalia_very23... Genitalia very large, as large as remainder of abdomen. Very long-haired species, abdomen laterally with long, outstanding foxy vestiture. Gonocoxite and aedeagus characteristic............................♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFB28FA05FA6A03F2" ID-CoL="8FL6X" authority="Macquart" authorityName="Macquart" box="[1276,1436,1487,1509]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="vestita">
<emphasis id="13B5EAECFFDBFF9EFB28FA05FACC03F3" bold="true" box="[1276,1338,1487,1509]" italics="true" pageId="14" pageNumber="15">vestita</emphasis>
Macquart
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43FA27FA6D0009" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43FA27FA6D0009" pageId="14" pageNumber="15">- Genitalia smaller, not as large as remainder of abdomen. Shorter-haired species, abdomen with shaggy yellowish-white hair. Gonocoxite and aedeagus different....................................................................... 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF43F9EDFA6D0082" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43F9EDFA6A004C" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F9EDFA6A004C" pageId="14" pageNumber="15">
24. Wing_membrane24.Wing membrane clear, faintly yellow tinged basally, veins dark brown, subcosta and base of stem of R yellow. Genitalia with yellow areas, tip of gonocoxite with brush of golden hairs (
<figureCitation id="B9FA2A7BFFDBFF9EFC00F98FFBC5004C" box="[980,1075,1605,1626]" captionStart="FIGURE 14" captionStartId="15.[151,250,869,891]" captionTargetId="figure@15.[358,1228,193,848]" captionTargetPageId="15" captionText="FIGURE 14. ♂ U. maghrebensis gonocoxite lateral view." figureDoi="http://doi.org/10.5281/zenodo.6376825" httpUri="https://zenodo.org/record/6376825/files/figure.png" pageId="14" pageNumber="15">Figure 14</figureCitation>
)... ♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFB55F98EFABF004C" ID-CoL="8FL8D" authority="Gibbs" authorityName="Gibbs" box="[1153,1353,1604,1626]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maghrebensis" status="sp. nov.">
<emphasis id="13B5EAECFFDBFF9EFB55F98EFAFC004C" bold="true" box="[1153,1290,1604,1626]" italics="true" pageId="14" pageNumber="15">maghrebensis</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFDBFF9EFA9BF98EFA6A004C" box="[1359,1436,1604,1626]" pageId="14" pageNumber="15" rank="species">
<emphasis id="13B5EAECFFDBFF9EFA9BF98EFA6A004C" bold="true" box="[1359,1436,1604,1626]" pageId="14" pageNumber="15">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43F9A8FA6D0082" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F9A8FA6D0082" pageId="14" pageNumber="15">
- Wing membrane basally infuscated, veins brown, subcosta and base of stem of R blackish. Genitalia entirely black, tip of gonocoxite lacking differentiated hairs......................................................♂ 
<taxonomicName id="E6C14D7DFFDBFF9EFB26F9B5FA6D0082" ID-CoL="8FL5X" authority="Fabricius" authorityName="Fabricius" box="[1266,1435,1663,1685]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="13B5EAECFFDBFF9EFB26F9B5FADA0083" bold="true" box="[1266,1324,1663,1685]" italics="true" pageId="14" pageNumber="15">atrata</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF43F957FA6D00D9" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43F957FA6D00A4" blockId="14.[151,1437,1223,2037]" box="[151,1435,1693,1714]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F957FA6D00A4" box="[151,1435,1693,1714]" pageId="14" pageNumber="15">25. At least knees contrastingly yellow.......................................................................26</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43F970FA6D00D9" blockId="14.[151,1437,1223,2037]" box="[151,1435,1722,1743]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F970FA6D00D9" box="[151,1435,1722,1743]" pageId="14" pageNumber="15">- Legs entirely black...................................................................................27</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF43F91DFA6D0152" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43F91DFA6D011C" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F91DFA6D011C" pageId="14" pageNumber="15">
26. Lower half of frons and oral margin yellow, strongly contrasting with black hind part (
<figureCitation id="B9FA2A7BFFDBFF9EFBC2F91DFB8300FA" box="[1046,1141,1751,1772]" captionStart="FIGURE 15" captionStartId="15.[151,250,1617,1639]" captionTargetId="figure@15.[358,1228,941,1596]" captionTargetPageId="15" captionText="FIGURE 15. ♀ U. notata frons dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376827" httpUri="https://zenodo.org/record/6376827/files/figure.png" pageId="14" pageNumber="15">Figure 15</figureCitation>
). Legs black other than yellow knees, tergites all dark, sternites all yellow.......................................................♀ 
<taxonomicName id="E6C14D7DFFDBFF9EFAF5F93EFA6D011C" ID-CoL="8FL6L" authority="Loew" authorityName="Loew" box="[1313,1435,1780,1802]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="13B5EAECFFDBFF9EFAF5F93EFAA9011C" bold="true" box="[1313,1375,1780,1802]" italics="true" pageId="14" pageNumber="15">notata</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43F8D8FA6D0152" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F8D8FA6D0152" pageId="14" pageNumber="15">
- Frons and oral margin black in ground colour. Tibia and basal tarsal segments yellow, tergites with yellow apical margins, sternites partly dark.................................................................. ♀ 
<taxonomicName id="E6C14D7DFFDBFF9EFB78F8E5FA6D0152" ID-CoL="8FL6T" authority="Paramonov" authorityName="Paramonov" box="[1196,1435,1839,1861]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="13B5EAECFFDBFF9EFB78F8E5FADE0153" bold="true" box="[1196,1320,1839,1861]" italics="true" pageId="14" pageNumber="15">transcaspica</emphasis>
Paramonov
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDBFF9EFF43F887FE9501E2" pageId="14" pageNumber="15">
<paragraph id="217E36FEFFDBFF9EFF43F887FA6D0169" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F887FA6D0169" pageId="14" pageNumber="15">27. Occiput dusted such that ground colour essentially obscured, triangle behind vertex often more thinly dusted so subshining..................................................................................................... 28</keyLead>
</paragraph>
<paragraph id="217E36FEFFDBFF9EFF43F84DFE9501E2" blockId="14.[151,1437,1223,2037]" pageId="14" pageNumber="15">
<keyLead id="9A3095CBFFDBFF9EFF43F84DFE9501E2" pageId="14" pageNumber="15">- Occiput either entirely undusted and shining, or with strip behind eyes shining and undusted or very thinly dusted so shining black cuticle clearly evident............................................................................ 37 [From here on many characters are prone to variation so any specimens that fail to fit description and distribution well must be dissected.]</keyLead>
</paragraph>
</keyStep>
<caption id="75BE6676FFDAFF9FFF43FCAFFD15056D" ID-DOI="http://doi.org/10.5281/zenodo.6376825" ID-Zenodo-Dep="6376825" box="[151,739,869,891]" httpUri="https://zenodo.org/record/6376825/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,869,891]" targetBox="[358,1227,193,847]" targetPageId="15">
<paragraph id="217E36FEFFDAFF9FFF43FCAFFD15056D" blockId="15.[151,739,869,891]" box="[151,739,869,891]" pageId="15" pageNumber="16">
<emphasis id="13B5EAECFFDAFF9FFF43FCAFFEE9056D" bold="true" box="[151,287,869,891]" pageId="15" pageNumber="16">FIGURE 14.</emphasis>
♂ 
<taxonomicName id="E6C14D7DFFDAFF9FFEEBFCACFE1C056D" ID-CoL="8FL8D" box="[319,490,869,891]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="maghrebensis">
<emphasis id="13B5EAECFFDAFF9FFEEBFCACFE1C056D" box="[319,490,869,891]" italics="true" pageId="15" pageNumber="16">U. maghrebensis</emphasis>
</taxonomicName>
gonocoxite lateral view.
</paragraph>
</caption>
<caption id="75BE6676FFDAFF9FFF43F99BFDD20071" ID-DOI="http://doi.org/10.5281/zenodo.6376827" ID-Zenodo-Dep="6376827" box="[151,548,1617,1639]" httpUri="https://zenodo.org/record/6376827/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1617,1639]" targetBox="[358,1227,941,1595]" targetPageId="15">
<paragraph id="217E36FEFFDAFF9FFF43F99BFDD20071" blockId="15.[151,548,1617,1639]" box="[151,548,1617,1639]" pageId="15" pageNumber="16">
<emphasis id="13B5EAECFFDAFF9FFF43F99BFEE90071" bold="true" box="[151,287,1617,1639]" pageId="15" pageNumber="16">FIGURE 15.</emphasis>
♀ 
<taxonomicName id="E6C14D7DFFDAFF9FFEEBF998FE690071" ID-CoL="8FL6L" box="[319,415,1618,1639]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="notata">
<emphasis id="13B5EAECFFDAFF9FFEEBF998FE690071" box="[319,415,1618,1639]" italics="true" pageId="15" pageNumber="16">U. notata</emphasis>
</taxonomicName>
frons dorsal.
</paragraph>
</caption>
<keyStep id="9A352E5BFFDAFF9FFF43F944FA6D010F" pageId="15" pageNumber="16">
<paragraph id="217E36FEFFDAFF9FFF43F944FA6D00C8" blockId="15.[151,1436,1678,2022]" pageId="15" pageNumber="16">
<keyLead id="9A3095CBFFDAFF9FFF43F944FA6D00C8" pageId="15" pageNumber="16">
28. Wing with well demarcated pattern, basally yellowish, from level of humeral crossvein to approximately level with crossvein r-m dark brown, beyond which wing membrane is clear, this pattern often best seen with the naked eye (e.g. 
<figureCitation id="B9FA2A7BFFDAFF9FFB24F966FAC900D7" box="[1264,1343,1708,1729]" captionStart="FIGURE 3" captionStartId="9.[151,250,869,891]" captionTargetId="figure@9.[358,1228,193,848]" captionTargetPageId="9" captionText="FIGURE 3. ♂ U. aenea wing of long-haired form." figureDoi="http://doi.org/10.5281/zenodo.6376801" httpUri="https://zenodo.org/record/6376801/files/figure.png" pageId="15" pageNumber="16">Figure 3</figureCitation>
)(in some small specimens the wing pattern can be very poorly developed and only dissection will permit identification)........... 29
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDAFF9FFF43F92CFA6D010F" blockId="15.[151,1436,1678,2022]" pageId="15" pageNumber="16">
<keyLead id="9A3095CBFFDAFF9FFF43F92CFA6D010F" pageId="15" pageNumber="16">
- Wing either clear, or if infuscated AND with a yellowish base (
<taxonomicName id="E6C14D7DFFDAFF9FFC8EF92FFC3700ED" ID-CoL="8FL82" box="[858,961,1765,1787]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="greatheadi">
<emphasis id="13B5EAECFFDAFF9FFC8EF92FFC3700ED" box="[858,961,1765,1787]" italics="true" pageId="15" pageNumber="16">greatheadi</emphasis>
</taxonomicName>
), with brown infuscation fading out so no clear demarcation between brown area and clear apical part of wing................................................. 31
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFDAFF9FFF43F8EBFA6D01DF" pageId="15" pageNumber="16">
<paragraph id="217E36FEFFDAFF9FFF43F8EBFA6D0167" blockId="15.[151,1436,1678,2022]" pageId="15" pageNumber="16">
<keyLead id="9A3095CBFFDAFF9FFF43F8EBFA6D0167" pageId="15" pageNumber="16">
29. Mesonotum very short haired, practically naked, those setulae that are present are most readily seen on notopleuron and sides of mesonotum above wing bases and are dark brown to blackish spicules. Similarly setae on ocellar tubercle dark and very short, no longer than diameter of an ocellus............................................. ..♀ 
<taxonomicName id="E6C14D7DFFDAFF9FFB13F891FA6D0167" ID-CoL="8FL5S" authority="Paramonov" authorityName="Paramonov" box="[1223,1435,1883,1905]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="aeneoides">
<emphasis id="13B5EAECFFDAFF9FFB13F891FADE0167" bold="true" box="[1223,1320,1883,1905]" italics="true" pageId="15" pageNumber="16">aeneoides</emphasis>
Paramonov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFDAFF9FFF43F8B3FA6D01DF" blockId="15.[151,1436,1678,2022]" pageId="15" pageNumber="16">
<keyLead id="9A3095CBFFDAFF9FFF43F8B3FA6D01DF" pageId="15" pageNumber="16">- Mesonotum longer haired, hairs, even in rubbed specimens, clearly visible on notopleuron and sides of mesonotum above wing bases and are pale yellowish white. Hairs on ocellar tubercle yellowish white and as long as the distance between lateral ocelli...................................................................................................30</keyLead>
</paragraph>
</keyStep>
<paragraph id="217E36FEFFDAFF80FF43F81BFA6A074A" blockId="15.[151,1436,1678,2022]" lastBlockId="16.[151,1437,151,349]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<keyStep id="9A352E5BFFDAFF80FF43F81BFA6A074A" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<keyLead id="9A3095CBFFDAFF80FF43F81BFA6A074A" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
30. Upper half of frons with series of relatively deep, diagonal grooves either side meeting in middle (
<figureCitation id="B9FA2A7BFFDAFF9FFB5CF81BFB1101F0" box="[1160,1255,2001,2022]" captionStart="FIGURE 16" captionStartId="16.[151,250,1116,1138]" captionTargetBox="[181,1405,397,1095]" captionTargetId="figure@16.[181,1405,397,1095]" captionTargetPageId="16" captionText="FIGURE 16. ♀ U. similis (left) &amp; U. aenea (right) frons dorsally." httpUri="https://zenodo.org/record/286091/files/figure.png" pageId="15" pageNumber="16">Figure 16</figureCitation>
). Occiput immediately behind vertex dusted, only slightly more thinly so than rest of occiput. Disc of mesonotum with strongly stellate hair insertions which in places coalesce to form a reticulate surface sculpture. Oral margin broader than apical diameter of palp. Apical sternite transverse, furca characteristic...............................................♀ 
<taxonomicName id="E6C14D7DFFC5FF80FB3DFF18FA6D06F1" ID-CoL="8FL6Q" authority="Paramonov" authorityName="Paramonov" box="[1257,1435,210,232]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="13B5EAECFFC5FF80FB3DFF18FADF06FE" bold="true" box="[1257,1321,210,232]" italics="true" pageId="16" pageNumber="17">similis</emphasis>
Paramonov
</taxonomicName>
- Upper half of frons smooth and shining, no more than shallow wrinkles medially (
<figureCitation id="B9FA2A7BFFC5FF80FC3BFF25FBB90712" box="[1007,1103,239,260]" captionStart="FIGURE 16" captionStartId="16.[151,250,1116,1138]" captionTargetBox="[181,1405,397,1095]" captionTargetId="figure@16.[181,1405,397,1095]" captionTargetPageId="16" captionText="FIGURE 16. ♀ U. similis (left) &amp; U. aenea (right) frons dorsally." httpUri="https://zenodo.org/record/286091/files/figure.png" pageId="16" pageNumber="17">Figure 16</figureCitation>
). Occiput immediately behind vertex with a shining undusted triangular area, often continuing as a black line to neck. Disc of mesonotum with hair insertions, while often stellate, not coalescing to form a reticulate surface sculpture. Oral margin no broader than apical diameter of palp. Apical sternite longer than broad, furca characteristic.............................................♀ 
<taxonomicName id="E6C14D7DFFC5FF80FAC2FE8DFA6A074A" ID-CoL="8FL5Q" authority="Rossi" authorityName="Rossi" box="[1302,1436,327,349]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="13B5EAECFFC5FF80FAC2FE8DFAA7074B" bold="true" box="[1302,1361,327,349]" italics="true" pageId="16" pageNumber="17">aenea</emphasis>
(Rossi)
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<caption id="75BE6676FFC5FF80FF43FB96FCC20264" box="[151,820,1116,1139]" httpUri="https://zenodo.org/record/286091/files/figure.png" pageId="16" pageNumber="17" targetBox="[181,1405,397,1095]" targetPageId="16">
<paragraph id="217E36FEFFC5FF80FF43FB96FCC20264" blockId="16.[151,820,1116,1139]" box="[151,820,1116,1139]" pageId="16" pageNumber="17">
<emphasis id="13B5EAECFFC5FF80FF43FB96FEE90264" bold="true" box="[151,287,1116,1138]" pageId="16" pageNumber="17">FIGURE 16.</emphasis>
♀ 
<taxonomicName id="E6C14D7DFFC5FF80FEEBFB94FE560264" ID-CoL="8FL6Q" box="[319,416,1116,1139]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="13B5EAECFFC5FF80FEEBFB94FE560264" box="[319,416,1116,1139]" italics="true" pageId="16" pageNumber="17">U. similis</emphasis>
</taxonomicName>
(left) &amp; 
<taxonomicName id="E6C14D7DFFC5FF80FE2DFB94FDA20265" ID-CoL="8FL5Q" box="[505,596,1118,1139]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="13B5EAECFFC5FF80FE2DFB94FDA20265" box="[505,596,1118,1139]" italics="true" pageId="16" pageNumber="17">U. aenea</emphasis>
</taxonomicName>
(right) frons dorsally.
</paragraph>
</caption>
<keyStep id="9A352E5BFFC5FF80FF43FB52FA6D02FE" pageId="16" pageNumber="17">
<paragraph id="217E36FEFFC5FF80FF43FB52FA6A02DD" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43FB52FA6A02DD" pageId="16" pageNumber="17">
31. Frons forward of anterior ocellus with a line of long hairs close to eye margins, at least half as long as those on ocellar tubercle and extending to about mid-point of frons (see 
<figureCitation id="B9FA2A7BFFC5FF80FD70FB7CFD0202DD" box="[676,756,1206,1227]" captionStart="FIGURE 1" captionStartId="7.[151,250,2010,2032]" captionTargetId="figure@7.[450,1129,1370,1980]" captionTargetPageId="7" captionText="FIGURE 1. ♂ U. bicolor frons." figureDoi="http://doi.org/10.5281/zenodo.6376797" httpUri="https://zenodo.org/record/6376797/files/figure.png" pageId="16" pageNumber="17">Figure 1</figureCitation>
)......................................................32
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC5FF80FF43FB19FA6D02FE" blockId="16.[151,1436,1176,2019]" box="[151,1435,1235,1256]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43FB19FA6D02FE" box="[151,1435,1235,1256]" pageId="16" pageNumber="17">- Frons forward of anterior ocellus bare or with very short, strongly proclinate, setulae only........................... 33</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC5FF80FF43FB3AFA6A036D" pageId="16" pageNumber="17">
<paragraph id="217E36FEFFC5FF80FF43FB3AFA6A0356" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43FB3AFA6A0356" pageId="16" pageNumber="17">
32. A shorter-haired, neater species with a ‘clipped’ appearance, ocellar hairs no longer than separation of eyes at vertex (this latter character can be difficult to interpret in the absence of reference material so examination of furca and apical sternite reccommended), tip of furca membranous (Plate III)................................................. ♀ 
<taxonomicName id="E6C14D7DFFC5FF80FB20FAE1FA6A0356" ID-CoL="8FL5Y" authority="Macquart" authorityName="Macquart" box="[1268,1436,1323,1345]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="13B5EAECFFC5FF80FB20FAE1FACF0357" bold="true" box="[1268,1337,1323,1345]" italics="true" pageId="16" pageNumber="17">bicolor</emphasis>
Macquart
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC5FF80FF43FA82FA6A036D" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43FA82FA6A036D" pageId="16" pageNumber="17">
- A longer-haired, rather untidy looking species, ocellar hairs longer than separation of eyes at vertex, but this is variable and reference to furca and apical sternite required, tip of furca pigmented (Plate VI)..................... ..♀ 
<taxonomicName id="E6C14D7DFFC5FF80FAD7FAAFFA6A036D" ID-CoL="8FL63" authority="Becker" authorityName="Becker" box="[1283,1436,1381,1403]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="crinipes">
<emphasis id="13B5EAECFFC5FF80FAD7FAAFFAA7036D" bold="true" box="[1283,1361,1381,1403]" italics="true" pageId="16" pageNumber="17">crinipes</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC5FF80FF43FA49FA6A001B" pageId="16" pageNumber="17">
<paragraph id="217E36FEFFC5FF80FF43FA49FA6D03A3" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43FA49FA6D03A3" pageId="16" pageNumber="17">33. Short-haired species, marginal hairs on scutellum less than half length of scutellum, ventral hairs on hind femora shorter than width of femora. Wing often infuscated or tinged with brown and sometimes with yellower base...................... 34</keyLead>
</paragraph>
<paragraph id="217E36FEFFC5FF80FF43FA74FA6A001B" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43FA74FA6A001B" pageId="16" pageNumber="17">- Longer haired species with marginal hairs of scutellum exceeding half length of scutellum, sometimes almost as long as scutellum medially, ventral hairs of hind femora longer than width of femora. Wing either clear, uniformly pale brownish or faintly yellow tinged........................................................................................35</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC5FF80FF43F9DCFA6A00EE" pageId="16" pageNumber="17">
<paragraph id="217E36FEFFC5FF80FF43F9DCFA6D0095" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43F9DCFA6D0095" pageId="16" pageNumber="17">
34. A very short-haired species, ocellar hairs shorter than distance between lateral ocelli, sometimes barely visible. Hairs on disc of mesonotum barely longer than distance between them. Mesonotum between hair insertions smooth and shining so that hair punctures easily visible (see male 
<figureCitation id="B9FA2A7BFFC5FF80FDE7F99AFD700073" box="[563,646,1616,1637]" captionStart="FIGURE 8" captionStartId="11.[151,250,1736,1758]" captionTargetId="figure@11.[358,1228,1060,1715]" captionTargetPageId="11" captionText="FIGURE 8. ♂ U. lata head and mesonotum dorsal." figureDoi="http://doi.org/10.5281/zenodo.6376812" httpUri="https://zenodo.org/record/6376812/files/figure.png" pageId="16" pageNumber="17">Figure 8</figureCitation>
). Wing usually with strong brown infuscation fading out in apical third of wing (very variable)..............................................................................♀ 
<taxonomicName id="E6C14D7DFFC5FF80FAEEF9A7FA6D0095" ID-CoL="8FL6G" authority="Loew" authorityName="Loew" box="[1338,1435,1645,1667]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="13B5EAECFFC5FF80FAEEF9A7FAA90095" bold="true" box="[1338,1375,1645,1667]" italics="true" pageId="16" pageNumber="17">lata</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC5FF80FF43F941FA6A00EE" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43F941FA6A00EE" pageId="16" pageNumber="17">
- A longer haired species, ocellar hairs always apparent and about as long as separation of eyes at vertex or longer. Mesonotal hairs relatively long, almost twice as long as the distance between them. Mesonotum between hair insertions with roughened, matt surface such that hair punctures not easily discerned. Wing faintly brown tinged with yellower base...............................................................................................♀ 
<taxonomicName id="E6C14D7DFFC5FF80FB75F929FABF00EE" ID-CoL="8FL82" authority="Gibbs" authorityName="Gibbs" box="[1185,1353,1763,1785]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="greatheadi" status="sp. nov.">
<emphasis id="13B5EAECFFC5FF80FB75F929FAFC00EF" bold="true" box="[1185,1290,1763,1785]" italics="true" pageId="16" pageNumber="17">greatheadi</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFC5FF80FA9BF928FA6A00EE" box="[1359,1436,1762,1784]" pageId="16" pageNumber="17" rank="species">
<emphasis id="13B5EAECFFC5FF80FA9BF928FA6A00EE" bold="true" box="[1359,1436,1762,1784]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC5FF80FF43F8CAFA6D019D" pageId="16" pageNumber="17">
<paragraph id="217E36FEFFC5FF80FF43F8CAFA6A0146" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43F8CAFA6A0146" pageId="16" pageNumber="17">
35. Larger species, often with greenish reflections, rather densely covered with long, wavy yellowish hairs giving it shaggy appearance. Unique among 
<taxonomicName id="E6C14D7DFFC5FF80FE38F8D4FDEF0125" ID-CoL="63VJK" box="[492,537,1822,1843]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC5FF80FE38F8D4FDEF0125" box="[492,537,1822,1843]" italics="true" pageId="16" pageNumber="17">Usia</emphasis>
</taxonomicName>
in having almost spherical spermathecal reservoirs (Plate XX, Figure g). 
<collectingCountry id="59D6766EFFC5FF80FB08F8D4FAD60125" box="[1244,1312,1822,1843]" name="France" pageId="16" pageNumber="17">France</collectingCountry>
, 
<collectingCountry id="59D6766EFFC5FF80FAFDF8D4FAA00125" box="[1321,1366,1822,1843]" name="Italy" pageId="16" pageNumber="17">Italy</collectingCountry>
, 
<collectingCountry id="59D6766EFFC5FF80FA8BF8D4FA600125" box="[1375,1430,1822,1843]" name="Spain" pageId="16" pageNumber="17">Spain</collectingCountry>
........................................................................................♀ 
<taxonomicName id="E6C14D7DFFC5FF80FAC2F8F1FA6A0146" ID-CoL="8FQ4W" authority="Strobl" authorityName="Strobl" box="[1302,1436,1851,1873]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pubera">
<emphasis id="13B5EAECFFC5FF80FAC2F8F1FAAD0147" bold="true" box="[1302,1371,1851,1873]" italics="true" pageId="16" pageNumber="17">pubera</emphasis>
Strobl
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC5FF80FF43F892FA6D019D" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43F892FA6D019D" pageId="16" pageNumber="17">- Small black species, vestiture rather sparse, long, straight or slightly curved, white hairs. Spermathecal reservoirs elongate as normal. Central Asia..................................................................................36</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC5FF81FF43F859FA6A05E7" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<paragraph id="217E36FEFFC5FF80FF43F859FA6D01F5" blockId="16.[151,1436,1176,2019]" pageId="16" pageNumber="17">
<keyLead id="9A3095CBFFC5FF80FF43F859FA6D01F5" pageId="16" pageNumber="17">
36. Front of mesonotum densely grey dusted laterally back to level of rear of post pronotal lobe, post pronotal lobe itself and notopleuron entirely dusted. Frons dusted right across immediately above antennae, slightly more thinly so centrally but clearly obscuring integument (
<figureCitation id="B9FA2A7BFFC5FF80FDD1F804FD9201F5" box="[517,612,1998,2019]" captionStart="FIGURE 17" captionStartId="17.[151,250,869,891]" captionTargetId="figure@17.[358,1228,193,848]" captionTargetPageId="17" captionText="FIGURE 17. ♀ U. unicolor head and mesonotum anterodorsally." figureDoi="http://doi.org/10.5281/zenodo.6376831" httpUri="https://zenodo.org/record/6376831/files/figure.png" pageId="16" pageNumber="17">Figure 17</figureCitation>
)......................................................♀ 
<taxonomicName id="E6C14D7DFFC5FF80FADFF807FA6D01F5" ID-CoL="8FL6V" authority="Loew" authorityName="Loew" box="[1291,1435,1997,2019]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="13B5EAECFFC5FF80FADFF807FAA901F5" bold="true" box="[1291,1375,1997,2019]" italics="true" pageId="16" pageNumber="17">unicolor</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<caption id="75BE6676FFC4FF81FF43FCAFFCD8056D" ID-DOI="http://doi.org/10.5281/zenodo.6376831" ID-Zenodo-Dep="6376831" box="[151,814,869,891]" httpUri="https://zenodo.org/record/6376831/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,869,891]" targetBox="[358,1227,193,847]" targetPageId="17">
<paragraph id="217E36FEFFC4FF81FF43FCAFFCD8056D" blockId="17.[151,814,869,891]" box="[151,814,869,891]" pageId="17" pageNumber="18">
<emphasis id="13B5EAECFFC4FF81FF43FCAFFEE9056D" bold="true" box="[151,287,869,891]" pageId="17" pageNumber="18">FIGURE 17.</emphasis>
♀ 
<taxonomicName id="E6C14D7DFFC4FF81FEEBFCACFE42056D" ID-CoL="8FL6V" box="[319,436,869,891]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="13B5EAECFFC4FF81FEEBFCACFE42056D" box="[319,436,869,891]" italics="true" pageId="17" pageNumber="18">U. unicolor</emphasis>
</taxonomicName>
head and mesonotum anterodorsally.
</paragraph>
</caption>
<paragraph id="217E36FEFFC4FF81FF43FC6BFA6A05E7" blockId="17.[151,1436,929,1009]" pageId="17" pageNumber="18">
<keyLead id="9A3095CBFFC4FF81FF43FC6BFA6A05E7" pageId="17" pageNumber="18">
- Anterior slope of mesonotum largely shining black, grey dusting close to suture of post pronotal lobe. Anterolateral dust spots on frons widely separated, area between them shining black. Post pronotal lobe ventrally shining, notopleuron with no more than a narrow line of thin dusting ventrally (
<figureCitation id="B9FA2A7BFFC4FF81FD5CFC16FD1005E7" box="[648,742,988,1009]" captionStart="FIGURE 18" captionStartId="17.[151,250,1734,1756]" captionTargetId="figure@17.[358,1228,1058,1713]" captionTargetPageId="17" captionText="FIGURE 18. ♀ [? U. incognita] head and mesonotum anterodorsally." figureDoi="http://doi.org/10.5281/zenodo.6376833" httpUri="https://zenodo.org/record/6376833/files/figure.png" pageId="17" pageNumber="18">Figure 18</figureCitation>
)................♀ 
<taxonomicName id="E6C14D7DFFC4FF81FC01FC11FB5305E7" ID-CoL="8FL6D" authority="Paramonov" authorityName="Paramonov" box="[981,1189,987,1009]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="13B5EAECFFC4FF81FC01FC11FBC405E7" bold="true" box="[981,1074,987,1009]" italics="true" pageId="17" pageNumber="18">incognita</emphasis>
Paramonov
</taxonomicName>
[no certain females seen]
</keyLead>
</paragraph>
</keyStep>
<caption id="75BE6676FFC4FF81FF43F90CFCAC00CA" ID-DOI="http://doi.org/10.5281/zenodo.6376833" ID-Zenodo-Dep="6376833" box="[151,858,1734,1757]" httpUri="https://zenodo.org/record/6376833/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1734,1756]" targetBox="[358,1227,1058,1712]" targetPageId="17">
<paragraph id="217E36FEFFC4FF81FF43F90CFCAC00CA" blockId="17.[151,858,1734,1757]" box="[151,858,1734,1757]" pageId="17" pageNumber="18">
<emphasis id="13B5EAECFFC4FF81FF43F90CFEE900CA" bold="true" box="[151,287,1734,1756]" pageId="17" pageNumber="18">FIGURE 18.</emphasis>
♀ [? 
<taxonomicName id="E6C14D7DFFC4FF81FE8DF902FE2E00CB" ID-CoL="8FL6D" box="[345,472,1736,1757]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="incognita">
<emphasis id="13B5EAECFFC4FF81FE8DF902FE2E00CB" box="[345,472,1736,1757]" italics="true" pageId="17" pageNumber="18">U. incognita</emphasis>
</taxonomicName>
] head and mesonotum anterodorsally.
</paragraph>
</caption>
<keyStep id="9A352E5BFFC4FF81FF43F8EBFA6A01BD" pageId="17" pageNumber="18">
<paragraph id="217E36FEFFC4FF81FF43F8EBFA6A0145" blockId="17.[151,1436,1825,2023]" pageId="17" pageNumber="18">
<keyLead id="9A3095CBFFC4FF81FF43F8EBFA6A0145" pageId="17" pageNumber="18">37. Acrostichals proclinate; rarely (in European specimens) a few hairs nearest head erect to reclinate, then katepisternum entirely dusted and dust spots on frons almost reaching level of anterior ocellus..........................................38</keyLead>
</paragraph>
<paragraph id="217E36FEFFC4FF81FF43F896FA6A01BD" blockId="17.[151,1436,1825,2023]" pageId="17" pageNumber="18">
<keyLead id="9A3095CBFFC4FF81FF43F896FA6A01BD" pageId="17" pageNumber="18">- Acrostichals with at least anterior hairs reclinate or too short to judge; in specimens from Middle East with both proclinate and reclinate acrostichals or anterior reclinate acrostichals missing then katepisternum subshining and undusted on ventral half and dust spots on frons extending only half way to anterior ocellus.................................................40</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC4FF82FF43F87EFA6D06DC" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<paragraph id="217E36FEFFC4FF81FF43F87EFA6A01F1" blockId="17.[151,1436,1825,2023]" pageId="17" pageNumber="18">
<keyLead id="9A3095CBFFC4FF81FF43F87EFA6A01F1" pageId="17" pageNumber="18">
38. Occiput with thin coating of dust on and around occipital tumescence. Katepisternum with ventral undusted shining patch. 
<collectingCountry id="59D6766EFFC4FF81FAA6F87EFEF501F0" name="Turkey" pageId="17" pageNumber="18">Turkey</collectingCountry>
..........................................................................♀ 
<taxonomicName id="E6C14D7DFFC4FF81FB47F81BFABF01F0" ID-CoL="8FL72" authority="Gibbs" authorityName="Gibbs" box="[1171,1353,2001,2023]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="anatoliensis" status="sp. nov.">
<emphasis id="13B5EAECFFC4FF81FB47F81BFAFC01F1" bold="true" box="[1171,1290,2001,2023]" italics="true" pageId="17" pageNumber="18">anatoliensis</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFC4FF81FA9BF81BFA6A01F1" box="[1359,1436,2001,2023]" pageId="17" pageNumber="18" rank="species">
<emphasis id="13B5EAECFFC4FF81FA9BF81BFA6A01F1" bold="true" box="[1359,1436,2001,2023]" pageId="17" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC7FF82FF43FF5DFA6D06DC" blockId="18.[151,1436,151,613]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43FF5DFA6D06DC" pageId="18" pageNumber="19">- Occiput entirely shining, any thin dusting confined to occipital tumescence. Katepisternum uniformly grey dusted obscuring ground colour....................................................................................... 39</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC7FF82FF43FF18FA6A0712" pageId="18" pageNumber="19">
<paragraph id="217E36FEFFC7FF82FF43FF18FA6D06F1" blockId="18.[151,1436,151,613]" box="[151,1435,210,232]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43FF18FA6D06F1" box="[151,1435,210,232]" pageId="18" pageNumber="19">
39. Wing membrane with uniform brownish tinge, mesonotal hairs somewhat brownish. 
<collectingCountry id="59D6766EFFC7FF82FBDAFF18FBCF06F1" box="[1038,1081,210,231]" name="Italy" pageId="18" pageNumber="19">Italy</collectingCountry>
................ ♀ 
<taxonomicName id="E6C14D7DFFC7FF82FAC8FF18FA6D06F1" ID-CoL="8FL6H" authority="Loew" authorityName="Loew" box="[1308,1435,210,232]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="manca">
<emphasis id="13B5EAECFFC7FF82FAC8FF18FAA806FE" bold="true" box="[1308,1374,210,232]" italics="true" pageId="18" pageNumber="19">manca</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC7FF82FF43FF25FA6A0712" blockId="18.[151,1436,151,613]" box="[151,1436,239,261]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43FF25FA6A0712" box="[151,1436,239,261]" pageId="18" pageNumber="19">
- Wing membrane clear, hardly tinged, mesonotal hairs white. 
<collectingCountry id="59D6766EFFC7FF82FCFDFF25FC960712" box="[809,864,239,260]" name="Spain" pageId="18" pageNumber="19">Spain</collectingCountry>
and 
<collectingCountry id="59D6766EFFC7FF82FC5AFF25FC260712" box="[910,976,239,260]" name="France" pageId="18" pageNumber="19">France</collectingCountry>
........................ ♀ 
<taxonomicName id="E6C14D7DFFC7FF82FAC6FF25FA6A0712" ID-CoL="8FL6P" authority="Becker" authorityName="Becker" box="[1298,1436,239,261]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFC7FF82FAC6FF25FAA40713" bold="true" box="[1298,1362,239,261]" italics="true" pageId="18" pageNumber="19">putilla</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC7FF82FF43FEC7FA6A076C" pageId="18" pageNumber="19">
<paragraph id="217E36FEFFC7FF82FF43FEC7FA6A0756" blockId="18.[151,1436,151,613]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43FEC7FA6A0756" pageId="18" pageNumber="19">
40. Occiput almost entirely shining black, even on ventral part adjacent to lower edge of oral opening. Most anterior acrostichals reclinate, remainder proclinate......................................................♀ 
<taxonomicName id="E6C14D7DFFC7FF82FB7EFEE0FABF0729" ID-CoL="8FL74" authority="Gibbs" authorityName="Gibbs" box="[1194,1353,298,320]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="annetteae" status="sp. nov.">
<emphasis id="13B5EAECFFC7FF82FB7EFEE0FAFC0756" bold="true" box="[1194,1290,298,320]" italics="true" pageId="18" pageNumber="19">annetteae</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFC7FF82FA9BFEE0FA6A0756" box="[1359,1436,298,320]" pageId="18" pageNumber="19" rank="species">
<emphasis id="13B5EAECFFC7FF82FA9BFEE0FA6A0756" bold="true" box="[1359,1436,298,320]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC7FF82FF43FE8DFA6A076C" blockId="18.[151,1436,151,613]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43FE8DFA6A076C" pageId="18" pageNumber="19">- At least ventral part of head dusted, even if only thinly so, often only a strip behind eyes shining and undusted. All acrostichals reclinate or too short to judge...........................................................................41</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC7FF82FF43FE48FA6D041A" pageId="18" pageNumber="19">
<paragraph id="217E36FEFFC7FF82FF43FE48FA6A07A2" blockId="18.[151,1436,151,613]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43FE48FA6A07A2" pageId="18" pageNumber="19">41. Very short-haired species, marginal hairs of scutellum barely even quarter mid length of scutellum. [Anepisternum at least partly undusted AND anal lobe at least as wide as anal cell]...................................................42</keyLead>
</paragraph>
<paragraph id="217E36FEFFC7FF82FF43FE77FA6D041A" blockId="18.[151,1436,151,613]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43FE77FA6D041A" pageId="18" pageNumber="19">- Longer-haired species, marginal hairs of scutellum from about half length of scutellum to equal to length of scutellum, at least more than a quarter scutellar mid-length. In African species with this feature unclear, then anepisternum entirely grey dusted AND anal lobe narrower than anal cell....................................................................44</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC7FF82FF49FDDFFA6D010E" pageId="18" pageNumber="19">
<paragraph id="217E36FEFFC7FF82FF49FDDFFA6D0472" blockId="18.[151,1436,151,613]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF49FDDFFA6D0472" pageId="18" pageNumber="19">
42. Mesonotum covered with very short hairs, most barely longer than distance between hair insertions (
<figureCitation id="B9FA2A7BFFC7FF82FB78FDDFFAF8043C" box="[1196,1294,533,554]" captionStart="FIGURE 19" captionStartId="18.[151,250,1647,1669]" captionTargetBox="[151,1436,662,1626]" captionTargetId="figure@18.[151,1436,662,1626]" captionTargetPageId="18" captionText="FIGURE 19. ♀ U. calva lectotype mesonotum." httpUri="https://zenodo.org/record/286092/files/figure.png" pageId="18" pageNumber="19">Figure 19</figureCitation>
). Hairs arising from relatively large punctures which, in anterior part of disc, separated by little more than diameter of puncture. Vaginal plate strongly sclerotized. 
<collectingCountry id="59D6766EFFC7FF82FE7DFD85FE1B0472" box="[425,493,591,612]" name="Turkey" pageId="18" pageNumber="19">Turkey</collectingCountry>
...................................................................♀ 
<taxonomicName id="E6C14D7DFFC7FF82FAFFFD85FA6D0472" ID-CoL="8FL5Z" authority="Loew" authorityName="Loew" box="[1323,1435,591,613]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="calva">
<emphasis id="13B5EAECFFC7FF82FAFFFD85FAA90473" bold="true" box="[1323,1375,591,613]" italics="true" pageId="18" pageNumber="19">calva</emphasis>
Loew
</taxonomicName>
</keyLead>
</paragraph>
<caption id="75BE6676FFC7FF82FF43F9A5FD8A0093" box="[151,636,1647,1670]" httpUri="https://zenodo.org/record/286092/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1436,662,1626]" targetPageId="18">
<paragraph id="217E36FEFFC7FF82FF43F9A5FD8A0093" blockId="18.[151,636,1647,1670]" box="[151,636,1647,1670]" pageId="18" pageNumber="19">
<emphasis id="13B5EAECFFC7FF82FF43F9A5FEE90093" bold="true" box="[151,287,1647,1669]" pageId="18" pageNumber="19">FIGURE 19.</emphasis>
♀ 
<taxonomicName id="E6C14D7DFFC7FF82FEEBF9BBFE630093" ID-CoL="8FL5Z" box="[319,405,1647,1670]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="calva">
<emphasis id="13B5EAECFFC7FF82FEEBF9BBFE630093" box="[319,405,1647,1670]" italics="true" pageId="18" pageNumber="19">U. calva</emphasis>
</taxonomicName>
lectotype mesonotum.
</paragraph>
</caption>
<paragraph id="217E36FEFFC7FF82FF43F903FA6D010E" blockId="18.[151,1436,1737,1875]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF43F903FA6D010E" pageId="18" pageNumber="19">- Mesonotum covered with longer hairs, most clearly longer than distance between hair insertions. Hairs arising from relatively small punctures which in anterior part of disc clearly separated by more than diameter of puncture, usually much more. Vaginal plate membranous. Africa.............................................................................. 43</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC7FF83FF49F8EBFA6A05E7" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<paragraph id="217E36FEFFC7FF82FF49F8EBFA6A0145" blockId="18.[151,1436,1737,1875]" pageId="18" pageNumber="19">
<keyLead id="9A3095CBFFC7FF82FF49F8EBFA6A0145" pageId="18" pageNumber="19">
43. Anterior slope of mesonotum between post pronotal lobe and paramedian vittae dusted (
<figureCitation id="B9FA2A7BFFC7FF82FB83F8EBFB4D0120" box="[1111,1211,1825,1846]" captionStart="FIGURE 20" captionStartId="19.[151,250,869,891]" captionTargetId="figure@19.[358,1228,193,848]" captionTargetPageId="19" captionText="FIGURE 20. ♀ U. accola anterior slope of mesonotum." figureDoi="http://doi.org/10.5281/zenodo.6376835" httpUri="https://zenodo.org/record/6376835/files/figure.png" pageId="18" pageNumber="19">Figure 20</figureCitation>
), anepisternum mostly dusted, shining only on anterodorsal corner. Tip of furca membranous, unpigmented................... ♀ 
<taxonomicName id="E6C14D7DFFC7FF82FAC6F8F7FA6A0145" ID-CoL="8FL5P" authority="Becker" authorityName="Becker" box="[1298,1436,1853,1875]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="accola">
<emphasis id="13B5EAECFFC7FF82FAC6F8F7FAA40145" bold="true" box="[1298,1362,1853,1875]" italics="true" pageId="18" pageNumber="19">accola</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
<caption id="75BE6676FFC6FF83FF43FCAFFD27056D" ID-DOI="http://doi.org/10.5281/zenodo.6376835" ID-Zenodo-Dep="6376835" box="[151,721,869,891]" httpUri="https://zenodo.org/record/6376835/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,869,891]" targetBox="[358,1228,193,847]" targetPageId="19">
<paragraph id="217E36FEFFC6FF83FF43FCAFFD27056D" blockId="19.[151,721,869,891]" box="[151,721,869,891]" pageId="19" pageNumber="20">
<emphasis id="13B5EAECFFC6FF83FF43FCAFFED6056D" bold="true" box="[151,288,869,891]" pageId="19" pageNumber="20">FIGURE 20.</emphasis>
♀ 
<taxonomicName id="E6C14D7DFFC6FF83FEEBFCACFE54056D" ID-CoL="8FL5P" box="[319,418,869,891]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="accola">
<emphasis id="13B5EAECFFC6FF83FEEBFCACFE54056D" box="[319,418,869,891]" italics="true" pageId="19" pageNumber="20">U. accola</emphasis>
</taxonomicName>
anterior slope of mesonotum.
</paragraph>
</caption>
<paragraph id="217E36FEFFC6FF83FF43FC6BFA6A05E7" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FC6BFA6A05E7" pageId="19" pageNumber="20">
- Anterior slope of mesonotum between post pronotal lobe and paramedian vittae shining except narrow band adjacent to post pronotal lobe, anepisternum mostly shining, dusted only along posterior margin. Tip of furca pointed, dark pigmented......................................................................................♀ 
<taxonomicName id="E6C14D7DFFC6FF83FB7EFC11FABF05E7" ID-CoL="8FL7C" authority="Gibbs" authorityName="Gibbs" box="[1194,1353,987,1009]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cornigera" status="sp. nov.">
<emphasis id="13B5EAECFFC6FF83FB7EFC11FAFC05E7" bold="true" box="[1194,1290,987,1009]" italics="true" pageId="19" pageNumber="20">cornigera</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFC6FF83FA9BFC11FA6A05E7" box="[1359,1436,987,1009]" pageId="19" pageNumber="20" rank="species">
<emphasis id="13B5EAECFFC6FF83FA9BFC11FA6A05E7" bold="true" box="[1359,1436,987,1009]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC6FF83FF43FC33FA6D0270" pageId="19" pageNumber="20">
<paragraph id="217E36FEFFC6FF83FF43FC33FA6A023D" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FC33FA6A023D" pageId="19" pageNumber="20">44. Ventral hairs on hind femora longer than greatest width of femora, if only just as long then with long hairs on frons extending at least half way from anterior ocellus to front of frons.......................................................45</keyLead>
</paragraph>
<paragraph id="217E36FEFFC6FF83FF43FBFEFA6D0270" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FBFEFA6D0270" pageId="19" pageNumber="20">- Ventral hairs on hind femora no longer than greatest width of femora, usually shorter, no long hairs on frons much beyond anterior ocellus...................................................................................... 46</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC6FF83FF43FBA4FA6D0300" pageId="19" pageNumber="20">
<paragraph id="217E36FEFFC6FF83FF43FBA4FA6A02A8" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FBA4FA6A02A8" pageId="19" pageNumber="20">
45. Very long-haired species, ventral hairs on hind femora clearly longer than width of femora. Hairs on lateral margins of frons hardly extending beyond anterior ocellus. Anepisternum dusted. No sclerotised vaginal plate, furca with elongated tip.............................................................................................♀ 
<taxonomicName id="E6C14D7DFFC6FF83FB28FB62FA6A02A8" ID-CoL="8FL6X" authority="Macquart" authorityName="Macquart" box="[1276,1436,1192,1214]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="vestita">
<emphasis id="13B5EAECFFC6FF83FB28FB62FACC02A8" bold="true" box="[1276,1338,1192,1214]" italics="true" pageId="19" pageNumber="20">vestita</emphasis>
Macquart
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC6FF83FF43FB0CFA6D0300" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FB0CFA6D0300" pageId="19" pageNumber="20">
- Shorter-haired species, ventral hairs on hind femora a little longer than width of femora. Frons with long hairs extending at least half way from anterior ocellus to front of frons. Anepisternum shining. V-shaped sclerotized vaginal plate, tip of furca simply rounded........................................................................♀ 
<taxonomicName id="E6C14D7DFFC6FF83FB26FACAFA6D0300" ID-CoL="8FL5X" authority="Fabricius" authorityName="Fabricius" box="[1266,1435,1280,1302]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="13B5EAECFFC6FF83FB26FACAFADA0300" bold="true" box="[1266,1324,1280,1302]" italics="true" pageId="19" pageNumber="20">atrata</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC6FF83FF43FAD4FA6A039D" pageId="19" pageNumber="20">
<paragraph id="217E36FEFFC6FF83FF43FAD4FA6A0347" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FAD4FA6A0347" pageId="19" pageNumber="20">
46. A small species, anal lobe narrower than anal cell; katepisternum glabrous. Frons parallel in hind half, abruptly diverging in anterior half. Arms of furca falcate and set at right angles to main part......................... ..♀ 
<taxonomicName id="E6C14D7DFFC6FF83FB00FAF1FA6A0347" ID-CoL="8FL5V" authority="Becker" authorityName="Becker" box="[1236,1436,1339,1361]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFC6FF83FB00FAF1FAA70347" bold="true" box="[1236,1361,1339,1361]" italics="true" pageId="19" pageNumber="20">angustifrons</emphasis>
Becker
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC6FF83FF43FA93FA6A039D" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FA93FA6A039D" pageId="19" pageNumber="20">- Larger species, anal cell broadly convex, as wide as anal cell; katepisternum with a few hairs centrally (can be difficult to see, check both sides). Frons diverging almost uniformly from narrowest part to front..................................47</keyLead>
</paragraph>
</keyStep>
<keyStep id="9A352E5BFFC6FF83FF43FA5EFA6D002D" pageId="19" pageNumber="20">
<paragraph id="217E36FEFFC6FF83FF43FA5EFA6A03F2" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FA5EFA6A03F2" pageId="19" pageNumber="20">
47. Anepisternum almost entirely dusted. Ocellar hairs longer than width across lateral ocelli, anterior slope of mesonotum dusted between paramedian vittae and post pronotal lobe. Basal spermathecal ducts coming to simple confluence.......................................................................................... ..♀ 
<taxonomicName id="E6C14D7DFFC6FF83FB55FA04FABF03F5" ID-CoL="8FL8D" authority="Gibbs" authorityName="Gibbs" box="[1153,1353,1486,1508]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="maghrebensis" status="sp. nov.">
<emphasis id="13B5EAECFFC6FF83FB55FA04FAFC03F2" bold="true" box="[1153,1290,1486,1508]" italics="true" pageId="19" pageNumber="20">maghrebensis</emphasis>
Gibbs
</taxonomicName>
<taxonomicNameLabel id="08865797FFC6FF83FA9BFA04FA6A03F2" box="[1359,1436,1486,1508]" pageId="19" pageNumber="20" rank="species">
<emphasis id="13B5EAECFFC6FF83FA9BFA04FA6A03F2" bold="true" box="[1359,1436,1486,1508]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="217E36FEFFC6FF83FF43FA26FA6D002D" blockId="19.[151,1436,929,1596]" pageId="19" pageNumber="20">
<keyLead id="9A3095CBFFC6FF83FF43FA26FA6D002D" pageId="19" pageNumber="20">
- Anepisternum largely shining. Ocellar hairs shorter than distance between lateral ocelli, dusting on anterior of mesonotum confined to narrow strip above post pronotal lobe. Basal spermathecal ducts joining at distinct vaginal plate, apical sternite characteristic...............................................................................♀ 
<taxonomicName id="E6C14D7DFFC6FF83FB26F9ECFA6D002D" ID-CoL="8FL69" authority="Fabricius" authorityName="Fabricius" box="[1266,1435,1574,1596]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="florea">
<emphasis id="13B5EAECFFC6FF83FB26F9ECFADB002A" bold="true" box="[1266,1325,1574,1596]" italics="true" pageId="19" pageNumber="20">florea</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection id="69DB6575FFC6FF83FF43F946FE7C00B0" box="[151,394,1676,1702]" pageId="19" pageNumber="20" type="multiple">
<paragraph id="217E36FEFFC6FF83FF43F946FE7C00B0" blockId="19.[151,394,1676,1702]" box="[151,394,1676,1702]" pageId="19" pageNumber="20">
<heading id="7A368192FFC6FF83FF43F946FE7C00B0" bold="true" box="[151,394,1676,1702]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="E6C14D7DFFC6FF83FF43F946FE7C00B0" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[151,394,1676,1702]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FF43F946FE7C00B0" bold="true" box="[151,394,1676,1702]" pageId="19" pageNumber="20">
<emphasis id="13B5EAECFFC6FF83FF43F946FF3800B0" bold="true" box="[151,206,1676,1702]" italics="true" pageId="19" pageNumber="20">Usia</emphasis>
Latreille, 1802
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFC6FF84FF43F918FE2A06DA" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="217E36FEFFC6FF83FF43F918FB2C0111" blockId="19.[151,1436,1746,1983]" pageId="19" pageNumber="20">
<taxonomicName id="E6C14D7DFFC6FF83FF43F918FE2D00FE" authority="Fabricius, 1794: 412" authorityName="Fabricius" authorityPageNumber="412" authorityYear="1794" box="[151,475,1746,1768]" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FF43F918FEF700FE" box="[151,257,1746,1768]" italics="true" pageId="19" pageNumber="20">Voluccella</emphasis>
Fabricius, 1794: 412
</taxonomicName>
. 
<typeStatus id="FE7A885CFFC6FF83FE31F919FDE100FE" box="[485,535,1747,1768]" pageId="19" pageNumber="20">Type</typeStatus>
species: 
<taxonomicName id="E6C14D7DFFC6FF83FDA1F918FC3E00FE" authority="Fabricius, 1794" authorityName="Fabricius" authorityYear="1794" box="[629,968,1746,1768]" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="florea">
<emphasis id="13B5EAECFFC6FF83FDA1F918FCD700FE" box="[629,801,1746,1768]" italics="true" pageId="19" pageNumber="20">Voluccella florea</emphasis>
Fabricius, 1794
</taxonomicName>
, automatic (by designation of 
<typeStatus id="FE7A885CFFC6FF83FB28F919FADE00FE" box="[1276,1320,1747,1768]" pageId="19" pageNumber="20">type</typeStatus>
species for 
<taxonomicName id="E6C14D7DFFC6FF83FF13F938FEA10111" ID-CoL="63VJK" authority="Latreille" authorityName="Latreille" box="[199,343,1777,1799]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FF13F938FF010111" box="[199,247,1778,1799]" italics="true" pageId="19" pageNumber="20">Usia</emphasis>
Latreille
</taxonomicName>
). [
<taxonomicName id="E6C14D7DFFC6FF83FEA4F93BFE2F0111" authority="Fabricius, 1794: 412" authorityName="Fabricius" authorityPageNumber="412" authorityYear="1794" box="[368,473,1777,1799]" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FEA4F93BFE2F0111" box="[368,473,1777,1799]" italics="true" pageId="19" pageNumber="20">Voluccella</emphasis>
</taxonomicName>
not preoccupied by 
<taxonomicName id="E6C14D7DFFC6FF83FD7CF93BFC5A0111" authority="Geoffroy, 1762" authorityName="Geoffroy" authorityYear="1762" box="[680,940,1777,1799]" class="Insecta" family="Bombyliidae" genus="Volucella" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FD7CF93BFCF00111" box="[680,774,1777,1799]" italics="true" pageId="19" pageNumber="20">Volucella</emphasis>
Geoffroy, 1762
</taxonomicName>
.] 
<emphasis id="13B5EAECFFC6FF83FC68F938FBA10111" box="[956,1111,1777,1799]" italics="true" pageId="19" pageNumber="20">Nomen oblitum</emphasis>
(see below).
</paragraph>
<paragraph id="217E36FEFFC6FF83FF43F8DAFB380130" blockId="19.[151,1436,1746,1983]" box="[151,1230,1808,1830]" pageId="19" pageNumber="20">
<emphasis id="13B5EAECFFC6FF83FF43F8DAFF0D0130" box="[151,251,1808,1830]" italics="true" pageId="19" pageNumber="20">Volvicella</emphasis>
, incorrect original spelling of 
<taxonomicName id="E6C14D7DFFC6FF83FDFAF8DAFD6E0130" authority="Fabricius, 1794: 412" authorityName="Fabricius" authorityPageNumber="412" authorityYear="1794" box="[558,664,1808,1830]" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FDFAF8DAFD6E0130" box="[558,664,1808,1830]" italics="true" pageId="19" pageNumber="20">Voluccella</emphasis>
</taxonomicName>
(Fabricius, 1794: [5] [last unnumbered page of index]).
</paragraph>
<paragraph id="217E36FEFFC6FF83FF43F8FAFB6E0175" blockId="19.[151,1436,1746,1983]" pageId="19" pageNumber="20">
<taxonomicName id="E6C14D7DFFC6FF83FF43F8FAFE6C0152" ID-CoL="63VJK" authority="Latreille, 1802: 430" authorityName="Latreille" authorityPageNumber="430" authorityYear="1802" box="[151,410,1838,1861]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FF43F8FAFF310153" box="[151,199,1840,1861]" italics="true" pageId="19" pageNumber="20">Usia</emphasis>
Latreille, 1802: 430
</taxonomicName>
(unjustified new replacement name for 
<taxonomicName id="E6C14D7DFFC6FF83FCE3F8E4FBBD0152" authority="Fabricius, 1794" authorityName="Fabricius" authorityYear="1794" box="[823,1099,1838,1860]" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FCE3F8E4FC570152" box="[823,929,1838,1860]" italics="true" pageId="19" pageNumber="20">Voluccella</emphasis>
Fabricius, 1794
</taxonomicName>
). 
<typeStatus id="FE7A885CFFC6FF83FB89F8E5FB790152" box="[1117,1167,1839,1860]" pageId="19" pageNumber="20">Type</typeStatus>
species: 
<taxonomicName id="E6C14D7DFFC6FF83FB3BF8E4FE910175" authority="Fabricius, 1794" authorityName="Fabricius" authorityYear="1794" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="florea">
<emphasis id="13B5EAECFFC6FF83FB3BF8E4FA6A0152" box="[1263,1436,1838,1860]" italics="true" pageId="19" pageNumber="20">Voluccella florea</emphasis>
Fabricius, 1794
</taxonomicName>
, by subsequent designation (Latreille, 1810: 443). 
<emphasis id="13B5EAECFFC6FF83FCBCF884FBED0175" box="[872,1051,1870,1891]" italics="true" pageId="19" pageNumber="20">Nomen protectum</emphasis>
(see below)
</paragraph>
<paragraph id="217E36FEFFC6FF83FF13F8A7FBC201A9" blockId="19.[151,1436,1746,1983]" pageId="19" pageNumber="20">
[
<emphasis id="13B5EAECFFC6FF83FF1BF8A7FE720194" box="[207,388,1901,1922]" italics="true" pageId="19" pageNumber="20">Nomen protectum</emphasis>
status for 
<taxonomicName id="E6C14D7DFFC6FF83FE21F8A7FDD30194" ID-CoL="63VJK" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[501,549,1901,1922]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FE21F8A7FDD30194" box="[501,549,1901,1922]" italics="true" pageId="19" pageNumber="20">Usia</emphasis>
</taxonomicName>
was established in 
<bibRefCitation id="45504B0FFFC6FF83FD26F8A6FBD40194" author="Evenhuis, N. L. &amp; Greathead, D. J." box="[754,1058,1900,1922]" firstAuthor="Evenhuis" journalOrPublisher="Zootaxa" pageId="19" pageNumber="20" pagination="1 - 64" part="300" refId="ref58625" refString="Evenhuis, N. L. &amp; Greathead, D. J. (2003) World Catalog of Bee Flies (Diptera: Bombyliidae) Corrigenda and Addenda. Zootaxa, 300, 1 - 64." title="World Catalog of Bee Flies (Diptera: Bombyliidae) Corrigenda and Addenda" type="journal article" year="2003">Evenhuis &amp; Greathead, 2003</bibRefCitation>
). By virtue of I.C.Z.N. 
<emphasis id="13B5EAECFFC6FF83FACCF8A6FABB0194" box="[1304,1357,1900,1922]" italics="true" pageId="19" pageNumber="20">Code</emphasis>
Article 23.9.1, whereby the criteria of both Articles 23.9.1.1 and 23.9.1.2 are met, 
<taxonomicName id="E6C14D7DFFC6FF83FC30F841FB4001B6" ID-CoL="63VJK" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[996,1206,1930,1952]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FC30F841FBE201B6" box="[996,1044,1931,1952]" italics="true" pageId="19" pageNumber="20">Usia</emphasis>
Latreille, 1802
</taxonomicName>
is valid and a 
<emphasis id="13B5EAECFFC6FF83FA8CF841FEDB01A9" italics="true" pageId="19" pageNumber="20">nomen protectum</emphasis>
by action of Article 23.9.2. 
<taxonomicName id="E6C14D7DFFC6FF83FD9FF863FCAC01A9" authority="Fabricius, 1794" authorityName="Fabricius" authorityYear="1794" box="[587,858,1961,1983]" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC6FF83FD9FF863FD4301A9" box="[587,693,1961,1983]" italics="true" pageId="19" pageNumber="20">Voluccella</emphasis>
Fabricius, 1794
</taxonomicName>
, is a 
<emphasis id="13B5EAECFFC6FF83FC59F860FBD201A9" box="[909,1060,1961,1983]" italics="true" pageId="19" pageNumber="20">nomen oblitum</emphasis>
.]
</paragraph>
<paragraph id="217E36FEFFC1FF84FF43FF52FE2A06DA" blockId="20.[151,1436,152,204]" pageId="20" pageNumber="21">
<taxonomicName id="E6C14D7DFFC1FF84FF43FF52FE1F06B8" authority="Rafinesque, 1815: 130" authorityName="Rafinesque" authorityPageNumber="130" authorityYear="1815" box="[151,489,152,174]" class="Insecta" family="Papilionidae" genus="Zerinthia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC1FF84FF43FF52FF0306B8" box="[151,245,152,174]" italics="true" pageId="20" pageNumber="21">Zerinthia</emphasis>
Rafinesque, 1815: 130
</taxonomicName>
(unnecessary replacement name for 
<taxonomicName id="E6C14D7DFFC1FF84FCBAFF53FBB206B8" ID-CoL="63VJK" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[878,1092,152,174]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC1FF84FCBAFF53FC6806B8" box="[878,926,153,174]" italics="true" pageId="20" pageNumber="21">Usia</emphasis>
Latreille, 1802
</taxonomicName>
). 
<typeStatus id="FE7A885CFFC1FF84FB8CFF53FB7C06B8" box="[1112,1162,153,174]" pageId="20" pageNumber="21">Type</typeStatus>
species: 
<taxonomicName id="E6C14D7DFFC1FF84FB39FF52FE9106DA" authority="Fabricius, 1794" authorityName="Fabricius" authorityYear="1794" class="Insecta" family="Syrphidae" genus="Voluccella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="florea">
<emphasis id="13B5EAECFFC1FF84FB39FF52FA6A06B8" box="[1261,1436,152,174]" italics="true" pageId="20" pageNumber="21">Voluccella florea</emphasis>
Fabricius, 1794
</taxonomicName>
, automatic.
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFC1FF84FF43FF33FDFA0707" box="[151,524,249,274]" pageId="20" pageNumber="21" type="etymology">
<paragraph id="217E36FEFFC1FF84FF43FF33FDFA0707" blockId="20.[151,524,249,274]" box="[151,524,249,274]" pageId="20" pageNumber="21">
<emphasis id="13B5EAECFFC1FF84FF43FF33FEE80704" bold="true" box="[151,286,249,274]" pageId="20" pageNumber="21">Etymology.</emphasis>
Not known for 
<taxonomicName id="E6C14D7DFFC1FF84FE04FF30FDF30707" box="[464,517,250,273]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="13B5EAECFFC1FF84FE04FF30FDF30707" box="[464,517,250,273]" italics="true" pageId="20" pageNumber="21">Usia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>