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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.221.98571" ID-Pensoft-Pub="1314-2003-221-61" ID-Pensoft-UUID="ED3480ABD44C590FB7F956151B065674" ModsDocID="1314-2003-221-61" checkinTime="1678295818264" checkinUser="pensoft" docAuthor="Siti-Munirah, Mat Yunoh &amp; Dome, Nikong" docDate="2023" docId="A9675D795A925D66B1454F4E197D535E" docLanguage="en" docName="PhytoKeys 221: 61-72" docOrigin="PhytoKeys 221" docPubDate="2023-03-07" docSource="http://dx.doi.org/10.3897/phytokeys.221.98571" docTitle="Thismia kenyirensis Siti-Munirah &amp; Dome 2023, sp. nov." docType="treatment" docVersion="2" id="ED3480ABD44C590FB7F956151B065674" lastPageNumber="61" masterDocId="ED3480ABD44C590FB7F956151B065674" masterDocTitle="Thismia kenyirensis (Thismiaceae), a new species from Taman Negeri Kenyir, Terengganu, Peninsular Malaysia" masterLastPageNumber="72" masterPageNumber="61" pageNumber="61" updateTime="1678296118906" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Thismia kenyirensis (Thismiaceae), a new species from Taman Negeri Kenyir, Terengganu, Peninsular Malaysia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Siti-Munirah, Mat Yunoh</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5062-9988</mods:nameIdentifier>
<mods:affiliation>Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia</mods:affiliation>
<mods:nameIdentifier type="email">sitimunirahfrim1@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dome, Nikong</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2620-7766</mods:nameIdentifier>
<mods:affiliation>DigitalDome Photography, 21500 Permaisuri, Terengganu, Malaysia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>221</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>61</mods:start>
<mods:end>72</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.221.98571</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.221.98571</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-221-61</mods:identifier>
<mods:identifier type="Pensoft-UUID">ED3480ABD44C590FB7F956151B065674</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:A9675D795A925D66B1454F4E197D535E" httpUri="http://treatment.plazi.org/id/A9675D795A925D66B1454F4E197D535E" lastPageNumber="61" pageId="0" pageNumber="61">
<subSubSection pageId="0" pageNumber="61" type="nomenclature">
<paragraph pageId="0" pageNumber="61">
<taxonomicName LSID="A9675D79-5A92-5D66-B145-4F4E197D535E" authority="Siti-Munirah &amp; Dome" authorityName="Siti-Munirah &amp; Dome" authorityYear="2023" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="species" species="kenyirensis" status="sp. nov.">Thismia kenyirensis Siti-Munirah &amp; Dome</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="61">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">
<figureCitation captionStart="Figure 1" captionStartId="F3" captionText="Figure 1. Thismia kenyirensis A plant with flower and roots B top view of flower showing tepals C top view of ovary and stigma D stamen (i, side view; ii, inner view) E outer view of three pendulous stamens F longitudinal section of ovary G style and stigma H bract I leaf. All from FRI 91122 a (spirit material). Drawings by Mohamad Aidil Noordin." figureDoi="10.3897/phytokeys.221.98571.figure1" httpUri="https://binary.pensoft.net/fig/821041" pageId="0" pageNumber="61">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F4" captionText="Figure 2. Thismia kenyirensis A mature plant with flowers and roots, scale in centimetres (cm) (FRI 91122 b) B mature plant in natural habitat (FRI 91122 a) C vermiform roots (FRI 91122 c) D young flower with mature ovary and stigma (FRI 91122 c). Photos by Siti-Munirah MY (A); Dome Nikong (B-D)." figureDoi="10.3897/phytokeys.221.98571.figure2" httpUri="https://binary.pensoft.net/fig/821042" pageId="0" pageNumber="61">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F5" captionText="Figure 3. Thismia kenyirensis A bracts with ovary and pistil, lateral view B top view of stigma C flower, side view D longitudinal section of floral tube and stamens E flower, top view F annulus G stamens, outer view. Photos by Siti-Munirah MY (A-C, E, F; All from FRI 91122 b); Dome Nikong (D, G; No specimens)." figureDoi="10.3897/phytokeys.221.98571.figure3" httpUri="https://binary.pensoft.net/fig/821043" pageId="0" pageNumber="61">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="diagnosis">
<paragraph pageId="0" pageNumber="61">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="61">
Distantly similar to
<taxonomicName class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia hongkongensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="species" species="hongkongensis">
<emphasis italics="true" pageId="0" pageNumber="61">Thismia hongkongensis</emphasis>
</taxonomicName>
Mar &amp; Saunders but differs by petaloid, ovate outer tepals which are ca. 8 mm long, narrowly lanceolate-linear, ca. 8 mm long inner tepals which are not forming a lose-dome, up to 28 mm long appendages of inner tepals, orange floral tube, which lacks reticulate pattern on its inner surface and connective apex with 3 long appendages.
</paragraph>
<caption doi="10.3897/phytokeys.221.98571.figure1" httpUri="https://binary.pensoft.net/fig/821041" pageId="0" pageNumber="61" start="Figure 1" startId="F3">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 1.</emphasis>
<taxonomicName authorityName="Siti-Munirah &amp; Dome" authorityYear="2023" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">Thismia kenyirensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
plant with flower and roots
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
top view of flower showing tepals
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
top view of ovary and stigma
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
stamen (i, side view; ii, inner view)
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
outer view of three pendulous stamens
<emphasis bold="true" pageId="0" pageNumber="61">F</emphasis>
longitudinal section of ovary
<emphasis bold="true" pageId="0" pageNumber="61">G</emphasis>
style and stigma
<emphasis bold="true" pageId="0" pageNumber="61">H</emphasis>
bract
<emphasis bold="true" pageId="0" pageNumber="61">I</emphasis>
leaf. All from FRI91122a (spirit material). Drawings by Mohamad Aidil Noordin.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="materials_examined">
<paragraph pageId="0" pageNumber="61">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="61">
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
. Peninsular
<materialsCitation accessionNumber="FRI 91122" collectingDate="2020-09-08" country="Malaysia" county="Hulu" elevation="204" location="Sungai Cendana" municipality="Taman Negeri Kenyir" specimenCount="1" stateProvince="Terengganu" typeStatus="holotype">
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
,
<collectingCounty>Hulu</collectingCounty>
<collectingRegion country="Malaysia" name="Terengganu">Terengganu District</collectingRegion>
,
<collectingMunicipality>Taman Negeri Kenyir</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:A9675D795A925D66B1454F4E197D535E:AD81CDD9BBABE49D3C28F60FFEDD6B93" country="Malaysia" county="Hulu" municipality="Taman Negeri Kenyir" name="Sungai Cendana" stateProvince="Terengganu">Sungai Cendana</location>
, elev. ca
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.04" unit="m" value="204.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.04" unit="m" value="204.0">204 m</elevation>
</quantity>
,
<collectingDate value="2020-09-08">8 Sept 2020</collectingDate>
, Dome Nikong,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91122">FRI 91122</accessionNumber>
a (
<typeStatus>holotype</typeStatus>
KEP!, spirit collection, barcode no. SC12015)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/phytokeys.221.98571.figure2" httpUri="https://binary.pensoft.net/fig/821042" pageId="0" pageNumber="61" start="Figure 2" startId="F4">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 2.</emphasis>
<taxonomicName authorityName="Siti-Munirah &amp; Dome" authorityYear="2023" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">Thismia kenyirensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
mature plant with flowers and roots, scale in centimetres (cm) (
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91122">FRI91122</accessionNumber>
b)
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
mature plant in natural habitat (
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91122">FRI91122</accessionNumber>
a)
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
vermiform roots (
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91122">FRI91122</accessionNumber>
c)
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
young flower with mature ovary and stigma (
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI91122">FRI91122</accessionNumber>
c). Photos by Siti-Munirah MY (
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
); Dome Nikong (
<emphasis bold="true" pageId="0" pageNumber="61">B-D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="description">
<paragraph pageId="0" pageNumber="61">Description.</paragraph>
<paragraph pageId="0" pageNumber="61">
Achlorophyllous herb, up to 130 mm tall.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Roots</emphasis>
vermiform, unbranched, ca. 1 mm in diameter, light brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Stem</emphasis>
erect, up to 90 mm long, 1.5-2 mm in diameter, pale brownish-orange, bearing 1-2 flowers.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Leaves</emphasis>
2-4, spirally arranged, triangular to narrowly triangular, scale-like, acute, margin almost entire but slightly irregularly serrate, 2-6 mm long, ca. 1 mm wide at base, pale brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Involucral bracts</emphasis>
3, similar to leaves but slightly larger, spirally arranged, triangular to narrowly triangular, scale-like, acute, entire, 8 mm long, ca. 1.5-2.5 mm wide at base, pale brown or pale orange.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Pedicel</emphasis>
up to 6 mm long at anthesis, elongating to ca. 10 mm long after anthesis, pale brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Flowers</emphasis>
terminal, actinomorphic, ca. 52 mm long (including ovary, floral tube and inner tepal with appendage);
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">floral tube</emphasis>
urceolate, 12-15 mm long, ca. 5 mm wide at base, ca. 6 mm wide at middle, ca. 7 mm wide distally;
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">outer surface</emphasis>
glabrous, orange to brownish-orange, with 12 darker orange longitudinal ribs alternating with 12 paler longitudinal lines,
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">inner surface</emphasis>
smooth or rough, almost similar to outer surface, without transverse bars or reticulate ornamentation;
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">outer tepals</emphasis>
3, free, petaloid, ovate, apex acute, 8 mm long, ca. 4-5.5 mm wide (ca. 4 mm at base, ca. 5.5 mm above middle), glabrous, bright orange;
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">inner tepals</emphasis>
3, free, narrowly lanceolate-linear, 8 mm long, ca. 1 mm wide, glabrous, dark orange, apically bearing a tentacle-like appendage; appendage narrowing towards apex, ca. 27-28 mm long, 0.5 mm wide, dark orange at base and bright orange at apex;
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">annulus</emphasis>
dark orange, glabrous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Stamens</emphasis>
6, pendent from the annulus, outer side greenish-orange or greenish-brown, inner side dark-brown to blackish and paler,
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">filaments</emphasis>
free, ca. 1 mm long, curved downwards;
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">connectives</emphasis>
flattened at inner surface, laterally connate to form a tube, ca. 4-5 mm long, narrow at base (ca. 0.8 mm wide) and broad at apex (ca. 1.7 mm wide), connective apex with 3 appendages, slightly curved inside, each appendage ca. 1.5 mm long; outer side of connective bearing a skirt-like lateral appendage protruding towards floral tube; lateral appendage not exceeding the tip of the connective appendages, with short translucent trichomes on margin,
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">interstaminal glands</emphasis>
greenish translucent, inserted on the line of fusion between connectives at the level of attachment of lateral appendages.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Ovary</emphasis>
inferior, unilocular;
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">placentas</emphasis>
3, free, forming columns and arising from the bottom of the ovary; ovules numerous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Style</emphasis>
dark brown-orangish, ca. 1 mm long;
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">stigma</emphasis>
ca. 1 mm long, papillose, 3-lobed, with lobes
<normalizedToken originalValue="±">+/-</normalizedToken>
rectangular, bifurcate at apex, pale brown or orangish to whitish.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Fruit</emphasis>
cup-shaped, 3-5 mm in height, 4-6 in diameter, pale orange (or white to creamy orange), darker at upper part.
<emphasis bold="true" italics="true" pageId="0" pageNumber="61">Seeds</emphasis>
unknown.
</paragraph>
<caption doi="10.3897/phytokeys.221.98571.figure3" httpUri="https://binary.pensoft.net/fig/821043" pageId="0" pageNumber="61" start="Figure 3" startId="F5">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 3.</emphasis>
<taxonomicName authorityName="Siti-Munirah &amp; Dome" authorityYear="2023" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">Thismia kenyirensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="61">A</emphasis>
bracts with ovary and pistil, lateral view
<emphasis bold="true" pageId="0" pageNumber="61">B</emphasis>
top view of stigma
<emphasis bold="true" pageId="0" pageNumber="61">C</emphasis>
flower, side view
<emphasis bold="true" pageId="0" pageNumber="61">D</emphasis>
longitudinal section of floral tube and stamens
<emphasis bold="true" pageId="0" pageNumber="61">E</emphasis>
flower, top view
<emphasis bold="true" pageId="0" pageNumber="61">F</emphasis>
annulus
<emphasis bold="true" pageId="0" pageNumber="61">G</emphasis>
stamens, outer view. Photos by Siti-Munirah MY (
<emphasis bold="true" pageId="0" pageNumber="61">A-C, E, F</emphasis>
; All from FRI 91122b); Dome Nikong (
<emphasis bold="true" pageId="0" pageNumber="61">D, G</emphasis>
; No specimens).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="additional specimen examined (">
<paragraph pageId="0" pageNumber="61">Additional specimen examined (paratype).</paragraph>
<paragraph pageId="0" pageNumber="61">Malaysia. Peninsular Malaysia: Terengganu, Hulu Terengganu District, Taman Negeri Kenyir, Sungai Cendana, elev. ca 204 m, 8 Sept 2020, Dome Nikong, FRI 91122b (KEP, spirit collection, barcode no. SC12016), FRI 91122c (KEP, spirit collection, barcode no. SC12017); Taman Negeri Kenyir, Sungai Cendana, elev. ca. 156 m, 16 June 2022, Siti-Munirah MY, FRI98678 (KEP, spirit collection, barcode no. SC12018).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="distribution">
<paragraph pageId="0" pageNumber="61">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="61">
Endemic to Terengganu, Peninsular Malaysia. Currently, the occurrence is only from the type locality, Sungai Cendana area (Map
<figureCitation captionStart="Map 1" captionStartId="F1" captionText="Map 1. Taman Negeri Kenyir, Terengganu (black circle), the type locality of Thismia kenyirensis." figureDoi="10.3897/phytokeys.221.98571.map1" httpUri="https://binary.pensoft.net/fig/821039" pageId="0" pageNumber="61">1</figureCitation>
). In addition,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
was observed in two other populations in the vicinity of Tasik Kenyir. It was sighted near Sungai Saok and on the way to Gunung Gagau (Wong Pui May personal communication), both in the Tasik Kenyir region. However, there are still no specimens and GPS information to verify these findings. This means that only the occurrence of the population in Sungai Cendana has been confirmed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="ecology">
<paragraph pageId="0" pageNumber="61">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="61">
In moist shady areas of lowland dipterocarp forest on moist soil at elevations of 150-220 m a.s.l. Flowering and fruiting mostly from September to April, but also recorded to flower in June. Historically, the type locality was botanized as early as 2007. Based on the results of
<bibRefCitation author="Lim, CL" journalOrPublisher="Rheedea" pageId="0" pageNumber="61" refId="B8" refString="Lim, CL, Chan, YC, Julius, A, Yao, TL, Chua, LSL, Kiew, R, 2008. A checklist of vascular plants for the Sungai Cicir Area, Tasik Kenyir, Terengganu, Peninsular Malaysia. In: Elagupillay S et al. (Eds) Proceedings of Seminar Biodiversiti Kebangsaan 2007. 27-29 November 2007, Kuala Lumpur, 135-154." title="A checklist of vascular plants for the Sungai Cicir Area, Tasik Kenyir, Terengganu, Peninsular Malaysia. In: Elagupillay S et al. (Eds) Proceedings of Seminar Biodiversiti Kebangsaan 2007. 27 - 29 November 2007, Kuala Lumpur, 135 - 154." year="2008">Lim et al. (2008)</bibRefCitation>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. alba" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="alba">
<emphasis italics="true" pageId="0" pageNumber="61">T. alba</emphasis>
</taxonomicName>
was also recorded during the survey in the Sungai Cendana area. This is in contrast to
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
, which was not mentioned, implying that it was not seen or collected at that time. During our recent visit, we were also able to find
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. alba" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="alba">
<emphasis italics="true" pageId="0" pageNumber="61">T. alba</emphasis>
</taxonomicName>
not far from the original population of
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="etymology">
<paragraph pageId="0" pageNumber="61">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="61">The epithet refers to Lake Kenyir, Kenyir State Park (Taman Negeri Kenyir), where the species was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="conservation status">
<paragraph pageId="0" pageNumber="61">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="61">
According to the IUCN standards (
<bibRefCitation author="Imhof, S" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="61" publicationUrl="http://www.iucnredlist.org/resources/redlistguidelines" refId="B4" refString="2022. . http://www.iucnredlist.org/resources/redlistguidelines" url="http://www.iucnredlist.org/resources/redlistguidelines" year="2022">IUCN 2022</bibRefCitation>
), we propose to classify the preliminary conservation status of the species as Least Concern (LC). This is because the species is likely to be widespread in TNK, where it has been sighted a few times at the type locality (Sungai Cendana) and also by other observers at two other sites, and its entire habitat is in a fully protected forest area. In addition, nearby forests in Terengganu National Park (Taman Negara Terengganu) are also fully protected. As mentioned earlier, it is very likely that there are other populations in the vicinity of Tasik Kenyir. According to the second
<normalizedToken originalValue="authors">author's</normalizedToken>
observations, four individuals were sighted near Sungai Saok and one on the way to Gunung Gagau on 22 November 2022. However, no specimens are available for confirmation.
<taxonomicName authorityName="Siti-Munirah &amp; Dome" authorityYear="2023" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">Thismia kenyirensis</emphasis>
</taxonomicName>
occurs in intact habitats in forested areas in TNK. However, the species is very common only during the flowering season and its populations are mostly scattered, as few individuals or solitary individuals were observed during the flowering season. During the first check of the type locality, the population in Sungai Cendana in 2020, about 20 adult individuals were found. However, during a revisit in 2022, only a single individual was observed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="61" type="notes">
<paragraph pageId="0" pageNumber="61">Notes.</paragraph>
<paragraph pageId="0" pageNumber="61">
<taxonomicName authorityName="Siti-Munirah &amp; Dome" authorityYear="2023" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">Thismia kenyirensis</emphasis>
</taxonomicName>
is easily recognised by the following combination of characteristics: vermiform roots, almost uniform light-to-dark orange flower coloration, petaloid ovate outer tepals, narrowly lanceolate-linear inner tepals with long appendages and each stamen with 3 appendages at its apex. Within the infrageneric classification by
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.319.3.2" author="Kumar, P" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="61" pagination="225 - 240" refId="B6" refString="Kumar, P, Gale, SW, Li, JH, Bouamanivong, S, Fischer, GA, 2017. Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos, and a key to subgeneric classification. Phytotaxa 319 (3): 225 - 240, DOI: https://doi.org/10.11646/phytotaxa.319.3.2" title="Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos, and a key to subgeneric classification." url="https://doi.org/10.11646/phytotaxa.319.3.2" volume="319" year="2017">Kumar et al. (2017)</bibRefCitation>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
resembles species of
<taxonomicName authority="Thismia" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia (Thismia) subsp. subgenus" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="subGenus" subGenus="Thismia">Thismia subgenus Thismia</taxonomicName>
section
<taxonomicName authority="Brunonithismia" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Thismia subsp. subsection" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="subSection" subSection="Brunonithismia">Thismia subsection Brunonithismia</taxonomicName>
Jonker, especially by its free and unequal tepals. Eight species are placed in
<taxonomicName authority="Brunonithismia" class="Liliopsida" family="Thismiaceae" genus="Thismia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. subsection" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Tracheophyta" rank="subSection" subSection="Brunonithismia">subsection Brunonithismia</taxonomicName>
by
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.319.3.2" author="Kumar, P" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="61" pagination="225 - 240" refId="B6" refString="Kumar, P, Gale, SW, Li, JH, Bouamanivong, S, Fischer, GA, 2017. Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos, and a key to subgeneric classification. Phytotaxa 319 (3): 225 - 240, DOI: https://doi.org/10.11646/phytotaxa.319.3.2" title="Thismia nigricoronata, a new species of Burmanniaceae (Thismieae, Dioscoreales) from Vang Vieng, Vientiane Province, Laos, and a key to subgeneric classification." url="https://doi.org/10.11646/phytotaxa.319.3.2" volume="319" year="2017">Kumar et al. (2017)</bibRefCitation>
:
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. arachnites" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="arachnites">
<emphasis italics="true" pageId="0" pageNumber="61">T. arachnites</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. brunonis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="brunonis">
<emphasis italics="true" pageId="0" pageNumber="61">T. brunonis</emphasis>
</taxonomicName>
Griff,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. gardneriana" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="gardneriana">
<emphasis italics="true" pageId="0" pageNumber="61">T. gardneriana</emphasis>
</taxonomicName>
Hook.f. ex Thwaites,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. hongkongensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="hongkongensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. hongkongensis</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. javanica" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="javanica">
<emphasis italics="true" pageId="0" pageNumber="61">T. javanica</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. labiata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="labiata">
<emphasis italics="true" pageId="0" pageNumber="61">T. labiata</emphasis>
</taxonomicName>
J.J.Sm.,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. neptunis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="neptunis">
<emphasis italics="true" pageId="0" pageNumber="61">T. neptunis</emphasis>
</taxonomicName>
Becc. (proved to be only distantly related to the other species of the subsection by
<bibRefCitation DOI="https://doi.org/10.1093/botlinnean/boaa017" author="Shepeleva, EA" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="61" pagination="287 - 315" refId="B12" refString="Shepeleva, EA, Schelkunov, MI, Hrones, M, Sochor, M, Dancak, M, Merckx, VSFT, Kikuchi, IABS, Chantanaorrapint, S, Suetsugu, K, Tsukaya, H, Mar, SS, Luu, HT, Li, H-Q, Logacheva, MD, Nuraliev, MS, 2020. Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits. Botanical Journal of the Linnean Society 193 (3): 287 - 315, DOI: https://doi.org/10.1093/botlinnean/boaa017" title="Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits." url="https://doi.org/10.1093/botlinnean/boaa017" volume="193" year="2020">Shepeleva et al. 2020</bibRefCitation>
) and
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. tentaculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="tentaculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. tentaculata</emphasis>
</taxonomicName>
K.Larsen &amp; Aver. Of them,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. arachnites" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="arachnites">
<emphasis italics="true" pageId="0" pageNumber="61">T. arachnites</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. javanica" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="javanica">
<emphasis italics="true" pageId="0" pageNumber="61">T. javanica</emphasis>
</taxonomicName>
occur in Peninsular Malaysia, including the state of Terengganu. They both are readily distinguished from
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F6" captionText="Figure 4. Morphologically close species to T. kenyirensis in Peninsular Malaysia A-C T. arachnites (Taman Negeri Kenyir) D-F T. javanica (Telemong Forest Reserve). Photos by Dome Nikong (A-C, F) and Siti-Munirah MY (D, E, G)." figureDoi="10.3897/phytokeys.221.98571.figure4" httpUri="https://binary.pensoft.net/fig/821044" pageId="0" pageNumber="61">4</figureCitation>
, Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison of T. kenyirensis with similar species T. arachnites (Chantanaorrapint 2018), T. bokorensis (Suetsugu et al. 2018), T. breviappendiculata (Tanaka et al. 2018), T. brunonis (Jonker 1938), T. gardneriana (Jonker 1938; Bandara et al. 2020), T. hongkongensis (Mar et al. 2015), T. javanica (Siti-Munirah and Dome 2019; Nuraliev et al. 2020) and T. tentaculata (Larsen and Averyanov 2007; Nuraliev et al. 2020)." httpUri="http://table.plazi.org/id/43591C2C7E9F8866CEDF1AB0FD5CC1D5" pageId="0" pageNumber="61" tableUuid="43591C2C7E9F8866CEDF1AB0FD5CC1D5">1</tableCitation>
). In addition,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. breviappendiculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="breviappendiculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. breviappendiculata</emphasis>
</taxonomicName>
Nob. Tanaka (
<bibRefCitation author="Tanaka, N" journalOrPublisher="Bulletin of the National Museum of Nature and Science, Series B, Botany" pageId="0" pageNumber="61" pagination="67 - 72" refId="B19" refString="Tanaka, N, Aung, MM, Latt, MM, 2018. Thismia breviappendiculata (Thismiaceae), a new mycoheterotrophic plant from southern Myanmar. Bulletin of the National Museum of Nature and Science, Series B, Botany 44: 67 - 72" title="Thismia breviappendiculata (Thismiaceae), a new mycoheterotrophic plant from southern Myanmar." volume="44" year="2018">Tanaka et al. 2018</bibRefCitation>
) from Myanmar,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. bokorensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="bokorensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. bokorensis</emphasis>
</taxonomicName>
Suetsugu &amp; Tsukaya (
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.334.1.10" author="Suetsugu, K" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="61" pagination="65 - 69" refId="B18" refString="Suetsugu, K, Tsukaya, H, Tagane, S, Chhang, P, Yukawa, T, Yahara, T, 2018. Flora of Bokor National Park VII: Thismia bokorensis (Burmanniaceae), a new species representing a new generic record. Phytotaxa 334 (1): 65 - 69, DOI: https://doi.org/10.11646/phytotaxa.334.1.10" title="Flora of Bokor National Park VII: Thismia bokorensis (Burmanniaceae), a new species representing a new generic record." url="https://doi.org/10.11646/phytotaxa.334.1.10" volume="334" year="2018">Suetsugu et al. 2018</bibRefCitation>
) from Cambodia,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. gardneriana" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="gardneriana">
<emphasis italics="true" pageId="0" pageNumber="61">T. gardneriana</emphasis>
</taxonomicName>
from Sri Lanka and
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. tentaculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="tentaculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. tentaculata</emphasis>
</taxonomicName>
from Hong Kong and Vietnam are similar to
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
. In a molecular phylogenetic reconstruction provided by
<bibRefCitation DOI="https://doi.org/10.1093/botlinnean/boaa017" author="Shepeleva, EA" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="61" pagination="287 - 315" refId="B12" refString="Shepeleva, EA, Schelkunov, MI, Hrones, M, Sochor, M, Dancak, M, Merckx, VSFT, Kikuchi, IABS, Chantanaorrapint, S, Suetsugu, K, Tsukaya, H, Mar, SS, Luu, HT, Li, H-Q, Logacheva, MD, Nuraliev, MS, 2020. Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits. Botanical Journal of the Linnean Society 193 (3): 287 - 315, DOI: https://doi.org/10.1093/botlinnean/boaa017" title="Phylogenetics of the mycoheterotrophic genus Thismia (Thismiaceae: Dioscoreales) with a focus on the Old World taxa: Delineation of novel natural groups and insights into the evolution of morphological traits." url="https://doi.org/10.1093/botlinnean/boaa017" volume="193" year="2020">Shepeleva et al. (2020)</bibRefCitation>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. javanica" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="javanica">
<emphasis italics="true" pageId="0" pageNumber="61">T. javanica</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. hongkongensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="hongkongensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. hongkongensis</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. gardneriana" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="gardneriana">
<emphasis italics="true" pageId="0" pageNumber="61">T. gardneriana</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. tentaculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="tentaculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. tentaculata</emphasis>
</taxonomicName>
occupied a relatively isolated and unstable position. All these species (and also
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
) are characterised by free appendaged inner tepals and outer tepals without appendages. A more detailed phylogenetic analysis is required to test the monophyly of this morphological group, and the relationships of each of these species including
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/phytokeys.221.98571.figure4" httpUri="https://binary.pensoft.net/fig/821044" pageId="0" pageNumber="61" start="Figure 4" startId="F6">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Figure 4.</emphasis>
Morphologically close species to
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
in Peninsular Malaysia
<emphasis bold="true" pageId="0" pageNumber="61">A-C</emphasis>
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. arachnites" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="arachnites">
<emphasis italics="true" pageId="0" pageNumber="61">T. arachnites</emphasis>
</taxonomicName>
(Taman Negeri Kenyir)
<emphasis bold="true" pageId="0" pageNumber="61">D-F</emphasis>
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. javanica" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="javanica">
<emphasis italics="true" pageId="0" pageNumber="61">T. javanica</emphasis>
</taxonomicName>
(Telemong Forest Reserve). Photos by Dome Nikong (
<emphasis bold="true" pageId="0" pageNumber="61">A-C, F</emphasis>
) and Siti-Munirah MY (
<emphasis bold="true" pageId="0" pageNumber="61">D, E, G</emphasis>
).
</paragraph>
</caption>
<caption ID-Table-UUID="43591C2C7E9F8866CEDF1AB0FD5CC1D5" httpUri="http://table.plazi.org/id/43591C2C7E9F8866CEDF1AB0FD5CC1D5" pageId="0" pageNumber="61" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="61">
<emphasis bold="true" pageId="0" pageNumber="61">Table 1.</emphasis>
Morphological comparison of
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
with similar species
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. arachnites" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="arachnites">
<emphasis italics="true" pageId="0" pageNumber="61">T. arachnites</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1007/s12225-018-9765-2" author="Chantanaorrapint, S" journalOrPublisher="Kew Bulletin" pageId="0" pageNumber="61" pagination=": 43" refId="B2" refString="Chantanaorrapint, S, 2018. Lectotypification of Thismia arachnites (Thismiaceae), a mysterious species newly reported for Thailand. Kew Bulletin 73 (3): 43, DOI: https://doi.org/10.1007/s12225-018-9765-2" title="Lectotypification of Thismia arachnites (Thismiaceae), a mysterious species newly reported for Thailand." url="https://doi.org/10.1007/s12225-018-9765-2" volume="73" year="2018">Chantanaorrapint 2018</bibRefCitation>
),
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. bokorensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="bokorensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. bokorensis</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.334.1.10" author="Suetsugu, K" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="61" pagination="65 - 69" refId="B18" refString="Suetsugu, K, Tsukaya, H, Tagane, S, Chhang, P, Yukawa, T, Yahara, T, 2018. Flora of Bokor National Park VII: Thismia bokorensis (Burmanniaceae), a new species representing a new generic record. Phytotaxa 334 (1): 65 - 69, DOI: https://doi.org/10.11646/phytotaxa.334.1.10" title="Flora of Bokor National Park VII: Thismia bokorensis (Burmanniaceae), a new species representing a new generic record." url="https://doi.org/10.11646/phytotaxa.334.1.10" volume="334" year="2018">Suetsugu et al. 2018</bibRefCitation>
),
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. breviappendiculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="breviappendiculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. breviappendiculata</emphasis>
</taxonomicName>
(
<bibRefCitation author="Tanaka, N" journalOrPublisher="Bulletin of the National Museum of Nature and Science, Series B, Botany" pageId="0" pageNumber="61" pagination="67 - 72" refId="B19" refString="Tanaka, N, Aung, MM, Latt, MM, 2018. Thismia breviappendiculata (Thismiaceae), a new mycoheterotrophic plant from southern Myanmar. Bulletin of the National Museum of Nature and Science, Series B, Botany 44: 67 - 72" title="Thismia breviappendiculata (Thismiaceae), a new mycoheterotrophic plant from southern Myanmar." volume="44" year="2018">Tanaka et al. 2018</bibRefCitation>
),
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. brunonis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="brunonis">
<emphasis italics="true" pageId="0" pageNumber="61">T. brunonis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Jonker, FP" journalOrPublisher="Mededeelinger van het Botanisch Museum en Herbarium van de Rijks Universiteit te Utrecht" pageId="0" pageNumber="61" pagination="1 - 279" refId="B5" refString="Jonker, FP, 1938. A monograph of the Burmanniaceae. Mededeelinger van het Botanisch Museum en Herbarium van de Rijks Universiteit te Utrecht 51: 1 - 279" title="A monograph of the Burmanniaceae." volume="51" year="1938">Jonker 1938</bibRefCitation>
),
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. gardneriana" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="gardneriana">
<emphasis italics="true" pageId="0" pageNumber="61">T. gardneriana</emphasis>
</taxonomicName>
(
<bibRefCitation author="Jonker, FP" journalOrPublisher="Mededeelinger van het Botanisch Museum en Herbarium van de Rijks Universiteit te Utrecht" pageId="0" pageNumber="61" pagination="1 - 279" refId="B5" refString="Jonker, FP, 1938. A monograph of the Burmanniaceae. Mededeelinger van het Botanisch Museum en Herbarium van de Rijks Universiteit te Utrecht 51: 1 - 279" title="A monograph of the Burmanniaceae." volume="51" year="1938">Jonker 1938</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.436.1.5" author="Bandara, C" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="61" pagination="55 - 62" refId="B1" refString="Bandara, C, Priyankara, T, Atthanagoda, AG, Lakkana, T, Ediriweera, S, Kumar, P, 2020. Gastrodia gunatillekeorum (Gastrodieae, Epidendroideae, Orchidaceae), a new species from a lowland rainforest of Sri Lanka. Phytotaxa 436 (1): 55 - 62, DOI: https://doi.org/10.11646/phytotaxa.436.1.5" title="Gastrodia gunatillekeorum (Gastrodieae, Epidendroideae, Orchidaceae), a new species from a lowland rainforest of Sri Lanka." url="https://doi.org/10.11646/phytotaxa.436.1.5" volume="436" year="2020">Bandara et al. 2020</bibRefCitation>
),
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. hongkongensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="hongkongensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. hongkongensis</emphasis>
</taxonomicName>
(Mar et al. 2015),
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. javanica" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="javanica">
<emphasis italics="true" pageId="0" pageNumber="61">T. javanica</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.124.34728" author="Siti-Munirah, MY" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="61" pagination="123 - 137" refId="B13" refString="Siti-Munirah, MY, Dome, N, 2019. Thismia domei and T. terengganuensis (Thismiaceae), two new species, and T. javanica, a new record from Terengganu, Peninsular Malaysia. PhytoKeys 124: 123 - 137, DOI: https://doi.org/10.3897/phytokeys.124.34728" title="Thismia domei and T. terengganuensis (Thismiaceae), two new species, and T. javanica, a new record from Terengganu, Peninsular Malaysia." url="https://doi.org/10.3897/phytokeys.124.34728" volume="124" year="2019">Siti-Munirah and Dome 2019</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.441.3.1" author="Nuraliev, MS" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="61" pagination="229 - 250" refId="B10" refString="Nuraliev, MS, Yudina, SV, Truong, BV, Do, TX, Luu, HT, Kuznetsov, AN, Kuznetsova, SP, 2020. A revision of the family Thismiaceae (Dioscoreales) in Cambodia, Laos and Vietnam. Phytotaxa 441 (3): 229 - 250, DOI: https://doi.org/10.11646/phytotaxa.441.3.1" title="A revision of the family Thismiaceae (Dioscoreales) in Cambodia, Laos and Vietnam." url="https://doi.org/10.11646/phytotaxa.441.3.1" volume="441" year="2020">Nuraliev et al. 2020</bibRefCitation>
) and
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. tentaculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="tentaculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. tentaculata</emphasis>
</taxonomicName>
(
<bibRefCitation author="Larsen, K" journalOrPublisher="Rheedea" pageId="0" pageNumber="61" pagination="13 - 19" refId="B7" refString="Larsen, K, Averyanov, LV, 2007. Thismia annamensis and Thismia tentaculata, two new species of Thismiaceae from central Vietnam. Rheedea 17 (1-2): 13 - 19" title="Thismia annamensis and Thismia tentaculata, two new species of Thismiaceae from central Vietnam." volume="17" year="2007">Larsen and Averyanov 2007</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.441.3.1" author="Nuraliev, MS" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="61" pagination="229 - 250" refId="B10" refString="Nuraliev, MS, Yudina, SV, Truong, BV, Do, TX, Luu, HT, Kuznetsov, AN, Kuznetsova, SP, 2020. A revision of the family Thismiaceae (Dioscoreales) in Cambodia, Laos and Vietnam. Phytotaxa 441 (3): 229 - 250, DOI: https://doi.org/10.11646/phytotaxa.441.3.1" title="A revision of the family Thismiaceae (Dioscoreales) in Cambodia, Laos and Vietnam." url="https://doi.org/10.11646/phytotaxa.441.3.1" volume="441" year="2020">Nuraliev et al. 2020</bibRefCitation>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="61">
<table pageId="0" pageNumber="61">
<tr pageId="0" pageNumber="61">
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. kenyirensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="kenyirensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. kenyirensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. arachnites" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="arachnites">
<emphasis italics="true" pageId="0" pageNumber="61">T. arachnites</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. bokorensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="bokorensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. bokorensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. breviappendiculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="breviappendiculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. breviappendiculata</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. brunonis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="brunonis">
<emphasis italics="true" pageId="0" pageNumber="61">T. brunonis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. gardneriana" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="gardneriana">
<emphasis italics="true" pageId="0" pageNumber="61">T. gardneriana</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. hongkongensis" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="hongkongensis">
<emphasis italics="true" pageId="0" pageNumber="61">T. hongkongensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. javanica" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="javanica">
<emphasis italics="true" pageId="0" pageNumber="61">T. javanica</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="61" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Thismiaceae" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="T. tentaculata" order="Dioscoreales" pageId="0" pageNumber="61" phylum="Magnoliophyta" rank="species" species="tentaculata">
<emphasis italics="true" pageId="0" pageNumber="61">T. tentaculata</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Colour of floral tube</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Orange to brownish-orange with 12 darker orange longitudinal ribs</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">White to orange with 6 or 12 reddish vertical streaks in the upper part</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Pure white</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">White, with 12 translucent longitudinal ridges</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Yellowish</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Yellowish-orange</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Pinkish-white with 12 dark red vertical ribs</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">White to pale orange or light orangish-red</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Pure white, sometimes with antetepalous veins indistinctly tinged with red</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Presence of transverse bars/ ornamentation on inner side of the floral tube</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Absent/absent</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Present/absent</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Absent/absent</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Present/absent</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Present/absent</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Absent/ Not known</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Absent/present</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Present/absent</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Absent/present</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Outer tepal Shape</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Ovate, petaloid</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Broadly ovate</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Broadly triangular, apex broadly obtuse or rounded</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Ovate, obtuse</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Broadly ovate, obtuse</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Broad, rotundate</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Triangular</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Ovate-triangular</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Isosceles triangular, broadly obtuse or rounded</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Length (mm)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">5-6.5</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.2</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Not known</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.5</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.8</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1-3.5</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.8-2.4</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Colour</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Bright orange</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Yellow to orange.</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Light yellow</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Yellow</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Not known</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Yellow</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Dark red</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Orange to red or yellow, translucent</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Light yellow or tinged with red, translucent</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Inner tepal (without appendage) Shape</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Narrowly lanceolate-linear</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Triangular</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Narrowly triangular</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Isosceles triangular</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Triangular at the base, caudate in long, thick, filiform tails</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Subulate</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Spatulate, adaxially concave</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Narrowly triangular</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Triangular with broad base</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Length (mm)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">6.5-9</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">2.6</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Not known</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Not known</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">3.3</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">1.5-3.3</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="61" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Colour</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Bright orange</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Orange to reddish-orange at the base</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Light yellow</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Yellow</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Not known</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Yellowish-orange</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Dark red</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Orange to red or yellow and usually dark red towards apex</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Light yellow</td>
</tr>
<tr pageId="0" pageNumber="61">
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="61">Stamen Apex</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">3 long, slightly curved appendages</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">3-toothed, each tooth bearing a distinct stiff hair</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">3-toothed, each tooth narrowly triangular, with ca. 1 mm long</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">Lobed</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">With numerous teeth</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">2-toothed, each tapering into a stiff hair</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">2-toothed, adorned with trichomes</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">3-toothed, each tooth bearing hair</td>
<td colspan="1" pageId="0" pageNumber="61" rowspan="1">2-toothed, each tooth tapering into hair, sometimes with additional hairs between teeth</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>