183 lines
21 KiB
XML
183 lines
21 KiB
XML
<document id="9C690238D218F094E423E2E0E0F3F848" ID="10.11646/zootaxa.4175.4.1" ID-DOI="10.11646/zootaxa.4175.4.1" ID-GBIF-Dataset="55c6a2a3-cf2b-41df-bb10-ee4ba1d423c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="256776" ID-ZooBank="5C11185B-E219-4214-8A01-517FBB8C99BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476684690294" checkinUser="plazi" docAuthor="Ruta, Rafał" docDate="2016" docId="A8548C3CFFDFFFF5C7F3FCF4FC98FE0A" docLanguage="en" docName="zootaxa.4175.4.1.pdf" docOrigin="Zootaxa 4175 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anticyphon santanderensis Ruta, 2016, sp. nov." docType="treatment" docUuid="86A42E98-9E25-43AF-938E-44E76BFC5A22" docUuidSource="ZooBank" docVersion="8" lastPageNumber="315" masterDocId="546DF444FFD1FFFBC764FFB0FFBDF965" masterDocTitle="Anticyphon gen. nov., a new genus of Scirtidae (Coleoptera: Scirtoidea) inhabiting high altitude Andean cloud forests and páramo formation" masterLastPageNumber="318" masterPageNumber="301" pageNumber="315" updateTime="1698389907452" updateUser="plazi">
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<mods:title id="5F6D505DB7D212670B7E07739C058358">Anticyphon gen. nov., a new genus of Scirtidae (Coleoptera: Scirtoidea) inhabiting high altitude Andean cloud forests and páramo formation</mods:title>
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<mods:name id="0957CA7962E33489A3277B33A5716849" type="personal">
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<mods:namePart id="7755FCAC23A9665AEAAECDC06BB371D2">Ruta, Rafał</mods:namePart>
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<mods:typeOfResource id="BB598380C1604ED39F70CA88048B727D">text</mods:typeOfResource>
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<mods:title id="5715F1F1C2F0AA5B952041D26B77A33F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F4BA7AC129751E2260C390BBF1AFA2BC">
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<mods:date id="C7B3A34291145836530DAD82EC8B6B34">2016</mods:date>
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<mods:detail id="9CBCCB07DCFBD40C7FCDEE7D6A7154AE" type="volume">
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<mods:number id="B02D4D2331D012C0542BDDCD4E17F7E1">4175</mods:number>
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<mods:number id="DB733EAA2968DD43262560BB978F64A0">4</mods:number>
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<mods:classification id="6F3569E3AC891484C6E467B47A986519">journal article</mods:classification>
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<mods:identifier id="21431042F4F0D11BB70FE25B6C8D7559" type="DOI">10.11646/zootaxa.4175.4.1</mods:identifier>
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<mods:identifier id="332B582B3C79BB801CCE66262839E506" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A8548C3CFFDFFFF5C7F3FCF4FC98FE0A" ID-DOI="http://doi.org/10.5281/zenodo.6060619" ID-GBIF-Taxon="124996808" ID-Zenodo-Dep="6060619" LSID="urn:lsid:zoobank.org:act:86A42E98-9E25-43AF-938E-44E76BFC5A22" httpUri="http://treatment.plazi.org/id/A8548C3CFFDFFFF5C7F3FCF4FC98FE0A" lastPageNumber="315" pageId="14" pageNumber="315">
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<subSubSection id="68E76EA1FFDFFFF5C7F3FCF4FE90FA1A" pageId="14" pageNumber="315" type="nomenclature">
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<paragraph id="20423D2AFFDFFFF5C7F3FCF4FDF9FA3B" blockId="14.[151,580,836,895]" box="[151,580,836,862]" pageId="14" pageNumber="315">
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<heading id="7B0A8A46FFDFFFF5C7F3FCF4FDF9FA3B" bold="true" box="[151,580,836,862]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
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<emphasis id="1289E138FFDFFFF5C7F3FCF4FDF9FA3B" bold="true" box="[151,580,836,862]" pageId="14" pageNumber="315">
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<taxonomicName id="E7FD46A9FFDFFFF5C7F3FCF4FE63FA3B" box="[151,478,836,862]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="santanderensis" status="sp. nov.">
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<emphasis id="1289E138FFDFFFF5C7F3FCF4FE63FA3B" bold="true" box="[151,478,836,862]" italics="true" pageId="14" pageNumber="315">Anticyphon santanderensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="09BA5C43FFDFFFF5C682FCF4FDF9FA3B" box="[486,580,836,862]" pageId="14" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="20423D2AFFDFFFF5C7F3FCD6FE90FA1A" blockId="14.[151,580,836,895]" box="[151,301,870,895]" pageId="14" pageNumber="315">
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(
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<figureCitation id="B8C621AFFFDFFFF5C7FBFCD6FF5FFA1A" box="[159,226,870,895]" captionStart="FIGURE 1" captionStartId="2.[151,250,1122,1144]" captionTargetBox="[151,1436,193,1099]" captionTargetId="figure@2.[151,1436,193,1101]" captionTargetPageId="2" captionText="FIGURE 1. Anticyphon gen. nov., habitus. A) A. davidsoni sp. nov., B) A. ecuadorensis sp. nov., C) A. oyonensis sp. nov., female, D) A. oyonensis sp. nov., male E) A. paramoensis sp. nov., light form, F) A. paramoensis sp. nov., dark form, G) A. peruvianus sp. nov., H) A. santanderensis sp. nov. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/256777/files/figure.png" pageId="14" pageNumber="315">Figs 1</figureCitation>
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H, 13)
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</paragraph>
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</subSubSection>
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<subSubSection id="68E76EA1FFDFFFF5C7F3FC1EFA21FAA2" box="[151,1436,942,967]" pageId="14" pageNumber="315" type="materials_examined">
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<paragraph id="20423D2AFFDFFFF5C7F3FC1EFA21FAA2" blockId="14.[151,1437,942,1903]" box="[151,1436,942,967]" pageId="14" pageNumber="315">
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<emphasis id="1289E138FFDFFFF5C7F3FC1EFEF9FAA2" bold="true" box="[151,324,942,967]" pageId="14" pageNumber="315">Type material.</emphasis>
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Holotype, male (NHMB): “Bucaramanga Nord \ Santandar
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\ Columbien \
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<date id="54431BEAFFDFFFF5C3BFFC1EFA85FAA2" box="[1243,1336,942,967]" pageId="14" pageNumber="315" value="1960-11">XI.1960</date>
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G. Frey”
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</paragraph>
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</subSubSection>
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<subSubSection id="68E76EA1FFDFFFF5C7A3FC62FA93FD56" pageId="14" pageNumber="315" type="diagnosis">
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<paragraph id="20423D2AFFDFFFF5C7A3FC62FA93FD56" blockId="14.[151,1437,942,1903]" pageId="14" pageNumber="315">
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<emphasis id="1289E138FFDFFFF5C7A3FC62FEFCFA8E" bold="true" box="[199,321,978,1003]" pageId="14" pageNumber="315">Diagnosis.</emphasis>
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The second largest species of the genus, pronotal disc with normal, not granulate punctures. Resembling
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<taxonomicName id="E7FD46A9FFDFFFF5C647FC47FE12FD6B" box="[291,431,1014,1038]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="davidsoni" status="sp. nov.">
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<emphasis id="1289E138FFDFFFF5C647FC47FE12FD6B" box="[291,431,1014,1038]" italics="true" pageId="14" pageNumber="315">A. davidsoni</emphasis>
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</taxonomicName>
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<emphasis id="1289E138FFDFFFF5C6D1FC46FDB3FD6A" bold="true" box="[437,526,1014,1039]" pageId="14" pageNumber="315">
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<taxonomicNameLabel id="09BA5C43FFDFFFF5C6D1FC46FDB3FD6A" box="[437,526,1014,1039]" pageId="14" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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in size and pronotal punctation. It can be identified on the basis of male genitalia: trigonium is distinctly shorter than parameroids (almost as long as parameroids in
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<taxonomicName id="E7FD46A9FFDFFFF5C355FBABFB00FD56" box="[1073,1213,1051,1075]" class="Insecta" family="Scirtidae" genus="Anticyphon" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="davidsoni" status="sp. nov.">
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<emphasis id="1289E138FFDFFFF5C355FBABFB00FD56" box="[1073,1213,1051,1075]" italics="true" pageId="14" pageNumber="315">A. davidsoni</emphasis>
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</taxonomicName>
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<emphasis id="1289E138FFDFFFF5C3A0FBAAFAA0FD56" bold="true" box="[1220,1309,1050,1075]" pageId="14" pageNumber="315">
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<taxonomicNameLabel id="09BA5C43FFDFFFF5C3A0FBAAFAA0FD56" box="[1220,1309,1050,1075]" pageId="14" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="68E76EA1FFDFFFF5C7A3FB8EFD05FE42" pageId="14" pageNumber="315" type="description">
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<emphasis id="1289E138FFDFFFF5C7A3FB8EFEE4FD32" bold="true" box="[199,345,1086,1111]" pageId="14" pageNumber="315">Description.</emphasis>
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Male. Body oval, moderately large, slightly flattened, clothed with brownish procumbent setae. Colouration of dorsum dark brown, antennae and legs dark brown. Head small, 1.7× wider than interocular space, covered with subtle granulate punctures; eyes relatively big, protuberant; frons with two shallow depressions. Mandibles without denticles on inner margins. Antennae as in genus description. Pronotum small, transversely rectangular, sides slightly curved, widest at posterior angles, anterolateral angles broadly rounded, not produced; disc moderately convex. Punctation of pronotum subtly granulate on lateral portions, very subtle and sparse in central portion. Pronotum without distinct pits along basal margin. Each elytron with 3 well marked longitudinal carinae. Elytral punctation irregular, relatively sparse; punctures shallow, separated by ca. 1.0 diameter. Penis (
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A) large (L
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,
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mm), parameroids long, diverging apically, pointed at apices; trigonium subtriangular, without lateral processes, 2× shorter than parameroids, pala slightly longer than parameroids, narrow, with membranous dorsal processes of penis; tegmen (
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<figureCitation id="B8C621AFFFDFFFF5C40CFA16FC01FCDA" box="[872,956,1446,1471]" captionStart="FIGURE 13" captionStartId="15.[151,250,1090,1112]" captionTargetBox="[160,1423,176,1067]" captionTargetId="figure@15.[160,1425,131,1068]" captionTargetPageId="15" captionText="FIGURE 13. Anticyphon santanderensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256788/files/figure.png" pageId="14" pageNumber="315">Fig. 13</figureCitation>
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B) large (L
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W
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mm), with narrow, subtriangular parameres, pointed at apices; sternite VIII (
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<figureCitation id="B8C621AFFFDFFFF5C40AFA7AFC00FC86" box="[878,957,1482,1507]" captionStart="FIGURE 13" captionStartId="15.[151,250,1090,1112]" captionTargetBox="[160,1423,176,1067]" captionTargetId="figure@15.[160,1425,131,1068]" captionTargetPageId="15" captionText="FIGURE 13. Anticyphon santanderensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256788/files/figure.png" pageId="14" pageNumber="315">Fig. 13</figureCitation>
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C) small (L
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<quantity id="E70590CFFFDFFFF5C320FA7BFB17FC87" box="[1092,1194,1483,1507]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="14" pageNumber="315" unit="mm" value="0.27">0.27 mm</quantity>
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W
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mm), widely V-shaped, with sparse setae in apical portions; sternite IX (
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<figureCitation id="B8C621AFFFDFFFF5C45EFA5EFC36FF62" box="[826,907,1518,1543]" captionStart="FIGURE 13" captionStartId="15.[151,250,1090,1112]" captionTargetBox="[160,1423,176,1067]" captionTargetId="figure@15.[160,1425,131,1068]" captionTargetPageId="15" captionText="FIGURE 13. Anticyphon santanderensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256788/files/figure.png" pageId="14" pageNumber="315">Fig. 13</figureCitation>
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D) relatively small (L
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,
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W
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mm), consisting of two subtriangular hemisternites, lightly sclerotized, with setose apical portion; tergite VIII (
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<figureCitation id="B8C621AFFFDFFFF5C255F9A2FA3CFF4E" box="[1329,1409,1554,1579]" captionStart="FIGURE 13" captionStartId="15.[151,250,1090,1112]" captionTargetBox="[160,1423,176,1067]" captionTargetId="figure@15.[160,1425,131,1068]" captionTargetPageId="15" captionText="FIGURE 13. Anticyphon santanderensis sp. nov., male genitalia. A) penis, B) tegmen, C) sternite VIII, D) sternite IX, E) tergite VIII, F) tergite IX. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/256788/files/figure.png" pageId="14" pageNumber="315">Fig. 13</figureCitation>
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E) (L
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,
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W
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mm) with transversely rectangular apical plate, apical portion darker, covered with microsetae, apical margin with row of dense, short setae intermixed with sparse, longer ones, basal portion of apical plate with transverse rod, apodemes shorter than apical portion; tergite IX (
|
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F) (L
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,
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W
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mm) distinctly narrower than tergite VIII, central portion lightly sclerotized.
|
||
</paragraph>
|
||
<paragraph id="20423D2AFFDFFFF5C7A3F976FE20FFBA" blockId="14.[151,1437,942,1903]" box="[199,413,1734,1759]" pageId="14" pageNumber="315">Female. Unknown.</paragraph>
|
||
<paragraph id="20423D2AFFDFFFF5C7A3F95AFD05FE42" blockId="14.[151,1437,942,1903]" pageId="14" pageNumber="315">
|
||
<emphasis id="1289E138FFDFFFF5C7A3F95AFE43FE66" bold="true" box="[199,510,1770,1795]" pageId="14" pageNumber="315">Measurements and ratios.</emphasis>
|
||
Male (n=1): TL
|
||
<quantity id="E70590CFFFDFFFF5C5DAF95AFCABFE67" box="[702,790,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="14" pageNumber="315" unit="mm" value="6.5">6.5 mm</quantity>
|
||
, PL
|
||
<quantity id="E70590CFFFDFFFF5C42FF95AFC0CFE67" box="[843,945,1770,1795]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="315" unit="mm" value="0.95">0.95 mm</quantity>
|
||
, PW
|
||
<quantity id="E70590CFFFDFFFF5C48BF95BFBE8FE67" box="[1007,1109,1771,1795]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="315" unit="mm" value="2.1">2.10 mm</quantity>
|
||
, EL
|
||
<quantity id="E70590CFFFDFFFF5C3EFF95AFB4FFE67" box="[1163,1266,1770,1795]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="14" pageNumber="315" unit="mm" value="5.7">5.70 mm</quantity>
|
||
, EW
|
||
<quantity id="E70590CFFFDFFFF5C255F95BFA2AFE67" box="[1329,1431,1771,1795]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="315" unit="mm" value="3.8">3.80 mm</quantity>
|
||
, TL/EW 1.7, PW/PL 2.2, EL/EW 1.5, EL/PL 6.0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68E76EA1FFDFFFF5C7A3F882FC6CFE2E" box="[199,977,1842,1867]" pageId="14" pageNumber="315" type="distribution">
|
||
<paragraph id="20423D2AFFDFFFF5C7A3F882FC6CFE2E" blockId="14.[151,1437,942,1903]" box="[199,977,1842,1867]" pageId="14" pageNumber="315">
|
||
<emphasis id="1289E138FFDFFFF5C7A3F882FEDDFE2E" bold="true" box="[199,352,1842,1867]" pageId="14" pageNumber="315">Distribution.</emphasis>
|
||
<collectingCountry id="58EA7DBAFFDFFFF5C603F882FE65FE2E" box="[359,472,1842,1867]" name="Colombia" pageId="14" pageNumber="315">Colombia</collectingCountry>
|
||
, known only from the
|
||
<typeStatus id="FF468388FFDFFFF5C5B2F883FCBBFE2E" box="[726,774,1843,1867]" pageId="14" pageNumber="315">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="B8C621AFFFDFFFF5C415F882FC79FE2E" box="[881,964,1842,1867]" captionStart="FIGURE 14" captionStartId="16.[151,250,1430,1452]" captionTargetBox="[341,1251,195,1398]" captionTargetId="figure@16.[331,1251,195,1398]" captionTargetPageId="16" captionText="FIGURE 14. Distribution of Anticyphon gen. nov. Please note that precise locality of A. peruvianus sp. nov. is unknown (see note under the species description)." httpUri="https://zenodo.org/record/256789/files/figure.png" pageId="14" pageNumber="315">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68E76EA1FFDFFFF5C7A3F8E6FC98FE0A" box="[199,805,1878,1903]" pageId="14" pageNumber="315" type="etymology">
|
||
<paragraph id="20423D2AFFDFFFF5C7A3F8E6FC98FE0A" blockId="14.[151,1437,942,1903]" box="[199,805,1878,1903]" pageId="14" pageNumber="315">
|
||
<emphasis id="1289E138FFDFFFF5C7A3F8E6FEF3FE0A" bold="true" box="[199,334,1878,1903]" pageId="14" pageNumber="315">Etymology.</emphasis>
|
||
After
|
||
<collectingRegion id="E239F3C8FFDFFFF5C6F2F8E6FD36FE0A" box="[406,651,1878,1903]" country="Colombia" name="Santander" pageId="14" pageNumber="315">Santander department</collectingRegion>
|
||
in
|
||
<collectingCountry id="58EA7DBAFFDFFFF5C5CBF8E6FC9FFE0A" box="[687,802,1878,1903]" name="Colombia" pageId="14" pageNumber="315">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |