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<document id="2FCD01A0FE84876F1A09A8E4F946278E" ID-CLB-Dataset="27896" ID-DOI="10.11646/zootaxa.4526.1.8" ID-GBIF-Dataset="6873630c-26b6-487c-84a4-d4605a4ec073" ID-ISSN="1175-5326" ID-Zenodo-Dep="2611464" ID-ZooBank="0C1BB4E3-2372-45A5-97B0-FE3D4307A3A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553697198732" checkinUser="plazi" docAuthor="Silva, Fabio Laurindo Da &amp; Song, Chao" docDate="2018" docId="A817EE29FFAF0F2235F8D9050704A9AF" docLanguage="en" docName="zootaxa.4526.1.8.pdf" docOrigin="Zootaxa 4526 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oleia fieldsi Silva &amp; Song 2018" docType="treatment" docVersion="8" lastPageNumber="98" masterDocId="542E9651FFAE0F20356FDC6B0249AF09" masterDocTitle="Oleia fieldsi sp. n., a new Orthocladiinae (Diptera: Chironomidae) species from the Brazilian Atlantic rainforest" masterLastPageNumber="100" masterPageNumber="96" pageNumber="97" updateTime="1698714210208" updateUser="ExternalLinkService">
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<mods:title id="24C3B0079ACE1E4E6D106027CE7FAC95">Oleia fieldsi sp. n., a new Orthocladiinae (Diptera: Chironomidae) species from the Brazilian Atlantic rainforest</mods:title>
</mods:titleInfo>
<mods:name id="BCA32F2C19D28512EC73DAD36EA58CE2" type="personal">
<mods:role id="83DF2D567773EADE3AAFDB33FB9D437B">
<mods:roleTerm id="136A3CDAEDA1E6A12EAAEE31489434EC">Author</mods:roleTerm>
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<mods:namePart id="164E2B11CC05A855DBE41C327483CA47">Silva, Fabio Laurindo Da</mods:namePart>
</mods:name>
<mods:name id="F3FF4659894AF3D91F42E36495897BEF" type="personal">
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<mods:namePart id="3B4B2ED500C8BFF931786D32EDF4BF46">Song, Chao</mods:namePart>
</mods:name>
<mods:typeOfResource id="6377E4F31097AA8A8E6DD6BF146F1C4C">text</mods:typeOfResource>
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<mods:title id="9A6D8DE64D5CBB0CF70632C73FAFF1FA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A7C6384583A8CD40E8067F5141FF88BA">
<mods:date id="2A3D29A5836AE276E4EBE27D976E1C27">2018</mods:date>
<mods:detail id="AA2A65ECB9F90E21EC3B34F132D1EE2C" type="pubDate">
<mods:number id="2B5D7017E2AEE8177FB7F32FDDDBCFA7">2018-11-28</mods:number>
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<mods:detail id="5831753A792A6763A443F60E452AD634" type="volume">
<mods:number id="6DF1A1AB69D092F69A0D664BCBC2F16D">4526</mods:number>
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<mods:number id="03EB495B388CA50B7E621E48FFC05C44">1</mods:number>
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<mods:classification id="6233E9ABDCDEB6F3485EF992E8A4134A">journal article</mods:classification>
<mods:identifier id="2EAF05AE25CAF6B7036F4B054D128CD5" type="CLB-Dataset">27896</mods:identifier>
<mods:identifier id="421768BB095F4CD7A0BAA2318D4496E3" type="DOI">10.11646/zootaxa.4526.1.8</mods:identifier>
<mods:identifier id="5EBC987C80DDEF22CDC69E9B15F40E97" type="GBIF-Dataset">6873630c-26b6-487c-84a4-d4605a4ec073</mods:identifier>
<mods:identifier id="13CC25E667E262AA454AB534A5A2998A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="84C84058E4413EB277D83C1BF7FC70C0" type="Zenodo-Dep">2611464</mods:identifier>
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</mods:mods>
<treatment id="A817EE29FFAF0F2235F8D9050704A9AF" ID-DOI="http://doi.org/10.5281/zenodo.6492174" ID-GBIF-Taxon="154507532" ID-Zenodo-Dep="6492174" LSID="urn:lsid:plazi:treatment:A817EE29FFAF0F2235F8D9050704A9AF" httpUri="http://treatment.plazi.org/id/A817EE29FFAF0F2235F8D9050704A9AF" lastPageId="2" lastPageNumber="98" pageId="1" pageNumber="97">
<subSubSection id="68A40CB4FFAF0F2135F8D9050053AA81" box="[151,538,1390,1416]" pageId="1" pageNumber="97" type="nomenclature">
<paragraph id="20015F3FFFAF0F2135F8D9050053AA81" blockId="1.[151,538,1390,1450]" box="[151,538,1390,1416]" pageId="1" pageNumber="97">
<heading id="7B49E853FFAF0F2135F8D9050053AA81" bold="true" box="[151,538,1390,1416]" fontSize="11" level="1" pageId="1" pageNumber="97" reason="1">
<emphasis id="12CA832DFFAF0F2135F8D9050053AA81" bold="true" box="[151,538,1390,1416]" pageId="1" pageNumber="97">
<taxonomicName id="E7BE24BCFFAF0F2135F8D9050327AA81" authority="Silva" authorityName="Silva &amp; Song" authorityYear="2018" box="[151,366,1390,1416]" class="Insecta" family="Chironomidae" genus="Oleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="97" phylum="Arthropoda" rank="species" species="fieldsi">
<emphasis id="12CA832DFFAF0F2135F8D9050361AA81" bold="true" box="[151,296,1390,1416]" italics="true" pageId="1" pageNumber="97">Oleia fieldsi</emphasis>
Silva
</taxonomicName>
et Song sp. n.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="68A40CB4FFAF0F2135F8D9FA02A2AAA3" box="[151,235,1425,1450]" pageId="1" pageNumber="97" type="description">
<paragraph id="20015F3FFFAF0F2135F8D9FA02A2AAA3" blockId="1.[151,538,1390,1450]" box="[151,235,1425,1450]" pageId="1" pageNumber="97">
(
<figureCitation id="B88543BAFFAF0F2135F0D9FA02A9AAA3" box="[159,224,1425,1450]" captionStart="FIGURES 1" captionStartId="3.[151,264,1230,1253]" captionTargetBox="[151,1435,301,1207]" captionTargetId="figure@3.[151,1435,267,1207]" captionTargetPageId="3" captionText="FIGURES 1A–D. Oleia fieldsi Silva et Song sp. n., adult male. A. Wing (scale bar = 500 µm). B. Tergite IX and dorsal aspect of gonocoxite, anal point and sternapodeme in detail (scale bar = 50 µm). C. Dorsal aspect of gonocoxite, phallapodeme in detail (scale bar = 50 µm). D. Ventral aspect of gonocoxite, gonostylar anterior branch in detail (scale bar = 50 µm)." httpUri="https://zenodo.org/record/2611466/files/figure.png" pageId="1" pageNumber="97">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="68A40CB4FFAF0F2135F8D9B20149A933" pageId="1" pageNumber="97" type="materials_examined">
<paragraph id="20015F3FFFAF0F2135F8D9B20149A933" blockId="1.[151,1437,1497,2026]" pageId="1" pageNumber="97">
<emphasis id="12CA832DFFAF0F2135F8D9B2031EAAFB" bold="true" box="[151,343,1497,1522]" pageId="1" pageNumber="97">Type examined.</emphasis>
<materialsCitation id="90D65562FFAF0F21340DD9B207D0A91F" ID-GBIF-Occurrence="2029159968" collectingDate="2017-10-13" collectionCode="NTNU-VM" collectorName="A. Kiszewski &amp; Kiszewski, E. S &amp; Dias, F. L &amp; R. C. Oliveira" country="Brazil" latitude="-25.6201" location="Iguacu National Park" longLatPrecision="1" longitude="-54.47547" municipality="Foz do Iguacu" pageId="1" pageNumber="97" specimenCount="1" specimenCount-male="1" stateProvince="Parana" typeStatus="holotype">
<typeStatus id="FF05E19DFFAF0F21340DD9B20384AAFB" box="[354,461,1497,1522]" pageId="1" pageNumber="97" type="holotype">Holotype</typeStatus>
,
<specimenCount id="36B894B6FFAF0F2134B2D9B10066AAFB" box="[477,559,1497,1522]" pageId="1" pageNumber="97" type="male">1 male</specimenCount>
,
<collectingCountry id="58A91FAFFFAF0F213750D9B100E0AAFB" box="[575,681,1498,1522]" name="Brazil" pageId="1" pageNumber="97">BRAZIL</collectingCountry>
:
<collectingRegion id="E27A91DDFFAF0F2137D8D9B1014FAAFB" box="[695,774,1498,1522]" country="Brazil" name="Parana" pageId="1" pageNumber="97">Paraná</collectingRegion>
,
<collectingMunicipality id="C065C545FFAF0F21367AD9B10188AAFB" box="[789,961,1497,1522]" pageId="1" pageNumber="97">Foz do Iguaçu</collectingMunicipality>
,
<location id="256109E4FFAF0F2136BFD9B10683AAFB" LSID="urn:lsid:plazi:treatment:A817EE29FFAF0F2235F8D9050704A9AF:256109E4FFAF0F2136BFD9B10683AAFB" box="[976,1226,1497,1522]" country="Brazil" latitude="-25.6201" longLatPrecision="1" longitude="-54.47547" municipality="Foz do Iguacu" name="Iguacu National Park" pageId="1" pageNumber="97" stateProvince="Parana">Iguaçu National Park</location>
,
<geoCoordinate id="458A39F8FFAF0F2131B6D9B20721AAFB" box="[1241,1384,1497,1522]" direction="south" orientation="latitude" pageId="1" pageNumber="97" precision="1" value="-25.6201">25.62010° S</geoCoordinate>
and
<geoCoordinate id="458A39F8FFAF0F2135F8D9960379A91C" box="[151,304,1533,1558]" direction="west" orientation="longitude" pageId="1" pageNumber="97" precision="1" value="-54.47547">54.47547° W</geoCoordinate>
,
<date id="540079FFFFAF0F213451D99503F8A91F" box="[318,433,1534,1558]" pageId="1" pageNumber="97" value="2017-10-13">
<collectingDate id="44448017FFAF0F213451D99503F8A91F" box="[318,433,1534,1558]" pageId="1" pageNumber="97" value="2017-10-13">13.x.2017</collectingDate>
</date>
, leg.
<collectorName id="8D4B3AE9FFAF0F21349CD99500D9A91F" box="[499,656,1533,1558]" pageId="1" pageNumber="97">A. Kiszewski</collectorName>
, E.S.
<collectorName id="8D4B3AE9FFAF0F2137B8D996010DA91C" box="[727,836,1533,1558]" pageId="1" pageNumber="97">Dias, F.L</collectorName>
. Silva &amp;
<collectorName id="8D4B3AE9FFAF0F2136D9D9950619A91F" box="[950,1104,1533,1558]" pageId="1" pageNumber="97">R.C. Oliveira</collectorName>
(
<collectionCode id="46AFC7FAFFAF0F21310ED99506A3A91F" box="[1121,1258,1534,1558]" country="Norway" httpUri="http://grbio.org/cool/pnpi-f6fs" name="Norwegian University of Science and Technology - University Museum" pageId="1" pageNumber="97" type="Museum">NTNU-VM</collectionCode>
, slide 201776)
</materialsCitation>
.
<materialsCitation id="90D65562FFAF0F2135F8DA4900B5A933" ID-GBIF-Occurrence="2029160273" box="[151,764,1569,1594]" collectingDate="2017-10-13" collectionCode="MZSP" collectorName="A. Kiszewski &amp; Kiszewski, E. S &amp; Dias, F. L &amp; R. C. Oliveira" country="Brazil" latitude="-25.6201" location="Iguacu National Park" longLatPrecision="1" longitude="-54.47547" municipality="Foz do Iguacu" pageId="1" pageNumber="97" specimenCount="3" specimenCount-male="3" stateProvince="Parana" typeStatus="paratype">
<typeStatus id="FF05E19DFFAF0F2135F8DA490340A933" box="[151,265,1570,1594]" pageId="1" pageNumber="97" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="36B894B6FFAF0F21347CDA490322A933" box="[275,363,1569,1594]" pageId="1" pageNumber="97" type="male">3 males</specimenCount>
, same data as for holotype (
<collectionCode id="46AFC7FAFFAF0F2137CBDA4900BDA933" box="[676,756,1570,1594]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="97" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="68A40CB4FFAF0F2135A8DA2E0725A9C0" pageId="1" pageNumber="97" type="etymology">
<paragraph id="20015F3FFFAF0F2135A8DA2E001EA98B" blockId="1.[151,1437,1497,2026]" pageId="1" pageNumber="97">
<emphasis id="12CA832DFFAF0F2135A8DA2E0307A957" bold="true" box="[199,334,1605,1630]" pageId="1" pageNumber="97">Etymology.</emphasis>
Named in honor of Rogério Campos ‘Fields’ de Oliveira, researcher, colleague and friend, for assistance in collecting the new species.
</paragraph>
<paragraph id="20015F3FFFAF0F2135A8DAE60725A9C0" blockId="1.[151,1437,1497,2026]" pageId="1" pageNumber="97">
<emphasis id="12CA832DFFAF0F2135A8DAE60387A9AF" bold="true" box="[199,462,1677,1702]" pageId="1" pageNumber="97">Diagnostic characters.</emphasis>
The male adult of
<taxonomicName id="E7BE24BCFFAF0F2137C2DAE60170A9AF" authority="Silva &amp; Song, 2018" authorityName="Silva &amp; Song" authorityYear="2018" box="[685,825,1677,1702]" class="Insecta" family="Chironomidae" genus="Oleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="97" phylum="Arthropoda" rank="species" species="fieldsi" status="sp. nov.">
<emphasis id="12CA832DFFAF0F2137C2DAE60170A9AF" box="[685,825,1677,1702]" italics="true" pageId="1" pageNumber="97">Oleia fieldsi</emphasis>
</taxonomicName>
<emphasis id="12CA832DFFAF0F21362BDAE601CEA9AF" bold="true" box="[836,903,1677,1702]" pageId="1" pageNumber="97">
<taxonomicNameLabel id="09F93E56FFAF0F21362BDAE601CEA9AF" box="[836,903,1677,1702]" pageId="1" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from that of other
<taxonomicName id="E7BE24BCFFAF0F2131DADAE606BBA9AF" authorityName="Andersen et Mendes" authorityYear="2007" box="[1205,1266,1677,1702]" class="Insecta" family="Chironomidae" genus="Oleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="97" phylum="Arthropoda" rank="genus">
<emphasis id="12CA832DFFAF0F2131DADAE606BBA9AF" box="[1205,1266,1677,1702]" italics="true" pageId="1" pageNumber="97">Oleia</emphasis>
</taxonomicName>
species by the combination of the following characters: anal point basally broad; gonostylar anterior branch with 5 strong setae.
</paragraph>
</subSubSection>
<subSubSection id="68A40CB4FFAF0F2235A8DABE0019A91F" lastPageId="2" lastPageNumber="98" pageId="1" pageNumber="97" type="description">
<paragraph id="20015F3FFFAF0F2135A8DABE010FA81B" blockId="1.[151,1437,1497,2026]" pageId="1" pageNumber="97">
<emphasis id="12CA832DFFAF0F2135A8DABE0310A9E7" bold="true" box="[199,345,1749,1774]" pageId="1" pageNumber="97">Description.</emphasis>
<emphasis id="12CA832DFFAF0F21340EDABE03D2A9E7" box="[353,411,1749,1774]" italics="true" pageId="1" pageNumber="97">Male</emphasis>
(n = 4).
<emphasis id="12CA832DFFAF0F21376ADABD007BA9E4" box="[517,562,1750,1773]" italics="true" pageId="1" pageNumber="97">Size</emphasis>
. Total length
<quantity id="E746F2DAFFAF0F2137BBDABE0135A9E4" box="[724,892,1749,1774]" metricMagnitude="-3" metricUnit="m" metricValue="2.16" metricValueMax="2.36" metricValueMin="1.96" pageId="1" pageNumber="97" unit="mm" value="2.16" valueMax="2.36" valueMin="1.96">1.96–2.36 mm</quantity>
. Wing length
<quantity id="E746F2DAFFAF0F213170DABE068EA9E4" box="[1055,1223,1749,1774]" metricMagnitude="-3" metricUnit="m" metricValue="1.2149999999999999" metricValueMax="1.25" metricValueMin="1.18" pageId="1" pageNumber="97" unit="mm" value="1.2149999999999999" valueMax="1.25" valueMin="1.18">1.18–1.25 mm</quantity>
. Total length/wing length 1.66–1.89. Wing length/length of profemur 2.34–2.99.
</paragraph>
<paragraph id="20015F3FFFAF0F2135A8DB7601BBA8AB" blockId="1.[151,1437,1497,2026]" pageId="1" pageNumber="97">
<emphasis id="12CA832DFFAF0F2135A8DB76030BA83F" box="[199,322,1821,1846]" italics="true" pageId="1" pageNumber="97">Coloration</emphasis>
. Head brown, maxillary palp pale brown. Thorax pale brown, with dark brown vittae, postnotum, median anepisternum, preepisternum, and epimeron II. Wing clear with membrane transparent and crossveins pale, without spots. All legs pale brown. Abdominal tergites brown with bases of abdominal setae and anterior and posterior bands pale brown. Hypopygium pale brown with brown anal point.
</paragraph>
<paragraph id="20015F3FFFAF0F2235A8DBC6003FAE14" blockId="1.[151,1437,1497,2026]" lastBlockId="2.[151,1437,151,692]" lastPageId="2" lastPageNumber="98" pageId="1" pageNumber="97">
<emphasis id="12CA832DFFAF0F2135A8DBC6034DA8CF" box="[199,260,1965,1990]" italics="true" pageId="1" pageNumber="97">Head</emphasis>
. Eye bare, reniform, without dorsomedian extension. Antennal flagellum missing, diameter of pedicel 91–110 µm. Temporal setae 8–10, irregularly uniserial, inner verticals weak, outer verticals and postorbitals strong. Frontal tubercle absent. Tentorium 117–142 µm long. Clypeus 51–70 µm long, 53–71 µm wide at largest part, bearing 5–6 setae. Cibarial pump 117–142 µm long, with anterior margin concave. Lengths of palpomeres 1–5 (in µm): 24–29 (3); 29–33 (3); 50–64 (3); 44–46 (3); 69–81 (3). Third palpomere with 3 lanceolate sensilla clavata grouped at apical sensillum coeloconicum.
</paragraph>
<paragraph id="20015F3FFFAC0F2235A8DD4C0652AE8E" blockId="2.[151,1437,151,692]" pageId="2" pageNumber="98">
<emphasis id="12CA832DFFAC0F2235A8DD4C0351AE49" box="[199,280,295,320]" italics="true" pageId="2" pageNumber="98">Thorax</emphasis>
. Antepronotal lobes narrowly separated, antepronotum with 3 setae. Acrostichals 14–19, decumbent, uniserial to biserial, starting at some distance from antepronotum. Dorsocentrals 6–8, simple, uniserial. Prealars 4, 3 posterior and 1 anterior. Supraalar 1. Scutellum with few 4 setae in single row.
</paragraph>
<paragraph id="20015F3FFFAC0F2235A8DDFE03B0AEFE" blockId="2.[151,1437,151,692]" pageId="2" pageNumber="98">
<emphasis id="12CA832DFFAC0F2235A8DDFE02B6AEA5" box="[199,255,405,428]" italics="true" pageId="2" pageNumber="98">Wing</emphasis>
(
<figureCitation id="B88543BAFFAC0F22347EDDFF0325AEA5" box="[273,364,404,429]" captionStart="FIGURES 1" captionStartId="3.[151,264,1230,1253]" captionTargetBox="[151,1435,301,1207]" captionTargetId="figure@3.[151,1435,267,1207]" captionTargetPageId="3" captionText="FIGURES 1A–D. Oleia fieldsi Silva et Song sp. n., adult male. A. Wing (scale bar = 500 µm). B. Tergite IX and dorsal aspect of gonocoxite, anal point and sternapodeme in detail (scale bar = 50 µm). C. Dorsal aspect of gonocoxite, phallapodeme in detail (scale bar = 50 µm). D. Ventral aspect of gonocoxite, gonostylar anterior branch in detail (scale bar = 50 µm)." httpUri="https://zenodo.org/record/2611466/files/figure.png" pageId="2" pageNumber="98">Fig. 1A</figureCitation>
). Width
<quantity id="E746F2DAFFAC0F2234A0DDFF0013AEA5" box="[463,602,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="98" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
, membrane without setae with fine punctation. Anal lobe protruding. VR 1.43–1.52. WW 0.29–0.30. Costa 1.00–1.10 µm long. R
<subScript id="BC3A5D7AFFAC0F223669DDAD0157AEDD" attach="left" box="[774,798,454,468]" fontSize="6" pageId="2" pageNumber="98">4+5</subScript>
ending above M
<subScript id="BC3A5D7AFFAC0F2236B8DDAD01A6AEDD" attach="left" box="[983,1007,454,468]" fontSize="6" pageId="2" pageNumber="98">3+4</subScript>
, FCu distal to RM. Brachiolum with 3 setae. Veins bare. Squama bare.
</paragraph>
<paragraph id="20015F3FFFAC0F2235A8DE6F01FEADBA" blockId="2.[151,1437,151,692]" pageId="2" pageNumber="98">
<emphasis id="12CA832DFFAC0F2235A8DE6F02B5AD12" box="[199,252,516,539]" italics="true" pageId="2" pageNumber="98">Legs</emphasis>
(in µm). Fore leg: width at apex of tibia 19–29, tibia with single, apical spur 34–46; ta
<subScript id="BC3A5D7AFFAC0F2231AEDE7A069FAD16" attach="left" box="[1217,1238,529,543]" fontSize="6" pageId="2" pageNumber="98">1-4</subScript>
without preapical pseudospurs. Mid leg: width at apex of tibia 23–29, tibia with two apical spurs 11–15; 13–27; ta
<subScript id="BC3A5D7AFFAC0F2231AEDE53069FAD4F" attach="left" box="[1217,1238,568,582]" fontSize="6" pageId="2" pageNumber="98">1-4</subScript>
without preapical pseudospurs. Hind leg: width at apex of tibia 26–40, tibia with two apical spurs 17–23; 39–43; comb with 10–12 bristles; ta
<subScript id="BC3A5D7AFFAC0F223464DEE80369AD98" attach="left" box="[267,288,643,657]" fontSize="6" pageId="2" pageNumber="98">1-4</subScript>
without preapical pseudospurs. Sensilla chaetica absent. Claws slender, distally recurved and pointed. Pulvilli vestigial. Lengths and proportion of leg segments as in
<tableCitation id="6D3C6A84FFAC0F223632DEF001F8ADBA" box="[861,945,667,692]" captionStart="TABLE 1" captionStartId="2.[151,239,735,758]" captionText="TABLE 1. Lengths (in µm) and proportions of prothoracic (p 1), mesothoracic (p 2) and metathoracic (p 3) legs of Oleia fieldsi Silva et Song sp. n., male (n = 1–3)." pageId="2" pageNumber="98">Table 1</tableCitation>
.
</paragraph>
<caption id="74C10FB7FFAC0F2235F8DEB40028AC11" pageId="2" pageNumber="98">
<paragraph id="20015F3FFFAC0F2235F8DEB40028AC11" blockId="2.[151,1436,734,792]" pageId="2" pageNumber="98">
<emphasis id="12CA832DFFAC0F2235F8DEB40345ADFF" bold="true" box="[151,268,735,758]" pageId="2" pageNumber="98">TABLE 1.</emphasis>
Lengths (in µm) and proportions of prothoracic (p
<subScript id="BC3A5D7AFFAC0F223657DE840176ADF3" attach="left" box="[824,831,751,762]" fontSize="5" pageId="2" pageNumber="98">1</subScript>
), mesothoracic (p
<subScript id="BC3A5D7AFFAC0F22316BDE840642ADF3" attach="left" box="[1028,1035,751,762]" fontSize="5" pageId="2" pageNumber="98">2</subScript>
) and metathoracic (p
<subScript id="BC3A5D7AFFAC0F22319ADE8406B5ADF3" attach="left" box="[1269,1276,751,762]" fontSize="5" pageId="2" pageNumber="98">3</subScript>
) legs of
<taxonomicName id="E7BE24BCFFAC0F22300EDEB5035FAC11" authority="Silva" authorityName="Silva &amp; Song" authorityYear="2018" class="Insecta" family="Chironomidae" genus="Oleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="98" phylum="Arthropoda" rank="species" species="fieldsi">
<emphasis id="12CA832DFFAC0F22300EDEB50291AC11" italics="true" pageId="2" pageNumber="98">Oleia fieldsi</emphasis>
Silva
</taxonomicName>
et Song
<emphasis id="12CA832DFFAC0F22341BDF6A03E6AC11" bold="true" box="[372,431,769,792]" pageId="2" pageNumber="98">sp. n.</emphasis>
, male (n = 1–3).
</paragraph>
</caption>
<paragraph id="20015F3FFFAC0F223445DF5F06A0AB5E" pageId="2" pageNumber="98">
<table id="52BEAD9FFFACF0DF35CEDF5F06A0AB52" box="[161,1257,820,1115]" gridcols="6" gridrows="8" pageId="2" pageNumber="98">
<tr id="9E8E5D7DFFACF0DF35CEDF5F06A0AC47" box="[161,1257,820,846]" gridrow="0" pageId="2" pageNumber="98" rowspan-0="1">
<th id="DD5F3401FFACF0DF3445DF5F03CAAC47" box="[298,387,820,846]" gridcol="1" gridrow="0" pageId="2" pageNumber="98">fe</th>
<th id="DD5F3401FFACF0DF3760DF5F002EAC47" box="[527,615,820,846]" gridcol="2" gridrow="0" pageId="2" pageNumber="98">ti</th>
<th id="DD5F3401FFACF0DF379BDF5F0169AC47" box="[756,800,820,846]" gridcol="3" gridrow="0" pageId="2" pageNumber="98">ta1</th>
<th id="DD5F3401FFACF0DF36B7DF5F064DAC47" box="[984,1028,820,846]" gridcol="4" gridrow="0" pageId="2" pageNumber="98">ta2</th>
<th id="DD5F3401FFACF0DF31D2DF5F06A0AC47" box="[1213,1257,820,846]" gridcol="5" gridrow="0" pageId="2" pageNumber="98">ta3</th>
</tr>
<tr id="9E8E5D7DFFACF0DF35CEDF3006A0AC7D" box="[161,1257,859,884]" gridrow="1" pageId="2" pageNumber="98">
<th id="DD5F3401FFACF0DF35CEDF3002FDAC7D" box="[161,180,859,884]" gridcol="0" gridrow="1" pageId="2" pageNumber="98">p1</th>
<td id="DD5F3401FFACF0DF3445DF3003CAAC7D" box="[298,387,859,884]" gridcol="1" gridrow="1" pageId="2" pageNumber="98">417–505</td>
<td id="DD5F3401FFACF0DF3760DF30002EAC7D" box="[527,615,859,884]" gridcol="2" gridrow="1" pageId="2" pageNumber="98">471–504</td>
<td id="DD5F3401FFACF0DF379BDF300169AC7D" box="[756,800,859,884]" gridcol="3" gridrow="1" pageId="2" pageNumber="98">256</td>
<td id="DD5F3401FFACF0DF36B7DF30064DAC7D" box="[984,1028,859,884]" gridcol="4" gridrow="1" pageId="2" pageNumber="98">136</td>
<td id="DD5F3401FFACF0DF31D2DF3006A0AC7D" box="[1213,1257,859,884]" gridcol="5" gridrow="1" pageId="2" pageNumber="98">85</td>
</tr>
<tr id="9E8E5D7DFFACF0DF35CEDFEA06A0AC92" box="[161,1257,897,923]" gridrow="2" pageId="2" pageNumber="98">
<th id="DD5F3401FFACF0DF35CEDFEA02FDAC92" box="[161,180,897,923]" gridcol="0" gridrow="2" pageId="2" pageNumber="98">p2</th>
<td id="DD5F3401FFACF0DF3445DFEA03CAAC92" box="[298,387,897,923]" gridcol="1" gridrow="2" pageId="2" pageNumber="98">405–457</td>
<td id="DD5F3401FFACF0DF3760DFEA002EAC92" box="[527,615,897,923]" gridcol="2" gridrow="2" pageId="2" pageNumber="98">429–454</td>
<td id="DD5F3401FFACF0DF379BDFEA0169AC92" box="[756,800,897,923]" gridcol="3" gridrow="2" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF36B7DFEA064DAC92" box="[984,1028,897,923]" gridcol="4" gridrow="2" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF31D2DFEA06A0AC92" box="[1213,1257,897,923]" gridcol="5" gridrow="2" pageId="2" pageNumber="98">–</td>
</tr>
<tr id="9E8E5D7DFFACF0DF35CEDFCC06A0ACC8" box="[161,1257,935,961]" gridrow="3" pageId="2" pageNumber="98">
<th id="DD5F3401FFACF0DF35CEDFCC02FDACC8" box="[161,180,935,961]" gridcol="0" gridrow="3" pageId="2" pageNumber="98">p3</th>
<td id="DD5F3401FFACF0DF3445DFCC03CAACC8" box="[298,387,935,961]" gridcol="1" gridrow="3" pageId="2" pageNumber="98">436–469</td>
<td id="DD5F3401FFACF0DF3760DFCC002EACC8" box="[527,615,935,961]" gridcol="2" gridrow="3" pageId="2" pageNumber="98">441–503</td>
<td id="DD5F3401FFACF0DF379BDFCC0169ACC8" box="[756,800,935,961]" gridcol="3" gridrow="3" pageId="2" pageNumber="98">182</td>
<td id="DD5F3401FFACF0DF36B7DFCC064DACC8" box="[984,1028,935,961]" gridcol="4" gridrow="3" pageId="2" pageNumber="98">102</td>
<td id="DD5F3401FFACF0DF31D2DFCC06A0ACC8" box="[1213,1257,935,961]" gridcol="5" gridrow="3" pageId="2" pageNumber="98">138</td>
</tr>
<tr id="9E8E5D7DFFACF0DF35CEDFA506A0ACE1" box="[161,1257,974,1000]" gridrow="4" pageId="2" pageNumber="98" rowspan-0="1">
<td id="DD5F3401FFACF0DF3445DFA503CAACE1" box="[298,387,974,1000]" gridcol="1" gridrow="4" pageId="2" pageNumber="98">ta4</td>
<td id="DD5F3401FFACF0DF3760DFA5002EACE1" box="[527,615,974,1000]" gridcol="2" gridrow="4" pageId="2" pageNumber="98">ta5</td>
<td id="DD5F3401FFACF0DF379BDFA50169ACE1" box="[756,800,974,1000]" gridcol="3" gridrow="4" pageId="2" pageNumber="98">LR</td>
<td id="DD5F3401FFACF0DF36B7DFA5064DACE1" box="[984,1028,974,1000]" gridcol="4" gridrow="4" pageId="2" pageNumber="98">BV</td>
<td id="DD5F3401FFACF0DF31D2DFA506A0ACE1" box="[1213,1257,974,1000]" gridcol="5" gridrow="4" pageId="2" pageNumber="98">SV</td>
</tr>
<tr id="9E8E5D7DFFACF0DF35CEDF9F06A0AB07" box="[161,1257,1012,1038]" gridrow="5" pageId="2" pageNumber="98">
<th id="DD5F3401FFACF0DF35CEDF9F02FDAB07" box="[161,180,1012,1038]" gridcol="0" gridrow="5" pageId="2" pageNumber="98">p1</th>
<td id="DD5F3401FFACF0DF3445DF9F03CAAB07" box="[298,387,1012,1038]" gridcol="1" gridrow="5" pageId="2" pageNumber="98">60</td>
<td id="DD5F3401FFACF0DF3760DF9F002EAB07" box="[527,615,1012,1038]" gridcol="2" gridrow="5" pageId="2" pageNumber="98">54</td>
<td id="DD5F3401FFACF0DF379BDF9F0169AB07" box="[756,800,1012,1038]" gridcol="3" gridrow="5" pageId="2" pageNumber="98">0.51</td>
<td id="DD5F3401FFACF0DF36B7DF9F064DAB07" box="[984,1028,1012,1038]" gridcol="4" gridrow="5" pageId="2" pageNumber="98">3.77</td>
<td id="DD5F3401FFACF0DF31D2DF9F06A0AB07" box="[1213,1257,1012,1038]" gridcol="5" gridrow="5" pageId="2" pageNumber="98">3.95</td>
</tr>
<tr id="9E8E5D7DFFACF0DF35CED87706A0AB3D" box="[161,1257,1052,1076]" gridrow="6" pageId="2" pageNumber="98">
<th id="DD5F3401FFACF0DF35CED87702FDAB3D" box="[161,180,1052,1076]" gridcol="0" gridrow="6" pageId="2" pageNumber="98">p2</th>
<td id="DD5F3401FFACF0DF3445D87703CAAB3D" box="[298,387,1052,1076]" gridcol="1" gridrow="6" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF3760D877002EAB3D" box="[527,615,1052,1076]" gridcol="2" gridrow="6" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF379BD8770169AB3D" box="[756,800,1052,1076]" gridcol="3" gridrow="6" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF36B7D877064DAB3D" box="[984,1028,1052,1076]" gridcol="4" gridrow="6" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF31D2D87706A0AB3D" box="[1213,1257,1052,1076]" gridcol="5" gridrow="6" pageId="2" pageNumber="98">–</td>
</tr>
<tr id="9E8E5D7DFFACF0DF35CED82A06A0AB52" box="[161,1257,1089,1115]" gridrow="7" pageId="2" pageNumber="98">
<th id="DD5F3401FFACF0DF35CED82A02FDAB52" box="[161,180,1089,1115]" gridcol="0" gridrow="7" pageId="2" pageNumber="98">p3</th>
<td id="DD5F3401FFACF0DF3445D82A03CAAB52" box="[298,387,1089,1115]" gridcol="1" gridrow="7" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF3760D82A002EAB52" box="[527,615,1089,1115]" gridcol="2" gridrow="7" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF379BD82A0169AB52" box="[756,800,1089,1115]" gridcol="3" gridrow="7" pageId="2" pageNumber="98">0.38</td>
<td id="DD5F3401FFACF0DF36B7D82A064DAB52" box="[984,1028,1089,1115]" gridcol="4" gridrow="7" pageId="2" pageNumber="98">–</td>
<td id="DD5F3401FFACF0DF31D2D82A06A0AB52" box="[1213,1257,1089,1115]" gridcol="5" gridrow="7" pageId="2" pageNumber="98">5.24</td>
</tr>
</table>
</paragraph>
<paragraph id="20015F3FFFAC0F2235A8D8FD01FCAAFB" blockId="2.[151,1437,1172,1702]" pageId="2" pageNumber="98">
<emphasis id="12CA832DFFAC0F2235A8D8FD031FABA4" box="[199,342,1174,1197]" italics="true" pageId="2" pageNumber="98">Hypopygium</emphasis>
(
<figureCitation id="B88543BAFFAC0F223407D8FF03E4ABA4" box="[360,429,1172,1197]" captionStart="FIGURES 1" captionStartId="3.[151,264,1230,1253]" captionTargetBox="[151,1435,301,1207]" captionTargetId="figure@3.[151,1435,267,1207]" captionTargetPageId="3" captionText="FIGURES 1A–D. Oleia fieldsi Silva et Song sp. n., adult male. A. Wing (scale bar = 500 µm). B. Tergite IX and dorsal aspect of gonocoxite, anal point and sternapodeme in detail (scale bar = 50 µm). C. Dorsal aspect of gonocoxite, phallapodeme in detail (scale bar = 50 µm). D. Ventral aspect of gonocoxite, gonostylar anterior branch in detail (scale bar = 50 µm)." httpUri="https://zenodo.org/record/2611466/files/figure.png" pageId="2" pageNumber="98">Figs 1</figureCitation>
B–D). Tergite IX with few setae to each side of anal point. Laterosternite IX with 4–5 setae. Anal point narrow and elongate, 53–57 (2) µm long, 16–18 (2) µm wide at base, with 14–16 (2) strong setae (
<figureCitation id="B88543BAFFAC0F2235F0D8B602B1ABFC" box="[159,248,1245,1270]" captionStart="FIGURES 1" captionStartId="3.[151,264,1230,1253]" captionTargetBox="[151,1435,301,1207]" captionTargetId="figure@3.[151,1435,267,1207]" captionTargetPageId="3" captionText="FIGURES 1A–D. Oleia fieldsi Silva et Song sp. n., adult male. A. Wing (scale bar = 500 µm). B. Tergite IX and dorsal aspect of gonocoxite, anal point and sternapodeme in detail (scale bar = 50 µm). C. Dorsal aspect of gonocoxite, phallapodeme in detail (scale bar = 50 µm). D. Ventral aspect of gonocoxite, gonostylar anterior branch in detail (scale bar = 50 µm)." httpUri="https://zenodo.org/record/2611466/files/figure.png" pageId="2" pageNumber="98">Fig. 1B</figureCitation>
). Phallapodeme well developed, with curved oral projections, 79–86 µm long; transverse sternapodeme anteriorly arched, oral projections barely indicated, 106–127 µm long. Penis cavity with horse-shoe shaped basal sclerite, 11–14 µm wide (
<figureCitation id="B88543BAFFAC0F2234DAD94E0044AA34" box="[437,525,1317,1342]" captionStart="FIGURES 1" captionStartId="3.[151,264,1230,1253]" captionTargetBox="[151,1435,301,1207]" captionTargetId="figure@3.[151,1435,267,1207]" captionTargetPageId="3" captionText="FIGURES 1A–D. Oleia fieldsi Silva et Song sp. n., adult male. A. Wing (scale bar = 500 µm). B. Tergite IX and dorsal aspect of gonocoxite, anal point and sternapodeme in detail (scale bar = 50 µm). C. Dorsal aspect of gonocoxite, phallapodeme in detail (scale bar = 50 µm). D. Ventral aspect of gonocoxite, gonostylar anterior branch in detail (scale bar = 50 µm)." httpUri="https://zenodo.org/record/2611466/files/figure.png" pageId="2" pageNumber="98">Fig. 1C</figureCitation>
); virga consisting of several separate sclerites. Gonocoxite 97–109 µm long, 39– 45 µm wide with low, rounded inferior volsella. Gonostylus trifid as in figure 1D, posterior branch 63–74 µm long, separated to base, with 10–12 (3) marginal setae, and 21–24 (2) µm long subapical megaseta; median branch oblong, 53–58 µm long, with row of 11 strong, curved marginal setae, longest 16–21 µm long, anteriomedian corner with weaker setae, longest 21–22 (2) µm long; anterior hyaline branch spoon-shaped, with 5 moderately strong setae, longest 14–27 (3) µm long. HR 2.26–2.59, HV 1.36–1.63.
</paragraph>
<paragraph id="20015F3FFFAC0F2235A8D9960019A91F" blockId="2.[151,1437,1172,1702]" box="[199,592,1533,1558]" pageId="2" pageNumber="98">
<emphasis id="12CA832DFFAC0F2235A8D9960392A91F" box="[199,475,1533,1558]" italics="true" pageId="2" pageNumber="98">Female, pupa and larva.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="68A40CB4FFAC0F2235A8DA4A0704A9AF" pageId="2" pageNumber="98" type="biology_ecology">
<paragraph id="20015F3FFFAC0F2235A8DA4A0704A9AF" blockId="2.[151,1437,1172,1702]" pageId="2" pageNumber="98">
<emphasis id="12CA832DFFAC0F2235A8DA4A036DA933" bold="true" box="[199,292,1569,1594]" pageId="2" pageNumber="98">Ecology</emphasis>
. Adult males of
<taxonomicName id="E7BE24BCFFAC0F2234B2DA4A002FA933" authority="Silva &amp; Song, 2018" authorityName="Silva &amp; Song" authorityYear="2018" box="[477,614,1569,1594]" class="Insecta" family="Chironomidae" genus="Oleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="98" phylum="Arthropoda" rank="species" species="fieldsi" status="sp. nov.">
<emphasis id="12CA832DFFAC0F2234B2DA4A002FA933" box="[477,614,1569,1594]" italics="true" pageId="2" pageNumber="98">Oleia fieldsi</emphasis>
</taxonomicName>
<emphasis id="12CA832DFFAC0F223702DA4A00E5A933" bold="true" box="[621,684,1569,1594]" pageId="2" pageNumber="98">
<taxonomicNameLabel id="09F93E56FFAC0F223702DA4A00E5A933" box="[621,684,1569,1594]" pageId="2" pageNumber="98" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were collected near a reservoir in a fragment of secondary forest in the Brazilian Atlantic Forest, with canopy intercepting 20% of sunlight, at Iguaçu National Park, on the border of the Argentinian province of
<collectingRegion id="E27A91DDFFAC0F2234BADA020075A98B" box="[469,572,1641,1666]" country="Argentina" name="Misiones" pageId="2" pageNumber="98">Misiones</collectingRegion>
and the Brazilian state of
<collectingRegion id="E27A91DDFFAC0F22360ADA0201FCA988" box="[869,949,1641,1665]" country="Brazil" name="Parana" pageId="2" pageNumber="98">Paraná</collectingRegion>
. According to
<bibRefCitation id="442F22CEFFAC0F223135DA0207D1A988" author="Andersen, T. &amp; Mendes, H. F." box="[1114,1432,1641,1666]" pageId="2" pageNumber="98" pagination="17 - 52" refId="ref3062" refString="Andersen, T. &amp; Mendes, H. F. (2007) Five enigmatic new orthoclad genera from Brazil (Diptera: Chironomidae, Orthocladiinae). Contributions to the Systematics and Ecology of Aquatic Diptera-A tribute to Ole A. Saether. The Caddis Press, Columbus, pp. 17 - 52." type="book chapter" year="2007">Andersen &amp; Mendes (2007)</bibRefCitation>
, the immature stages of
<taxonomicName id="E7BE24BCFFAC0F2234F3DAE60393A9AF" authorityName="Andersen et Mendes" authorityYear="2007" box="[412,474,1677,1702]" class="Insecta" family="Chironomidae" genus="Oleia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="12CA832DFFAC0F2234F3DAE60393A9AF" box="[412,474,1677,1702]" italics="true" pageId="2" pageNumber="98">Oleia</emphasis>
</taxonomicName>
might be terrestrial or semiterrestrial given the proven difficult to locate them.
</paragraph>
</subSubSection>
</treatment>
</document>
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