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<document ID-CLB-Dataset="27191" ID-DOI="http://dx.doi.org/10.3897/zookeys.1146.94607" ID-GBIF-Dataset="8259c245-8f4e-465c-85f4-040039359601" ID-PMC="PMC10208602" ID-Pensoft-Pub="1313-2970-1146-87" ID-Pensoft-UUID="E4986F2FF30351588CF5C50C253222E4" ID-PubMed="37234291" ID-ZooBank="9F4ADF001BE748789B98FB23B72B4435" ModsDocID="1313-2970-1146-87" checkinTime="1675846373446" checkinUser="pensoft" docAuthor="Marra Santos, Fidelis Junio" docDate="2023" docId="A7FC4D63F7815CC1BA31DD6FA74986DA" docLanguage="en" docName="ZooKeys 1146: 87-114" docOrigin="ZooKeys 1146" docPubDate="2023-02-07" docSource="http://dx.doi.org/10.3897/zookeys.1146.94607" docTitle="Liotyphlops palauophis Marra Santos 2023, sp. nov." docType="treatment" docUuid="0CD30C35-2263-491C-BB62-2494019740C8" docUuidSource="ZooBank" docVersion="6" id="E4986F2FF30351588CF5C50C253222E4" lastPageNumber="87" masterDocId="E4986F2FF30351588CF5C50C253222E4" masterDocTitle="A new species of the genus Liotyphlops Peters, 1881 (Serpentes, Anomalepididae) from Colombia and the synonymization of Liotyphlops beui (Amaral, 1924) with Liotyphlops ternetzii (Boulenger, 1896)" masterLastPageNumber="114" masterPageNumber="87" pageNumber="87" updateTime="1732826855592" updateUser="ExternalLinkService">
<mods:mods id="8D4480167B1C6E0EA8AD7DD216F38A8F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="445DD11CD9590E1E118EA3FD88B5C57A">
<mods:title id="713EF07B2E37185C6A9DD1C694117E11">A new species of the genus Liotyphlops Peters, 1881 (Serpentes, Anomalepididae) from Colombia and the synonymization of Liotyphlops beui (Amaral, 1924) with Liotyphlops ternetzii (Boulenger, 1896)</mods:title>
</mods:titleInfo>
<mods:name id="9B118925AEB8EBE7DE9DFB9F3DDB6D5A" type="personal">
<mods:role id="DB6FA37F103981314C8D21A7EB441DDE">
<mods:roleTerm id="A9F03D5F6991CBC6F02CE1549CCAD276">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E378A0F6AE79581A51945D6991BFAD63">Marra Santos, Fidelis Junio</mods:namePart>
<mods:nameIdentifier id="E7E284696C470D1D0F9B18CF29646311" type="ORCID">https://orcid.org/0000-0002-5140-2830</mods:nameIdentifier>
<mods:affiliation id="8D0CAB198D325AC20132C3CD991DF956">Pontificia Universidade Catolica do Rio Grande do Sul (PUCRS), Laboratory of Vertebrate Systematics. Av. Ipiranga, 6681 Partenon; 90619 - 900, Porto Alegre, Rio Grande do Sul, Brazil</mods:affiliation>
<mods:nameIdentifier id="D26C18FA8DB567D3D9F186B4D947B119" type="email">fidelismarra@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B4A6E772BDB6029CF67E0806FCA23DA3">text</mods:typeOfResource>
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<mods:title id="EABB1D288A0115E9C77EC572CCF3D440">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="0A899127657962A8B326AF8746533D34">
<mods:date id="5F60059C772A033DE1DC5084F3D82F1E">2023</mods:date>
<mods:detail id="88C7FA889785838C6EB80798F130C2DA" type="pubDate">
<mods:number id="0A322E9E5A1F4175A39B00B96E5D127E">2023-02-07</mods:number>
</mods:detail>
<mods:detail id="D890E5F64AD41C39568E6B0585555806" type="volume">
<mods:number id="3A12845A67C3F7684356E46D6A598AEA">1146</mods:number>
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<mods:start id="8002C31AE8E79745BDCDC1B8CDBB6F55">87</mods:start>
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<mods:url id="50FF0BC864A068DF19B92829CF9C6205">http://dx.doi.org/10.3897/zookeys.1146.94607</mods:url>
</mods:location>
<mods:classification id="C1A4D3E95120E423EDE408CA5FCE0971">journal article</mods:classification>
<mods:identifier id="2A4F0F69A935B00C5990F1A2F79B2EC0" type="DOI">http://dx.doi.org/10.3897/zookeys.1146.94607</mods:identifier>
<mods:identifier id="AFA33C21FE48F85B46DA8711F6179EC5" type="Pensoft-Pub">1313-2970-1146-87</mods:identifier>
<mods:identifier id="F67227D51BB74C3AD983CE218B2D25CF" type="ZooBank">9F4ADF001BE748789B98FB23B72B4435</mods:identifier>
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<treatment id="A7FC4D63F7815CC1BA31DD6FA74986DA" ID-GBIF-Taxon="209283673" LSID="urn:lsid:zoobank.org:act:0CD30C35-2263-491C-BB62-2494019740C8" httpUri="http://treatment.plazi.org/id/A7FC4D63F7815CC1BA31DD6FA74986DA" lastPageNumber="87" pageId="0" pageNumber="87" scope_family="Anomalepididae">
<subSubSection id="8494EFE628DA12DC1B3B4295B256E3B6" pageId="0" pageNumber="87" type="nomenclature">
<paragraph id="8BF57712E57BD85D5C3CA79D02BA53EE" pageId="0" pageNumber="87">
<taxonomicName id="070CF57B6B311143EA8A85AE24CA4EA2" LSID="https://zoobank.org/0CD30C35-2263-491C-BB62-2494019740C8" authority="Marra Santos, 2023" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis" status="sp. nov.">Liotyphlops palauophis</taxonomicName>
<taxonomicNameLabel id="3EF74169E9C61902F9C3EF50F952F2DF" pageId="0" pageNumber="87">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="BB634828F499FE32D3FF640F8925F842" pageId="0" pageNumber="87" type="description">
<paragraph id="75490F4AA8B87A485A5055A862B835A5" pageId="0" pageNumber="87">
<figureCitation id="4115A99C0C582216140023358E030D1A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype of Liotyphlops palauophis sp. nov., AMNH R- 9550, 361.2 mm TL, Colombia, neighborhood of Bogota. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1146.94607.figure1" httpUri="https://binary.pensoft.net/fig/806304" pageId="0" pageNumber="87">Figs 1</figureCitation>
<figureCitation id="08260E3BB700ADB531BAFC4EE2455A51" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal (top), lateral (center), and ventral (bottom) views of the head of Liotyphlops palauophis sp. nov., AMNH R- 9550, holotype. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1146.94607.figure2" httpUri="https://binary.pensoft.net/fig/806305" pageId="0" pageNumber="87">, 2</figureCitation>
<figureCitation id="A22696DF5783B28DCB34318A9B6A93CA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Three-dimensional reconstruction of the skull of Liotyphlops palauophis sp. nov., AMNH R- 9550, holotype, based on HRXCT data. A lateral view B dorsal view C ventral view with lower jaw partially digitally removed D anterior view E posterior view. Scale bar: 1 mm. Anatomical abbreviations: a: angular; bo: basioccipital; cb: compound bone; co: coronoid; d: dentary; ec: ectopterygoid; en: external naris; f: frontal; fo: foramen; fv: fenestra vomeronasalis; m: maxilla; mf: mental foramen; n: nasal; oc: occipital condyle; ooc: otico-occipital (fused prootic + opisthotic + exoccipital); p: parietal; pa: palatine; pbs: parabasisphenoid; pe: postorbital element; pf: prefrontal; pg: pterygoid; pm: premaxilla; q: quadrate; rp: retroarticular process; sf: surangular foramen; sm: septomaxilla; st: supratemporal; so: supraoccipital; tf: trigeminal foramen; v: vomer; vf: vomerine foramen." figureDoi="10.3897/zookeys.1146.94607.figure3" httpUri="https://binary.pensoft.net/fig/806306" pageId="0" pageNumber="87">, 3</figureCitation>
<figureCitation id="683C852DAF8CCD1C43226DA8229D7837" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Three-dimensional reconstruction of the skull of Liotyphlops palauophis sp. nov., AMNH R- 9550, holotype, based on HRXCT data. A transversal view B frontal view C sagittal view. Scale bar: 1 mm. Anatomical abbreviations: a: angular; an: acoustic nerve foramen; bo: basioccipital; cb: compound bone; cbp: compound bone prearticular component; cbs: compound bone surangular component; ci: conchal invagination; co: coronoid; cv: cavum vestibuli; d: dentary; dp: descensus parietalis; ec: ectopterygoid; en: external naris; f: frontal; fl: frontal laterally descending flange; fo: foramen; fpb: facial nerve palatine branch foramen; fs: frontal subolfactory process; fv: fenestra vomeronasalis; m: maxilla; Mc: Meckel's canal; mf: mental foramen; n: nasal; nl: nasal lateral flange; ns: medial nasal septum; oc: occipital condyle; on: optic nerve foramen; ooc: otico-occipital (fused prootic + opisthotic + exoccipital); p: parietal; pa: palatine; pbc: parabasal (Vidian) canal; pbs: parabasisphenoid; pe: postorbital element; pf: prefrontal; pg: pterygoid; pm: premaxilla; pvs: posterior vertical semicircular canal; q: quadrate; rp: retroarticular process; rstm: recessus scalae tympani medial aperture; s: stapes; sf: surangular foramen; sm: septomaxilla; sml: septomaxilla lateral flange; st: supratemporal; so: supraoccipital; tf: trigeminal foramen; trc: trigeminofacialis chamber; v: vomer; vc: vomeronasal cupola; vf: vomerine foramen; vn: vomeronasal nerve passage." figureDoi="10.3897/zookeys.1146.94607.figure4" httpUri="https://binary.pensoft.net/fig/806307" pageId="0" pageNumber="87">, 4</figureCitation>
<figureCitation id="6C346C4085FEEE7675F329744F1EABEA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Three-dimensional reconstruction of the lower jaw of Liotyphlops palauophis sp. nov., AMNH R- 9550, holotype, based on HRXCT data. A lateral view B medial view C dorsal view D ventral view E anterior view F posterior view. Scale bar: 1 mm. Anatomical abbreviations: a: angular; arf: articular fossa; cb: compound bone; cbp: compound bone prearticular component; cbs: compound bone surangular component; co: coronoid; d: dentary; Mc: Meckel's canal; mf: mental foramen; rp: retroarticular process; sf: surangular foramen." figureDoi="10.3897/zookeys.1146.94607.figure5" httpUri="https://binary.pensoft.net/fig/806308" pageId="0" pageNumber="87">, 5</figureCitation>
<figureCitation id="2326B8792E8EF56C44220565542655AF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Location of the holotype of Liotyphlops palauophis sp. nov. (black star), lectotype of Liotyphlops anops (white dot), and specimens of L. anops examined in this study (green dot).? = lack of detailed information about the type locality of L. palauophis sp. nov. and L. anops. This locality is based on information provided by Cope (1899)." figureDoi="10.3897/zookeys.1146.94607.figure6" httpUri="https://binary.pensoft.net/fig/806309" pageId="0" pageNumber="87">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="728782CA0B3B871506BCD54C340DE01E" pageId="0" pageNumber="87" type="reference_group">
<paragraph id="42DF30E4C49405BDE1713D1412D3651E" pageId="0" pageNumber="87">
<taxonomicName id="3D6F5907E1F19C8A4BA3C6FD3250946D" authority="Marra Santos, 2023" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Helminthophis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Helminthophis anops" order="Squamata" pageId="0" pageNumber="87" rank="species" species="anops" status="sp. nov.">Helminthophis anops</taxonomicName>
Cope, 1899 (in part). Syntype of
<taxonomicName id="3322A0EEB0556E9482485805CD1C57AF" authority="Marra Santos, 2023" authorityName="Marra Santos" authorityYear="2023" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="H. anops" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="anops" status="sp. nov.">H. anops</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BD6DF6815756E266933D2BCC1DD85623" pageId="0" pageNumber="87" type="type material">
<paragraph id="9DCE33F419C52C30F162E8A6DDD974A1" pageId="0" pageNumber="87">Type material.</paragraph>
<paragraph id="55E9EB5D36ED56C11654953FAB3B7EE8" pageId="0" pageNumber="87">
<emphasis id="CF6689D020FDD3A66D0D461F242BBC64" bold="true" pageId="0" pageNumber="87">
<emphasis id="D1DE6C9308342B69F0D0C037C40D0114" italics="true" pageId="0" pageNumber="87">Holotype</emphasis>
.
</emphasis>
AMNH R-9550, 361 mm TL, Colombia, neighborhood of Bogota, 1899.
</paragraph>
</subSubSection>
<subSubSection id="07FEEE5BA95B4D762A592BF0C2C23398" pageId="0" pageNumber="87" type="diagnosis">
<paragraph id="FFBEE268C60B75AC1DA9D76DACF2E6C3" pageId="0" pageNumber="87">Diagnosis.</paragraph>
<paragraph id="404F606F874284230B29E0944AF0E1EB" pageId="0" pageNumber="87">
<taxonomicName id="1381FC02614244F1792D1506543AE30C" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis">
<emphasis id="829C6999E6E7EAEFBD3158B8CC07EE43" italics="true" pageId="0" pageNumber="87">Liotyphlops palauophis</emphasis>
</taxonomicName>
sp. nov. is distinguished from all other
<taxonomicName id="543F96F8F68ACF1403C4A58F3FA66938" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops" order="Squamata" pageId="0" pageNumber="87" rank="genus">
<emphasis id="9E580C60540C2655915B0D978333B27C" italics="true" pageId="0" pageNumber="87">Liotyphlops</emphasis>
</taxonomicName>
by having the frontal scale divided (vs single) and a central foramen in the parabasisphenoid (vs foramen absent). It is further distinguished from
<taxonomicName id="D14EBA3A63B28A26A134BFAA97E33C05" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. albirostris" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="albirostris">
<emphasis id="3F05D1D979304B1479BC133D2B03DF8F" italics="true" pageId="0" pageNumber="87">L. albirostris</emphasis>
</taxonomicName>
,
<taxonomicName id="EA631A7A8C4F531FF0D61171855C0242" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. argaleus" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="argaleus">
<emphasis id="B56B86CC6DF73AF203AC3D1114CA142C" italics="true" pageId="0" pageNumber="87">L. argaleus</emphasis>
</taxonomicName>
,
<taxonomicName id="594956EBB6AE844252CDF52D70ED5264" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. bondensis" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="bondensis">
<emphasis id="3BDC2F56BB54025D88C052844173EEE2" italics="true" pageId="0" pageNumber="87">L. bondensis</emphasis>
</taxonomicName>
,
<taxonomicName id="65A1214C2EE6E9D5AF5A4FFFF7FE3911" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. caissara" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="caissara">
<emphasis id="0DE49162885772BD1B51A3FF138E6FED" italics="true" pageId="0" pageNumber="87">L. caissara</emphasis>
</taxonomicName>
,
<taxonomicName id="9F1DFDCCF647697814B431FE6F21890F" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. haadi" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="haadi">
<emphasis id="1EBF1930B4B580938DC465E2C37C4E44" italics="true" pageId="0" pageNumber="87">L. haadi</emphasis>
</taxonomicName>
,
<taxonomicName id="06A15281F6501F25104C235012287EEB" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. trefauti" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="trefauti">
<emphasis id="083C9B92A77B31442D8F1C5F273FB804" italics="true" pageId="0" pageNumber="87">L. trefauti</emphasis>
</taxonomicName>
, and
<taxonomicName id="7065BBE0949EC73FD10D0649A53281A8" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. wilderi" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="wilderi">
<emphasis id="129CBD255EC435E1273BFC57CB4B8FDD" italics="true" pageId="0" pageNumber="87">L. wilderi</emphasis>
</taxonomicName>
in having two scales (vs one scale) contacting the posterior edge of the nasal between the second supralabial and prefrontal. It is further distinguished from
<taxonomicName id="9F19263E27DFF59A929F0E8106C18063" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. beui" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="beui">
<emphasis id="D5797E8D4064B2F51364C130DC5AA777" italics="true" pageId="0" pageNumber="87">L. beui</emphasis>
</taxonomicName>
,
<taxonomicName id="98042C43CDC79989D1AB4CEE32C96D03" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. schubarti" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="schubarti">
<emphasis id="F1A36DAA2BA0B54B2CBF77B0623C76E2" italics="true" pageId="0" pageNumber="87">L. schubarti</emphasis>
</taxonomicName>
,
<taxonomicName id="B02A3A6721FB4EE2C917510136C753DF" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. taylori" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="taylori">
<emphasis id="6405FBD94BDDD3B8AF16BDD94BACD0FA" italics="true" pageId="0" pageNumber="87">L. taylori</emphasis>
</taxonomicName>
, and
<taxonomicName id="D26AF76F560AFC0CB8505EBD96CB9069" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. ternetzii" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="ternetzii">
<emphasis id="F18579282C547DEADF4FF7C0CB1C14D5" italics="true" pageId="0" pageNumber="87">L. ternetzii</emphasis>
</taxonomicName>
by having four (vs three) scales contacting the posterior edge of the prefrontal. It is distinguished from
<taxonomicName id="7EA271DF2A5D3D087284B9D2A3929ED9" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. anops" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="anops">
<emphasis id="2BE934948CC0782B3C6EF96B84905C2B" italics="true" pageId="0" pageNumber="87">L. anops</emphasis>
</taxonomicName>
by having 28/26/26 scales around the body and 19 subcaudal scales (vs 26/24/24 scales around the body and 12-14 subcaudal scales), and from
<taxonomicName id="01CF927C48261B7AB6EC85B741930AD6" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. sousai" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="sousai">
<emphasis id="8FB6413F771604305993A5A50C6726C0" italics="true" pageId="0" pageNumber="87">L. sousai</emphasis>
</taxonomicName>
in having 573 dorsal scales and 561 ventral scales (vs 439 dorsal scales and 427 ventral scales).
</paragraph>
<caption id="673553C27C5593B5D205BAC4A112FB1E" doi="10.3897/zookeys.1146.94607.figure1" httpUri="https://binary.pensoft.net/fig/806304" pageId="0" pageNumber="87" start="Figure 1" startId="F1">
<paragraph id="3B1460AD163236E2E10001E0373402D1" pageId="0" pageNumber="87">
<emphasis id="308D93636F9ED05852A5F7001A0CDB24" bold="true" pageId="0" pageNumber="87">Figure 1.</emphasis>
Holotype of
<taxonomicName id="BDAE9200CD8BFB2640173F06BA8BB27B" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis">
<emphasis id="BDAA2DEC6B775CBF26561BA9B52AFE3E" italics="true" pageId="0" pageNumber="87">Liotyphlops palauophis</emphasis>
</taxonomicName>
sp. nov., AMNH R-9550, 361.2 mm TL, Colombia, neighborhood of Bogota. Scale bar: 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="3D45684AA9361AD85CCF4FAC98FBCA7E" pageId="0" pageNumber="87" type="description">
<paragraph id="8195C62C11E441AA7C62375C9ED64E23" pageId="0" pageNumber="87">Description.</paragraph>
<paragraph id="72C4AAAE9707B9744747BBBBC80E9C0F" pageId="0" pageNumber="87">
Meristic data in Table
<tableCitation id="B5C798048717CB48D3CAC8E8BE16B8CA" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Meristic characters of species of Liotyphlops from the specimens examined in this study, presented as ranges with minimum, maximum, and mode in parentheses. SPEP = number of scales contacting posterior edge of prefrontal; SPEN = number of scales contacting posterior edge of nasal between second supralabial and prefrontal; SFVRD = number of scales in the first vertical row of dorsals; SL = number of supralabial scales; IL = number of infralabial scales; ASR = number of anterior scale rows around body; MSR = number of scale rows around the midbody; PSR = number of posterior scale rows around body; DSR = number of dorsal scale rows; VSR = number of ventral scales rows; SC = number of subcaudal scales. n = number of specimens examined in this study. a = number of specimens examined by Santos and Reis (2018). b = number of specimens examined by Centeno et al. (2010). c = number of specimens examined by Freire et al. (2007). d = number of specimens examined by Linares-Vargas et al. (2021)." httpUri="http://table.plazi.org/id/4D2439378301DDACB18EE1530225E3E6" pageId="0" pageNumber="87" tableUuid="4D2439378301DDACB18EE1530225E3E6">1</tableCitation>
. Total length 361.2 mm, head length 5.3 mm (1.5% TL), snout-vent length 353 mm (97.7% TL), tail length 8.2 mm (2.3% TL), head width 3.8 mm (71.7% HL), and head height 3.1 mm (58.5% HL). Body covered with cycloid scales. Rostral scale large, longer than wide, contacting nasals anterolaterally, prefrontals laterally, and divided frontal posteriorly. Pair of triangular prefrontals bordered anterolaterally by rostral, ventrally by large divided nasal, and dorsoposteriorly by frontal. Posterior edge of prefrontals passing posterior edge of rostral. Frontal scale divided. Nasal scale divided and bordered anteriorly by rostral, dorsally by prefrontal, ventrally by first and second supralabials, and posteriorly by two scales that lie between prefrontal and second supralabial. Eye spot not visible. Four scales contacting posterior edge of prefrontal (three cycloid scales + frontal). Two scales contacting posterior edge of nasal between second supralabial and prefrontal. Six scales in first vertical row of dorsal scales. Mental triangular, not divided, wider than long, contacting first infralabials. Supralabials four, infralabials three. Scales around body 28/26/26. Dorsal scales 573, vental scales 561, and subcaudal scales 19.
</paragraph>
<caption id="4D2439378301DDACB18EE1530225E3E6" ID-Table-UUID="4D2439378301DDACB18EE1530225E3E6" httpUri="http://table.plazi.org/id/4D2439378301DDACB18EE1530225E3E6" pageId="0" pageNumber="87" start="Table 1" startId="T1">
<paragraph id="DCE3083AF2A9A1A1CFD0A287F6B55732" pageId="0" pageNumber="87">
<emphasis id="7EEF472C2A9E005CA62D64A511BA20AF" bold="true" pageId="0" pageNumber="87">Table 1.</emphasis>
Meristic characters of species of
<taxonomicName id="F68BB64C557B5D2DC9309E008BEA177D" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops" order="Squamata" pageId="0" pageNumber="87" rank="genus">
<emphasis id="22EC84D542D3A460F7A064FE4707D368" italics="true" pageId="0" pageNumber="87">Liotyphlops</emphasis>
</taxonomicName>
from the specimens examined in this study, presented as ranges with minimum, maximum, and mode in parentheses.
<emphasis id="2A9963FACBAAB8ED26C5ADEC286633DF" bold="true" pageId="0" pageNumber="87">SPEP</emphasis>
= number of scales contacting posterior edge of prefrontal;
<emphasis id="526B4904AA407FDDAD707DE9C6AF80D3" bold="true" pageId="0" pageNumber="87">SPEN</emphasis>
= number of scales contacting posterior edge of nasal between second supralabial and prefrontal;
<emphasis id="B8FE744EF4B133CA8516BB992CAB6CF8" bold="true" pageId="0" pageNumber="87">SFVRD</emphasis>
= number of scales in the first vertical row of dorsals;
<emphasis id="9C53DDA0A2B4B7B877111506F2432A31" bold="true" pageId="0" pageNumber="87">SL</emphasis>
= number of supralabial scales;
<emphasis id="A44F835030FA6F7445D9F3D37369A9D3" bold="true" pageId="0" pageNumber="87">IL</emphasis>
= number of infralabial scales;
<emphasis id="109FA46B9E974BAD57A195E05428822B" bold="true" pageId="0" pageNumber="87">ASR</emphasis>
= number of anterior scale rows around body;
<emphasis id="025C404309232289D4BFFD1F191842DC" bold="true" pageId="0" pageNumber="87">MSR</emphasis>
= number of scale rows around the midbody;
<emphasis id="69022760DD0C4231599788446ED83431" bold="true" pageId="0" pageNumber="87">PSR</emphasis>
= number of posterior scale rows around body;
<emphasis id="ACAAAEFD0BCA9229D208DF1CBB5318D4" bold="true" pageId="0" pageNumber="87">DSR</emphasis>
= number of dorsal scale rows;
<emphasis id="C9A1459B7DE1A77212A69EA55BF24C97" bold="true" pageId="0" pageNumber="87">VSR</emphasis>
= number of ventral scales rows;
<emphasis id="33C4BC6002E9E6CC4A3DDA4B83F5862E" bold="true" pageId="0" pageNumber="87">SC</emphasis>
= number of subcaudal scales.
<emphasis id="82391631560D2F9A746018E006A29513" bold="true" pageId="0" pageNumber="87">n</emphasis>
= number of specimens examined in this study.
<emphasis id="967812DA69AD913A8C09ED1E78FFAFEB" bold="true" pageId="0" pageNumber="87">a</emphasis>
= number of specimens examined by
<bibRefCitation id="ABAE799E11253BCE5B86AADFFF424B2F" DOI="https://doi.org/10.1643/CH-18-081" author="Santos, FJM" journalOrPublisher="Copeia" pageId="0" pageNumber="87" pagination="507 - 514" refId="B19" refString="Santos, FJM, Reis, RE, 2018. Two new blind snake species of the genus Liotyphlops Peters, 1881 (Serpentes: Anomalepididae), from Central and South Brazil. Copeia 106 (3): 507 - 514, DOI: https://doi.org/10.1643/CH-18-081" title="Two new blind snake species of the genus Liotyphlops Peters, 1881 (Serpentes: Anomalepididae), from Central and South Brazil." url="https://doi.org/10.1643/CH-18-081" volume="106" year="2018">Santos and Reis (2018)</bibRefCitation>
.
<emphasis id="7DF65C044BD234CC6D3B035CCA3AFAE4" bold="true" pageId="0" pageNumber="87">b</emphasis>
= number of specimens examined by
<bibRefCitation id="82F926CBE8E156DECEF98406C14495AF" DOI="https://doi.org/10.1655/08-079.1" author="Centeno, FC" journalOrPublisher="Herpetologica" pageId="0" pageNumber="87" pagination="86 - 91" refId="B6" refString="Centeno, FC, Sawaya, RJ, Germano, VJ, 2010. A new species of Liotyphlops (Serpentes: Anomalepididae) from the Atlantic Coastal Forest in southeastern Brazil. Herpetologica 66 (1): 86 - 91, DOI: https://doi.org/10.1655/08-079.1" title="A new species of Liotyphlops (Serpentes: Anomalepididae) from the Atlantic Coastal Forest in southeastern Brazil." url="https://doi.org/10.1655/08-079.1" volume="66" year="2010">Centeno et al. (2010)</bibRefCitation>
.
<emphasis id="FFD55C4AF787CBD00FEA0CE215A6AA8A" bold="true" pageId="0" pageNumber="87">c</emphasis>
= number of specimens examined by
<bibRefCitation id="A28240A87B9D7EA91F545477845B5DE8" DOI="https://doi.org/10.11646/zootaxa.1393.1.2" author="Freire, EMX" journalOrPublisher="Zootaxa" pageId="0" pageNumber="87" pagination="19 - 26" refId="B10" refString="Freire, EMX, Caramaschi, U, Argolo, AJS, 2007. A new species of Liotyphlops (Serpentes: Anomalepididae) from the Atlantic Rain Forest of northeastern Brazil. Zootaxa 1393 (1): 19 - 26, DOI: https://doi.org/10.11646/zootaxa.1393.1.2" title="A new species of Liotyphlops (Serpentes: Anomalepididae) from the Atlantic Rain Forest of northeastern Brazil." url="https://doi.org/10.11646/zootaxa.1393.1.2" volume="1393" year="2007">Freire et al. (2007)</bibRefCitation>
.
<emphasis id="26F895B32AD1FB2878BC78ACDBA4EE15" bold="true" pageId="0" pageNumber="87">d</emphasis>
= number of specimens examined by
<bibRefCitation id="7AD5C1518ADFCF3758206D2EF1F1720F" DOI="https://doi.org/10.1002/ar.24730" author="Linares-Vargas, CA" journalOrPublisher="The Anatomical Record" pageId="0" pageNumber="87" pagination="2264 - 2278" refId="B12" refString="Linares-Vargas, CA, Bolivar-Garcia, W, Herrera-Martinez, A, Osorio-Dominguez, D, Ospina, OE, Thomas, R, Daza, JD, 2021. The status of the anomalepidid snake Liotyphlops albirostris and the revalidation of three taxa based on morphology and ecological niche models. The Anatomical Record 304 (10): 2264 - 2278, DOI: https://doi.org/10.1002/ar.24730" title="The status of the anomalepidid snake Liotyphlops albirostris and the revalidation of three taxa based on morphology and ecological niche models." url="https://doi.org/10.1002/ar.24730" volume="304" year="2021">Linares-Vargas et al. (2021)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="C4575EB2A579CA1E8754D13C9BEFE134" pageId="0" pageNumber="87">
<table id="ABC247D7AAE66481B3B957EC57D9157B" pageId="0" pageNumber="87">
<tr id="1EDA158B2C3787F502C983F9314563D9" pageId="0" pageNumber="87">
<th id="A8257CF32BD64468A4CEB7E65A06F72C" colspan="1" pageId="0" pageNumber="87" rowspan="1">Species/Count</th>
<th id="045847ECC8337EAC3874E4B41F062C54" colspan="1" pageId="0" pageNumber="87" rowspan="1">n</th>
<th id="88980FDC544DBF90E7A5BAA9C73169D2" colspan="1" pageId="0" pageNumber="87" rowspan="1">SPEP</th>
<th id="13699C20D9AB8D4C8C1E9F05A80E29B6" colspan="1" pageId="0" pageNumber="87" rowspan="1">SPEN</th>
<th id="3C2257E1F114770E67FF0597F4A7A51C" colspan="1" pageId="0" pageNumber="87" rowspan="1">SFVRD</th>
<th id="AED849032A7F06AAE1DA22915B30BC75" colspan="1" pageId="0" pageNumber="87" rowspan="1">SL</th>
<th id="484354DF4B72D49F6BA9930DB9D2939A" colspan="1" pageId="0" pageNumber="87" rowspan="1">IL</th>
<th id="CF20BCD09F3C7A186A7F4BCE2B298BBD" colspan="1" pageId="0" pageNumber="87" rowspan="1">ASR</th>
<th id="7B0B41C4D0BF43972AEBB7216B36A0BD" colspan="1" pageId="0" pageNumber="87" rowspan="1">MSR</th>
<th id="D2E14B5049BF566AAC5A6BEB1DF1B266" colspan="1" pageId="0" pageNumber="87" rowspan="1">PSR</th>
<th id="CF1CC3F7DFE4722353EC5152D0BCEDAA" colspan="1" pageId="0" pageNumber="87" rowspan="1">DSR</th>
<th id="96B919AFABDE96AD658A5383A8495CE1" colspan="1" pageId="0" pageNumber="87" rowspan="1">VSR</th>
<th id="9F9AF6CB43699EB8783D07888CCCA185" colspan="1" pageId="0" pageNumber="87" rowspan="1">SC</th>
</tr>
<tr id="757FD4D533134D95F8FEBD2F4E8CFD1B" pageId="0" pageNumber="87">
<td id="ECF78008810266BD3590876FAD1B1A26" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="B3A71A8A679FCB622A830B1E5E4E024F" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. albirostris" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="albirostris">
<emphasis id="11E9FF5F362A249D5718B4E263CF7953" italics="true" pageId="0" pageNumber="87">L. albirostris</emphasis>
</taxonomicName>
<emphasis id="F94A7A48A2115786953F9310879E41F5" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="584AFA716BD8E2C95471DF9BF17B8004" colspan="1" pageId="0" pageNumber="87" rowspan="1">6</td>
<td id="5F17C806C27F28369DF2A2425973A347" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="F5F4B798F3599163A4CA3F50327C6398" colspan="1" pageId="0" pageNumber="87" rowspan="1">1-1(1)</td>
<td id="5D3490D5678A7AD58556997782E714BE" colspan="1" pageId="0" pageNumber="87" rowspan="1">5-5(5)</td>
<td id="46FF154CD967296E7226F1392D009549" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="5CBD5C529075ED3EEDF1291A8C58EE07" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="8095C9C870364EF5C72D160C851CC9FF" colspan="1" pageId="0" pageNumber="87" rowspan="1">24-26(26)</td>
<td id="05A522A93A49C499D7F9549DBAFF4225" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-22(22)</td>
<td id="0BC7E7B34073B12C636AD4CD0589F324" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-22(22)</td>
<td id="26132B91E79E62684777EFA15FEAF6F0" colspan="1" pageId="0" pageNumber="87" rowspan="1">432-478</td>
<td id="757C8F47C8F056BE210F62C982932707" colspan="1" pageId="0" pageNumber="87" rowspan="1">417-453</td>
<td id="6B22ECE1F26E5545EE05FFFC0D30FF8C" colspan="1" pageId="0" pageNumber="87" rowspan="1">12-17(12)</td>
</tr>
<tr id="78487D90C4DEC83FAEF336DFA7D596A7" pageId="0" pageNumber="87">
<td id="0C05F5BB30E45A590936E18ECA6E5ED1" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="C8963345B637E480E6A048E3A18AD6AA" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. anops" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="anops">
<emphasis id="611E26BEA1770DE0EFA4E523E9BDD5F9" italics="true" pageId="0" pageNumber="87">L. anops</emphasis>
</taxonomicName>
<emphasis id="152C21BCC2C854CB9CD19A762C855D0D" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="C6378E8AE4B59AEBD63586488EB811F7" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="AD1CBB990CDC407744D8CCCFA871EC35" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="086D7159A952ADB91AEBCC6E3E785837" colspan="1" pageId="0" pageNumber="87" rowspan="1">2-2(2)</td>
<td id="9A32F2A98A5D66CEE8EDD17224737B8F" colspan="1" pageId="0" pageNumber="87" rowspan="1">5-6(5)</td>
<td id="3D675F85417A98EDA67CE5A6CECFA60E" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="9F302B593D8189593A925E3321366DA3" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="26DFF08788CBCCE9DE8F4ED17B9D8293" colspan="1" pageId="0" pageNumber="87" rowspan="1">26-26(26)</td>
<td id="9A3ADFC8FAA146981D8312B67840950A" colspan="1" pageId="0" pageNumber="87" rowspan="1">24-24(24)</td>
<td id="9598676AE14DAA7C1F7A0CD905201823" colspan="1" pageId="0" pageNumber="87" rowspan="1">24-24(24)</td>
<td id="81C373A8A68FAA1966D19A5883CD49D5" colspan="1" pageId="0" pageNumber="87" rowspan="1">562-597</td>
<td id="4BABD68DB468FDF423CFF32A6294AA6B" colspan="1" pageId="0" pageNumber="87" rowspan="1">531-572</td>
<td id="C46CFB2AEC6437CAC9135E6D3269F976" colspan="1" pageId="0" pageNumber="87" rowspan="1">12-14</td>
</tr>
<tr id="3A2A050557DBA7E89981B0F3950EE163" pageId="0" pageNumber="87">
<td id="8CBAE20D53E0A7767EC885D77F59B615" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="A9B59FADE145C57E83D53EE608BC8A5E" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. argaleus" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="argaleus">
<emphasis id="BB30A2875FBB48B395B5024B33A40E42" italics="true" pageId="0" pageNumber="87">L. argaleus</emphasis>
</taxonomicName>
<emphasis id="756C7A6193ABAEE50F7CD9E46140B818" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="9263F2F1F72DD5D13D7C302DD4F13D0A" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="9BEF82D96B9105118E85CC60CF76E185" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="38CC017D0EC39B0E9E5217F89E64DFCA" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="1A67748CAEA2AD7F2514D7E1A1538953" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="CCCAEC577CBD4A6A93F5DCA7BE37F9AD" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="C68761A7D3D830470BC2AD9D3C61236D" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="C6798A4794F2D60077B1FDC9F4E792AC" colspan="1" pageId="0" pageNumber="87" rowspan="1">25</td>
<td id="A78D46BFE5BE2AF014A1FABE0F9A83A8" colspan="1" pageId="0" pageNumber="87" rowspan="1">23</td>
<td id="F4E660F8A6BFAE0CB32EAEAA93C8578A" colspan="1" pageId="0" pageNumber="87" rowspan="1">22</td>
<td id="B30BD0C9CB7E5ADE9187C04BEC69045F" colspan="1" pageId="0" pageNumber="87" rowspan="1">497</td>
<td id="04EFA09BDB82720BBFE7D4881F88D8CB" colspan="1" pageId="0" pageNumber="87" rowspan="1">472</td>
<td id="8E13D9AC4706526DA0FEB6E6E0EFCF06" colspan="1" pageId="0" pageNumber="87" rowspan="1">16</td>
</tr>
<tr id="B4FD94A1E69A6F163DD13719D878B620" pageId="0" pageNumber="87">
<td id="33CA6865F121EDBED2CF5B5D6C8AABAD" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="A25C41BBBC9DDC3E5540810D27A1FC7E" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. beui" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="beui">
<emphasis id="B6A8F68BEFE3A065321820672193BB45" italics="true" pageId="0" pageNumber="87">L. beui</emphasis>
</taxonomicName>
<emphasis id="8836D7B583AC6E14E62C2D6C272BF793" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="64549CC590ED4303B13A8D8728C00632" colspan="1" pageId="0" pageNumber="87" rowspan="1">50</td>
<td id="C06D7A930A2F6F4B2553AA1F590FFEA1" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="854A1F9AB3E4961BA38B470004B46873" colspan="1" pageId="0" pageNumber="87" rowspan="1">2-2(2)</td>
<td id="2F6CF6BA058CCEF6000FFA89FB5C7133" colspan="1" pageId="0" pageNumber="87" rowspan="1">5-6(5)</td>
<td id="20DB54CCF3E26601BEF744A5C2738C39" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="0B743CBDD7C7C236C61BB8A232D18932" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="CC5D0328206523F784BBD88764E13295" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-26(22)</td>
<td id="C37D7FB71B40F5CF27F5B9B9EF2DC053" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-22(22)</td>
<td id="6C1E47B80116186D9146EC0B634BCE8C" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-22(20)</td>
<td id="BE14A2A3A2D5EA216AA4659CA592131F" colspan="1" pageId="0" pageNumber="87" rowspan="1">366-532(453)</td>
<td id="A916868C860AC969E1EA67143411C03C" colspan="1" pageId="0" pageNumber="87" rowspan="1">348-511(364)</td>
<td id="CE1EEFC5FC8079514B996515B1B45D78" colspan="1" pageId="0" pageNumber="87" rowspan="1">11-22(12)</td>
</tr>
<tr id="74FE540FC7041DB22B35050E1291E3F2" pageId="0" pageNumber="87">
<td id="9E81C7341F7C7CA7E5B824B496C41588" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="95AB316B2EF4C6232883E8F59925EA2C" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. bondensis" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="bondensis">
<emphasis id="FDC6A3F19AC1D1A5B0E8B4C559585A3B" italics="true" pageId="0" pageNumber="87">L. bondensis</emphasis>
</taxonomicName>
<emphasis id="33631E974C0856E7AFE9385058E2BCF0" bold="true" pageId="0" pageNumber="87">d</emphasis>
</td>
<td id="20447E1D23255DF21F476302DCE716A7" colspan="1" pageId="0" pageNumber="87" rowspan="1">17</td>
<td id="63EC4584A796C452BF82E4CE5E18097F" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="8CA19FE114FB70CF1BE40531E3010DA2" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="DB6B7C0CF899BF3C03882FDA36BE57AD" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="6FF384B532716375CFC8F65840A3A476" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="79AE6350EC5B723952D60FCC3D71BD66" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="59AF928FE64704226E11ED11BA66E459" colspan="1" pageId="0" pageNumber="87" rowspan="1">24</td>
<td id="EAC622696B20664CFDC928D52AB88924" colspan="1" pageId="0" pageNumber="87" rowspan="1">22</td>
<td id="77CB54EE7DF8F379FE79B498B0590628" colspan="1" pageId="0" pageNumber="87" rowspan="1">22</td>
<td id="423CE85453857FA88DBB24A1390AA4B0" colspan="1" pageId="0" pageNumber="87" rowspan="1">363-449</td>
<td id="DD863802F7FDB785AF7AE638610FD4EF" colspan="1" pageId="0" pageNumber="87" rowspan="1">347-434</td>
<td id="0DA91B8915ECF4278E1A7CD4A52458CD" colspan="1" pageId="0" pageNumber="87" rowspan="1">11-17</td>
</tr>
<tr id="4A0F8FDB2E95CAEA42A11C0B4D6F1BF8" pageId="0" pageNumber="87">
<td id="7FB5C41338E5107A10251555239B96D5" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="9477FD470FA0E108DA53436D24B42D55" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. caissara" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="caissara">
<emphasis id="301918B8881A1D4A63F79A16764EE543" italics="true" pageId="0" pageNumber="87">L. caissara</emphasis>
</taxonomicName>
<emphasis id="F09DB0A14A21894FD8772B486E58F240" bold="true" pageId="0" pageNumber="87">b</emphasis>
</td>
<td id="79A376B250E6A62880BF390A5D4B0006" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="D686A6BCAFB363A51B1B066805CB4030" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="70467B6DFB0C39CD155B19D47E1E81D5" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="B2273C174987E78F77C0982DE9D52503" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="0091024CB3221B8103161ACB2FCA8D2F" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="0E4C15E385FCCBC31A6B39F912856B5F" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="5620C7DAC8845AF63E998212900894AA" colspan="1" pageId="0" pageNumber="87" rowspan="1">22</td>
<td id="614F89D9ED2A9FA3A83372833E724509" colspan="1" pageId="0" pageNumber="87" rowspan="1">20</td>
<td id="6D7B3D433020829ADA83F75AF2E70191" colspan="1" pageId="0" pageNumber="87" rowspan="1">20</td>
<td id="F6E75A17F3D0300AE0B1873B60AEC88D" colspan="1" pageId="0" pageNumber="87" rowspan="1">326</td>
<td id="A62F9B1A94FDC6654445B0CC8682E122" colspan="1" pageId="0" pageNumber="87" rowspan="1">308</td>
<td id="22D552049BBE0555B28C6C00B0796757" colspan="1" pageId="0" pageNumber="87" rowspan="1">10</td>
</tr>
<tr id="565063A8E2CCBE53B4A6F4CB8CFCA4C7" pageId="0" pageNumber="87">
<td id="BD7631524AC9AF9DD4314E2A53C06144" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="2B3D920F8141D62531A637EBDD5A7AA4" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. haadi" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="haadi">
<emphasis id="806161D8D38001A104E9CD8945B79A3C" italics="true" pageId="0" pageNumber="87">L. haadi</emphasis>
</taxonomicName>
<emphasis id="268F9F5B7B21739953E6F35D0811DFD3" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="0F796DC3F56F43D28B3B41BAD7443082" colspan="1" pageId="0" pageNumber="87" rowspan="1">2</td>
<td id="A05369DF4592EF95BADE35B4FF9B05DC" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="BAE276544F7D690804075D53C6BD185A" colspan="1" pageId="0" pageNumber="87" rowspan="1">1-1(1)</td>
<td id="7BDC36411A2E7CE9CCF90B43328FD185" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="0E84153603310BE8F1A7B8A571B4C38B" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="FBE0405A65523DAAAB8B94FBC975ACF2" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="DE5FC5040B275D1A85C54227FB06AAB7" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-20(20)</td>
<td id="DD1896376A5F7D2990B9DB7BBA135E9C" colspan="1" pageId="0" pageNumber="87" rowspan="1">19-20</td>
<td id="D4062FC591101C486EA970473F071A89" colspan="1" pageId="0" pageNumber="87" rowspan="1">18-20</td>
<td id="98DBEB0BE271BC10C1200B74378B951C" colspan="1" pageId="0" pageNumber="87" rowspan="1">333-384</td>
<td id="5E52DE863CB9C9FEF5123B11581946E9" colspan="1" pageId="0" pageNumber="87" rowspan="1">309-348</td>
<td id="D2244EB6E60C45386A18CBDD99B452FD" colspan="1" pageId="0" pageNumber="87" rowspan="1">11-12</td>
</tr>
<tr id="33F94C0EAAA944B177291634BF0DC89E" pageId="0" pageNumber="87">
<td id="4DFF16E4EAD7E030DDA1CAF914BD1F71" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="185186BADE59767EDCECDF38DA11210B" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. palauophis" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="palauophis">
<emphasis id="EA0947A5F3330607D1A0A84D80460A62" italics="true" pageId="0" pageNumber="87">L. palauophis</emphasis>
</taxonomicName>
</td>
<td id="FD73F5C506B20416BCC82BD6FD0A158C" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="85C1D97F92704F419D6961AAA9A4E898" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="116E0B66087E0A304B7EB9BFE81DE05D" colspan="1" pageId="0" pageNumber="87" rowspan="1">2</td>
<td id="AFA1FB5F803887760CFB86EEA6A47811" colspan="1" pageId="0" pageNumber="87" rowspan="1">6</td>
<td id="BB45BB5CE46295A393D123839823A473" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="45CF1836C7AD63E5F0A7CBFB06143ACA" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="372185BE804FA3FBC8AFDCD28524E58C" colspan="1" pageId="0" pageNumber="87" rowspan="1">28</td>
<td id="762906142C3612FA7A64E10593399EE3" colspan="1" pageId="0" pageNumber="87" rowspan="1">26</td>
<td id="801A796B3B7179C8D32C3821A59EC471" colspan="1" pageId="0" pageNumber="87" rowspan="1">26</td>
<td id="EE6A70C662D1C53E76245BF876831B06" colspan="1" pageId="0" pageNumber="87" rowspan="1">573</td>
<td id="8DECB35864E80D491A98F74E7100842F" colspan="1" pageId="0" pageNumber="87" rowspan="1">561</td>
<td id="D3B42390F3E9418956E01028F34EBAC2" colspan="1" pageId="0" pageNumber="87" rowspan="1">19</td>
</tr>
<tr id="AE9695FA0B00B94CC2BB818CBEEE26D7" pageId="0" pageNumber="87">
<td id="EE7C7980823513D5DA46E1C956E5BCDD" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="EBDCC150BF976BD43409B241F1CDA899" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. schubarti" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="schubarti">
<emphasis id="BCD5E5BC4BEFAD4A3138BBAF3C4AE6A3" italics="true" pageId="0" pageNumber="87">L. schubarti</emphasis>
</taxonomicName>
<emphasis id="65FA5E8D5DC5074A1B44F108817C059B" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="1FA9CC35624D8B8F090226FF474EECBF" colspan="1" pageId="0" pageNumber="87" rowspan="1">5</td>
<td id="490BC5AC9A8D212353AE29194F3266F3" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="F2D388350B49B82ABE9B2E7BBA4FA2E4" colspan="1" pageId="0" pageNumber="87" rowspan="1">2-2(2)</td>
<td id="B13CE2898600C73958E3FB9D6A80B34A" colspan="1" pageId="0" pageNumber="87" rowspan="1">5-5(5)</td>
<td id="8A6FE8BEA6888F405B7BD1AD1FE0395A" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="E3A82E45B40E3DB70A514D28159ED91A" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="D13BB47BCB64C124DCB08FAEB12F423F" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-24(22)</td>
<td id="CFD7517AC4D5DB36B771BA796740838E" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-22(20)</td>
<td id="97B3503F9BF15B0511B3501251E871D0" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-20(20)</td>
<td id="6C7A703DBBFB714373298D0F13F226DE" colspan="1" pageId="0" pageNumber="87" rowspan="1">417-463</td>
<td id="072237F577FBBF68D93029088D5A005C" colspan="1" pageId="0" pageNumber="87" rowspan="1">398-451</td>
<td id="F90BC339218F1A1836443A0E953F1117" colspan="1" pageId="0" pageNumber="87" rowspan="1">11-14(13)</td>
</tr>
<tr id="320ED321A605F65F6D7A264665E6AB2A" pageId="0" pageNumber="87">
<td id="254FEADF55642080E21FA87CA79F845B" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="5FBCA9A2BDFDF6FA746ABBCB58231F95" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. sousai" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="sousai">
<emphasis id="E30EB1FA7B4DAA65F1A3EDCF0FA2F951" italics="true" pageId="0" pageNumber="87">L. sousai</emphasis>
</taxonomicName>
<emphasis id="6FC939A7AB5BEDFCBC848CBE76936DDA" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="A112778F8AEAC1C4276E92B7A34C1AE1" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="7C077054C897B887E7CFA432B6625A77" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="F47C292A78F0CA7A67362B0776EDA558" colspan="1" pageId="0" pageNumber="87" rowspan="1">2</td>
<td id="51C5AC9CEB1DF8CB3AC4B065C0CAAC43" colspan="1" pageId="0" pageNumber="87" rowspan="1">6</td>
<td id="2C1D4CD51D87F4B61B18550B74637324" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="802138D45D90B16464DE446232ECCC7C" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="9F1E1EBA7916865835180640C6C62D77" colspan="1" pageId="0" pageNumber="87" rowspan="1">24</td>
<td id="FD7BE2B64CB4B777D24C74838042B650" colspan="1" pageId="0" pageNumber="87" rowspan="1">22</td>
<td id="2279273925E50EEE3A31E2CDFEF05604" colspan="1" pageId="0" pageNumber="87" rowspan="1">20</td>
<td id="470B80856492B4875985B802D829235D" colspan="1" pageId="0" pageNumber="87" rowspan="1">439</td>
<td id="8399B2B403A1953AAA1F52624B8E60E0" colspan="1" pageId="0" pageNumber="87" rowspan="1">427</td>
<td id="C6D246C734987CF5B17D905CBEAA242C" colspan="1" pageId="0" pageNumber="87" rowspan="1">13</td>
</tr>
<tr id="31E9187006617BF72043C16708E844CF" pageId="0" pageNumber="87">
<td id="E0F53B8C23C4503A2181138DB18E099D" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="497B5452FDCED7D41BB4B61872B3C44C" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. taylori" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="taylori">
<emphasis id="9836BD8E123052C4BEB0CB7821780712" italics="true" pageId="0" pageNumber="87">L. taylori</emphasis>
</taxonomicName>
<emphasis id="50BA7E0489BB32B3EF7493B5D5E212D3" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="AA5A7A459F1AD4891ECCA71906657860" colspan="1" pageId="0" pageNumber="87" rowspan="1">1</td>
<td id="F8C7509DC85C62FA4CD23A6C11817D8F" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="E7424484BE2C76DBB98953540BBDE35C" colspan="1" pageId="0" pageNumber="87" rowspan="1">2</td>
<td id="168E4B90D486B8EAD59A53976E62D0D6" colspan="1" pageId="0" pageNumber="87" rowspan="1">5</td>
<td id="31E1E2B2D12E2CB5319E7D586E70CF36" colspan="1" pageId="0" pageNumber="87" rowspan="1">4</td>
<td id="0D4D4A650B202BFADBFEF9176756C9D7" colspan="1" pageId="0" pageNumber="87" rowspan="1">2</td>
<td id="3F6317C7C292721FD729F1168DFBA036" colspan="1" pageId="0" pageNumber="87" rowspan="1">22</td>
<td id="79DD0DE25D143E78081C6EFA60E0B368" colspan="1" pageId="0" pageNumber="87" rowspan="1">20</td>
<td id="BF769F8935DEE3733B86A9BB527EDBAB" colspan="1" pageId="0" pageNumber="87" rowspan="1">20</td>
<td id="775B4F94FE6F0FECA6085B4CFB255E00" colspan="1" pageId="0" pageNumber="87" rowspan="1">455</td>
<td id="48C662783FF1BF35924B8686B7EECCF8" colspan="1" pageId="0" pageNumber="87" rowspan="1">441</td>
<td id="9BB4F95E6A1303A5C5FBCF37FCA03B19" colspan="1" pageId="0" pageNumber="87" rowspan="1">14</td>
</tr>
<tr id="95D5BC510053CEA184E22775C685C891" pageId="0" pageNumber="87">
<td id="6D10AE26E2DC7086FFA11E7C8E97610E" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="51291092DB570EFAE98AC23CC4587C67" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. ternetzii" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="ternetzii">
<emphasis id="F344A220647EC9E869D08ECAD18171F6" italics="true" pageId="0" pageNumber="87">L. ternetzii</emphasis>
</taxonomicName>
<emphasis id="23818DD915EDB3129341F7BA0CC507BA" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="7F7BEAB247C4594EA4A86EA53944C266" colspan="1" pageId="0" pageNumber="87" rowspan="1">50</td>
<td id="51D3D130E1FA6A49DE8F5E3C1ABE093D" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="DB5DCC39C3B2865FBE8F402F3CEED12E" colspan="1" pageId="0" pageNumber="87" rowspan="1">2-2(2)</td>
<td id="0DCAD4870C219A6CE1CC49BB59C6536A" colspan="1" pageId="0" pageNumber="87" rowspan="1">5-6(5)</td>
<td id="31323F13FC45C6CB86045CD79733CE6B" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="BB04E5DA2DE1ADE397407546B239B3AC" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="BA9054831FEB2B7852D5FD03A8B7CE16" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-26(22)</td>
<td id="A9F7800E2CE29551564DC4938B59C038" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-23(20)</td>
<td id="E444B5A45BBB6CFD4762951C019B34B8" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-22(20)</td>
<td id="7BE8FA13205CD9C3202A7353B1E53EB3" colspan="1" pageId="0" pageNumber="87" rowspan="1">353-539(417)</td>
<td id="AEE208B10CC959019110B3A93010B2E2" colspan="1" pageId="0" pageNumber="87" rowspan="1">341-514(381)</td>
<td id="C0507D94A88BD28ED849F1494CF1B5C5" colspan="1" pageId="0" pageNumber="87" rowspan="1">11-22(15)</td>
</tr>
<tr id="91357C651FF1A5DAE4DAD2B9A1AF5623" pageId="0" pageNumber="87">
<td id="ECA15243412253E7995A484AFA02CEA5" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="0F6CCAB257C06B2897F628FB696F6B92" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. trefauti" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="trefauti">
<emphasis id="1D9A266C49352158890B99A0D09EF463" italics="true" pageId="0" pageNumber="87">L. trefauti</emphasis>
</taxonomicName>
<emphasis id="84239B0A4A4FD03490AA6369B6B9C8DB" bold="true" pageId="0" pageNumber="87">c</emphasis>
</td>
<td id="0B15C7B6DA1C6028993990BBE2F7B964" colspan="1" pageId="0" pageNumber="87" rowspan="1">2</td>
<td id="595834C31D049AB77AAB8FA18FA77191" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="DEA715A7D12838BDD19251E9E2F89FBE" colspan="1" pageId="0" pageNumber="87" rowspan="1">1-1(1)</td>
<td id="FDAA75835EDB6E3041A99C25E6011A30" colspan="1" pageId="0" pageNumber="87" rowspan="1">5-5(5)</td>
<td id="B4292ED7F5E54564E7AA0A665BBE64B4" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="288E28101287420E8630F653892E8924" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="BBE8225A0321D70F9D0AA9C3E4B1346E" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-22(22)</td>
<td id="41C3DD7073B5D1528062D22C92EED50B" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-22(22)</td>
<td id="097B3071B8BD194BF3949A814FDF6B12" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-22(22)</td>
<td id="76EDFCE58EFA13D1C1A82A5FE2CBD54A" colspan="1" pageId="0" pageNumber="87" rowspan="1">520-543</td>
<td id="02AA8E724A3B7A774BAC8AA0843FA9EE" colspan="1" pageId="0" pageNumber="87" rowspan="1">499-531</td>
<td id="9C070F01F947BA990B8B18A8AD61150B" colspan="1" pageId="0" pageNumber="87" rowspan="1">8(8)</td>
</tr>
<tr id="91B63442D957C9AB97FCC310A02CC995" pageId="0" pageNumber="87">
<td id="57C130CC8FE5E49DCCB04383985CCDB8" colspan="1" pageId="0" pageNumber="87" rowspan="1">
<taxonomicName id="5CBBDBFC69F2808314FA5E301C3BADE8" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. wilderi" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="wilderi">
<emphasis id="BC09406305A3DF5357E76AE7FD2ECDF0" italics="true" pageId="0" pageNumber="87">L. wilderi</emphasis>
</taxonomicName>
<emphasis id="0EAEE533A1FE94ED04C42A264FDC9B27" bold="true" pageId="0" pageNumber="87">a</emphasis>
</td>
<td id="427031AFF189C69B636A186129E891A2" colspan="1" pageId="0" pageNumber="87" rowspan="1">3</td>
<td id="E72000AE6D0DFBB9EEE72D41480F9878" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="4FDA9429151A2A0DDC97631D379F8CC8" colspan="1" pageId="0" pageNumber="87" rowspan="1">1-1(1)</td>
<td id="372EBE3C4D9EA2CE3A64C06DC6CBA935" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="5CA223F3B5BEC1FE7D35C477A1F793A9" colspan="1" pageId="0" pageNumber="87" rowspan="1">4-4(4)</td>
<td id="9FD17EA998182706E1127E18A83FD956" colspan="1" pageId="0" pageNumber="87" rowspan="1">3-3(3)</td>
<td id="BB04554BAD257352B91EDF735A139854" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-24(22)</td>
<td id="596DD3E0ADB19AFFCB8AA2A98CF4D994" colspan="1" pageId="0" pageNumber="87" rowspan="1">22-22(22)</td>
<td id="E75FFD203F968AEC5C1EBCE91FA70908" colspan="1" pageId="0" pageNumber="87" rowspan="1">20-21(20)</td>
<td id="25356F0D1B2746A1960CEE87C26F0D2A" colspan="1" pageId="0" pageNumber="87" rowspan="1">385-402</td>
<td id="37AE58CFE40DA7F49C0D77D5071FBA51" colspan="1" pageId="0" pageNumber="87" rowspan="1">371-383</td>
<td id="AAFF0D461478995BA4145BFAD516C180" colspan="1" pageId="0" pageNumber="87" rowspan="1">12-19(12)</td>
</tr>
</table>
</paragraph>
<caption id="246F2FD796A46416738545D9A454F4D7" doi="10.3897/zookeys.1146.94607.figure2" httpUri="https://binary.pensoft.net/fig/806305" pageId="0" pageNumber="87" start="Figure 2" startId="F2">
<paragraph id="9FE7D30885D9FB4A2A10E4020D7EC58D" pageId="0" pageNumber="87">
<emphasis id="8C3DF2B3A8B8913353FC6C00B31160F0" bold="true" pageId="0" pageNumber="87">Figure 2.</emphasis>
Dorsal (top), lateral (center), and ventral (bottom) views of the head of
<taxonomicName id="124A20F518859A3386DEA37AD5D7F5F4" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis">
<emphasis id="768EAD13B871AFD46085DC11120B50DF" italics="true" pageId="0" pageNumber="87">Liotyphlops palauophis</emphasis>
</taxonomicName>
sp. nov., AMNH R-9550, holotype. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7837B3256C923FBECE81FA8AC01B362E" pageId="0" pageNumber="87" type="description">
<paragraph id="6013E69DED43147DD709C113C4775712" pageId="0" pageNumber="87">Description of skull.</paragraph>
<paragraph id="8E8545808A64301CFE866A4D2D07B6BD" pageId="0" pageNumber="87">
High-resolution x-ray computed tomography of skull bones in Figs
<figureCitation id="F0D3B8A7146B04C4654A21507F714EED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Three-dimensional reconstruction of the skull of Liotyphlops palauophis sp. nov., AMNH R- 9550, holotype, based on HRXCT data. A lateral view B dorsal view C ventral view with lower jaw partially digitally removed D anterior view E posterior view. Scale bar: 1 mm. Anatomical abbreviations: a: angular; bo: basioccipital; cb: compound bone; co: coronoid; d: dentary; ec: ectopterygoid; en: external naris; f: frontal; fo: foramen; fv: fenestra vomeronasalis; m: maxilla; mf: mental foramen; n: nasal; oc: occipital condyle; ooc: otico-occipital (fused prootic + opisthotic + exoccipital); p: parietal; pa: palatine; pbs: parabasisphenoid; pe: postorbital element; pf: prefrontal; pg: pterygoid; pm: premaxilla; q: quadrate; rp: retroarticular process; sf: surangular foramen; sm: septomaxilla; st: supratemporal; so: supraoccipital; tf: trigeminal foramen; v: vomer; vf: vomerine foramen." figureDoi="10.3897/zookeys.1146.94607.figure3" httpUri="https://binary.pensoft.net/fig/806306" pageId="0" pageNumber="87">3</figureCitation>
-
<figureCitation id="12EFA34C5976C01AD652AF423F487F6A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Three-dimensional reconstruction of the lower jaw of Liotyphlops palauophis sp. nov., AMNH R- 9550, holotype, based on HRXCT data. A lateral view B medial view C dorsal view D ventral view E anterior view F posterior view. Scale bar: 1 mm. Anatomical abbreviations: a: angular; arf: articular fossa; cb: compound bone; cbp: compound bone prearticular component; cbs: compound bone surangular component; co: coronoid; d: dentary; Mc: Meckel's canal; mf: mental foramen; rp: retroarticular process; sf: surangular foramen." figureDoi="10.3897/zookeys.1146.94607.figure5" httpUri="https://binary.pensoft.net/fig/806308" pageId="0" pageNumber="87">5</figureCitation>
. Main body of premaxilla on ventral surface of snout. Maxilla-premaxilla contact widely separated. Lateral maxillary foramina absent. Maxilla alveolar row oriented transversely. Nasal fused. Nasal-frontal boundary convex posteriorly in a shallow W-shaped suture. Prefrontal separated from nasal. Prefrontal moveably articulated to frontal. Postorbital element present. Posterior orbital margin incomplete. Frontals gradually tapering anteriorly. Frontal paired. Frontal-parietal contact (dorsal aspect) mostly straight and transverse, median notch in frontals slight at most. Parietal paired. Posterior border of parietal without median projection over supraoccipital. Supratemporal present. Posteromedial flange of septomaxilla short, not contacting frontal. Septomaxilla with lateral flange contributing to posterior border of external naris. Fenestra for duct of
<normalizedToken id="83EBD9D6920D27BD8A24464ED84EF6E3" originalValue="Jacobson’s">Jacobson's</normalizedToken>
organ posteroventrally positioned. Palatine not in contact with vomer, maxilla, or pterygoid. Central foramen present in parabasisphenoid. Ectopterygoid present. Supraoccipital present and single not participating in internal sidewall of neurocranium. External surface (dorsoposterior) of supraoccipital without transverse ridge. Supraoccipital-prootic contact narrow, less than half supraoccipital-parietal contact. Splenial not present as discrete element. Coronoid and angular separated by prearticular portion of compound bone. Retroarticular process long, longer than articular facet. Teeth present in maxilla, but lacking in dentary, premaxilla, palatine, and pterygoid.
</paragraph>
<caption id="60E372ED9069CF83CF428880BCB1FA69" doi="10.3897/zookeys.1146.94607.figure3" httpUri="https://binary.pensoft.net/fig/806306" pageId="0" pageNumber="87" start="Figure 3" startId="F3">
<paragraph id="FFDFCC1F4CD4C2461B7F882F191B49FE" pageId="0" pageNumber="87">
<emphasis id="FA43C534DE92D9F496B73EE2DE41EAC3" bold="true" pageId="0" pageNumber="87">Figure 3.</emphasis>
Three-dimensional reconstruction of the skull of
<taxonomicName id="F60ACAFE6185B97CD25DC86ECB0F453B" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis">
<emphasis id="DEE43105378D70DF4D6B7FB53EA8F7F8" italics="true" pageId="0" pageNumber="87">Liotyphlops palauophis</emphasis>
</taxonomicName>
sp. nov., AMNH R-9550, holotype, based on HRXCT data.
<emphasis id="10DE50318A547B4531E6A15538B0C393" bold="true" pageId="0" pageNumber="87">A</emphasis>
lateral view
<emphasis id="0648DC19E6A2ED6960B329457DBC22E2" bold="true" pageId="0" pageNumber="87">B</emphasis>
dorsal view
<emphasis id="0724999B3FEFBB971914CA024B53E589" bold="true" pageId="0" pageNumber="87">C</emphasis>
ventral view with lower jaw partially digitally removed
<emphasis id="07DC1C9890768374DB6583D3C3C79F68" bold="true" pageId="0" pageNumber="87">D</emphasis>
anterior view
<emphasis id="3AA42AD35C82FF0FCF0A50E9808BB633" bold="true" pageId="0" pageNumber="87">E</emphasis>
posterior view. Scale bar: 1 mm. Anatomical abbreviations:
<emphasis id="5E4B2AA778E8DB3B8545529A263CCBCE" bold="true" pageId="0" pageNumber="87">a</emphasis>
: angular;
<emphasis id="A2EB6391DC2D3C5CEA176FF00632D027" bold="true" pageId="0" pageNumber="87">bo</emphasis>
: basioccipital;
<emphasis id="0E54B1D93C085682C91EA064B85761F3" bold="true" pageId="0" pageNumber="87">cb</emphasis>
: compound bone;
<emphasis id="E45C7BCAFD3C8CD44ACADEE06068E8F5" bold="true" pageId="0" pageNumber="87">co</emphasis>
: coronoid;
<emphasis id="39674C2D07F54391544C6A5BE1EDBFC1" bold="true" pageId="0" pageNumber="87">d</emphasis>
: dentary;
<emphasis id="7FDE73136719EF3CC31F8AB39E6FD61F" bold="true" pageId="0" pageNumber="87">ec</emphasis>
: ectopterygoid;
<emphasis id="93627312353FF2B51318899B79D2707C" bold="true" pageId="0" pageNumber="87">en</emphasis>
: external naris;
<emphasis id="7EEACE873E488D481C4AF0B0C8D10C0C" bold="true" pageId="0" pageNumber="87">f</emphasis>
: frontal;
<emphasis id="FD2FFD7FE8BC70FF3C3A671C7DFEFA94" bold="true" pageId="0" pageNumber="87">fo</emphasis>
: foramen;
<emphasis id="7CF9F2536E6A98C76C29BA5861CBFBF1" bold="true" pageId="0" pageNumber="87">fv</emphasis>
: fenestra vomeronasalis;
<emphasis id="4547982627F29D237C79B6E86279F6E2" bold="true" pageId="0" pageNumber="87">m</emphasis>
: maxilla;
<emphasis id="2F5A23C9B33C1589EB5E772F10F7F5A5" bold="true" pageId="0" pageNumber="87">mf</emphasis>
: mental foramen;
<emphasis id="4F1BE4B208F78BA09379B7AE3008B65D" bold="true" pageId="0" pageNumber="87">n</emphasis>
: nasal;
<emphasis id="CC6C810E29A324FABD039154D33D7270" bold="true" pageId="0" pageNumber="87">oc</emphasis>
: occipital condyle;
<emphasis id="225A2D1DE6DA6F0BF2A1B3A4EB9FFB3C" bold="true" pageId="0" pageNumber="87">ooc</emphasis>
: otico-occipital (fused prootic + opisthotic + exoccipital);
<emphasis id="F33A7B5971B3A868590FD9EC4809EDD0" bold="true" pageId="0" pageNumber="87">p</emphasis>
: parietal;
<emphasis id="A328D81CC39579992F31BFBCFF9A8A9A" bold="true" pageId="0" pageNumber="87">pa</emphasis>
: palatine;
<emphasis id="F47DA0C7AAC34F0A8A504A65F4C1B129" bold="true" pageId="0" pageNumber="87">pbs</emphasis>
: parabasisphenoid;
<emphasis id="F4280B9A407C6CDC586FF6309FB79B7C" bold="true" pageId="0" pageNumber="87">pe</emphasis>
: postorbital element;
<emphasis id="9DFFC2C1D27E64B03119D78997833772" bold="true" pageId="0" pageNumber="87">pf</emphasis>
: prefrontal;
<emphasis id="C37A2A4BD633CCA98307A96160D0685C" bold="true" pageId="0" pageNumber="87">pg</emphasis>
: pterygoid;
<emphasis id="E969C5FE4FF9F62E3B877369E2270DCE" bold="true" pageId="0" pageNumber="87">pm</emphasis>
: premaxilla;
<emphasis id="039519543526EA441A804C097D66681B" bold="true" pageId="0" pageNumber="87">q</emphasis>
: quadrate;
<emphasis id="2F69A909B3D3207684719B1836825624" bold="true" pageId="0" pageNumber="87">rp</emphasis>
: retroarticular process;
<emphasis id="D1027B00F7893CADC9E00BC47FE1F36F" bold="true" pageId="0" pageNumber="87">sf</emphasis>
: surangular foramen;
<emphasis id="C04A8E6FF6529F6989FA0D3A3CEC509F" bold="true" pageId="0" pageNumber="87">sm</emphasis>
: septomaxilla;
<emphasis id="86034B940DD8BB340C705AE74B9A5BC3" bold="true" pageId="0" pageNumber="87">st</emphasis>
: supratemporal;
<emphasis id="02E2D76EBC86F15C2CC6866F6E139128" bold="true" pageId="0" pageNumber="87">so</emphasis>
: supraoccipital;
<emphasis id="3D198D2A8EDD2F2FB7A3E6B3BDB1B887" bold="true" pageId="0" pageNumber="87">tf</emphasis>
: trigeminal foramen;
<emphasis id="7AF39C476D71A451E8D6A085E905C5D5" bold="true" pageId="0" pageNumber="87">v</emphasis>
: vomer;
<emphasis id="436EEFACD43AB1F00F21B6D55E97EFFB" bold="true" pageId="0" pageNumber="87">vf</emphasis>
: vomerine foramen.
</paragraph>
</caption>
<caption id="3C26A1DC0C75CBEC33FA733DDD68C067" doi="10.3897/zookeys.1146.94607.figure4" httpUri="https://binary.pensoft.net/fig/806307" pageId="0" pageNumber="87" start="Figure 4" startId="F4">
<paragraph id="57E10F1D2033A1E0390231882E893349" pageId="0" pageNumber="87">
<emphasis id="6CD79663AC3B8EDD1621C7A59AB54C0B" bold="true" pageId="0" pageNumber="87">Figure 4.</emphasis>
Three-dimensional reconstruction of the skull of
<taxonomicName id="B34CABEE647793A6C2CA1ED9B5CD07CD" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis">
<emphasis id="F591B872AB8DEA13FFF80D4A4E08CECE" italics="true" pageId="0" pageNumber="87">Liotyphlops palauophis</emphasis>
</taxonomicName>
sp. nov., AMNH R-9550, holotype, based on HRXCT data.
<emphasis id="E00910BA8A9D609E7120E5461C17A175" bold="true" pageId="0" pageNumber="87">A</emphasis>
transversal view
<emphasis id="99DB0C1DA71C8EBCAD06A4ACC6AE945B" bold="true" pageId="0" pageNumber="87">B</emphasis>
frontal view
<emphasis id="92D412CC2239C45619705A34A2D025D8" bold="true" pageId="0" pageNumber="87">C</emphasis>
sagittal view. Scale bar: 1 mm. Anatomical abbreviations:
<emphasis id="687F0692A3EF9058CF80015CFB562354" bold="true" pageId="0" pageNumber="87">a</emphasis>
: angular;
<emphasis id="217432CED6B55900AAD174674D7B7C04" bold="true" pageId="0" pageNumber="87">an</emphasis>
: acoustic nerve foramen;
<emphasis id="23795D05E54FFDEECB5D8B7BF1388FE5" bold="true" pageId="0" pageNumber="87">bo</emphasis>
: basioccipital;
<emphasis id="C83CE0C0B1F4A961638A4066023243BA" bold="true" pageId="0" pageNumber="87">cb</emphasis>
: compound bone;
<emphasis id="9B30E9C8773F5D2BA505DFBD8EAF6545" bold="true" pageId="0" pageNumber="87">cbp</emphasis>
: compound bone prearticular component;
<emphasis id="3D9584EE5F8E0F1D7588090A72886613" bold="true" pageId="0" pageNumber="87">cbs</emphasis>
: compound bone surangular component;
<emphasis id="E7A64CD9C3D11806F2BAE1496C011693" bold="true" pageId="0" pageNumber="87">ci</emphasis>
: conchal invagination;
<emphasis id="806AED9B2EF94F7BB2B163F4E900D65B" bold="true" pageId="0" pageNumber="87">co</emphasis>
: coronoid;
<emphasis id="4FBC96FCF76C6E011CDD303DC317A1C9" bold="true" pageId="0" pageNumber="87">cv</emphasis>
: cavum vestibuli;
<emphasis id="6077BAC13C41365F2D54895DA1468064" bold="true" pageId="0" pageNumber="87">d</emphasis>
: dentary;
<emphasis id="C2D8215E5828761D72CE81FFFE7D1AAC" bold="true" pageId="0" pageNumber="87">dp</emphasis>
: descensus parietalis;
<emphasis id="018874DC683B247E1DC47431E431A945" bold="true" pageId="0" pageNumber="87">ec</emphasis>
: ectopterygoid;
<emphasis id="45EBDED35921C684857DF868409F02A0" bold="true" pageId="0" pageNumber="87">en</emphasis>
: external naris;
<emphasis id="DA8D3BD6093DC5BA9E11878160814793" bold="true" pageId="0" pageNumber="87">f</emphasis>
: frontal;
<emphasis id="F4DFB3A672D396E27891D4C17C509282" bold="true" pageId="0" pageNumber="87">
<normalizedToken id="2616D825397B1ED01085725E8A0D0E59" originalValue="fl">fl</normalizedToken>
</emphasis>
: frontal laterally descending
<normalizedToken id="C7C37B9CEED03E8E7CEF589391DD5EE6" originalValue="flange">flange</normalizedToken>
;
<emphasis id="F19B3B017D5D64BA0CECB64D7968180D" bold="true" pageId="0" pageNumber="87">fo</emphasis>
: foramen;
<emphasis id="796DA0FB097B72793B412393F7925B67" bold="true" pageId="0" pageNumber="87">fpb</emphasis>
: facial nerve palatine branch foramen;
<emphasis id="55D34F1E073490B728AA6A9C83D773BC" bold="true" pageId="0" pageNumber="87">fs</emphasis>
: frontal subolfactory process;
<emphasis id="1848D0C8EA7ACEE3D9C43E4C51A2E35A" bold="true" pageId="0" pageNumber="87">fv</emphasis>
: fenestra vomeronasalis;
<emphasis id="18228B879E200A773369A8EE2B872C82" bold="true" pageId="0" pageNumber="87">m</emphasis>
: maxilla;
<emphasis id="B5C0CF6329BA1C8B586F3FB765472305" bold="true" pageId="0" pageNumber="87">Mc</emphasis>
:
<normalizedToken id="01EF5FC11AAEA83DCA822F75C75D5D02" originalValue="Meckel’s">Meckel's</normalizedToken>
canal;
<emphasis id="43F38B609B102D81B1602CFC78D1F526" bold="true" pageId="0" pageNumber="87">mf</emphasis>
: mental foramen;
<emphasis id="72BA7F9D681F25DAC973C84DA06C0957" bold="true" pageId="0" pageNumber="87">n</emphasis>
: nasal;
<emphasis id="98B9BBE16AD34A83924C7CCD304119CF" bold="true" pageId="0" pageNumber="87">nl</emphasis>
: nasal lateral
<normalizedToken id="A35110B254A4C421839A7BD327685457" originalValue="flange">flange</normalizedToken>
;
<emphasis id="C310967ACBF2D2168E3CAF4EA9F16A14" bold="true" pageId="0" pageNumber="87">ns</emphasis>
: medial nasal septum;
<emphasis id="05660CB8AEE6A7EBD7072F2684F18DEF" bold="true" pageId="0" pageNumber="87">oc</emphasis>
: occipital condyle;
<emphasis id="1E5CAB773456A0F6F5797DBE6ECFFC63" bold="true" pageId="0" pageNumber="87">on</emphasis>
: optic nerve foramen;
<emphasis id="26BEB37475CA1DBA043DE24856AA94AB" bold="true" pageId="0" pageNumber="87">ooc</emphasis>
: otico-occipital (fused prootic + opisthotic + exoccipital);
<emphasis id="AC1A2A6670BF73ECA6C8D029EF724E49" bold="true" pageId="0" pageNumber="87">p</emphasis>
: parietal;
<emphasis id="4996EF1273E3817EF34B9FFD74C388B1" bold="true" pageId="0" pageNumber="87">pa</emphasis>
: palatine;
<emphasis id="8C38B3DD856F7A228CFFFC406664D21B" bold="true" pageId="0" pageNumber="87">pbc</emphasis>
: parabasal (Vidian) canal;
<emphasis id="B1EFF88F65CC5792D23CAC059ECB8882" bold="true" pageId="0" pageNumber="87">pbs</emphasis>
: parabasisphenoid;
<emphasis id="9D0AB93D45F454263025974BA12E6AE2" bold="true" pageId="0" pageNumber="87">pe</emphasis>
: postorbital element;
<emphasis id="27FE8D9F05437E8063F0F9CE3BF10C33" bold="true" pageId="0" pageNumber="87">pf</emphasis>
: prefrontal;
<emphasis id="DF996145CEADC454B001BA6FA48C1CB7" bold="true" pageId="0" pageNumber="87">pg</emphasis>
: pterygoid;
<emphasis id="0C0DC2663986C9A3D162C6860234A99D" bold="true" pageId="0" pageNumber="87">pm</emphasis>
: premaxilla;
<emphasis id="DB630241A65080E8D05821C5F4DAC60C" bold="true" pageId="0" pageNumber="87">pvs</emphasis>
: posterior vertical semicircular canal;
<emphasis id="724E07536C42BFFD9D15147706568C0B" bold="true" pageId="0" pageNumber="87">q</emphasis>
: quadrate;
<emphasis id="A271BECB9077FCD2C7CB83451BC806F4" bold="true" pageId="0" pageNumber="87">rp</emphasis>
: retroarticular process;
<emphasis id="0781900A2BFDF43489A404C6ED8F644E" bold="true" pageId="0" pageNumber="87">rstm</emphasis>
: recessus scalae tympani medial aperture;
<emphasis id="56DD121A5A2E3C2FC259BA8D4453BB50" bold="true" pageId="0" pageNumber="87">s</emphasis>
: stapes;
<emphasis id="BF8E885F850BAAD684764C783A0CD651" bold="true" pageId="0" pageNumber="87">sf</emphasis>
: surangular foramen;
<emphasis id="52B758F0298909814883A66E0603ACFF" bold="true" pageId="0" pageNumber="87">sm</emphasis>
: septomaxilla;
<emphasis id="8EF8C468BA02C3C1E59641D3EB2CA919" bold="true" pageId="0" pageNumber="87">sml</emphasis>
: septomaxilla lateral
<normalizedToken id="7AC7E4C0B963AE56414E3CD37B858083" originalValue="flange">flange</normalizedToken>
;
<emphasis id="5349A40F19392098FC17D55BB86A62F9" bold="true" pageId="0" pageNumber="87">st</emphasis>
: supratemporal;
<emphasis id="B03DD8A744B01BC795AE3F11EA833E40" bold="true" pageId="0" pageNumber="87">so</emphasis>
: supraoccipital;
<emphasis id="48CC614F2929242C5BA25DA3E7E93260" bold="true" pageId="0" pageNumber="87">tf</emphasis>
: trigeminal foramen;
<emphasis id="360DC73E48A0DB86EB2B5765FE9D15DD" bold="true" pageId="0" pageNumber="87">trc</emphasis>
: trigeminofacialis chamber;
<emphasis id="AB72A83FCD1C732D7D7A7E82C4BBF684" bold="true" pageId="0" pageNumber="87">v</emphasis>
: vomer;
<emphasis id="EB76AF41E01468714F7A9B593C8F63E6" bold="true" pageId="0" pageNumber="87">vc</emphasis>
: vomeronasal cupola;
<emphasis id="76DE375CA25332CCF37DE706A53A14E1" bold="true" pageId="0" pageNumber="87">vf</emphasis>
: vomerine foramen;
<emphasis id="C2E42B17EC91ED0EFD77697B313FFC02" bold="true" pageId="0" pageNumber="87">vn</emphasis>
: vomeronasal nerve passage.
</paragraph>
</caption>
<caption id="55104AEE53E4AD6769F7885E3B7B18CD" doi="10.3897/zookeys.1146.94607.figure5" httpUri="https://binary.pensoft.net/fig/806308" pageId="0" pageNumber="87" start="Figure 5" startId="F5">
<paragraph id="2CFD6EDF1534222C4864BFC8F47BB206" pageId="0" pageNumber="87">
<emphasis id="8145D619FFF30CD7C7C5D47926193C7A" bold="true" pageId="0" pageNumber="87">Figure 5.</emphasis>
Three-dimensional reconstruction of the lower jaw of
<taxonomicName id="8D50F86E8B44F0D481B37A428850B294" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis">
<emphasis id="30CE5845D2B29CB4277506F5CFCFC39C" italics="true" pageId="0" pageNumber="87">Liotyphlops palauophis</emphasis>
</taxonomicName>
sp. nov., AMNH R-9550, holotype, based on HRXCT data.
<emphasis id="ABEE3CDBC96D7335EFC39F2BD9012B9D" bold="true" pageId="0" pageNumber="87">A</emphasis>
lateral view
<emphasis id="7770EAF54925502C3B44352D651A51B4" bold="true" pageId="0" pageNumber="87">B</emphasis>
medial view
<emphasis id="463EE8A5750A646484FD802802E894E4" bold="true" pageId="0" pageNumber="87">C</emphasis>
dorsal view
<emphasis id="38A738DD4BFE5322292D14FDFFE6983B" bold="true" pageId="0" pageNumber="87">D</emphasis>
ventral view
<emphasis id="05D781A76B355C373567CC8B1D07E6FA" bold="true" pageId="0" pageNumber="87">E</emphasis>
anterior view
<emphasis id="32E4AC3D2F7964CF66F361CBD3CD7DBF" bold="true" pageId="0" pageNumber="87">F</emphasis>
posterior view. Scale bar: 1 mm. Anatomical abbreviations:
<emphasis id="CA6130CA181EE69AFB04EDAD0DF99689" bold="true" pageId="0" pageNumber="87">a</emphasis>
: angular;
<emphasis id="62A85E454B9B8B4B9F08DEC6FCF19470" bold="true" pageId="0" pageNumber="87">arf</emphasis>
: articular fossa;
<emphasis id="2330C42A69F131D19592F4308E3016D6" bold="true" pageId="0" pageNumber="87">cb</emphasis>
: compound bone;
<emphasis id="6397210B6D5DB9B672A0943992408F50" bold="true" pageId="0" pageNumber="87">cbp</emphasis>
: compound bone prearticular component;
<emphasis id="7148DE2836003888A83C90A50C2C30AA" bold="true" pageId="0" pageNumber="87">cbs</emphasis>
: compound bone surangular component;
<emphasis id="02D12D9381722C457AC80BC14764B0B1" bold="true" pageId="0" pageNumber="87">co</emphasis>
: coronoid;
<emphasis id="E25555977B71D519DA61BA7F426E21AC" bold="true" pageId="0" pageNumber="87">d</emphasis>
: dentary;
<emphasis id="62BA915AC91B20010D2E3149B4FE1BAE" bold="true" pageId="0" pageNumber="87">Mc</emphasis>
:
<normalizedToken id="16137DCACFBA866DE5178C628981C3AD" originalValue="Meckel’s">Meckel's</normalizedToken>
canal;
<emphasis id="457733F6EC6E73070C9142617833892F" bold="true" pageId="0" pageNumber="87">mf</emphasis>
: mental foramen;
<emphasis id="D11A688321F63B3EF584B5E06023AD91" bold="true" pageId="0" pageNumber="87">rp</emphasis>
: retroarticular process;
<emphasis id="FAFBFEC766DE53ACF29FCCBCA5F26779" bold="true" pageId="0" pageNumber="87">sf</emphasis>
: surangular foramen.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B0A8319229DA3BC194891160DA1A4433" pageId="0" pageNumber="87" type="coloration in alcohol">
<paragraph id="7FFA38F0459A9B1AA3DEF1C0F6A1B477" pageId="0" pageNumber="87">Coloration in alcohol.</paragraph>
<paragraph id="16F6C7BA47B1BF9D15BA7FF09163964C" pageId="0" pageNumber="87">Dorsal and ventral body pale cream with brown pigmentation points along dorsal region of body.</paragraph>
</subSubSection>
<subSubSection id="56D14937B32F9A35F2847FA85EB752E8" pageId="0" pageNumber="87" type="distribution">
<paragraph id="FFC3D0C5E2553DC3ACA3984D2CFF02B8" pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph id="4D8932D1671B0AFA9B2D1C4E41FD6500" pageId="0" pageNumber="87">
Known only from the type locality in the neighborhood of Bogota, Colombia (Fig.
<figureCitation id="36B1AB874E7783470AD257F78AFBFBB2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Location of the holotype of Liotyphlops palauophis sp. nov. (black star), lectotype of Liotyphlops anops (white dot), and specimens of L. anops examined in this study (green dot).? = lack of detailed information about the type locality of L. palauophis sp. nov. and L. anops. This locality is based on information provided by Cope (1899)." figureDoi="10.3897/zookeys.1146.94607.figure6" httpUri="https://binary.pensoft.net/fig/806309" pageId="0" pageNumber="87">6</figureCitation>
), according to the information provided by
<bibRefCitation id="631FCD02769833FD9D7FC31D61164777" DOI="https://doi.org/10.5962/bhl.title.54674" author="Cope, ED" journalOrPublisher="The Philadelphia Museums Scientific Bulletin" pageId="0" pageNumber="87" pagination="3 - 26" refId="B7" refString="Cope, ED, 1899. Contributions to the herpetology of New Granada and Argentina with descriptions of new forms. The Philadelphia Museums Scientific Bulletin 1: 3 - 26, DOI: https://doi.org/10.5962/bhl.title.54674" title="Contributions to the herpetology of New Granada and Argentina with descriptions of new forms." url="https://doi.org/10.5962/bhl.title.54674" volume="1" year="1899">Cope (1899)</bibRefCitation>
.
</paragraph>
<caption id="CCA966D121E028204EFBDC5587D7B815" doi="10.3897/zookeys.1146.94607.figure6" httpUri="https://binary.pensoft.net/fig/806309" pageId="0" pageNumber="87" start="Figure 6" startId="F6">
<paragraph id="585C2BEC3C864A597F78C223BB43F2FF" pageId="0" pageNumber="87">
<emphasis id="8BD8B196AF99BFEE719F6E8D8FB5EC12" bold="true" pageId="0" pageNumber="87">Figure 6.</emphasis>
Location of the holotype of
<taxonomicName id="D5916617C65F0D252612A5149A7A191E" authorityName="Marra Santos" authorityYear="2023" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops palauophis" order="Squamata" pageId="0" pageNumber="87" rank="species" species="palauophis">
<emphasis id="DB6C03D7C6FE9772F8EEEAA1B55BECA8" italics="true" pageId="0" pageNumber="87">Liotyphlops palauophis</emphasis>
</taxonomicName>
sp. nov. (black star), lectotype of
<taxonomicName id="7B2F25B3DF77F9E3BA88E84A5F0724E4" baseAuthorityName="Cope" baseAuthorityYear="1899" family="Anomalepididae" genus="Liotyphlops" kingdom="Animalia" lsidName="Liotyphlops anops" order="Squamata" pageId="0" pageNumber="87" rank="species" species="anops">
<emphasis id="8D2020F84B697FAE67E7CB066E525623" italics="true" pageId="0" pageNumber="87">Liotyphlops anops</emphasis>
</taxonomicName>
(white dot), and specimens of
<taxonomicName id="A497D6BB07BEBA319304AFC363ECB24B" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. anops" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="anops">
<emphasis id="94246E316E72D308CFFAAD0250DC88E9" italics="true" pageId="0" pageNumber="87">L. anops</emphasis>
</taxonomicName>
examined in this study (green dot).? = lack of detailed information about the type locality of
<taxonomicName id="8C9E91F76F2F485A25BF7D2763A206DB" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. palauophis" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="palauophis">
<emphasis id="9083ADD4B1BD969646EC94CAAC83DD75" italics="true" pageId="0" pageNumber="87">L. palauophis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="6E6D293346823724160985A8E1E2CC17" class="Squamata" family="Anomalepididae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="L. anops" order="Squamata" pageId="0" pageNumber="87" phylum="Chordata" rank="species" species="anops">
<emphasis id="550F01B1A2626EBDC9835EA9E507BEEC" italics="true" pageId="0" pageNumber="87">L. anops</emphasis>
</taxonomicName>
. This locality is based on information provided by
<bibRefCitation id="8F01D70FEE662CBA7B3F309E2CF24746" DOI="https://doi.org/10.5962/bhl.title.54674" author="Cope, ED" journalOrPublisher="The Philadelphia Museums Scientific Bulletin" pageId="0" pageNumber="87" pagination="3 - 26" refId="B7" refString="Cope, ED, 1899. Contributions to the herpetology of New Granada and Argentina with descriptions of new forms. The Philadelphia Museums Scientific Bulletin 1: 3 - 26, DOI: https://doi.org/10.5962/bhl.title.54674" title="Contributions to the herpetology of New Granada and Argentina with descriptions of new forms." url="https://doi.org/10.5962/bhl.title.54674" volume="1" year="1899">Cope (1899)</bibRefCitation>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="BF68D87C50DDA693EA46FCD5E68046E8" pageId="0" pageNumber="87" type="etymology">
<paragraph id="0F58431CDE56F1A299644FD8D84DC5CB" pageId="0" pageNumber="87">Etymology.</paragraph>
<paragraph id="1D6201DCFB8807BE45329F19D112FA15" pageId="0" pageNumber="87">
The species name is in honor of Alfredo
<taxonomicName id="E69E57AB1B2AE929EF22C0F3F12D3875" genus="Palau" lsidName="Palau" pageId="0" pageNumber="87" rank="genus">Palau</taxonomicName>
<normalizedToken id="F4149FE975338CCE9BD37F26A1424CA9" originalValue="Peña">Pena</normalizedToken>
(June 10, 1969-August 8, 2020), a Brazilian herpetologist and my friend, who was killed by the COVID-19 virus. A combination of his name
<taxonomicName id="621172E5358B6DC786E121B98540E905" genus="Palau" lsidName="Palau" pageId="0" pageNumber="87" rank="genus">
<emphasis id="6CE8DDE0FF2CE028744FEDE9F89D0304" italics="true" pageId="0" pageNumber="87">Palau</emphasis>
</taxonomicName>
and the Greek
<emphasis id="B1CF2CFA189BC176AAAC7178CE7F315D" italics="true" pageId="0" pageNumber="87">ophis</emphasis>
, meaning snake.
</paragraph>
</subSubSection>
</treatment>
</document>
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