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<document id="8D501681B95FCDB033E08000FE12015D" ID-DOI="10.11646/phytotaxa.671.2.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="14521284" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734559178251" checkinUser="felipe" docAuthor="Swanepoel, Wessel &amp; Van Wyk, Abraham E." docDate="2024" docId="A74387E1FFD60426FF1D4F3347CB9F0C" docLanguage="en" docName="phytotaxa.671.2.128-138.pdf" docOrigin="Phytotaxa 671 (2)" docSource="https://doi.org/10.11646/phytotaxa.671.2.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Petalidium namibense Swanepoel &amp; A. E. van Wyk 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="134" masterDocId="5B7AFF99FFD70420FF954B4746629864" masterDocTitle="Petalidium namibense (Acanthaceae), a new species from Namibia" masterLastPageNumber="138" masterPageNumber="128" pageNumber="129" updateTime="1734563815495" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CE828DC72ACAE00F87466BF024371154">Petalidium namibense (Acanthaceae), a new species from Namibia</mods:title>
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<mods:namePart id="569F36F16F0AC56E214B43E68814CB83">Swanepoel, Wessel</mods:namePart>
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<mods:namePart id="5396C603031DCA63E7642E20B6F34CAF">Van Wyk, Abraham E.</mods:namePart>
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<mods:title id="5D23BFD82CB28F9CFF5F2271B72C0529">Phytotaxa</mods:title>
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<mods:date id="EC4C8BE8F2A0773AD7E455FBF44E5D6A">2024</mods:date>
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<mods:number id="D11BD85BF3B22812F67E43CFF105FBEC">2024-11-12</mods:number>
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<mods:number id="508FF5E5942B28E05F9E2727E794D0A5">671</mods:number>
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<mods:classification id="9B601F0B7C38AA8658B6307EB153EB3C">journal article</mods:classification>
<mods:identifier id="3CA484AB2FDE681A7EE4EBA78766467B" type="DOI">10.11646/phytotaxa.671.2.2</mods:identifier>
<mods:identifier id="506F7D23AF577A12A1A6C4273E1B4976" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="3ACE935625F1CDDF3A7E29F59FD748C7" type="Zenodo-Dep">14521284</mods:identifier>
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<treatment id="A74387E1FFD60426FF1D4F3347CB9F0C" ID-DOI="http://doi.org/10.5281/zenodo.14521286" ID-Zenodo-Dep="14521286" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C" httpUri="http://treatment.plazi.org/id/A74387E1FFD60426FF1D4F3347CB9F0C" lastPageId="6" lastPageNumber="134" pageId="1" pageNumber="129">
<subSubSection id="67F0657CFFD60421FF1D4F3345C09CEA" box="[136,930,1140,1167]" pageId="1" pageNumber="129" type="nomenclature">
<paragraph id="2F5536F7FFD60421FF1D4F3345C09CEA" blockId="1.[136,930,1140,1167]" box="[136,930,1140,1167]" pageId="1" pageNumber="129">
<heading id="741D819BFFD60421FF1D4F3345C09CEA" box="[136,930,1140,1167]" fontSize="11" level="2" pageId="1" pageNumber="129" reason="3">
<taxonomicName id="E8EA4D74FFD60421FF1D4F3344AA9CEA" authority="Swanepoel &amp; A.E.van Wyk" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[136,712,1140,1167]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="129" phylum="Tracheophyta" rank="species" species="namibense" status="sp. nov.">
<emphasis id="1D9EEAE5FFD60421FF1D4F3347EB9CEB" bold="true" box="[136,393,1140,1167]" italics="true" pageId="1" pageNumber="129">Petalidium namibense</emphasis>
Swanepoel &amp; A.E.van Wyk
</taxonomicName>
,
<emphasis id="1D9EEAE5FFD60421FD464F32454A9CEA" box="[723,808,1141,1166]" italics="true" pageId="1" pageNumber="129">
<taxonomicNameLabel id="06AD579EFFD60421FD464F32454A9CEA" box="[723,808,1141,1166]" pageId="1" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B7D12A72FFD60421FCA34F3345FB9CEA" box="[822,921,1140,1166]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[136,229,2031,2053]" captionStartId-1="4.[136,229,1952,1974]" captionStartId-2="5.[136,229,1952,1974]" captionTargetBox-0="[222,1366,425,2007]" captionTargetBox-1="[165,1422,190,1927]" captionTargetBox-2="[165,1422,190,1928]" captionTargetId-0="figure-130@3.[222,1366,425,2007]" captionTargetId-1="figure-18@4.[165,1422,190,1928]" captionTargetId-2="figure-17@5.[165,1422,190,1928]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Petalidium namibense, habitat and habit. A. Several plants (dark grey dwarf shrubs) growing in the bed of an ephemeral drainage line. B. Ancient windswept plant sprouting from persistent woody stems with corky bark. Photographs by W. Swanepoel." captionText-1="FIGURE 2. Petalidium namibense, habitat and habit. A. Mature plant (ca. 0.5 m high) with greyish appearance, growing among rocks. B. Multiple stems (thickest ca. 80 mm in diam.) from base of a relatively old plant, each covered with thick corky bark. Photographs by W. Swanepoel." captionText-2="FIGURE 3. Petalidium namibense, morphology of leaves and flowers. A. Flower, reduced shoots, and congested leaves; leaves with dense greyish indumentum. B. Flower in the process of fading as indicated by its pale colours. C. Newly opened flower with bracteoles in oblique lateral view; note brownish abaxial colour of posterior corolla lobes. D. Flower in front view. E. Flower with bracteoles in lateral view. F. Flower in dorsal view. Photographs: W. Swanepoel." figureDoi-0="http://doi.org/10.5281/zenodo.14521288" figureDoi-1="http://doi.org/10.5281/zenodo.14521290" figureDoi-2="http://doi.org/10.5281/zenodo.14521292" httpUri-0="https://zenodo.org/record/14521288/files/figure.png" httpUri-1="https://zenodo.org/record/14521290/files/figure.png" httpUri-2="https://zenodo.org/record/14521292/files/figure.png" pageId="1" pageNumber="129">Figs 1–3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67F0657CFFD60421FF1D4FF844E49DEE" pageId="1" pageNumber="129" type="diagnosis">
<paragraph id="2F5536F7FFD60421FF1D4FF844E49DEE" blockId="1.[136,1452,1215,1490]" pageId="1" pageNumber="129">
<emphasis id="1D9EEAE5FFD60421FF1D4FF846919CB1" bold="true" box="[136,243,1215,1237]" pageId="1" pageNumber="129">Diagnosis:</emphasis>
—A woody dwarf shrub up to
<quantity id="E8129B12FFD60421FD804F8744589CB1" box="[533,570,1216,1237]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="129" unit="m" value="1.0">1 m</quantity>
tall, morphologically most similar to
<emphasis id="1D9EEAE5FFD60421FC374F8742C09CB1" box="[930,1186,1216,1237]" italics="true" pageId="1" pageNumber="129">
<taxonomicName id="E8EA4D74FFD60421FC374F8742FC9CB1" authorityName="Swanepoel &amp; Manzitto-Tripp" authorityYear="2022" box="[930,1182,1216,1237]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="129" phylum="Tracheophyta" rank="species" species="sesfonteinense">Petalidium sesfonteinense</taxonomicName>
,
</emphasis>
differing by having the leaf lamina ovate, elliptic, suborbicular or orbicular (
<emphasis id="1D9EEAE5FFD60421FD184FA344CA9C9D" box="[653,680,1252,1273]" italics="true" pageId="1" pageNumber="129">vs.</emphasis>
ovate, elliptic or oblanceolate); bracteoles with trichomes dendritic, interspersed with glandular ones (
<emphasis id="1D9EEAE5FFD60421FE1E4E4F47FD9D79" box="[395,415,1288,1309]" italics="true" pageId="1" pageNumber="129">vs</emphasis>
. trichomes glandular with in addition appressed simple ones towards base); corolla shorter, 15.0–
<quantity id="E8129B12FFD60421FAC34E4F43C99D79" box="[1366,1451,1288,1310]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="129" unit="mm" value="17.5">17.5 mm</quantity>
(
<emphasis id="1D9EEAE5FFD60421FF504E6B46BB9D25" box="[197,217,1324,1345]" italics="true" pageId="1" pageNumber="129">vs</emphasis>
.
<quantity id="E8129B12FFD60421FF704E6B47299D25" box="[229,331,1324,1345]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="1" pageNumber="129" unit="mm" value="22.0" valueMax="24.0" valueMin="20.0">20–24 mm</quantity>
long), upper lobes rectangular, smaller, 4.9–5.2 ×
<quantity id="E8129B12FFD60421FCB94E6B45FD9D25" box="[812,927,1324,1345]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="1" pageNumber="129" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2 mm</quantity>
(
<emphasis id="1D9EEAE5FFD60421FC384E6B45A39D25" box="[941,961,1324,1345]" italics="true" pageId="1" pageNumber="129">vs</emphasis>
. obovate, 5.8–8.2 ×
<quantity id="E8129B12FFD60421FB104E6B42999D25" box="[1157,1275,1324,1346]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="1" pageNumber="129" unit="mm" value="3.9" valueMax="4.3" valueMin="3.5">3.5–4.3 mm</quantity>
), upper and lateral lobes differently coloured than anterior lobe with nectar guides absent or inconspicuous (
<emphasis id="1D9EEAE5FFD60421FBA74E1742249D01" box="[1074,1094,1360,1381]" italics="true" pageId="1" pageNumber="129">vs</emphasis>
. all lobes similarly coloured, nectar guides on upper and lateral lobes conspicuous).
</paragraph>
</subSubSection>
<subSubSection id="67F0657CFFD60422FF1D4EDF42289C7C" lastPageId="2" lastPageNumber="130" pageId="1" pageNumber="129" type="materials_examined">
<paragraph id="2F5536F7FFD60421FF1D4EDF43759DB5" blockId="1.[136,1452,1215,1490]" pageId="1" pageNumber="129">
<emphasis id="1D9EEAE5FFD60421FF1D4EDF46A09DC9" bold="true" box="[136,194,1432,1453]" pageId="1" pageNumber="129">Type:</emphasis>
—
<materialsCitation id="9F823CAAFFD60421FF4E4EDF43719DB5" collectingDate="2022-05-23" collectionCode="WIND, PRE, PRU" country="Namibia" county="Namib Desert" elevation="377" location="Skeleton Coast Park" municipality="Khumib River" pageId="1" pageNumber="129" specimenCount="2" stateProvince="Kunene" typeStatus="holotype">
<collectingCountry id="57FD7667FFD60421FF4E4EDF47259DC9" box="[219,327,1432,1453]" name="Namibia" pageId="1" pageNumber="129">NAMIBIA</collectingCountry>
.
<collectingRegion id="ED2EF815FFD60421FEC44EDF478F9DCA" box="[337,493,1432,1454]" country="Namibia" name="Kunene" pageId="1" pageNumber="129">Kunene Region</collectingRegion>
: 1812 (Sanitatas),
<collectingCounty id="C6344E7BFFD60421FD314EDF454C9DC9" box="[676,814,1432,1454]" pageId="1" pageNumber="129">Namib Desert</collectingCounty>
,
<quantity id="E8129B12FFD60421FCAC4EDF450E9DCA" box="[825,876,1432,1454]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="129" unit="km" value="3.0">3 km</quantity>
north of
<collectingMunicipality id="CF31AC8DFFD60421FC5C4EDF42359DCA" box="[969,1111,1432,1454]" pageId="1" pageNumber="129">Khumib River</collectingMunicipality>
,
<quantity id="E8129B12FFD60421FBF74EDF42F79DCA" box="[1122,1173,1432,1454]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="129" unit="km" value="4.0">4 km</quantity>
east of
<location id="2A35602CFFD60421FB704EDF43C99DCA" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C:2A35602CFFD60421FB704EDF43C99DCA" box="[1253,1451,1432,1454]" country="Namibia" county="Namib Desert" municipality="Khumib River" name="Skeleton Coast Park" pageId="1" pageNumber="129" stateProvince="Kunene">Skeleton Coast Park</location>
boundary, gneiss ridge (–DA),
<elevation id="A4C7D1C4FFD60421FE734EFB44359DB6" box="[486,599,1468,1490]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="1" pageNumber="129" unit="m" value="377.0">
<quantity id="E8129B12FFD60421FE734EFB44419DB5" box="[486,547,1468,1489]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="1" pageNumber="129" unit="m" value="377.0">377 m</quantity>
a.s.l.
</elevation>
,
<date id="5B541037FFD60421FDF74EFB44869DB5" box="[610,740,1468,1489]" pageId="1" pageNumber="129" value="2022-05-23">
<collectingDate id="4B10E9DFFFD60421FDF74EFB44869DB5" box="[610,740,1468,1489]" pageId="1" pageNumber="129" value="2022-05-23">23 May 2022</collectingDate>
</date>
,
<emphasis id="1D9EEAE5FFD60421FD7B4EFB45E29DB5" box="[750,896,1468,1489]" italics="true" pageId="1" pageNumber="129">Swanepoel 636</emphasis>
(
<typeStatus id="F0518855FFD60421FC194EFB45809DB6" box="[908,994,1468,1490]" pageId="1" pageNumber="129" type="holotype">holotype</typeStatus>
<collectionCode id="49FBAE32FFD60421FC724EFB42529DB5" box="[999,1072,1468,1489]" country="Namibia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13705" lsid="urn:lsid:biocol.org:col:13705" name="National Botanical Research Institute" pageId="1" pageNumber="129" type="Herbarium">WIND</collectionCode>
!;
<typeStatus id="F0518855FFD60421FBA84EFB42EE9DB6" box="[1085,1164,1468,1490]" pageId="1" pageNumber="129" type="isotype">isotypes</typeStatus>
<collectionCode id="49FBAE32FFD60421FB074EFB42A79DB5" box="[1170,1221,1468,1489]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="1" pageNumber="129" type="Herbarium">PRE</collectionCode>
!,
<collectionCode id="49FBAE32FFD60421FB474EFB436B9DB5" box="[1234,1289,1468,1489]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14594" lsid="urn:lsid:biocol.org:col:14594" name="University of Pretoria" pageId="1" pageNumber="129" type="Herbarium">PRU</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="2F5536F7FFD60422FF1D4D4642C59B90" blockId="1.[136,1452,1536,2067]" lastBlockId="2.[136,1452,158,2093]" lastPageId="2" lastPageNumber="130" pageId="1" pageNumber="129">
Erect, hemispherical or procumbent woody dwarf shrub up to
<quantity id="E8129B12FFD60421FCF14D4745C59E7F" box="[868,935,1536,1563]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="129" unit="m" value="0.8">0.8 m</quantity>
tall; all vegetative parts with a dense white indumentum of dendritic trichomes, the trichomes sparsely branched or bottlebrush-like, sometimes rebranching, appearing lanate, interspersed with few or scattered short-stalked glandular trichomes, the latter often concealed by the much longer dendritic ones.
<emphasis id="1D9EEAE5FFD60421FE574D2B44619EE2" box="[450,515,1644,1670]" italics="true" pageId="1" pageNumber="129">Stems</emphasis>
single or multi-stemmed from just below or above ground level from thick rootstock or main stem, up to
<quantity id="E8129B12FFD60421FEF04DD747A39ECF" box="[357,449,1680,1707]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="129" unit="mm" value="120.0">120 mm</quantity>
in diam., bark rough and fissured, corky in older plants, grey or blackish grey; older distal stems cylindrical, bark smooth, often longitudinally fissured or rough and corky, cream-white, cream-brown or grey; young stems quadrangular, green, becoming cream-white with age, glabrescent, cystoliths visible.
<emphasis id="1D9EEAE5FFD60421FB604D9E43219E96" box="[1269,1347,1753,1778]" italics="true" pageId="1" pageNumber="129">Leaves</emphasis>
opposite and decussate on new shoots, fascicled on older stems; petiole
<quantity id="E8129B12FFD60421FCF64DBB45B39F73" box="[867,977,1788,1815]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="14.0" metricValueMin="1.0" pageId="1" pageNumber="129" unit="mm" value="7.5" valueMax="14.0" valueMin="1.0">1–14 mm</quantity>
long; lamina ovate, elliptic, suborbicular or orbicular, flat, often sub-conduplicate towards apex, up to 42 ×
<quantity id="E8129B12FFD60421FC1C4C6745BF9F5F" box="[905,989,1824,1851]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="129" unit="mm" value="26.0">26 mm</quantity>
, green, apices rounded or acute, usually apiculate, bases cuneate, truncate or rounded, margins entire, often subinvolute, midrib and 3–6 principal lateral veins prominently raised adaxially, cystoliths not visible; indumentum on leaves with short bifurcate trichomes in addition.
<emphasis id="1D9EEAE5FFD60421FF1D4CCB46869FC2" box="[136,228,1932,1958]" italics="true" pageId="1" pageNumber="129">Flowers</emphasis>
in short axillary dichasia; bracts foliaceous, lanceolate (
<emphasis id="1D9EEAE5FFD60421FCE34CCA45D69FC2" box="[886,948,1933,1958]" italics="true" pageId="1" pageNumber="129">sensu</emphasis>
Lindley), linear-lanceolate or oblanceolate, sessile, 6–13 ×
<quantity id="E8129B12FFD60421FEA44CF747ED9FAF" box="[305,399,1968,1995]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="129" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
; pedicels (below bracteoles; “peduncle” of some authors) up to
<quantity id="E8129B12FFD60421FBD94CF742E99FAF" box="[1100,1163,1968,1995]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="129" unit="mm" value="1.0">1 mm</quantity>
long; bracteoles narrowly ovate, slightly asymmetrical, coriaceous, ca. 11 ×
<quantity id="E8129B12FFD60421FD384C93448D9F8B" box="[685,751,2004,2031]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="129" unit="mm" value="5.0">5 mm</quantity>
, connate proximally for up to
<quantity id="E8129B12FFD60421FBAC4C9342ED9F8B" box="[1081,1167,2004,2031]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="129" unit="mm" value="2.5">2.5 mm</quantity>
, apex acute or acuminate, pale green, cream-brown when dry, venation reticulate, indumentum abaxially similar to vegetative parts but trichomes longer and more robust, adaxially strigose towards apex and with additional short-stalked glandular trichomes or glabrous, margin lanate towards apex, cystoliths visible both sides, linear or curved, dense.
<taxonomicName id="E8EA4D74FFD50422FB0A4B85428298B8" box="[1183,1248,194,220]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="2" pageNumber="130" phylum="Porifera" rank="genus">
<emphasis id="1D9EEAE5FFD50422FB0A4B85428298B8" box="[1183,1248,194,220]" italics="true" pageId="2" pageNumber="130">Calyx</emphasis>
</taxonomicName>
<quantity id="E8129B12FFD50422FB7C4B85431398B9" box="[1257,1393,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="6.5" metricValueMin="5.9" pageId="2" pageNumber="130" unit="mm" value="6.2" valueMax="6.5" valueMin="5.9">5.9–6.5 mm</quantity>
long including basal tube of
<quantity id="E8129B12FFD50422FE1B4BA144779965" box="[398,533,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="2" pageNumber="130" unit="mm" value="1.2" valueMax="1.5" valueMin="0.9">0.9–1.5 mm</quantity>
deep, lobes 4, regular, narrowly triangular, acute, unequal, 3.9–5.6 ×
<quantity id="E8129B12FFD50422FA8B4BA143C59965" box="[1310,1447,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.4" metricValueMin="0.6" pageId="2" pageNumber="130" unit="mm" value="1.0" valueMax="1.4" valueMin="0.6">0.6–1.4 mm</quantity>
, anticous lobe indistinctly bifid; strigose both sides, scattered short-stalked glandular trichomes in addition abaxially.
<emphasis id="1D9EEAE5FFD50422FF1D4A6946BC992C" box="[136,222,302,328]" italics="true" pageId="2" pageNumber="130">Corolla</emphasis>
with narrow unexpanded portion of tube cylindrical, laterally slightly flattened, 15.0–
<quantity id="E8129B12FFD50422FB0C4A694299992D" box="[1177,1275,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="2" pageNumber="130" unit="mm" value="17.5">17.5 mm</quantity>
long with lobes straightened, narrow portion
<quantity id="E8129B12FFD50422FE444A15443B9909" box="[465,601,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" metricValueMax="8.2" metricValueMin="7.4" pageId="2" pageNumber="130" unit="mm" value="7.8" valueMax="8.2" valueMin="7.4">7.4–8.2 mm</quantity>
long, ca.
<quantity id="E8129B12FFD50422FD5F4A1545439909" box="[714,801,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="130" unit="mm" value="2.3">2.3 mm</quantity>
diam., expanded portion at a slight angle to anterior side of narrow portion,
<quantity id="E8129B12FFD50422FEF74A31478999F5" box="[354,491,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.5" metricValueMin="3.9" pageId="2" pageNumber="130" unit="mm" value="4.2" valueMax="4.5" valueMin="3.9">3.9–4.5 mm</quantity>
long, outside glabrous, sometimes sparingly puberulous on posterior side, inside of anticous portion towards mouth puberulous and with few long stiff white simple trichomes, inside otherwise glabrous; lobes patent with respect to corolla tube axis, anterior lobe obovate, margins entire, apices retuse or truncate, 5.0–5.7 ×
<quantity id="E8129B12FFD50422FF364AA547529999" box="[163,304,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" metricValueMax="5.5" metricValueMin="4.7" pageId="2" pageNumber="130" unit="mm" value="5.1" valueMax="5.5" valueMin="4.7">4.7–5.5 mm</quantity>
, lateral and upper lobes rectangular (upper lobes obovate when flattened), margins often revolute, apices retuse or truncate, lateral lobes 4.0–4.9 ×
<quantity id="E8129B12FFD50422FD304941454C9A45" box="[677,814,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.5" metricValueMin="2.6" pageId="2" pageNumber="130" unit="mm" value="3.05" valueMax="3.5" valueMin="2.6">2.6–3.5 mm</quantity>
, upper lobes 4.9–5.2 ×
<quantity id="E8129B12FFD50422FBA1494142DF9A45" box="[1076,1213,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="2" pageNumber="130" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2 mm</quantity>
, connate for 25–40% of their length, slightly overlapping, discolorous, abaxially light brown, adaxially vermillion, anterior lobe adaxially magenta with two narrowly triangular yellow nectar guides, the two lateral lobes adaxially vermillion or carmine distally, grading into vermillion towards throat, lacking nectar guides or nectar guides inconspicuous, slightly darker in colour than that of the associated lobes, lobes glabrous except for few long stiff white simple trichomes towards bases adaxially; palate prominently transversely 4- or 5-ribbed.
<emphasis id="1D9EEAE5FFD50422FCD949FC45CB9AB1" box="[844,937,699,725]" italics="true" pageId="2" pageNumber="130">Stamens</emphasis>
didynamous, inserted dorsally in throat, fused portion
<quantity id="E8129B12FFD50422FF77499947089A9D" box="[226,362,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.7" metricValueMin="1.1" pageId="2" pageNumber="130" unit="mm" value="1.4" valueMax="1.7" valueMin="1.1">1.1–1.7 mm</quantity>
long, free parts slightly tapering towards apex, glabrous, long filaments ca. 4.0 mm long, short filaments ca.
<quantity id="E8129B12FFD50422FEB7484547189B79" box="[290,378,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="130" unit="mm" value="2.6">2.6 mm</quantity>
long, outer filament with basal ridge from point of insertion on corolla (“trace”) decurrent to
<quantity id="E8129B12FFD50422FF1D4861476D9B25" box="[136,271,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.7" metricValueMin="4.2" pageId="2" pageNumber="130" unit="mm" value="4.95" valueMax="5.7" valueMin="4.2">4.2–5.7 mm</quantity>
from base of tube, puberulous; filament curtain reduced (
<emphasis id="1D9EEAE5FFD50422FC07486045B29B24" box="[914,976,807,832]" italics="true" pageId="2" pageNumber="130">sensu</emphasis>
terminology of
<bibRefCitation id="4B7B4B06FFD50422FB104861432A9B24" author="Manktelow, M." box="[1157,1352,806,832]" pageId="2" pageNumber="130" pagination="129 - 160" refId="ref5170" refString="Manktelow, M. (2000) The filament curtain: a structure important to systematics and pollination biology in the Acanthaceae. Botanical Journal of the Linnean Society 133 (2): 129 - 160. https: // doi. org / 10.1006 / bojl. 1999.0309" type="journal article" year="2000">Manktelow 2000</bibRefCitation>
); anthers 2-thecous, thecae linear-elliptic, equal, ca.
<quantity id="E8129B12FFD50422FDF2480D44DF9B01" box="[615,701,842,869]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="130" unit="mm" value="2.3">2.3 mm</quantity>
long including short basal spur, cream-brown with scattered short-stalked glandular trichomes.
<emphasis id="1D9EEAE5FFD50422FE46482844319BED" box="[467,595,879,905]" italics="true" pageId="2" pageNumber="130">Gynoecium</emphasis>
12.0–
<quantity id="E8129B12FFD50422FD09482945639BED" box="[668,769,878,905]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="130" unit="mm" value="13.8">13.8 mm</quantity>
long; ovary ovoid, laterally compressed, 1.1–1.7 ×
<quantity id="E8129B12FFD50422FAC2482946D49BC9" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="2" pageNumber="130" unit="mm" value="1.1" valueMax="1.4" valueMin="0.8">0.8–1.4 mm</quantity>
, inserted in fleshy disc, glabrous; style filiform, 10.0–
<quantity id="E8129B12FFD50422FCB248D545EE9BC9" box="[807,908,914,941]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="130" unit="mm" value="11.8">11.8 mm</quantity>
long, puberulous, stigma lobes linear, unequal, longer lobe ca.1.0 mm long, shorter lobe ca.
<quantity id="E8129B12FFD50422FD1648F144B89BB5" box="[643,730,950,977]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="130" unit="mm" value="0.7">0.7 mm</quantity>
long.
<emphasis id="1D9EEAE5FFD50422FCB748F1451C9BB4" box="[802,894,950,976]" italics="true" pageId="2" pageNumber="130">Capsule</emphasis>
flattened ovoid, ca. 5.5 ×
<quantity id="E8129B12FFD50422FB3D48F143639BB5" box="[1192,1281,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="130" unit="mm" value="3.1">3.1 mm</quantity>
, tawny, glossy, glabrous; seeds cordate, ca. 3.1 ×
<quantity id="E8129B12FFD50422FD96489D44399B91" box="[515,603,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="130" unit="mm" value="2.4">2.4 mm</quantity>
, densely covered with white hygroscopic trichomes.
</paragraph>
<paragraph id="2F5536F7FFD50422FF2848B942289C7C" blockId="2.[136,1452,158,2093]" box="[189,1098,1022,1049]" pageId="2" pageNumber="130">
<emphasis id="1D9EEAE5FFD50422FF2848B947219C7C" bold="true" box="[189,323,1022,1048]" pageId="2" pageNumber="130">Phenology:</emphasis>
—Flowers and fruit have been recorded from February to November.
</paragraph>
</subSubSection>
<subSubSection id="67F0657CFFD50422FF284F6544899E18" pageId="2" pageNumber="130" type="distribution">
<paragraph id="2F5536F7FFD50422FF284F6542E29DC1" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="130">
<emphasis id="1D9EEAE5FFD50422FF284F6547859C58" bold="true" box="[189,487,1058,1084]" pageId="2" pageNumber="130">Distribution and habitat:</emphasis>
—At present,
<taxonomicName id="E8EA4D74FFD50422FD174F6545189C58" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[642,890,1058,1084]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FD174F6545189C58" box="[642,890,1058,1084]" italics="true" pageId="2" pageNumber="130">Petalidium namibense</emphasis>
</taxonomicName>
is only known from the area to the southwest, west and northwest of Puros, between the lower Sechomib and Hoarusib rivers below the Great Escarpment in northwestern
<collectingCountry id="57FD7667FFD50422FF1D4F2D468A9CE0" box="[136,232,1130,1156]" name="Namibia" pageId="2" pageNumber="130">Namibia</collectingCountry>
(
<figureCitation id="B7D12A72FFD50422FF6D4F2D475F9CE0" box="[248,317,1130,1156]" captionStart="FIGURE 4" captionStartId="6.[136,229,1477,1499]" captionTargetBox="[151,1435,190,1453]" captionTargetId="figure-18@6.[151,1435,190,1453]" captionTargetPageId="6" captionText="FIGURE 4. Known distribution of Petalidium namibense (black dots; ●) based on specimens in Herbs PRE and WIND.Also depicted are the distribution range of P. sesfonteinense (shaded grey), based on Swanepoel &amp; Manzitto-Tripp (2022)." figureDoi="http://doi.org/10.5281/zenodo.14521294" httpUri="https://zenodo.org/record/14521294/files/figure.png" pageId="2" pageNumber="130">Fig. 4</figureCitation>
). The specimen
<emphasis id="1D9EEAE5FFD50422FE604F2C45639CE0" box="[501,769,1130,1157]" italics="true" pageId="2" pageNumber="130">Moss &amp; Jacobsen K312</emphasis>
from the Hoanib River Valley (southernmost point in
<figureCitation id="B7D12A72FFD50422FAF54F2D43C19CE0" box="[1376,1443,1130,1156]" captionStart="FIGURE 4" captionStartId="6.[136,229,1477,1499]" captionTargetBox="[151,1435,190,1453]" captionTargetId="figure-18@6.[151,1435,190,1453]" captionTargetPageId="6" captionText="FIGURE 4. Known distribution of Petalidium namibense (black dots; ●) based on specimens in Herbs PRE and WIND.Also depicted are the distribution range of P. sesfonteinense (shaded grey), based on Swanepoel &amp; Manzitto-Tripp (2022)." figureDoi="http://doi.org/10.5281/zenodo.14521294" httpUri="https://zenodo.org/record/14521294/files/figure.png" pageId="2" pageNumber="130">Fig. 4</figureCitation>
) is located ca.
<quantity id="E8129B12FFD50422FEB54FC947059CCC" box="[288,359,1166,1192]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="130" unit="km" value="70.0">70 km</quantity>
to the south of the known core range of
<taxonomicName id="E8EA4D74FFD50422FCB94FC845DD9CCC" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[812,959,1166,1192]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FCB94FC845DD9CCC" box="[812,959,1166,1192]" italics="true" pageId="2" pageNumber="130">P. namibense</emphasis>
</taxonomicName>
. It morphologically seems to conform in all respects to
<taxonomicName id="E8EA4D74FFD50422FE9F4FF447FD9CA8" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[266,415,1202,1228]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FE9F4FF447FD9CA8" box="[266,415,1202,1228]" italics="true" pageId="2" pageNumber="130">P. namibense</emphasis>
</taxonomicName>
and probably represents a second, outlier population of the species.
<taxonomicName id="E8EA4D74FFD50422FB254FF543CE9CA8" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[1200,1452,1202,1228]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FB254FF543CE9CA8" box="[1200,1452,1202,1228]" italics="true" pageId="2" pageNumber="130">Petalidium namibense</emphasis>
</taxonomicName>
occurs at the base of rocky outcrops, on arid hillsides, and along drainage lines at elevations of
<quantity id="E8129B12FFD50422FB344F91437D9C95" box="[1185,1311,1238,1265]" metricMagnitude="2" metricUnit="m" metricValue="4.2" metricValueMax="5.6" metricValueMin="2.8" pageId="2" pageNumber="130" unit="m" value="420.0" valueMax="560.0" valueMin="280.0">280–560 m</quantity>
a.s.l.,
<quantity id="E8129B12FFD50422FAF04F9146CE9D70" metricMagnitude="4" metricUnit="m" metricValue="3.75" metricValueMax="5.5" metricValueMin="2.0" pageId="2" pageNumber="130" unit="km" value="37.5" valueMax="55.0" valueMin="20.0">20–55 km</quantity>
inland from the Atlantic Ocean. Average annual rainfall in the area is less than
<quantity id="E8129B12FFD50422FB8F4FBD42149D71" box="[1050,1142,1274,1301]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="130" unit="mm" value="100.0">100 mm</quantity>
and falls mainly in summer (
<bibRefCitation id="4B7B4B06FFD50422FF054E5947BB9D5C" author="Atlas of Namibia Team" box="[144,473,1310,1337]" pageId="2" pageNumber="130" refId="ref4491" refString="Atlas of Namibia Team. (2022) Atlas of Namibia: its land, water and life. Namibia Nature Foundation, Windhoek, 390 pp." type="book" year="2022">
Atlas of
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Team 2022
</bibRefCitation>
). In certain parts of the Namib Desert, fog from the Atlantic Ocean is a regular and vital source of moisture, particularly for smaller animals (
<bibRefCitation id="4B7B4B06FFD50422FD594E0545C99D38" author="Mitchell, D. &amp; Henschel, J. R. &amp; Hetem, R. S. &amp; Wassenaar, T. D. &amp; Strauss, W. M. &amp; Hanrahan, S. A. &amp; Seely, M. K." box="[716,939,1346,1372]" pageId="2" pageNumber="130" refId="ref5364" refString="Mitchell, D., Henschel, J. R., Hetem, R. S., Wassenaar, T. D., Strauss, W. M., Hanrahan, S. A. &amp; Seely, M. K. (2020) Fog and fauna of the Namib Desert: past and future. Ecosphere 11 (1): e 02996 [40 pp.]. https: // doi. org / 10.1002 / ecs 2.2996" type="book" year="2020">
Mitchell
<emphasis id="1D9EEAE5FFD50422FCA74E04450B9D38" box="[818,873,1346,1372]" italics="true" pageId="2" pageNumber="130">et al.</emphasis>
2020
</bibRefCitation>
). Although the area in the distribution range of
<taxonomicName id="E8EA4D74FFD50422FF1D4E2047799DE4" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[136,283,1382,1408]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FF1D4E2047799DE4" box="[136,283,1382,1408]" italics="true" pageId="2" pageNumber="130">P. namibense</emphasis>
</taxonomicName>
experiences fog about 5–10% of the year (
<bibRefCitation id="4B7B4B06FFD50422FD634E21425F9DE4" author="Atlas of Namibia Team" box="[758,1085,1382,1409]" pageId="2" pageNumber="130" refId="ref4491" refString="Atlas of Namibia Team. (2022) Atlas of Namibia: its land, water and life. Namibia Nature Foundation, Windhoek, 390 pp." type="book" year="2022">
Atlas of
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Team 2022
</bibRefCitation>
), the poor condition of the plants during droughts suggests that they may not benefit significantly from this moisture source.
</paragraph>
<paragraph id="2F5536F7FFD50422FF284EE944899E18" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="130">
<emphasis id="1D9EEAE5FFD50422FF284EE947D19DAC" bold="true" box="[189,435,1454,1480]" pageId="2" pageNumber="130">Conservation status:</emphasis>
—
<taxonomicName id="E8EA4D74FFD50422FE5A4EE944AA9DAC" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[463,712,1454,1480]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FE5A4EE944AA9DAC" box="[463,712,1454,1480]" italics="true" pageId="2" pageNumber="130">Petalidium namibense</emphasis>
</taxonomicName>
has been recorded at several localities in an area of ca. 55 ×
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where it is occasional to locally common. Although a brief search at various other localities with seemingly suitable habitat did not reveal any plants, it is probably more widespread than currently known. Even though it is protected within the Skeleton Coast National Park, where its range slightly extends, a recent survey of its distribution area revealed numerous recently dead plants, likely due to extended droughts.
<taxonomicName id="E8EA4D74FFD50422FC514D7942DD9E3C" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[964,1215,1598,1624]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FC514D7942DD9E3C" box="[964,1215,1598,1624]" italics="true" pageId="2" pageNumber="130">Petalidium namibense</emphasis>
</taxonomicName>
is here provisionally ranked as Vulnerable VU B1a(iii), (b)iii (
<bibRefCitation id="4B7B4B06FFD50422FDC04D2544829E18" author="IUCN" box="[597,736,1634,1660]" pageId="2" pageNumber="130" refId="ref5066" refString="IUCN (2012) IUCN red list categories and criteria: Version 3.1. 2 nd edn. Gland, Switzerland and Cambridge U. K., iv + 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="67F0657CFFD50426FF284DC147CB9F0C" lastPageId="6" lastPageNumber="134" pageId="2" pageNumber="130" type="etymology">
<paragraph id="2F5536F7FFD50422FF284DC142E29E8D" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="130">
<emphasis id="1D9EEAE5FFD50422FF284DC1472A9EC4" bold="true" box="[189,328,1670,1696]" pageId="2" pageNumber="130">Etymology:</emphasis>
—The specific epithet refers to the Namib Desert to which
<taxonomicName id="E8EA4D74FFD50422FC674DC1428C9EC4" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[1010,1262,1670,1696]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FC674DC1428C9EC4" box="[1010,1262,1670,1696]" italics="true" pageId="2" pageNumber="130">Petalidium namibense</emphasis>
</taxonomicName>
is endemic. The Namib Desert in its broadest definition, stretches along the Atlantic Ocean from Saõ Nicolau (Bentiaba) in
<collectingCountry id="57FD7667FFD50422FACC4DED43C99EA0" box="[1369,1451,1706,1732]" name="Angola" pageId="2" pageNumber="130">Angola</collectingCountry>
through
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to the Olifants River in
<collectingCountry id="57FD7667FFD50422FDC24D8944849E8C" box="[599,742,1742,1768]" name="South Africa" pageId="2" pageNumber="130">South Africa</collectingCountry>
(
<bibRefCitation id="4B7B4B06FFD50422FD614D8945149E8C" author="Seely, M." box="[756,886,1742,1768]" pageId="2" pageNumber="130" refId="ref5649" refString="Seely, M. (2004) The Namib: natural history of an ancient desert. Desert Research Foundation of Namibia, Windhoek, 110 pp." type="book" year="2004">Seely 2004</bibRefCitation>
,
<bibRefCitation id="4B7B4B06FFD50422FC144D8942169E8D" author="Goudie, A. &amp; Viles, H." box="[897,1140,1742,1769]" pageId="2" pageNumber="130" refId="ref5022" refString="Goudie, A. &amp; Viles, H. (2015) Landscapes and land forms of Namibia. Springer, Dordrecht, 173 pp. https: // doi. org / 10.1007 / 978 - 94 - 017 - 8020 - 9" type="book" year="2015">Goudie &amp; Viles 2015</bibRefCitation>
).
</paragraph>
<paragraph id="2F5536F7FFD50422FF284DB543329F1C" blockId="2.[136,1452,158,2093]" pageId="2" pageNumber="130">
<emphasis id="1D9EEAE5FFD50422FF284DB5476B9F68" bold="true" box="[189,265,1778,1804]" pageId="2" pageNumber="130">Notes:</emphasis>
—
<taxonomicName id="E8EA4D74FFD50422FEB34DB544429F68" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[294,544,1778,1804]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FEB34DB544429F68" box="[294,544,1778,1804]" italics="true" pageId="2" pageNumber="130">Petalidium namibense</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="E8EA4D74FFD50422FC244DB4420D9F68" authorityName="Swanepoel &amp; Manzitto-Tripp" authorityYear="2022" box="[945,1135,1778,1804]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="1D9EEAE5FFD50422FC244DB4420D9F68" box="[945,1135,1778,1804]" italics="true" pageId="2" pageNumber="130">P. sesfonteinense</emphasis>
</taxonomicName>
, perhaps its closest relative. Hence these two species were compared in the diagnoses above. Some of the morphological features to distinguish between
<taxonomicName id="E8EA4D74FFD50422FF7E4C7C471F9F30" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[235,381,1850,1876]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD50422FF7E4C7C471F9F30" box="[235,381,1850,1876]" italics="true" pageId="2" pageNumber="130">P. namibense</emphasis>
</taxonomicName>
and
<taxonomicName id="E8EA4D74FFD50422FE244C7C440F9F30" authorityName="Swanepoel &amp; Manzitto-Tripp" authorityYear="2022" box="[433,621,1850,1876]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="1D9EEAE5FFD50422FE244C7C440F9F30" box="[433,621,1850,1876]" italics="true" pageId="2" pageNumber="130">P. sesfonteinense</emphasis>
</taxonomicName>
are provided in
<tableCitation id="6268034CFFD50422FC8B4C7D45129F30" box="[798,880,1850,1876]" captionStart="TABLE 1" captionStartId="8.[136,220,158,182]" captionText="TABLE 1. Morphological differences between Petalidium namibense and P. sesfonteinense." pageId="2" pageNumber="130">Table 1</tableCitation>
; also see
<figureCitation id="B7D12A72FFD50422FC4C4C7D42799F31" box="[985,1051,1850,1877]" captionStart="FIGURE 5" captionStartId="7.[136,229,1952,1974]" captionTargetBox="[165,1422,190,1928]" captionTargetId="figure-17@7.[165,1422,190,1928]" captionTargetPageId="7" captionText="FIGURE 5. Petalidium sesfonteinense, morphology of leaves and flowers. A. Flowers, shoots, and leaves; greyish leaves are still densely covered in trichomes; green leaves almost glabrous. B–E. Flowers in front view showing variation in corolla colour and lobe margins, with two yellow nectar guides. Photographs: W. Swanepoel. Republished from Swanepoel &amp; Manzitto-Tripp (2022)." figureDoi="http://doi.org/10.5281/zenodo.14521296" httpUri="https://zenodo.org/record/14521296/files/figure.png" pageId="2" pageNumber="130">Fig. 5</figureCitation>
. An identification key to distinguish between these two species, as well as
<taxonomicName id="E8EA4D74FFD50422FDA44C1844B89F1C" authorityName="Clarke" authorityYear="1899" baseAuthorityName="Schinz" baseAuthorityYear="1890" box="[561,730,1886,1912]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="englerianum">
<emphasis id="1D9EEAE5FFD50422FDA44C1844B89F1C" box="[561,730,1886,1912]" italics="true" pageId="2" pageNumber="130">P. englerianum</emphasis>
</taxonomicName>
,
<taxonomicName id="E8EA4D74FFD50422FD7D4C1845D79F1C" authorityName="Meyer" authorityYear="1961" box="[744,949,1887,1912]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="rossmannianum">
<emphasis id="1D9EEAE5FFD50422FD7D4C1845D79F1C" box="[744,949,1887,1912]" italics="true" pageId="2" pageNumber="130">P. rossmannianum</emphasis>
</taxonomicName>
, and
<taxonomicName id="E8EA4D74FFD50422FC674C1842119F1C" authorityName="Clarke" authorityYear="1899" box="[1010,1139,1886,1912]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="variabile">
<emphasis id="1D9EEAE5FFD50422FC674C1842119F1C" box="[1010,1139,1886,1912]" italics="true" pageId="2" pageNumber="130">P. variabile</emphasis>
</taxonomicName>
, is provided below.
</paragraph>
<paragraph id="2F5536F7FFD50423FF284CC5445199F4" blockId="2.[136,1452,158,2093]" lastBlockId="3.[136,1452,158,401]" lastPageId="3" lastPageNumber="131" pageId="2" pageNumber="130">
When not in flower and especially in the case of herbarium specimens, the new species can also be confused with several other species of
<taxonomicName id="E8EA4D74FFD50422FD954CE144199FA4" authorityName="Nees von Esenbeck" authorityYear="1832" box="[512,635,1958,1984]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="genus">
<emphasis id="1D9EEAE5FFD50422FD954CE144199FA4" box="[512,635,1958,1984]" italics="true" pageId="2" pageNumber="130">Petalidium</emphasis>
</taxonomicName>
from the
<collectingRegion id="ED2EF815FFD50422FC924CE045A19FA4" box="[775,963,1958,1985]" country="Namibia" name="Kunene" pageId="2" pageNumber="130">Kunene Region</collectingRegion>
,
<collectingCountry id="57FD7667FFD50422FC4D4CE142599FA4" box="[984,1083,1958,1984]" name="Namibia" pageId="2" pageNumber="130">Namibia</collectingCountry>
, with dense indumentum and similar inflorescences, notably
<taxonomicName id="E8EA4D74FFD50422FE7E4C8D45959F80" authority="Swanepoel (2020: 237)" authorityName="Swanepoel" authorityPageNumber="237" authorityYear="2020" box="[491,1015,1994,2020]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="1D9EEAE5FFD50422FE7E4C8D44869F80" box="[491,740,1994,2020]" italics="true" pageId="2" pageNumber="130">Petalidium kaokoense</emphasis>
<bibRefCitation id="4B7B4B06FFD50422FD794C8D45959F80" author="Swanepoel, W." box="[748,1015,1994,2020]" pageId="2" pageNumber="130" pagination="236 - 242" refId="ref5677" refString="Swanepoel, W. (2020) Petalidium kaokoense (Acanthaceae), a new species from Namibia. Phytotaxa 468 (3): 236 - 242. https: // doi. org / 10.11646 / phytotaxa. 468.3.1" type="journal article" year="2020">Swanepoel (2020: 237)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E8EA4D74FFD50422FB914C8C43E89F80" authority="Meyer (1973: 108)" authorityName="Meyer" authorityPageNumber="108" authorityYear="1973" box="[1028,1418,1994,2021]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="ohopohense">
<emphasis id="1D9EEAE5FFD50422FB914C8C42CA9F80" box="[1028,1192,1994,2020]" italics="true" pageId="2" pageNumber="130">P. ohopohense</emphasis>
<bibRefCitation id="4B7B4B06FFD50422FB254C8C43E89F80" author="Meyer, P. G." box="[1200,1418,1994,2021]" pageId="2" pageNumber="130" pagination="101 - 113" refId="ref5302" refString="Meyer, P. G. (1973) Neue und enig bekannte Acanthaceen aus dem Kaokoveld (Sudwestafrika). Mitteilungen der Botanischen Staatssammlung Munchen 11: 101 - 113. Available from: https: // www. biodiversitylibrary. org / item / 51768 # page / 111 / mode / 1 up (accessed 11 November 2024)" type="journal article" year="1973">Meyer (1973: 108)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="E8EA4D74FFD50422FA024C8C4758906C" authorityName="Meyer" authorityYear="1961" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="rossmannianum">
<emphasis id="1D9EEAE5FFD50422FA024C8C4758906C" italics="true" pageId="2" pageNumber="130">P. rossmannianum</emphasis>
</taxonomicName>
, and
<taxonomicName id="E8EA4D74FFD50422FEE24CA8448D906C" authority="Moore (1880: 227)" authorityName="Moore" authorityPageNumber="227" authorityYear="1880" box="[375,751,2030,2057]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis id="1D9EEAE5FFD50422FEE24CA84472906C" box="[375,528,2030,2056]" italics="true" pageId="2" pageNumber="130">P. welwitschii</emphasis>
<bibRefCitation id="4B7B4B06FFD50422FD824CA8448D906C" author="Moore, S." box="[535,751,2030,2057]" pageId="2" pageNumber="130" pagination="225 - 233" refId="ref5450" refString="Moore, S. (1880) Enumeratio Acanthacearum Herbarii Welwitschiani Angolensis. Journal of Botany, British and Foreign 18: 225 - 233. [https: // www. biodiversitylibrary. org / item / 108761 # page / 236 / mode / 1 up]" type="journal article" year="1880">Moore (1880: 227)</bibRefCitation>
</taxonomicName>
. However, the indumentum of
<taxonomicName id="E8EA4D74FFD50422FBDD4CA842BA906C" authorityName="Swanepoel" authorityYear="2020" box="[1096,1240,2030,2056]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="130" phylum="Tracheophyta" rank="species" species="kaokoense">
<emphasis id="1D9EEAE5FFD50422FBDD4CA842BA906C" box="[1096,1240,2030,2056]" italics="true" pageId="2" pageNumber="130">P. kaokoense</emphasis>
</taxonomicName>
consists of stellate trichomes interspersed with a few dendritic ones (
<emphasis id="1D9EEAE5FFD50422FD5A435444849048" box="[719,742,2067,2092]" italics="true" pageId="2" pageNumber="130">vs</emphasis>
. dendritic trichomes interspersed with scattered short-stalked glandular trichomes); some branches of the dendritic trichomes of
<taxonomicName id="E8EA4D74FFD40423FCF24BD8426598DC" authorityName="Meyer" authorityYear="1973" box="[871,1031,158,184]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="131" phylum="Tracheophyta" rank="species" species="ohopohense">
<emphasis id="1D9EEAE5FFD40423FCF24BD8426598DC" box="[871,1031,158,184]" italics="true" pageId="3" pageNumber="131">P. ohopohense</emphasis>
</taxonomicName>
bear a terminal gland and the corollas are pilose on the outside (
<emphasis id="1D9EEAE5FFD40423FE3B4B8447A798B8" box="[430,453,195,220]" italics="true" pageId="3" pageNumber="131">vs</emphasis>
. trichome branches eglandular and corolla glabrous outside); the dendritic trichomes on the leaves and bracteoles of
<taxonomicName id="E8EA4D74FFD40423FE5D4BA044F59964" authorityName="Meyer" authorityYear="1961" box="[456,663,231,256]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="131" phylum="Tracheophyta" rank="species" species="rossmannianum">
<emphasis id="1D9EEAE5FFD40423FE5D4BA044F59964" box="[456,663,231,256]" italics="true" pageId="3" pageNumber="131">P. rossmannianum</emphasis>
</taxonomicName>
are very small with short lateral branches, interspersed with scattered stellate trichomes, and the expanded part of the corolla tube is puberulous (
<emphasis id="1D9EEAE5FFD40423FB944A4C427A9940" box="[1025,1048,267,292]" italics="true" pageId="3" pageNumber="131">vs</emphasis>
. dendritic trichomes larger, longer branched, interspersed with scattered short-stalked glandular trichomes, corolla glabrous outside); the bracteoles of
<taxonomicName id="E8EA4D74FFD40423FA024A6847649908" authorityName="Moore" authorityYear="1880" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="131" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis id="1D9EEAE5FFD40423FA024A6847649908" italics="true" pageId="3" pageNumber="131">P. welwitschii</emphasis>
</taxonomicName>
are covered abaxially with scattered villose trichomes up to
<quantity id="E8129B12FFD40423FC364A1545819909" box="[931,995,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="131" unit="mm" value="4.0">4 mm</quantity>
long (
<emphasis id="1D9EEAE5FFD40423FBBF4A1442239908" box="[1066,1089,339,364]" italics="true" pageId="3" pageNumber="131">vs</emphasis>
. bracteoles appearing lanate due to dense layer of dendritic trichomes).
</paragraph>
<caption id="7B95667FFFD40423FF1D4CA8430F904E" ID-DOI="http://doi.org/10.5281/zenodo.14521288" ID-Zenodo-Dep="14521288" httpUri="https://zenodo.org/record/14521288/files/figure.png" pageId="3" pageNumber="131" startId="3.[136,229,2031,2053]" targetBox="[222,1366,425,2007]" targetPageId="3" targetType="figure">
<paragraph id="2F5536F7FFD40423FF1D4CA8430F904E" blockId="3.[136,1451,2031,2090]" pageId="3" pageNumber="131">
<emphasis id="1D9EEAE5FFD40423FF1D4CA847629060" bold="true" box="[136,256,2031,2053]" pageId="3" pageNumber="131">FIGURE 1.</emphasis>
<taxonomicName id="E8EA4D74FFD40423FE924CB747BD9061" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[263,479,2032,2053]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="131" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD40423FE924CB747BD9061" box="[263,479,2032,2053]" italics="true" pageId="3" pageNumber="131">Petalidium namibense</emphasis>
</taxonomicName>
, habitat and habit.
<emphasis id="1D9EEAE5FFD40423FD0A4CA844D49060" bold="true" box="[671,694,2031,2052]" pageId="3" pageNumber="131">A.</emphasis>
Several plants (dark grey dwarf shrubs) growing in the bed of an ephemeral drainage line.
<emphasis id="1D9EEAE5FFD40423FE8443534745904D" bold="true" box="[273,295,2068,2089]" pageId="3" pageNumber="131">B.</emphasis>
Ancient windswept plant sprouting from persistent woody stems with corky bark. Photographs by W. Swanepoel.
</paragraph>
</caption>
<caption id="7B95667FFFD30424FF1D4CE7477B9F9A" ID-DOI="http://doi.org/10.5281/zenodo.14521290" ID-Zenodo-Dep="14521290" httpUri="https://zenodo.org/record/14521290/files/figure.png" pageId="4" pageNumber="132" startId="4.[136,229,1952,1974]" targetBox="[165,1422,190,1927]" targetPageId="4" targetType="figure">
<paragraph id="2F5536F7FFD30424FF1D4CE7477B9F9A" blockId="4.[136,1452,1952,2046]" pageId="4" pageNumber="132">
<emphasis id="1D9EEAE5FFD30424FF1D4CE7469C9FD1" bold="true" box="[136,254,1952,1974]" pageId="4" pageNumber="132">FIGURE 2.</emphasis>
<taxonomicName id="E8EA4D74FFD30424FE904CE747BE9FD1" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[261,476,1952,1973]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="132" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD30424FE904CE747BE9FD1" box="[261,476,1952,1973]" italics="true" pageId="4" pageNumber="132">Petalidium namibense</emphasis>
</taxonomicName>
, habitat and habit.
<emphasis id="1D9EEAE5FFD30424FD024CE744CD9FD1" bold="true" box="[663,687,1952,1973]" pageId="4" pageNumber="132">A.</emphasis>
Mature plant (ca. 0.5 m high) with greyish appearance, growing among rocks.
<emphasis id="1D9EEAE5FFD30424FF1D4C8246FC9FBE" bold="true" box="[136,158,1989,2010]" pageId="4" pageNumber="132">B.</emphasis>
Multiple stems (thickest ca. 80 mm in diam.) from base of a relatively old plant, each covered with thick corky bark. Photographs by W. Swanepoel.
</paragraph>
</caption>
<caption id="7B95667FFFD20425FF1D4CE744A99046" ID-DOI="http://doi.org/10.5281/zenodo.14521292" ID-Zenodo-Dep="14521292" httpUri="https://zenodo.org/record/14521292/files/figure.png" pageId="5" pageNumber="133" startId="5.[136,229,1952,1974]" targetBox="[165,1422,190,1928]" targetPageId="5" targetType="figure">
<paragraph id="2F5536F7FFD20425FF1D4CE744A99046" blockId="5.[136,1452,1952,2082]" pageId="5" pageNumber="133">
<emphasis id="1D9EEAE5FFD20425FF1D4CE747629FD1" bold="true" box="[136,256,1952,1974]" pageId="5" pageNumber="133">FIGURE 3.</emphasis>
<taxonomicName id="E8EA4D74FFD20425FE9D4CE747839FD1" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[264,481,1952,1973]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="133" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD20425FE9D4CE747839FD1" box="[264,481,1952,1973]" italics="true" pageId="5" pageNumber="133">Petalidium namibense</emphasis>
</taxonomicName>
, morphology of leaves and flowers.
<emphasis id="1D9EEAE5FFD20425FCDB4CE745079FD1" bold="true" box="[846,869,1952,1973]" pageId="5" pageNumber="133">A.</emphasis>
Flower, reduced shoots, and congested leaves; leaves with dense greyish indumentum.
<emphasis id="1D9EEAE5FFD20425FE004C8247CE9FBE" bold="true" box="[405,428,1989,2010]" pageId="5" pageNumber="133">B.</emphasis>
Flower in the process of fading as indicated by its pale colours.
<emphasis id="1D9EEAE5FFD20425FB854C83424A9FBE" bold="true" box="[1040,1064,1988,2010]" pageId="5" pageNumber="133">C.</emphasis>
Newly opened flower with bracteoles in oblique lateral view; note brownish abaxial colour of posterior corolla lobes.
<emphasis id="1D9EEAE5FFD20425FCF94CAE45E69F9A" bold="true" box="[876,900,2025,2046]" pageId="5" pageNumber="133">D.</emphasis>
Flower in front view.
<emphasis id="1D9EEAE5FFD20425FBCC4CAE42129F9A" bold="true" box="[1113,1136,2025,2046]" pageId="5" pageNumber="133">E.</emphasis>
Flower with bracteoles in lateral view.
<emphasis id="1D9EEAE5FFD20425FF54434A46B19046" bold="true" box="[193,211,2061,2082]" pageId="5" pageNumber="133">F.</emphasis>
Flower in dorsal view. Photographs: W. Swanepoel.
</paragraph>
</caption>
<caption id="7B95667FFFD10426FF1D4E8242129D9A" ID-DOI="http://doi.org/10.5281/zenodo.14521294" ID-Zenodo-Dep="14521294" httpUri="https://zenodo.org/record/14521294/files/figure.png" pageId="6" pageNumber="134" startId="6.[136,229,1477,1499]" targetBox="[151,1435,190,1453]" targetPageId="6" targetType="figure">
<paragraph id="2F5536F7FFD10426FF1D4E8242129D9A" blockId="6.[136,1451,1476,1535]" pageId="6" pageNumber="134">
<emphasis id="1D9EEAE5FFD10426FF1D4E82469F9DBE" bold="true" box="[136,253,1477,1499]" pageId="6" pageNumber="134">FIGURE 4.</emphasis>
Known distribution of
<taxonomicName id="E8EA4D74FFD10426FE4E4E8244D39DBE" authorityName="Swanepoel &amp; A. E. van Wyk" authorityYear="2024" box="[475,689,1477,1498]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="134" phylum="Tracheophyta" rank="species" species="namibense">
<emphasis id="1D9EEAE5FFD10426FE4E4E8244D39DBE" box="[475,689,1477,1498]" italics="true" pageId="6" pageNumber="134">Petalidium namibense</emphasis>
</taxonomicName>
(black dots; ●) based on specimens in Herbs PRE and WIND. Also depicted are the distribution range of
<taxonomicName id="E8EA4D74FFD10426FEE04EAD447A9D9A" authorityName="Swanepoel &amp; Manzitto-Tripp" authorityYear="2022" box="[373,536,1513,1535]" class="Magnoliopsida" family="Acanthaceae" genus="Petalidium" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="134" phylum="Tracheophyta" rank="species" species="sesfonteinense">
<emphasis id="1D9EEAE5FFD10426FEE04EAD447A9D9A" box="[373,536,1513,1535]" italics="true" pageId="6" pageNumber="134">P. sesfonteinense</emphasis>
</taxonomicName>
(shaded grey), based on
<bibRefCitation id="4B7B4B06FFD10426FC924EAE420E9D9A" author="Swanepoel, W. &amp; Manzitto-Tripp, E." box="[775,1132,1513,1535]" pageId="6" pageNumber="134" pagination="127 - 135" refId="ref5785" refString="Swanepoel, W. &amp; Manzitto-Tripp, E. (2022) Petalidium sesfonteinense (Acanthaceae), a new species from the Kaokoveld, Namibia. Phytotaxa 549 (2): 127 - 135. https: // doi. org / 10.11646 / phytotaxa. 549.2.1" type="journal article" year="2022">Swanepoel &amp; Manzitto-Tripp (2022)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="2F5536F7FFD10426FF284D6A47F89E08" blockId="6.[136,1452,1581,1896]" pageId="6" pageNumber="134">
All the mentioned taxa are from the group composed of plants with irregular, four-parted calyces (
<bibRefCitation id="4B7B4B06FFD10426FABF4D6A46A19E0F" author="Obermeijer, A. A." pageId="6" pageNumber="134" pagination="151 - 162" refId="ref5587" refString="Obermeijer, A. A. (1936) The South African species of Petalidium. Annals of the Transvaal Museum 18: 151 - 162." type="journal article" year="1936">Obermeijer 1936</bibRefCitation>
,
<bibRefCitation id="4B7B4B06FFD10426FF5B4D1647EC9E08" author="Tripp, E. A. &amp; Tsai, Y. E. &amp; Zhuang, Y. &amp; Dexter, K. G." box="[206,398,1617,1644]" pageId="6" pageNumber="134" pagination="1 - 17" refId="ref6169" refString="Tripp, E. A., Tsai, Y. E., Zhuang, Y. &amp; Dexter, K. G. (2017) RADseq dataset with 90 % missing data fully resolves recent radiation of Petalidium (Acanthaceae) in the ultra-arid deserts of Namibia. Ecology and Evolution 7 (19): 1 - 17. https: // doi. org / 10.1002 / ece 3.3274" type="journal article" year="2017">
Tripp
<emphasis id="1D9EEAE5FFD10426FE874D1547219E0F" box="[274,323,1617,1643]" italics="true" pageId="6" pageNumber="134">et al</emphasis>
. 2017
</bibRefCitation>
).
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(
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):
</emphasis>
—
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,
<collectingRegion id="ED2EF815FFD10426FCFE4D3142799EEB" box="[875,1051,1653,1680]" country="Namibia" name="Kunene" pageId="6" pageNumber="134">Kunene Region</collectingRegion>
:
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southwest of
<location id="2A35602CFFD10426FB784D3143C59EEB" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C:2A35602CFFD10426FB784D3143C59EEB" box="[1261,1447,1653,1680]" country="Namibia" name="Ogams Fountain" pageId="6" pageNumber="134" stateProvince="Kunene">Ogams Fountain</location>
, 1812
<collectionCode id="49FBAE32FFD10426FF574DDD468A9ED0" box="[194,232,1690,1716]" country="Australia" name="The CB Rhizobium Collection" pageId="6" pageNumber="134">CB</collectionCode>
,
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,
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<collectingDate id="4B10E9DFFFD10426FED64DDE446E9ED0" box="[323,524,1689,1716]" pageId="6" pageNumber="134" value="2023-11-05">5 November 2023</collectingDate>
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,
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<collectorName id="821F5321FFD10426FD804DDE44EC9ED7" box="[533,654,1689,1715]" pageId="6" pageNumber="134">Swanepoel</collectorName>
639
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(
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!)
</materialsCitation>
;
<materialsCitation id="9F823CAAFFD10426FCAD4DDE44879EBC" collectingDate="1963-06-09" collectionCode="PRE, WIND" collectorName="Giess &amp; Leippert" country="Namibia" county="Namib-Kiesflache" location="Sarusas" municipality="Meilen" pageId="6" pageNumber="134" specimenCount="1">
<collectingCounty id="C6344E7BFFD10426FCAD4DDE42679ED7" box="[824,1029,1689,1715]" pageId="6" pageNumber="134">Namib-Kiesfläche</collectingCounty>
14
<collectingMunicipality id="CF31AC8DFFD10426FBBE4DDE42189ED7" box="[1067,1146,1689,1715]" pageId="6" pageNumber="134">Meilen</collectingMunicipality>
nördlich
<location id="2A35602CFFD10426FB774DDD43589ED0" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C:2A35602CFFD10426FB774DDD43589ED0" box="[1250,1338,1690,1716]" country="Namibia" county="Namib-Kiesflache" municipality="Meilen" name="Sarusas" pageId="6" pageNumber="134">Sarusas</location>
, 1812DA,
<date id="5B541037FFD10426FF1D4DF947779EB3" box="[136,277,1725,1752]" pageId="6" pageNumber="134" value="1963-06-09">
<collectingDate id="4B10E9DFFFD10426FF1D4DF947779EB3" box="[136,277,1725,1752]" pageId="6" pageNumber="134" value="1963-06-09">9 June 1963</collectingDate>
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,
<emphasis id="1D9EEAE5FFD10426FEB74DF944449EBC" box="[290,550,1726,1752]" italics="true" pageId="6" pageNumber="134">
<collectorName id="821F5321FFD10426FEB74DF947029EBC" box="[290,352,1726,1752]" pageId="6" pageNumber="134">Giess</collectorName>
&amp;
<collectorName id="821F5321FFD10426FE124DF947849EB3" box="[391,486,1726,1751]" pageId="6" pageNumber="134">Leippert</collectorName>
7453
</emphasis>
(
<collectionCode id="49FBAE32FFD10426FDA24DF944119EBC" box="[567,627,1726,1752]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="6" pageNumber="134" type="Herbarium">PRE</collectionCode>
!,
<collectionCode id="49FBAE32FFD10426FD164DF944BB9EBC" box="[643,729,1726,1752]" country="Namibia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13705" lsid="urn:lsid:biocol.org:col:13705" name="National Botanical Research Institute" pageId="6" pageNumber="134" type="Herbarium">WIND</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="9F823CAAFFD10426FD664DF947A49E98" collectingDate="1966-04-26" collectionCode="PRE, WIND" collectorName="Giess" country="Namibia" location="Sarusas Ost" municipality="Meilen" pageId="6" pageNumber="134" specimenCount="1">
20
<collectingMunicipality id="CF31AC8DFFD10426FC824DFA45049EB3" box="[791,870,1725,1751]" pageId="6" pageNumber="134">Meilen</collectingMunicipality>
nordoestlich
<location id="2A35602CFFD10426FB974DF942E89EBC" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C:2A35602CFFD10426FB974DF942E89EBC" box="[1026,1162,1726,1752]" country="Namibia" municipality="Meilen" name="Sarusas Ost" pageId="6" pageNumber="134">Sarusas Ost</location>
, 1812DA,
<date id="5B541037FFD10426FA934DFA43C59EB3" box="[1286,1447,1725,1751]" pageId="6" pageNumber="134" value="1966-04-26">
<collectingDate id="4B10E9DFFFD10426FA934DFA43C59EB3" box="[1286,1447,1725,1751]" pageId="6" pageNumber="134" value="1966-04-26">26 April 1966</collectingDate>
</date>
,
<emphasis id="1D9EEAE5FFD10426FF1D4DA547659E9F" box="[136,263,1761,1788]" italics="true" pageId="6" pageNumber="134">
<collectorName id="821F5321FFD10426FF1D4DA546A49E98" box="[136,198,1762,1788]" pageId="6" pageNumber="134">Giess</collectorName>
9416
</emphasis>
(
<collectionCode id="49FBAE32FFD10426FE8D4DA547369E98" box="[280,340,1762,1788]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="6" pageNumber="134" type="Herbarium">PRE</collectionCode>
!,
<collectionCode id="49FBAE32FFD10426FEF14DA547D89E98" box="[356,442,1762,1788]" country="Namibia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13705" lsid="urn:lsid:biocol.org:col:13705" name="National Botanical Research Institute" pageId="6" pageNumber="134" type="Herbarium">WIND</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="9F823CAAFFD10426FE464DA544CB9F44" collectingDate="2018-05-30" collectionCode="WIND" collectorName="Swanepoel" country="Namibia" elevation="456" location="Skeleton Coast National Park" municipality="Puros" pageId="6" pageNumber="134" specimenCount="1">
<quantity id="E8129B12FFD10426FE464DA5447E9E9F" box="[467,540,1761,1788]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="134" unit="km" value="30.0">30 km</quantity>
west-northwest of
<collectingMunicipality id="CF31AC8DFFD10426FD6D4DA5455A9E98" box="[760,824,1762,1788]" pageId="6" pageNumber="134">Puros</collectingMunicipality>
along track to
<location id="2A35602CFFD10426FC704DA643559E9F" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C:2A35602CFFD10426FC704DA643559E9F" box="[997,1335,1761,1788]" country="Namibia" municipality="Puros" name="Skeleton Coast National Park" pageId="6" pageNumber="134">Skeleton Coast National Park</location>
, 1812DA,
<quantity id="E8129B12FFD10426FF1D4C4246B19F44" box="[136,211,1797,1824]" metricMagnitude="2" metricUnit="m" metricValue="4.56" pageId="6" pageNumber="134" unit="m" value="456.0">
<elevation id="A4C7D1C4FFD10426FF1D4C4246B19F44" box="[136,211,1797,1824]" metricMagnitude="2" metricUnit="m" metricValue="4.56" pageId="6" pageNumber="134" unit="m" value="456.0">456 m</elevation>
</quantity>
,
<date id="5B541037FFD10426FF754C41471E9F44" box="[224,380,1798,1824]" pageId="6" pageNumber="134" value="2018-05-30">
<collectingDate id="4B10E9DFFFD10426FF754C41471E9F44" box="[224,380,1798,1824]" pageId="6" pageNumber="134" value="2018-05-30">30 May 2018</collectingDate>
</date>
,
<emphasis id="1D9EEAE5FFD10426FE1C4C4244579F7B" box="[393,565,1797,1823]" italics="true" pageId="6" pageNumber="134">
<collectorName id="821F5321FFD10426FE1C4C4244609F7B" box="[393,514,1797,1823]" pageId="6" pageNumber="134">Swanepoel</collectorName>
635
</emphasis>
(
<collectionCode id="49FBAE32FFD10426FDD34C4144FF9F44" box="[582,669,1798,1824]" country="Namibia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13705" lsid="urn:lsid:biocol.org:col:13705" name="National Botanical Research Institute" pageId="6" pageNumber="134" type="Herbarium">WIND</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="9F823CAAFFD10426FD234C4147929F20" collectingDate="2023-11-05" collectionCode="WIND" collectorName="Swanepoel" country="Namibia" elevation="551" location="Ogams Fountain" pageId="6" pageNumber="134" specimenCount="1">
<quantity id="E8129B12FFD10426FD234C4144939F7B" box="[694,753,1797,1824]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="134" unit="km" value="8.0">8 km</quantity>
southeast of
<location id="2A35602CFFD10426FC1E4C41422B9F7B" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C:2A35602CFFD10426FC1E4C41422B9F7B" box="[907,1097,1797,1824]" country="Namibia" name="Ogams Fountain" pageId="6" pageNumber="134">Ogams Fountain</location>
, 1812DA,
<quantity id="E8129B12FFD10426FB534C4243739F44" box="[1222,1297,1797,1824]" metricMagnitude="2" metricUnit="m" metricValue="5.51" pageId="6" pageNumber="134" unit="m" value="551.0">
<elevation id="A4C7D1C4FFD10426FB534C4243739F44" box="[1222,1297,1797,1824]" metricMagnitude="2" metricUnit="m" metricValue="5.51" pageId="6" pageNumber="134" unit="m" value="551.0">551 m</elevation>
</quantity>
,
<date id="5B541037FFD10426FA8B4C4246A19F20" pageId="6" pageNumber="134" value="2023-11-05">
<collectingDate id="4B10E9DFFFD10426FA8B4C4246A19F20" pageId="6" pageNumber="134" value="2023-11-05">5 November 2023</collectingDate>
</date>
,
<emphasis id="1D9EEAE5FFD10426FF454C6E471E9F27" box="[208,380,1833,1859]" italics="true" pageId="6" pageNumber="134">
<collectorName id="821F5321FFD10426FF454C6E472B9F27" box="[208,329,1833,1859]" pageId="6" pageNumber="134">Swanepoel</collectorName>
638
</emphasis>
(
<collectionCode id="49FBAE32FFD10426FE1B4C6D47879F20" box="[398,485,1834,1860]" country="Namibia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13705" lsid="urn:lsid:biocol.org:col:13705" name="National Botanical Research Institute" pageId="6" pageNumber="134" type="Herbarium">WIND</collectionCode>
!)
</materialsCitation>
;
<materialsCitation id="9F823CAAFFD10426FE6B4C6D47C79F0C" collectingDate="2023-11-03" collectionCode="DD, WIND" collectorName="Swanepoel" country="Namibia" elevation="292" location="Hoarusib River" municipality="Puros" pageId="6" pageNumber="134" specimenCount="1">
<collectingMunicipality id="CF31AC8DFFD10426FE6B4C6D44239F20" box="[510,577,1834,1860]" pageId="6" pageNumber="134">Puros</collectingMunicipality>
, rocky terrain opposite
<location id="2A35602CFFD10426FCC54C6E42629F27" LSID="urn:lsid:plazi:treatment:A74387E1FFD60426FF1D4F3347CB9F0C:2A35602CFFD10426FCC54C6E42629F27" box="[848,1024,1833,1859]" country="Namibia" municipality="Puros" name="Hoarusib River" pageId="6" pageNumber="134">Hoarusib River</location>
, 1812
<collectionCode id="49FBAE32FFD10426FBD34C6D42129F20" box="[1094,1136,1834,1860]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="6" pageNumber="134" type="Herbarium">DD</collectionCode>
,
<quantity id="E8129B12FFD10426FBEB4C6D42AA9F20" box="[1150,1224,1834,1860]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="6" pageNumber="134" unit="m" value="292.0">
<elevation id="A4C7D1C4FFD10426FBEB4C6D42AA9F20" box="[1150,1224,1834,1860]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="6" pageNumber="134" unit="m" value="292.0">292 m</elevation>
</quantity>
,
<date id="5B541037FFD10426FB434C6D43C59F20" box="[1238,1447,1833,1860]" pageId="6" pageNumber="134" value="2023-11-03">
<collectingDate id="4B10E9DFFFD10426FB434C6D43C59F20" box="[1238,1447,1833,1860]" pageId="6" pageNumber="134" value="2023-11-03">3 November 2023</collectingDate>
</date>
,
<emphasis id="1D9EEAE5FFD10426FF1D4C0A47509F03" box="[136,306,1869,1895]" italics="true" pageId="6" pageNumber="134">
<collectorName id="821F5321FFD10426FF1D4C0A47639F03" box="[136,257,1869,1895]" pageId="6" pageNumber="134">Swanepoel</collectorName>
637
</emphasis>
(
<collectionCode id="49FBAE32FFD10426FED74C0947FB9F0C" box="[322,409,1870,1896]" country="Namibia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13705" lsid="urn:lsid:biocol.org:col:13705" name="National Botanical Research Institute" pageId="6" pageNumber="134" type="Herbarium">WIND</collectionCode>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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