465 lines
63 KiB
XML
465 lines
63 KiB
XML
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<mods:title id="D418D7AF13F66831610925B99BD48BD3">Agaporomorphus sharynae, a new species of diving beetle (Coleoptera: Dytiscidae: Copelatinae) from Venezuela</mods:title>
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<mods:name id="5F89D9BDA0916DB3772D3230B6739EBC" type="personal">
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<mods:namePart id="55FF7AD364A938E9A90DD3B3FC9C6574">Miller, Kelly B.</mods:namePart>
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<mods:title id="2B12F3F2EC325379B77481A7A16F0A1E">Zootaxa</mods:title>
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<mods:part id="F6795B2C5120BA1AF4F30D3A68AE3B5A">
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<mods:date id="788FE87131D581D64E0CDB40D3B3DF7A">2014</mods:date>
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<mods:number id="2F187941C90E816678B3F7C4C96F823D">3790</mods:number>
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<treatment id="A73D87A79F4ACA35FF5E31A3FE1FAD1A" ID-DOI="http://doi.org/10.5281/zenodo.6126364" ID-GBIF-Taxon="119423327" ID-Zenodo-Dep="6126364" LSID="urn:lsid:plazi:treatment:A73D87A79F4ACA35FF5E31A3FE1FAD1A" httpUri="http://treatment.plazi.org/id/A73D87A79F4ACA35FF5E31A3FE1FAD1A" lastPageId="4" lastPageNumber="181" pageId="1" pageNumber="178">
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<subSubSection id="678E653A9F4ACA30FF5E31A3FEEBAB74" pageId="1" pageNumber="178" type="nomenclature">
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<paragraph id="2F2B36B19F4ACA30FF5E31A3FDADAA90" blockId="1.[151,537,223,285]" box="[151,537,223,248]" pageId="1" pageNumber="178">
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<heading id="746381DD9F4ACA30FF5E31A3FDADAA90" bold="true" box="[151,537,223,248]" fontSize="10" level="2" pageId="1" pageNumber="178" reason="2">
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<emphasis id="1DE0EAA39F4ACA30FF5E31A3FDADAA90" bold="true" box="[151,537,223,248]" pageId="1" pageNumber="178">
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<taxonomicName id="E8944D329F4ACA30FF5E31A3FE66AA90" ID-CoL="8LX8L" box="[151,466,223,248]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
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<emphasis id="1DE0EAA39F4ACA30FF5E31A3FE66AA90" bold="true" box="[151,466,223,248]" italics="true" pageId="1" pageNumber="178">Agaporomorphus sharynae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="06D357D89F4ACA30FE1031A3FDADAA90" box="[473,537,223,248]" pageId="1" pageNumber="178" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2F2B36B19F4ACA30FF5E3078FEEBAB74" blockId="1.[151,537,223,285]" box="[151,351,260,285]" pageId="1" pageNumber="178">
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(
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<figureCitation id="B7AF2A349F4ACA30FF563078FEB7AB74" box="[159,259,260,285]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 16" captionStartId-0="1.[151,249,1986,2008]" captionStartId-1="2.[151,263,1692,1715]" captionTargetBox-0="[531,1056,1066,1945]" captionTargetBox-1="[151,1436,193,1645]" captionTargetId-0="figure@1.[521,1066,1057,1965]" captionTargetId-1="figure@2.[151,1436,193,1654]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Agaporomorphus sharynae sp. n., holotype specimen, dorsal habitus. Scale bar = 1 mm." captionText-1="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri-0="https://zenodo.org/record/225546/files/figure.png" httpUri-1="https://zenodo.org/record/225547/files/figure.png" pageId="1" pageNumber="178">Figs 1–4</figureCitation>
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,
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<figureCitation id="B7AF2A349F4ACA30FEC63078FEE3AB74" box="[271,343,260,284]" captionStart-0="FIGURES 17 – 18" captionStart-1="FIGURE 19" captionStart-2="FIGURES 20 – 23" captionStartId-0="3.[151,264,1600,1623]" captionStartId-1="5.[151,250,1062,1084]" captionStartId-2="6.[151,264,1718,1741]" captionTargetBox-0="[450,1140,601,1531]" captionTargetBox-1="[188,1398,234,1018]" captionTargetBox-2="[152,1435,195,1668]" captionTargetId-0="figure@3.[448,1140,601,1532]" captionTargetId-1="figure@5.[188,1398,229,1018]" captionTargetId-2="figure@6.[152,1437,195,1669]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURES 17 – 18. Agaporomorphus sharynae sp. n. 17) female genitalia, ventral aspect. 18) male right protarsal claws. Scale bars = 0.25 mm." captionText-1="FIGURE 19. Single most parsimonious cladogram of Agaporomorphus species (length = 15, CI = 86, RI = 92). Numbers above hash marks refer to characters numbers. Numbers below hash marks refer to derived character state transformations." captionText-2="FIGURES 20 – 23. Agaporomorphus sharynae sp. n., habitat and type locality." httpUri-0="https://zenodo.org/record/225548/files/figure.png" httpUri-1="https://zenodo.org/record/225549/files/figure.png" httpUri-2="https://zenodo.org/record/225550/files/figure.png" pageId="1" pageNumber="178">17–23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="678E653A9F4ACA30FF5E3030FC5DA855" pageId="1" pageNumber="178" type="materials_examined">
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<paragraph id="2F2B36B19F4ACA30FF5E3030FE99ABE0" blockId="1.[151,1437,332,1005]" pageId="1" pageNumber="178">
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<emphasis id="1DE0EAA39F4ACA30FF5E3030FE89AB0D" bold="true" box="[151,317,332,357]" pageId="1" pageNumber="178">
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<typeStatus id="F02F88139F4ACA30FF5E3030FF66AB0D" box="[151,210,332,357]" pageId="1" pageNumber="178">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="578376219F4ACA30FE853030FE76AB0D" box="[332,450,332,357]" name="Venezuela" pageId="1" pageNumber="178">Venezuela</collectingCountry>
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, Amazonas, Communidad Caño Gato, small tributary of Rio Sipapo,
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<geoCoordinate id="4AA050769F4ACA30FAD63030FA28AB0D" box="[1311,1436,332,357]" direction="north" orientation="latitude" pageId="1" pageNumber="178" precision="1" value="4.9806333">4º58.838'N</geoCoordinate>
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<geoCoordinate id="4AA050769F4ACA30FF5E3013FE9DABE0" box="[151,297,367,392]" direction="west" orientation="longitude" pageId="1" pageNumber="178" precision="1" value="-67.73901">67º44.341'W</geoCoordinate>
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.
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</paragraph>
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<paragraph id="2F2B36B19F4ACA30FF0E30E8FC5DA855" blockId="1.[151,1437,332,1005]" pageId="1" pageNumber="178">
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<emphasis id="1DE0EAA39F4ACA30FF0E30E8FE0AABC5" bold="true" box="[199,446,404,429]" pageId="1" pageNumber="178">Material examined.</emphasis>
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<typeStatus id="F02F88139F4ACA30FE1B30E8FD8AABC5" box="[466,574,404,429]" pageId="1" pageNumber="178" type="holotype">Holotype</typeStatus>
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: ♂ in
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<collectionCode id="4985AE749F4ACA30FD6030E8FD44ABC4" LSID="urn:lsid:biocol.org:col:33834" box="[681,752,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="1" pageNumber="178">MIZA</collectionCode>
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labeled, “
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<collectingCountry id="578376219F4ACA30FCB230E8FB9CABC4" box="[891,1064,404,428]" name="Venezuela" pageId="1" pageNumber="178">VENEZUELA</collectingCountry>
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: Amazonas State
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<geoCoordinate id="4AA050769F4ACA30FADF30E8FA2CABC5" box="[1302,1432,404,429]" direction="north" orientation="latitude" pageId="1" pageNumber="178" precision="1" value="4.9806333">4°58.838'N</geoCoordinate>
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,
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<geoCoordinate id="4AA050769F4ACA30FF5E30CBFE99ABB8" box="[151,301,439,464]" direction="west" orientation="longitude" pageId="1" pageNumber="178" precision="1" value="-67.73901">67°44.341'W</geoCoordinate>
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;
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<quantity id="E86C9B549F4ACA30FEF430CBFECFABA7" box="[317,379,439,464]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="178" unit="m" value="95.0">95 m</quantity>
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Communidad Caño Gato, on Rio Sipapo.
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<date id="5B2A10719F4ACA30FCBE30CBFC50ABB8" box="[887,996,439,464]" pageId="1" pageNumber="178" value="2009-01-16">16.i.2009</date>
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; leg. Short, Miller, Camacho, Joly & Garcia VZ09–0116–
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<date id="5B2A10719F4ACA30FE4D30A0FE18AB9D" box="[388,428,476,501]" pageId="1" pageNumber="178" value="2001-10">01x</date>
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||
: along stream/
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<typeStatus id="F02F88139F4ACA30FD9930A0FD53AB9C" box="[592,743,476,500]" pageId="1" pageNumber="178" type="holotype">HOLOTYPE</typeStatus>
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:
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<taxonomicName id="E8944D329F4ACA30FD3830A0FB18AB9D" authority="K.B. Miller" authorityName="K.B. Miller" box="[753,1196,476,501]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae">
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<emphasis id="1DE0EAA39F4ACA30FD3830A0FB97AB9C" box="[753,1059,476,500]" italics="true" pageId="1" pageNumber="178">Agaporomorphus sharynae</emphasis>
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K.B. Miller
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</taxonomicName>
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[red label with black line border]”.
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<typeStatus id="F02F88139F4ACA30FE98337CFE0AA870" box="[337,446,512,536]" pageId="1" pageNumber="178" type="paratype">Paratypes</typeStatus>
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in
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<collectionCode id="4985AE749F4ACA30FE36337CFDF2A870" LSID="urn:lsid:biocol.org:col:33834" box="[511,582,512,536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="1" pageNumber="178">MIZA</collectionCode>
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and
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<collectionCode id="4985AE749F4ACA30FD53337CFD58A870" box="[666,748,512,536]" pageId="1" pageNumber="178">MSBA</collectionCode>
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, 6♀, labeled same as
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<typeStatus id="F02F88139F4ACA30FBEE3083FB3DA870" box="[1063,1161,511,536]" pageId="1" pageNumber="178" type="holotype">holotype</typeStatus>
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except “
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<typeStatus id="F02F88139F4ACA30FACE337CFA2DA870" box="[1287,1433,512,536]" pageId="1" pageNumber="178" type="paratype">PARATYPE</typeStatus>
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:
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<taxonomicName id="E8944D329F4ACA30FF5E3358FDE4A855" authority="K.B. Miller" authorityName="K.B. Miller" box="[151,592,548,573]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae">
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<emphasis id="1DE0EAA39F4ACA30FF5E3358FE7CA854" box="[151,456,548,572]" italics="true" pageId="1" pageNumber="178">Agaporomorphus sharynae</emphasis>
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K.B. Miller
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</taxonomicName>
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[blue label with black line border].”
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</paragraph>
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</subSubSection>
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<subSubSection id="678E653A9F4ACA30FF0E333BFA8CA984" pageId="1" pageNumber="178" type="diagnosis">
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<paragraph id="2F2B36B19F4ACA30FF0E333BFA8CA984" blockId="1.[151,1437,332,1005]" pageId="1" pageNumber="178">
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<emphasis id="1DE0EAA39F4ACA30FF0E333BFEF5A808" bold="true" box="[199,321,583,608]" pageId="1" pageNumber="178">Diagnosis.</emphasis>
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<taxonomicName id="E8944D329F4ACA30FE993334FD3EA808" box="[336,650,584,608]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
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<emphasis id="1DE0EAA39F4ACA30FE993334FD3EA808" box="[336,650,584,608]" italics="true" pageId="1" pageNumber="178">Agaporomorphus sharynae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="06D357D89F4ACA30FD50333BFD54A808" box="[665,736,583,608]" pageId="1" pageNumber="178" rank="species">
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||
<emphasis id="1DE0EAA39F4ACA30FD50333BFD54A808" bold="true" box="[665,736,583,608]" pageId="1" pageNumber="178">sp. n.</emphasis>
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</taxonomicNameLabel>
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lacks a distinctive bursa copulatrix in the female which distinguishes it from other genera within
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<taxonomicName id="E8944D329F4ACA30FDA73310FD4CA8ED" box="[622,760,620,645]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="subFamily" subFamily="Copelatinae">Copelatinae</taxonomicName>
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||
. The new species is a member of the
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<taxonomicName id="E8944D329F4ACA30FB613310FABFA8EC" box="[1192,1291,620,644]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="knishi">
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<emphasis id="1DE0EAA39F4ACA30FB613310FABFA8EC" box="[1192,1291,620,644]" italics="true" pageId="1" pageNumber="178">A. knishi</emphasis>
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</taxonomicName>
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group
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<emphasis id="1DE0EAA39F4ACA30FA973311FA28A8EC" box="[1374,1436,621,644]" italics="true" pageId="1" pageNumber="178">sensu</emphasis>
|
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<bibRefCitation id="4B054B409F4ACA30FF5E33ECFEC2A8C0" author="Miller" box="[151,374,655,680]" pageId="1" pageNumber="178" refString="Miller, K. B. (2005) Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">K.B. Miller (2005)</bibRefCitation>
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||
. Members of this group have convoluted male genitalia with series of setae on the dorsal surface (
|
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<bibRefCitation id="4B054B409F4ACA30FF3033C8FE73A8A5" author="Miller" box="[249,455,692,717]" pageId="1" pageNumber="178" refString="Miller, K. B. (2005) Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">K.B. Miller, 2005</bibRefCitation>
|
||
). Male specimens of
|
||
<taxonomicName id="E8944D329F4ACA30FD7133C8FCF4A8A4" box="[696,832,692,716]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4ACA30FD7133C8FCF4A8A4" box="[696,832,692,716]" italics="true" pageId="1" pageNumber="178">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F4ACA30FC8E33C8FC3CA8A5" box="[839,904,692,717]" pageId="1" pageNumber="178" rank="species">
|
||
<emphasis id="1DE0EAA39F4ACA30FC8E33C8FC3CA8A5" bold="true" box="[839,904,692,717]" pageId="1" pageNumber="178">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differ from the other members of the
|
||
<taxonomicName id="E8944D329F4ACA30FAF033C8FA28A8A4" box="[1337,1436,692,716]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="knishi">
|
||
<emphasis id="1DE0EAA39F4ACA30FAF033C8FA28A8A4" box="[1337,1436,692,716]" italics="true" pageId="1" pageNumber="178">A. knishi</emphasis>
|
||
</taxonomicName>
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group by the combination of absence of modified antennae, an apparent stridulatory device, and the medio-apical spine on abdominal ventrite V.
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||
<taxonomicName id="E8944D329F4ACA30FE363380FBBDA97D" authority="K.B. Miller, 2005" authorityName="K.B. Miller" authorityYear="2005" box="[511,1033,764,789]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="1DE0EAA39F4ACA30FE363380FC81A97C" box="[511,821,764,788]" italics="true" pageId="1" pageNumber="178">Agaporomorphus silvaticus</emphasis>
|
||
<bibRefCitation id="4B054B409F4ACA30FCF73381FBBDA97D" author="Miller" box="[830,1033,764,789]" pageId="1" pageNumber="178" refString="Miller, K. B. (2005) Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">K.B. Miller, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
also lacks modified antennomeres, but that species has very different male genitalia (
|
||
<figureCitation id="B7AF2A349F4ACA30FD0E3263FCFCA950" box="[711,840,799,824]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="1" pageNumber="178">Figs 11–13</figureCitation>
|
||
). Males of
|
||
<taxonomicName id="E8944D329F4ACA30FC0E325CFBF9A950" box="[967,1101,800,824]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4ACA30FC0E325CFBF9A950" box="[967,1101,800,824]" italics="true" pageId="1" pageNumber="178">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F4ACA30FB9C3263FB21A950" box="[1109,1173,799,824]" pageId="1" pageNumber="178" rank="species">
|
||
<emphasis id="1DE0EAA39F4ACA30FB9C3263FB21A950" bold="true" box="[1109,1173,799,824]" pageId="1" pageNumber="178">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have modified anterior protarsal claws with a large basal expansion and subbasal emargination along the ventral margin with the posterior claw slightly shorter than the anterior (
|
||
<figureCitation id="B7AF2A349F4ACA30FD83321BFD29A9E8" box="[586,669,871,896]" captionStart="FIGURES 17 – 18" captionStartId="3.[151,264,1600,1623]" captionTargetBox="[450,1140,601,1531]" captionTargetId="figure@3.[448,1140,601,1532]" captionTargetPageId="3" captionText="FIGURES 17 – 18. Agaporomorphus sharynae sp. n. 17) female genitalia, ventral aspect. 18) male right protarsal claws. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/225548/files/figure.png" pageId="1" pageNumber="178">Fig. 18</figureCitation>
|
||
). Unique features of the male genitalia of
|
||
<taxonomicName id="E8944D329F4ACA30FBBF3214FB48A9E8" box="[1142,1276,872,896]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4ACA30FBBF3214FB48A9E8" box="[1142,1276,872,896]" italics="true" pageId="1" pageNumber="178">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F4ACA30FACA321BFAF7A9E8" box="[1283,1347,871,896]" pageId="1" pageNumber="178" rank="species">
|
||
<emphasis id="1DE0EAA39F4ACA30FACA321BFAF7A9E8" bold="true" box="[1283,1347,871,896]" pageId="1" pageNumber="178">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
include atypically large apical, transparent lobes and the setae on the median lobe present only as a series of sparse, short setae along the left side of the dorsal surface (
|
||
<figureCitation id="B7AF2A349F4ACA30FD6A32D3FD5DA9A0" box="[675,745,943,968]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="1" pageNumber="178">Fig. 2</figureCitation>
|
||
). The lateral lobes are characterized by a large, membranous subapical lobe and two large regions of setae with those in the medial cluster long and prominent (
|
||
<figureCitation id="B7AF2A349F4ACA30FB2C32A8FA9EA984" box="[1253,1322,980,1005]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="1" pageNumber="178">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7BEB66399F4ACA30FF5E36BEFB24ADB0" box="[151,1168,1986,2008]" httpUri="https://zenodo.org/record/225546/files/figure.png" pageId="1" pageNumber="178" targetBox="[531,1056,1066,1945]" targetPageId="1">
|
||
<paragraph id="2F2B36B19F4ACA30FF5E36BEFB24ADB0" blockId="1.[151,1168,1986,2008]" box="[151,1168,1986,2008]" pageId="1" pageNumber="178">
|
||
<emphasis id="1DE0EAA39F4ACA30FF5E36BEFEA7ADBF" bold="true" box="[151,275,1986,2008]" pageId="1" pageNumber="178">FIGURE 1.</emphasis>
|
||
<taxonomicName id="E8944D329F4ACA30FED136BEFD98ADBF" box="[280,556,1986,2007]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="178" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4ACA30FED136BEFD98ADBF" box="[280,556,1986,2007]" italics="true" pageId="1" pageNumber="178">Agaporomorphus sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F4ACA30FDFB36BEFDDFADBF" box="[562,619,1986,2007]" pageId="1" pageNumber="178" rank="species">
|
||
<emphasis id="1DE0EAA39F4ACA30FDFB36BEFDDFADBF" bold="true" box="[562,619,1986,2007]" pageId="1" pageNumber="178">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype specimen, dorsal habitus. Scale bar = 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7BEB66399F49CA33FF5E37E0FC21AC99" httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="2" pageNumber="179" targetBox="[151,1436,193,1645]" targetPageId="2">
|
||
<paragraph id="2F2B36B19F49CA33FF5E37E0FC21AC99" blockId="2.[151,1437,1692,1777]" pageId="2" pageNumber="179">
|
||
<emphasis id="1DE0EAA39F49CA33FF5E37E0FEFCACDA" bold="true" box="[151,328,1692,1715]" pageId="2" pageNumber="179">FIGURES 2–16.</emphasis>
|
||
<taxonomicName id="E8944D329F49CA33FE8637E1FE4BACDA" box="[335,511,1693,1714]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="179" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1DE0EAA39F49CA33FE8637E1FE4BACDA" box="[335,511,1693,1714]" italics="true" pageId="2" pageNumber="179">Agaporomorphus</emphasis>
|
||
</taxonomicName>
|
||
species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2–4)
|
||
<taxonomicName id="E8944D329F49CA33FBBD37C0FB44ACB9" box="[1140,1264,1724,1745]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="179" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F49CA33FBBD37C0FB44ACB9" box="[1140,1264,1724,1745]" italics="true" pageId="2" pageNumber="179">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F49CA33FB3337C0FA82ACB9" box="[1274,1334,1724,1745]" pageId="2" pageNumber="179" rank="species">
|
||
<emphasis id="1DE0EAA39F49CA33FB3337C0FA82ACB9" bold="true" box="[1274,1334,1724,1745]" pageId="2" pageNumber="179">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, (5–7)
|
||
<taxonomicName id="E8944D329F49CA33FA4F37C0FF52AC98" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="179" phylum="Arthropoda" rank="species" species="colberti">
|
||
<emphasis id="1DE0EAA39F49CA33FA4F37C0FF52AC98" italics="true" pageId="2" pageNumber="179">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
, (8–10)
|
||
<taxonomicName id="E8944D329F49CA33FEF537A7FE5EAC98" box="[316,490,1755,1776]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="179" phylum="Arthropoda" rank="species" species="tambopatensis">
|
||
<emphasis id="1DE0EAA39F49CA33FEF537A7FE5EAC98" box="[316,490,1755,1776]" italics="true" pageId="2" pageNumber="179">A. tambopatensis</emphasis>
|
||
</taxonomicName>
|
||
, (11–13)
|
||
<taxonomicName id="E8944D329F49CA33FD8537A7FD7DAC98" box="[588,713,1755,1776]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="179" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="1DE0EAA39F49CA33FD8537A7FD7DAC98" box="[588,713,1755,1776]" italics="true" pageId="2" pageNumber="179">A. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
, (14–16)
|
||
<taxonomicName id="E8944D329F49CA33FCE237A7FC3BAC98" box="[811,911,1755,1776]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="179" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="1DE0EAA39F49CA33FCE237A7FC3BAC98" box="[811,911,1755,1776]" italics="true" pageId="2" pageNumber="179">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="678E653A9F49CA35FF0E3661FBD2A855" lastPageId="4" lastPageNumber="181" pageId="2" pageNumber="179" type="description">
|
||
<paragraph id="2F2B36B19F49CA33FF0E3661FC46ADCA" blockId="2.[151,1436,1821,1954]" pageId="2" pageNumber="179">
|
||
<emphasis id="1DE0EAA39F49CA33FF0E3661FD00AD5E" bold="true" box="[199,692,1821,1846]" pageId="2" pageNumber="179">Description. Measurements and habitus</emphasis>
|
||
. TL =
|
||
<quantity id="E86C9B549F49CA33FCDB3661FC2BAD5D" box="[786,927,1821,1846]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="2" pageNumber="179" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3–3.5 mm</quantity>
|
||
(
|
||
<typeStatus id="F02F88139F49CA33FC7A3661FBA3AD5E" box="[947,1047,1821,1846]" pageId="2" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
=
|
||
<quantity id="E86C9B549F49CA33FB893661FB2BAD5E" box="[1088,1183,1821,1846]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="179" unit="mm" value="3.5">3.5 mm</quantity>
|
||
), GW =
|
||
<quantity id="E86C9B549F49CA33FAD93661FA28AD5D" box="[1296,1436,1821,1846]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="2" pageNumber="179" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6–1.7 mm</quantity>
|
||
(
|
||
<typeStatus id="F02F88139F49CA33FF57363DFEB6AD32" box="[158,258,1857,1882]" pageId="2" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
=
|
||
<quantity id="E86C9B549F49CA33FEE9363EFECEAD32" box="[288,378,1858,1882]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="179" unit="mm" value="1.7">1.7 mm</quantity>
|
||
), HW =
|
||
<quantity id="E86C9B549F49CA33FE13363EFDD5AD31" box="[474,609,1858,1882]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="179" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
|
||
(
|
||
<typeStatus id="F02F88139F49CA33FDB9363DFD60AD32" box="[624,724,1857,1882]" pageId="2" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
=
|
||
<quantity id="E86C9B549F49CA33FD38363EFCFFAD32" box="[753,843,1858,1882]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="179" unit="mm" value="0.8">0.8 mm</quantity>
|
||
), EW =
|
||
<quantity id="E86C9B549F49CA33FC61363DFB9BAD31" box="[936,1071,1857,1882]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="179" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
|
||
(
|
||
<typeStatus id="F02F88139F49CA33FBF4363DFB15AD32" box="[1085,1185,1857,1882]" pageId="2" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
=
|
||
<quantity id="E86C9B549F49CA33FB76363DFAADAD32" box="[1215,1305,1857,1882]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="179" unit="mm" value="0.5">0.5 mm</quantity>
|
||
), TL/GW = 2.0–2.1 (
|
||
<typeStatus id="F02F88139F49CA33FEC83619FED1AD16" box="[257,357,1893,1918]" pageId="2" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
= 2.1), HW/EW = 1.5–1.7 (
|
||
<typeStatus id="F02F88139F49CA33FD043619FC85AD16" box="[717,817,1893,1918]" pageId="2" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
= 1.6). Body shape elongate oval, lateral margins continuous between pronotum and elytron; body dorsoventrally compressed.
|
||
</paragraph>
|
||
<paragraph id="2F2B36B19F48CA32FF0E31EBFAAEAB74" blockId="3.[151,1437,151,536]" pageId="3" pageNumber="180">
|
||
<emphasis id="1DE0EAA39F48CA32FF0E31EBFEFCAAD8" bold="true" box="[199,328,151,176]" pageId="3" pageNumber="180">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="B7AF2A349F48CA32FE9431EBFE10AAD8" box="[349,420,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,249,1986,2008]" captionTargetBox="[531,1056,1066,1945]" captionTargetId="figure@1.[521,1066,1057,1965]" captionTargetPageId="1" captionText="FIGURE 1. Agaporomorphus sharynae sp. n., holotype specimen, dorsal habitus. Scale bar = 1 mm." httpUri="https://zenodo.org/record/225546/files/figure.png" pageId="3" pageNumber="180">Fig. 1</figureCitation>
|
||
). Head yellow-brown to brown, lighter anteriorly and darker around eyes. Pronotum yellow-brown to brown medially, yellow laterally, darker narrowly along anterior and posterior margins. Elytra with most of surface yellow-brown to brown; with yellow in broad, basal, diffusely-demarcated band; lateral margins diffusely yellow. Appendages and ventral surfaces yellow, darker on metatibia and metatarsus.
|
||
</paragraph>
|
||
<paragraph id="2F2B36B19F48CA32FF0E305BFEDCA870" blockId="3.[151,1437,151,536]" pageId="3" pageNumber="180">
|
||
<emphasis id="1DE0EAA39F48CA32FF0E305BFE50AB28" bold="true" box="[199,484,295,320]" pageId="3" pageNumber="180">Sculpture and structure</emphasis>
|
||
. Pronotum with microsculpture consisting of fine cells, with few very fine punctures interspersed; lateral pronotal bead fine, absent in anterior one-fifth. Elytron covered with extremely fine microstriae, otherwise smooth, impunctate or with few punctures in indistinct longitudinal lines. Prosternum medially strongly carinate, carina extending onto prosternal process; prosternal process moderately short, medially with a distinct, sharp longitudinal carina extending to apex, laterally with strongly beaded margins, apex pointed. Metacoxae smooth, impunctate; metacoxal lines closely approximated. Metafemur moderately broad, length about 3 × greatest width.
|
||
</paragraph>
|
||
<caption id="7BEB66399F48CA32FF5E373CFE89AC1D" httpUri="https://zenodo.org/record/225548/files/figure.png" pageId="3" pageNumber="180" targetBox="[450,1140,601,1531]" targetPageId="3">
|
||
<paragraph id="2F2B36B19F48CA32FF5E373CFE89AC1D" blockId="3.[151,1436,1600,1653]" pageId="3" pageNumber="180">
|
||
<emphasis id="1DE0EAA39F48CA32FF5E373CFEE0AC3D" bold="true" box="[151,340,1600,1623]" pageId="3" pageNumber="180">FIGURES 17–18.</emphasis>
|
||
<taxonomicName id="E8944D329F48CA32FE92373DFDDBAC3E" box="[347,623,1601,1622]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="180" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F48CA32FE92373DFDDBAC3E" box="[347,623,1601,1622]" italics="true" pageId="3" pageNumber="180">Agaporomorphus sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F48CA32FDBC373DFD1BAC3E" box="[629,687,1601,1622]" pageId="3" pageNumber="180" rank="species">
|
||
<emphasis id="1DE0EAA39F48CA32FDBC373DFD1BAC3E" bold="true" box="[629,687,1601,1622]" pageId="3" pageNumber="180">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
17) female genitalia, ventral aspect. 18) male right protarsal claws. Scale bars = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2F2B36B19F48CA32FF0E37E0FC89ADE5" blockId="3.[151,1437,1692,2006]" pageId="3" pageNumber="180">
|
||
<emphasis id="1DE0EAA39F48CA32FF0E37E0FEC6ACDD" bold="true" box="[199,370,1692,1717]" pageId="3" pageNumber="180">Male genitalia</emphasis>
|
||
. Median lobe in lateral aspect (
|
||
<figureCitation id="B7AF2A349F48CA32FD1037E0FCA9ACDD" box="[729,797,1692,1717]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="3" pageNumber="180">Fig. 2</figureCitation>
|
||
) moderately robust, with broad expansion ventrad, apex elongate triangular, with extremely large, distinct posteriorly-directed round, transparent lobe; left side of median lobe with elongate, linear series of fine, short setae, no setae on right side; in ventral aspect (
|
||
<figureCitation id="B7AF2A349F48CA32FB643799FB44AC95" box="[1197,1264,1765,1790]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="3" pageNumber="180">Fig. 3</figureCitation>
|
||
) robust, broad, with complicated asymmetrical folding and structures, apical lobes bilaterally symmetrical and prominent. Lateral lobes (
|
||
<figureCitation id="B7AF2A349F48CA32FF2D3651FE9DAD2D" box="[228,297,1837,1862]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="3" pageNumber="180">Fig. 4</figureCitation>
|
||
) bilaterally symmetrical, in lateral aspect long with elongate basal, curved process, with prominent cluster of long setae medially; apical half with large membranous lobe bearing distinct cluster of long setae (
|
||
<figureCitation id="B7AF2A349F48CA32FAA3362CFF1CADE5" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="3" pageNumber="180">Figs 4</figureCitation>
|
||
); apex very slender with few elongate apical setae (
|
||
<figureCitation id="B7AF2A349F48CA32FD233609FC84ADE5" box="[746,816,1909,1934]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="3" pageNumber="180">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2F2B36B19F48CA35FF0E36E4FC2BAABD" blockId="3.[151,1437,1692,2006]" lastBlockId="4.[151,1437,151,1005]" lastPageId="4" lastPageNumber="181" pageId="3" pageNumber="180">
|
||
<emphasis id="1DE0EAA39F48CA32FF0E36E4FE3FADD9" bold="true" box="[199,395,1944,1969]" pageId="3" pageNumber="180">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="B7AF2A349F48CA32FE5336E4FE58ADD9" box="[410,492,1944,1969]" captionStart="FIGURES 17 – 18" captionStartId="3.[151,264,1600,1623]" captionTargetBox="[450,1140,601,1531]" captionTargetId="figure@3.[448,1140,601,1532]" captionTargetPageId="3" captionText="FIGURES 17 – 18. Agaporomorphus sharynae sp. n. 17) female genitalia, ventral aspect. 18) male right protarsal claws. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/225548/files/figure.png" pageId="3" pageNumber="180">Fig. 17</figureCitation>
|
||
). Gonocoxa elongate and slender, apically slightly expanded and rounded with small cluster of fine, elongate sensory setae; laterotergite slender and apically curved. Bursa absent or extremely small; spermathecal duct slender and long, slightly expanded proximate to spermatheca which is elongate, expanded and coiled; fertilization duct short, very slender; vagina elongate, slender.
|
||
</paragraph>
|
||
<paragraph id="2F2B36B19F4FCA35FF0E31A3FBD2A855" blockId="4.[151,1437,151,1005]" pageId="4" pageNumber="181">
|
||
<emphasis id="1DE0EAA39F4FCA35FF0E31A3FDA1AA90" bold="true" box="[199,533,223,248]" pageId="4" pageNumber="181">Sexually dimorphic features</emphasis>
|
||
. Male protarsal claws modified, anterior claw with basal expansion and subbasal emargination on ventral margin (
|
||
<figureCitation id="B7AF2A349F4FCA35FDC73078FDD7AB74" box="[526,611,260,285]" captionStart="FIGURES 17 – 18" captionStartId="3.[151,264,1600,1623]" captionTargetBox="[450,1140,601,1531]" captionTargetId="figure@3.[448,1140,601,1532]" captionTargetPageId="3" captionText="FIGURES 17 – 18. Agaporomorphus sharynae sp. n. 17) female genitalia, ventral aspect. 18) male right protarsal claws. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/225548/files/figure.png" pageId="4" pageNumber="181">Fig. 18</figureCitation>
|
||
); male pro- and mesotarsal claws about 3/4 length of tarsomere V; male without apical lobe on mesotarsomere V; male protarsomeres I and II broadened, protarsomere I with two large adhesive setae, protarsomere II without adhesive setae; male mesotarsomeres I and II slightly broadened, mesotarsomere I with one large, medial adhesive seta and two large, apical adhesive setae, mesotarsomere II with two smaller, apical adhesive setae; female pro- and mesotarsomeres and claws unmodified. Male and female without triangular, posteriorly-directed prominence medially along posterior margin of visible abdominal ventrite V. Male with broad, elongate depression medially on abdominal ventrite VI; female without depression. Male and female without distinct parallel series of rugulosities on each side of midline on abdominal ventrite III. Male and female antennomeres V and VI not modified. Male and female metafemur unmodified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="678E653A9F4FCA35FF0E333BFD27A8CF" pageId="4" pageNumber="181" type="etymology">
|
||
<paragraph id="2F2B36B19F4FCA35FF0E333BFD27A8CF" blockId="4.[151,1437,151,1005]" pageId="4" pageNumber="181">
|
||
<emphasis id="1DE0EAA39F4FCA35FF0E333BFEFAA808" bold="true" box="[199,334,583,608]" pageId="4" pageNumber="181">Etymology.</emphasis>
|
||
This species is named to honor my valued friend, Sharyn Davidson and all her efforts as a volunteer in the Division of Arthropods at the Museum of Southwestern Biology (University of New
|
||
<collectingCountry id="578376219F4FCA35FACD3310FAEBA8ED" box="[1284,1375,620,645]" name="Mexico" pageId="4" pageNumber="181">Mexico</collectingCountry>
|
||
). The specific epithet is a noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="678E653A9F4FCA35FF0E33C8FE1FAD1A" pageId="4" pageNumber="181" type="distribution">
|
||
<paragraph id="2F2B36B19F4FCA35FF0E33C8FB9AA8A4" blockId="4.[151,1437,151,1005]" box="[199,1070,692,717]" pageId="4" pageNumber="181">
|
||
<emphasis id="1DE0EAA39F4FCA35FF0E33C8FED4A8A5" bold="true" box="[199,352,692,717]" pageId="4" pageNumber="181">Distribution.</emphasis>
|
||
This species is known only from the
|
||
<typeStatus id="F02F88139F4FCA35FCCD33C9FC80A8A5" box="[772,820,693,717]" pageId="4" pageNumber="181">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="B7AF2A349F4FCA35FC5733C8FB96A8A4" box="[926,1058,692,717]" captionStart="FIGURES 20 – 23" captionStartId="6.[151,264,1718,1741]" captionTargetBox="[152,1435,195,1668]" captionTargetId="figure@6.[152,1437,195,1669]" captionTargetPageId="6" captionText="FIGURES 20 – 23. Agaporomorphus sharynae sp. n., habitat and type locality." httpUri="https://zenodo.org/record/225550/files/figure.png" pageId="4" pageNumber="181">Figs 20–23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2F2B36B19F4FCA35FF0E33ABFB30A984" blockId="4.[151,1437,151,1005]" pageId="4" pageNumber="181">
|
||
<emphasis id="1DE0EAA39F4FCA35FF0E33ABFEF6A898" bold="true" box="[199,322,727,752]" pageId="4" pageNumber="181">Phylogeny</emphasis>
|
||
.
|
||
<emphasis id="1DE0EAA39F4FCA35FE9933ABFE74A898" bold="true" box="[336,448,727,752]" pageId="4" pageNumber="181">Methods.</emphasis>
|
||
Phylogenetic methods are the same as in K.B.
|
||
<bibRefCitation id="4B054B409F4FCA35FC1133ABFBC2A898" author="Miller" box="[984,1142,727,752]" pageId="4" pageNumber="181" refString="Miller, K. B. (2001 a) On the genus Agaporomorphus Zimmermann, 1921 (Coleoptera: Dytiscidae: Copelatinae). Annals of the Entomological Society of America, 94, 520 - 529. http: // dx. doi. org / 10.1603 / 0013 - 8746 (2001) 094 [0520: ROTGAZ] 2.0. CO; 2" type="journal article" year="2001" yearSuffix="a">Miller (2001a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4B054B409F4FCA35FB4B33ABFB08A898" author="Miller" box="[1154,1212,727,752]" pageId="4" pageNumber="181" refString="Miller, K. B. (2005) Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">2005</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="4B054B409F4FCA35FB3533A4FEF8A97C" author="Miller" pageId="4" pageNumber="181" refString="Miller, K. B. & Wheeler, Q. D. (2008) A new species of Agaporomorphus Zimmermann from Venezuela, and a review of the A. knischi species group (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 1859, 63 - 68." type="journal article" year="2008">K.B. Miller & Wheeler (2008)</bibRefCitation>
|
||
. The new species was added to the matrix (
|
||
<tableCitation id="6216030A9F4FCA35FC893380FC20A97C" box="[832,916,764,789]" captionStart="TABLE 1" captionStartId="4.[151,239,1088,1111]" captionText="TABLE 1. Character matrix for parsimony analysis. Characters 1 – 11 are described by K. B. Miller (2001 a, 2005) and K. B. Miller & Wheeler (2008). Character 12 is new to this analysis. All characters are treated as non-additive and equal in weight." pageId="4" pageNumber="181">Table 1</tableCitation>
|
||
) described by K.B.
|
||
<bibRefCitation id="4B054B409F4FCA35FBB73380FAA9A97C" author="Miller" box="[1150,1309,764,789]" pageId="4" pageNumber="181" refString="Miller, K. B. (2001 a) On the genus Agaporomorphus Zimmermann, 1921 (Coleoptera: Dytiscidae: Copelatinae). Annals of the Entomological Society of America, 94, 520 - 529. http: // dx. doi. org / 10.1603 / 0013 - 8746 (2001) 094 [0520: ROTGAZ] 2.0. CO; 2" type="journal article" year="2001" yearSuffix="a">Miller (2001a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4B054B409F4FCA35FAE33380FAD0A97D" author="Miller" box="[1322,1380,764,789]" pageId="4" pageNumber="181" refString="Miller, K. B. (2005) Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">2005</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="4B054B409F4FCA35FF5E325CFE46A950" author="Miller" box="[151,498,799,824]" pageId="4" pageNumber="181" refString="Miller, K. B. & Wheeler, Q. D. (2008) A new species of Agaporomorphus Zimmermann from Venezuela, and a review of the A. knischi species group (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 1859, 63 - 68." type="journal article" year="2008">K.B. Miller & Wheeler (2008)</bibRefCitation>
|
||
. Additionally, one new character (hyaline lobes present or absent at the apex of the median lobe) was included which helps resolve placement of
|
||
<taxonomicName id="E8944D329F4FCA35FC993238FC62A934" box="[848,982,836,860]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4FCA35FC993238FC62A934" box="[848,982,836,860]" italics="true" pageId="4" pageNumber="181">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1DE0EAA39F4FCA35FC163238FBABA935" bold="true" box="[991,1055,836,861]" pageId="4" pageNumber="181">
|
||
<taxonomicNameLabel id="06D357D89F4FCA35FC163238FBABA935" box="[991,1055,836,861]" pageId="4" pageNumber="181" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="B7AF2A349F4FCA35FBE63238FB22A934" box="[1071,1174,836,861]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="4" pageNumber="181">Figs 2–3</figureCitation>
|
||
). The male genitalia of the other four members of the
|
||
<taxonomicName id="E8944D329F4FCA35FE3F3214FD8CA9E8" box="[502,568,872,896]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="knishi">
|
||
<emphasis id="1DE0EAA39F4FCA35FE3F3214FD8CA9E8" box="[502,568,872,896]" italics="true" pageId="4" pageNumber="181">knishi</emphasis>
|
||
</taxonomicName>
|
||
group are given for comparison (
|
||
<figureCitation id="B7AF2A349F4FCA35FC72321BFB86A9E8" box="[955,1074,871,896]" captionStart="FIGURES 2 – 16" captionStartId="2.[151,263,1692,1715]" captionTargetBox="[151,1436,193,1645]" captionTargetId="figure@2.[151,1436,193,1654]" captionTargetPageId="2" captionText="FIGURES 2 – 16. Agaporomorphus species, male genitalia. (2, 5, 8, 11, 14) median lobe, right lateral aspect, (3, 6, 9, 12, 15) median lobe, ventral aspect, (4, 7, 10, 13, 16) right lateral lobe, right lateral aspect of: (2 – 4) A. sharynae sp. n., (5 – 7) A. colberti, (8 – 10) A. tambopatensis, (11 – 13) A. silvaticus, (14 – 16) A. knischi." httpUri="https://zenodo.org/record/225547/files/figure.png" pageId="4" pageNumber="181">Figs 5–16</figureCitation>
|
||
). Each member of the
|
||
<taxonomicName id="E8944D329F4FCA35FAF13214FA28A9E8" box="[1336,1436,872,896]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="knishi">
|
||
<emphasis id="1DE0EAA39F4FCA35FAF13214FA28A9E8" box="[1336,1436,872,896]" italics="true" pageId="4" pageNumber="181">A. knishi</emphasis>
|
||
</taxonomicName>
|
||
group and
|
||
<taxonomicName id="E8944D329F4FCA35FEDB32F0FE2DA9CC" box="[274,409,908,932]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4FCA35FEDB32F0FE2DA9CC" box="[274,409,908,932]" italics="true" pageId="4" pageNumber="181">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F4FCA35FE6832F0FE57A9CD" box="[417,483,908,933]" pageId="4" pageNumber="181" rank="species">
|
||
<emphasis id="1DE0EAA39F4FCA35FE6832F0FE57A9CD" bold="true" box="[417,483,908,933]" pageId="4" pageNumber="181">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have symmetrical, translucent lobes at the apex of the male median lobe. Although the median lobe in the
|
||
<taxonomicName id="E8944D329F4FCA35FE5F32CCFE4CA9A0" box="[406,504,944,968]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="knishi">
|
||
<emphasis id="1DE0EAA39F4FCA35FE5F32CCFE4CA9A0" box="[406,504,944,968]" italics="true" pageId="4" pageNumber="181">A. knishi</emphasis>
|
||
</taxonomicName>
|
||
group in general is conspicuously asymmetrical, the apex, including these lobes, is symmetrical in all
|
||
<taxonomicName id="E8944D329F4FCA35FEA132A8FD9FA984" box="[360,555,980,1004]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1DE0EAA39F4FCA35FEA132A8FD9FA984" box="[360,555,980,1004]" italics="true" pageId="4" pageNumber="181">Agaporomorphus</emphasis>
|
||
</taxonomicName>
|
||
species. The character matrix is presented in
|
||
<tableCitation id="6216030A9F4FCA35FBE332A8FBCAA984" box="[1066,1150,980,1005]" captionStart="TABLE 1" captionStartId="4.[151,239,1088,1111]" captionText="TABLE 1. Character matrix for parsimony analysis. Characters 1 – 11 are described by K. B. Miller (2001 a, 2005) and K. B. Miller & Wheeler (2008). Character 12 is new to this analysis. All characters are treated as non-additive and equal in weight." pageId="4" pageNumber="181">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="7BEB66399F4FCA35FF5E353CFEB6AEFF" pageId="4" pageNumber="181">
|
||
<paragraph id="2F2B36B19F4FCA35FF5E353CFEB6AEFF" blockId="4.[151,1436,1088,1175]" pageId="4" pageNumber="181">
|
||
<emphasis id="1DE0EAA39F4FCA35FF5E353CFEB8AE3F" bold="true" box="[151,268,1088,1111]" pageId="4" pageNumber="181">TABLE 1.</emphasis>
|
||
Character matrix for parsimony analysis. Characters 1–11 are described by K.B. Miller (2001a, 2005) and K.B. Miller & Wheeler (2008). Character 12 is new to this analysis. All characters are treated as non-additive and equal in weight.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2F2B36B19F4FCA35FE2035CEFE1FAD1A" blockId="4.[159,641,1202,1668]" lastBlockId="4.[151,1436,1737,1906]" pageId="4" pageNumber="181">
|
||
0 0 0 0 0 0 0 0 0 111 123456789012 Outgroup 0 0 0 0 0 0 0 0 0 0 0 0
|
||
<taxonomicName id="E8944D329F4FCA35FF563458FEBEAF51" box="[159,266,1316,1337]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="colberti">
|
||
<emphasis id="1DE0EAA39F4FCA35FF563458FEBEAF51" box="[159,266,1316,1337]" italics="true" pageId="4" pageNumber="181">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
0 0 0 0 0 0
|
||
<date id="5B2A10719F4FCA35FDF83458FDCFAF52" box="[561,635,1316,1338]" pageId="4" pageNumber="181" value="11-11-11">111111</date>
|
||
<taxonomicName id="E8944D329F4FCA35FF563432FEEFAF0A" box="[159,347,1357,1379]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="dolichodactylus">
|
||
<emphasis id="1DE0EAA39F4FCA35FF563432FEEFAF0A" box="[159,347,1357,1379]" italics="true" pageId="4" pageNumber="181">A. dolichodactylus</emphasis>
|
||
</taxonomicName>
|
||
211110000000
|
||
<taxonomicName id="E8944D329F4FCA35FF56340BFEE7AFE4" box="[159,339,1399,1420]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="grandisinuatus">
|
||
<emphasis id="1DE0EAA39F4FCA35FF56340BFEE7AFE4" box="[159,339,1399,1420]" italics="true" pageId="4" pageNumber="181">A. grandisinuatus</emphasis>
|
||
</taxonomicName>
|
||
101000000000
|
||
<taxonomicName id="E8944D329F4FCA35FF5634DCFEB7AFDD" box="[159,259,1440,1461]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="1DE0EAA39F4FCA35FF5634DCFEB7AFDD" box="[159,259,1440,1461]" italics="true" pageId="4" pageNumber="181">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
0 0 0 0 0 1111111
|
||
<taxonomicName id="E8944D329F4FCA35FF5634B6FE93AFB6" box="[159,295,1481,1503]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="mecolobus">
|
||
<emphasis id="1DE0EAA39F4FCA35FF5634B6FE93AFB6" box="[159,295,1481,1503]" italics="true" pageId="4" pageNumber="181">A. mecolobus</emphasis>
|
||
</taxonomicName>
|
||
211110000000
|
||
<taxonomicName id="E8944D329F4FCA35FF56348FFEB8AC60" box="[159,268,1523,1544]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="pereirai">
|
||
<emphasis id="1DE0EAA39F4FCA35FF56348FFEB8AC60" box="[159,268,1523,1544]" italics="true" pageId="4" pageNumber="181">A. pereirai</emphasis>
|
||
</taxonomicName>
|
||
0 0 0 0 0 1000000
|
||
<taxonomicName id="E8944D329F4FCA35FF563760FEACAC59" box="[159,280,1564,1585]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4FCA35FF563760FEACAC59" box="[159,280,1564,1585]" italics="true" pageId="4" pageNumber="181">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F4FCA35FED73760FEE3AC59" box="[286,343,1564,1585]" pageId="4" pageNumber="181" rank="species">
|
||
<emphasis id="1DE0EAA39F4FCA35FED73760FEE3AC59" bold="true" box="[286,343,1564,1585]" pageId="4" pageNumber="181">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
0 0 0 0 0 1100001
|
||
<taxonomicName id="E8944D329F4FCA35FF56373AFEAFAC32" box="[159,283,1605,1627]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="1DE0EAA39F4FCA35FF56373AFEAFAC32" box="[159,283,1605,1627]" italics="true" pageId="4" pageNumber="181">A. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
0 0 0 0 0 0 100001
|
||
<taxonomicName id="E8944D329F4FCA35FF563713FEF9ACEC" box="[159,333,1647,1668]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="tambopatensis">
|
||
<emphasis id="1DE0EAA39F4FCA35FF563713FEF9ACEC" box="[159,333,1647,1668]" italics="true" pageId="4" pageNumber="181">A. tambopatensis</emphasis>
|
||
</taxonomicName>
|
||
0 0 0 0 0 0
|
||
<date id="5B2A10719F4FCA35FDFB3712FDC9ACEC" box="[562,637,1646,1668]" pageId="4" pageNumber="181" value="01-11-11">111101</date>
|
||
<emphasis id="1DE0EAA39F4FCA35FF0E37B5FDBCAC8A" bold="true" box="[199,520,1737,1762]" pageId="4" pageNumber="181">Phylogenetic relationships.</emphasis>
|
||
The phylogenetic analysis resulted in a single parsimony tree (length = 15, CI = 86, RI = 92). The tree is the same as previous analyses (
|
||
<bibRefCitation id="4B054B409F4FCA35FCD13792FC45AD6E" author="Miller" box="[792,1009,1773,1798]" pageId="4" pageNumber="181" refString="Miller, K. B. (2001 a) On the genus Agaporomorphus Zimmermann, 1921 (Coleoptera: Dytiscidae: Copelatinae). Annals of the Entomological Society of America, 94, 520 - 529. http: // dx. doi. org / 10.1603 / 0013 - 8746 (2001) 094 [0520: ROTGAZ] 2.0. CO; 2" type="journal article" year="2001" yearSuffix="a">K.B. Miller, 2001a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4B054B409F4FCA35FC343791FB8DAD6E" author="Miller" box="[1021,1081,1773,1798]" pageId="4" pageNumber="181" refString="Miller, K. B. (2005) Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4B054B409F4FCA35FB8C3792FA21AD6E" author="Miller" box="[1093,1429,1773,1798]" pageId="4" pageNumber="181" refString="Miller, K. B. & Wheeler, Q. D. (2008) A new species of Agaporomorphus Zimmermann from Venezuela, and a review of the A. knischi species group (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 1859, 63 - 68." type="journal article" year="2008">K.B. Miller & Wheeler, 2008</bibRefCitation>
|
||
) with the addition of
|
||
<taxonomicName id="E8944D329F4FCA35FEBE366EFE49AD41" box="[375,509,1809,1833]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="sharynae" status="sp. nov.">
|
||
<emphasis id="1DE0EAA39F4FCA35FEBE366EFE49AD41" box="[375,509,1809,1833]" italics="true" pageId="4" pageNumber="181">A. sharynae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="06D357D89F4FCA35FDCA366DFDF6AD42" box="[515,578,1809,1834]" pageId="4" pageNumber="181" rank="species">
|
||
<emphasis id="1DE0EAA39F4FCA35FDCA366DFDF6AD42" bold="true" box="[515,578,1809,1834]" pageId="4" pageNumber="181">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in an unresolved position as part of the
|
||
<taxonomicName id="E8944D329F4FCA35FC34366EFBEAAD41" box="[1021,1118,1809,1833]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="knishi">
|
||
<emphasis id="1DE0EAA39F4FCA35FC34366EFBEAAD41" box="[1021,1118,1809,1833]" italics="true" pageId="4" pageNumber="181">A. knishi</emphasis>
|
||
</taxonomicName>
|
||
group with
|
||
<taxonomicName id="E8944D329F4FCA35FB2C366EFAD9AD41" box="[1253,1389,1809,1833]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="1DE0EAA39F4FCA35FB2C366EFAD9AD41" box="[1253,1389,1809,1833]" italics="true" pageId="4" pageNumber="181">A. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
and a clade containing
|
||
<taxonomicName id="E8944D329F4FCA35FEAE364AFD43AD26" authority="K.B. Miller, 2005" authorityName="K.B. Miller" authorityYear="2005" box="[359,759,1845,1870]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="tambopatensis">
|
||
<emphasis id="1DE0EAA39F4FCA35FEAE364AFD9CAD25" box="[359,552,1845,1869]" italics="true" pageId="4" pageNumber="181">A. tambopatensis</emphasis>
|
||
<bibRefCitation id="4B054B409F4FCA35FDE7364AFD43AD26" author="Miller" box="[558,759,1845,1870]" pageId="4" pageNumber="181" refString="Miller, K. B. (2005) Two new species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">K.B. Miller, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E8944D329F4FCA35FCC8364AFBFCAD26" authority="Zimmermann, 1921" authorityName="Zimmermann" authorityYear="1921" box="[769,1096,1845,1870]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="knishi">
|
||
<emphasis id="1DE0EAA39F4FCA35FCC8364AFCD6AD25" box="[769,866,1845,1869]" italics="true" pageId="4" pageNumber="181">A. knishi</emphasis>
|
||
<bibRefCitation id="4B054B409F4FCA35FCA03649FBFCAD26" author="Zimmermann" box="[873,1096,1845,1870]" pageId="4" pageNumber="181" refString="Zimmermann, A. (1921) Beitrage zur Kenntnis der sudamerikanischen Schwimmkaferfauna nebst 41 Neubeschreibungen. Archiv fur Naturgeschichte, 87 A (3), 181 - 206." type="journal article" year="1921">Zimmermann, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="E8944D329F4FCA35FB4A364AFE8FAD1A" authority="K.B. Miller & Wheeler, 2008" authorityName="K.B. Miller & Wheeler" authorityYear="2008" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="181" phylum="Arthropoda" rank="species" species="colberti">
|
||
<emphasis id="1DE0EAA39F4FCA35FB4A364AFB4CAD25" box="[1155,1272,1845,1869]" italics="true" pageId="4" pageNumber="181">A. colberti</emphasis>
|
||
<bibRefCitation id="4B054B409F4FCA35FB37364AFE8FAD1A" author="Miller" pageId="4" pageNumber="181" refString="Miller, K. B. & Wheeler, Q. D. (2008) A new species of Agaporomorphus Zimmermann from Venezuela, and a review of the A. knischi species group (Coleoptera: Dytiscidae: Copelatinae). Zootaxa, 1859, 63 - 68." type="journal article" year="2008">K.B. Miller & Wheeler, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="B7AF2A349F4FCA35FE833625FE29AD1A" box="[330,413,1881,1906]" captionStart="FIGURE 19" captionStartId="5.[151,250,1062,1084]" captionTargetBox="[188,1398,234,1018]" captionTargetId="figure@5.[188,1398,229,1018]" captionTargetPageId="5" captionText="FIGURE 19. Single most parsimonious cladogram of Agaporomorphus species (length = 15, CI = 86, RI = 92). Numbers above hash marks refer to characters numbers. Numbers below hash marks refer to derived character state transformations." httpUri="https://zenodo.org/record/225549/files/figure.png" pageId="4" pageNumber="181">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |