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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.983.52630" ID-GBIF-Dataset="d3b857f7-25c1-4b21-9d39-4510ab675fca" ID-PMC="PMC7655787" ID-Pensoft-Pub="1313-2970-983-1" ID-Pensoft-UUID="C890C7FD4B2D57A8B1A062305ED42D53" ID-PubMed="33244289" ID-ZooBank="7DED4CE2934C4539945F758930C927F9" ModsDocID="1313-2970-983-1" checkinTime="1604475085656" checkinUser="pensoft" docAuthor="Smith, Sarah M., Beaver, Roger A. &amp; Cognato, Anthony I." docDate="2020" docId="A6F7E14CFEF41E5E439973F46F87A786" docLanguage="en" docName="ZooKeys 983: 1-442" docOrigin="ZooKeys 983" docSource="http://dx.doi.org/10.3897/zookeys.983.52630" docTitle="Coptodryas amydra Smith &amp; Beaver &amp; Cognato 2020, sp. nov." docType="treatment" docUuid="AFAB64B8-8AF9-4595-803E-B146C73EEC5A" docUuidSource="ZooBank" docVersion="4" id="C890C7FD4B2D57A8B1A062305ED42D53" lastPageNumber="1" masterDocId="C890C7FD4B2D57A8B1A062305ED42D53" masterDocTitle="A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China" masterLastPageNumber="442" masterPageNumber="1" pageNumber="1" updateTime="1668169715782" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Sarah M.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5173-3736</mods:nameIdentifier>
<mods:nameIdentifier type="email">camptocerus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beaver, Roger A.</mods:namePart>
<mods:affiliation>161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cognato, Anthony I.</mods:namePart>
<mods:affiliation>Department of Entomology, Michigan State University, 288 Farm Lane, East Lansing, Michigan 48824, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>983</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>442</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.983.52630</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.983.52630</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-983-1</mods:identifier>
<mods:identifier type="ZooBank">7DED4CE2934C4539945F758930C927F9</mods:identifier>
<mods:identifier type="Pensoft-UUID">C890C7FD4B2D57A8B1A062305ED42D53</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168754590" LSID="urn:lsid:zoobank.org:act:AFAB64B8-8AF9-4595-803E-B146C73EEC5A" httpUri="http://treatment.plazi.org/id/A6F7E14CFEF41E5E439973F46F87A786" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/AFAB64B8-8AF9-4595-803E-B146C73EEC5A" authority="Smith &amp; Beaver &amp; Cognato, 2020" authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Coptodryas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodryas amydra" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="amydra" status="sp. nov.">Coptodryas amydra</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Dorsal, lateral and declivital view of Coptodryas amydra holotype, 1.9 mm (A, B, I), C. bella, 3.75 - 4.0 mm (C, D, J), C. carinata holotype, 2.7 mm (E, F, K), and C. concinna, 2.3 mm (G, H, L)." figureDoi="10.3897/zookeys.983.52630.figure35" httpUri="https://binary.pensoft.net/fig/470835" pageId="0" pageNumber="1">Fig. 35A, B, I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
, female, Vietnam: Ninh Binh, Cuc Phuong N.P., Mac Lake,
<geoCoordinate degrees="20" direction="north" minutes="15" orientation="latitude" precision="1" seconds="29.0" value="20.258055">20°15'29.0&quot;N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="42" orientation="longitude" precision="1" seconds="27.5" value="105.707634">105°42'27.5&quot;E</geoCoordinate>
, 155 m, 4-7.v.2009, J.B. Heppner, ex blacklight trap (FSCA).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
1.9 mm long (n = 1); 2.71
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. This species is distinguished by the dense tuft of setae along the elytral base extending to interstriae 4; discal striae punctate, interstriae impunctate; elytral disc and declivity shagreened; declivital face subconvex; declivital interstriae 1-4 costate, 5 and 6 subcarinate; declivital posterolateral margin carinate to interstriae 7; pronotum basic (type 0) when viewed laterally, basic (type 2) when viewed dorsally; and anterior margin of the pronotum without a distinct row of serrations.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="similar species">
<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Coptodryas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodryas carinata" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="1">Coptodryas carinata</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="C. concinna" pageId="0" pageNumber="1" rank="species" species="concinna">
<emphasis italics="true" pageId="0" pageNumber="1">C. concinna</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="C. elegans" pageId="0" pageNumber="1" rank="species" species="elegans">
<emphasis italics="true" pageId="0" pageNumber="1">C. elegans</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="C. nudipennis" pageId="0" pageNumber="1" rank="species" species="nudipennis">
<emphasis italics="true" pageId="0" pageNumber="1">C. nudipennis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">(female).</emphasis>
1.9 mm long (n = 1); 2.71
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. Pronotum, head, antennae, legs and elytral disc light brown, declivity dark brown.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: epistoma entire, transverse, with a row of hair-like setae. Frons weakly convex to upper level of eyes, shagreened, alutaceous, impunctate, glabrous. Eyes deeply emarginate just above antennal insertion, upper part smaller than lower part. Submentum narrowly triangular, slightly impressed. Antennal scape regularly thick, as long as club. Pedicel as wide as scape, shorter than funicle. Funicle 4-segmented, segment 1 shorter than pedicel. Club approximately circular, flattened, type 4; segment 1 corneous, small, convex; segment 2 larger than segment 1, narrow, transverse, corneous; segments 1-3 present on posterior face.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Pronotum</emphasis>
: 1.06
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide. In dorsal view basic and parallel-sided, type 2, sides parallel in basal 2/3, rounded anteriorly; anterior margin without serrations. In lateral view basic, type 0, disc flat, summit at midpoint. Anterior slope with densely spaced, broad asperities, becoming lower and more strongly transverse towards summit. Disc subshiny with dense minute punctures, glabrous, some longer hair-like setae at margins. Lateral margins obliquely costate. Base weakly bisinuate, posterior angles acutely rounded, almost subquadrate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Elytra</emphasis>
: 1.73
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, 1.63
<normalizedToken originalValue="×">x</normalizedToken>
as long as pronotum. Scutellum minute, convex, slightly raised above elytral surface. Elytral mycangium setal tuft along elytral base dense, extending to interstriae 4. Elytral base bisinuate, edge oblique, humeral angles rounded, parallel-sided in basal 4/5, narrowly rounded to apex. Disc flat, shagreened, striae not impressed, with minute shallow punctures separated by three diameters of a puncture, glabrous; interstriae flat, impunctate, glabrous. Declivity occupying approximately 2/5 of elytral length, rounded, face subconvex, strongly shagreened; striae flat, punctate, punctures much larger than those of disc and very shallow; interstriae 1-4 costate, five and six subcarinate, impunctate, feebly to moderately granulate. Posterolateral margin carinate to interstriae 7.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
: procoxae contiguous, prosternal coxal piece short, conical. Protibiae obliquely triangular, broadest at apical 1/3; posterior face smooth; apical 1/2 of outer margin with six moderate socketed denticles, their length as long as basal width. Meso- and metatibiae flattened, outer margins evenly rounded with eight and nine large socketed denticles, respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
G.
<emphasis italics="true" pageId="0" pageNumber="1">amydros</emphasis>
= indistinct. Named in reference to its uninteresting habitus. A Latinized adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Vietnam.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="host">
<paragraph pageId="0" pageNumber="1">Host plants.</paragraph>
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
<caption doi="10.3897/zookeys.983.52630.figure35" httpUri="https://binary.pensoft.net/fig/470835" pageId="0" pageNumber="1" start="Figure 35" startId="F35">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 35.</emphasis>
Dorsal, lateral and declivital view of
<taxonomicName authorityName="Smith &amp; Beaver &amp; Cognato" authorityYear="2020" class="Insecta" family="Curculionidae" genus="Coptodryas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptodryas amydra" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="amydra">
<emphasis italics="true" pageId="0" pageNumber="1">Coptodryas amydra</emphasis>
</taxonomicName>
holotype, 1.9 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, B, I</emphasis>
),
<taxonomicName lsidName="C. bella" pageId="0" pageNumber="1" rank="species" species="bella">
<emphasis italics="true" pageId="0" pageNumber="1">C. bella</emphasis>
</taxonomicName>
, 3.75-4.0 mm (
<emphasis bold="true" pageId="0" pageNumber="1">C, D, J</emphasis>
),
<taxonomicName lsidName="C. carinata" pageId="0" pageNumber="1" rank="species" species="carinata">
<emphasis italics="true" pageId="0" pageNumber="1">C. carinata</emphasis>
</taxonomicName>
holotype, 2.7 mm (
<emphasis bold="true" pageId="0" pageNumber="1">E, F, K</emphasis>
), and
<taxonomicName lsidName="C. concinna" pageId="0" pageNumber="1" rank="species" species="concinna">
<emphasis italics="true" pageId="0" pageNumber="1">C. concinna</emphasis>
</taxonomicName>
, 2.3 mm (
<emphasis bold="true" pageId="0" pageNumber="1">G, H, L</emphasis>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>