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<mods:title id="B7D383A01969DA5743FD4DA4E7CEE410">Trichoribates and Jugatala (Acari: Oribatida: Ceratozetidae) from the Central and Southern Alps, with notes on their distribution</mods:title>
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<mods:namePart id="8FF9D0A7EB78861CB2D5074B4421D306">Bayartogtokh, B.</mods:namePart>
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<mods:namePart id="1862F6683E97147FC2D80E94EFDC5C5F">Schatz, H.</mods:namePart>
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<treatment id="A6CAB06F4A14B5B4D8E5073259F92F46" ID-DOI="http://doi.org/10.5281/zenodo.6229541" ID-GBIF-Taxon="100113810" ID-Zenodo-Dep="6229541" LSID="urn:lsid:plazi:treatment:A6CAB06F4A14B5B4D8E5073259F92F46" httpUri="http://treatment.plazi.org/id/A6CAB06F4A14B5B4D8E5073259F92F46" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28">
<subSubSection id="6AA1008E9C272A0CF33D8BE05725B91F" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="B9FBD7F0D31698975FE399A3438878AF" pageId="27" pageNumber="28">
<taxonomicName id="49E55E804B85C9317E551182AE2EEFAF" class="Arachnida" family="Ceratozetidae" genus="Jugatala" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cribelliger">
<pageBreakToken id="392482C81254E142D257750E5BABAEE4" pageId="27" pageNumber="28" start="start">Jugatala</pageBreakToken>
cribelliger
</taxonomicName>
(Berlese, 1904)
</paragraph>
</subSubSection>
<subSubSection id="B29E6E0600A3B18BFA154905F002A9A2" pageId="27" pageNumber="28" type="description">
<paragraph id="2D7C55960DC95396B6899DD60AFDA96F" pageId="27" pageNumber="28">(Figs. 15-17)</paragraph>
</subSubSection>
<subSubSection id="4414CACC605C47D11F31FF05801D32C9" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="5A9CEA2DE73A854459F34A7892F96C16" pageId="27" pageNumber="28">
<taxonomicName id="8E9A0B0783902BF0705D30A46134D7AF" genus="Oribates" pageId="27" pageNumber="28" rank="species" species="cribelliger">Oribates cribelliger</taxonomicName>
Berlese 1904, p. 28, fig. 46.
</paragraph>
<paragraph id="A81AD0780CECCAD21B145EA3E2EAF219" pageId="27" pageNumber="28">
<taxonomicName id="05A9BCCFB8526A899C9893292AA8055C" genus="Oribates" pageId="27" pageNumber="28" rank="species" species="cribelliger">Oribates cribelliger</taxonomicName>
. Castanogli &amp; Pegazzano 1985, p. 96.
</paragraph>
<paragraph id="BC14A859AB51CD9E04421F61CDE1C27B" pageId="27" pageNumber="28">
<taxonomicName id="62EBBC88C41F9775728494A99EF47219" class="Arachnida" family="Ceratozetidae" genus="Ceratozetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cribelliger">Ceratozetes cribelliger</taxonomicName>
: Schweizer 1956, p. 313, fig. 258.
</paragraph>
<paragraph id="46C6D48CDFBB4ADD377BA03621A3CD3B" pageId="27" pageNumber="28">
<taxonomicName id="FA506440270C5146E4B355A06321CF31" class="Arachnida" family="Punctoribatidae" genus="Mycobates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cribelliger">Mycobates cribelliger</taxonomicName>
: Mahunka &amp; Mahunka-Papp 1995, p. 84, figs. 155, 156.
</paragraph>
<paragraph id="7F18468A2C0F7B9EAB3B69441E9A11A0" pageId="27" pageNumber="28">
<taxonomicName id="7F201412DFFCADCA9EFA1B9B573C83B4" class="Arachnida" family="Ceratozetidae" genus="Jugatala" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cribelliger">Jugatala cribelliger</taxonomicName>
:
<normalizedToken id="5035E369AB78A5CDB847D669D0CD8D73" originalValue="Subías" pageId="27" pageNumber="28">Subias</normalizedToken>
2000, p. 23.
</paragraph>
<paragraph id="BA0C938831CC0F8CACC6BCD50FBFF035" pageId="27" pageNumber="28">
<taxonomicName id="B81B01C088175636F888AF65C9CBF543" class="Arachnida" family="Punctoribatidae" genus="Mycobates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="cribelliger" subGenus="Calyptozetes">Mycobates (Calyptozetes) cribelliger</taxonomicName>
<normalizedToken id="B040CBFFE98DE455B06D84DCF00DBB91" originalValue="Subías" pageId="27" pageNumber="28">Subias</normalizedToken>
2004, 183; 2008, p. 344.
</paragraph>
<paragraph id="D729808831B6C381F6D278FCBBA4B5A9" pageId="27" pageNumber="28">
<taxonomicName id="BB785C04A5B0CCB22B7B123065DCA8AF" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="oxypterus">Trichoribates oxypterus</taxonomicName>
: Schatz 1989, p. 120.
</paragraph>
</subSubSection>
<subSubSection id="30B8F6FDD579DA3E2FC1FF1474D3BD8B" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="130C83BB9E39AC72580CACE7C5AB7540" pageId="27" pageNumber="28">Diagnosis. Rostrum broadly rounded, without lateral dens, with a nose-like protuberance dorsally; all prodorsal setae finely barbed; lamella relatively wide, with medium long, thin translamella; lamellar cusp with or without minute lateral dens; inner dens absent; sensillus short, with smooth clavate head; tutorium medium in size, its basal part approximately same in width, cusp widened proximally, but pointed distally; 10 pairs of notogastral setae of medium length, thin, smooth, dp absent; porose areas round to oval, outer margin poorly developed; lenticulus absent; all ventral setae thin, smooth.</paragraph>
</subSubSection>
<subSubSection id="9CA87D84969271DCE38597DCBD389159" pageId="27" pageNumber="28" type="description">
<paragraph id="4608EA416DF3242A1318965148354185" pageId="27" pageNumber="28">
Measurements. (n = 8) Body length 418-433 (425)
<normalizedToken id="69F38F8707F02AB538B29FEB94032753" originalValue="µm" pageId="27" pageNumber="28">ym</normalizedToken>
; length of notogaster 342-369 (358)
<normalizedToken id="263BBC085C9FF3B457C7D194A8DF010E" originalValue="µm" pageId="27" pageNumber="28">ym</normalizedToken>
; width of notogaster 266-281 (274)
<normalizedToken id="3242865C66F575093018AF604EAB0CC9" originalValue="µm" pageId="27" pageNumber="28">ym</normalizedToken>
.
</paragraph>
<paragraph id="A448A23FAA2AC5B7CBC4E554E52E747E" pageId="27" pageNumber="28">Integument. Body colour yellowish brown. With thin cerotegument, surface of dorsal and ventral plates nearly smooth.</paragraph>
<paragraph id="DE792D2417244101E992080C18E0545F" pageId="27" pageNumber="28">Prodorsum (Figs. 15A, B, E, 16A-D). Rostrum broadly rounded, smooth, without lateral dens, with a nose-like protuberance dorsally. All prodorsal setae finely barbed, subequal in length; interlamellar seta short, just reaching level of translamella. Lamella relatively wide, with long, but thin translamella; lamellar cusp with or without minute lateral dens; medial dens absent (Fig. 16B-D). Sensillus short, with smooth clavate head (Fig. 15B). Bothridium large, with strongly developed bothridial scale svm, latter not pointed, but rounded distally. Tutorium medium in size, extending to alveolus of rostral seta; width of its basal part approximately same; proximal part of cusp widened, but pointed distally (Fig. 15E).</paragraph>
<paragraph id="1CBD13345ADCE424DC8D16519D362C48" pageId="27" pageNumber="28">Notogaster (Figs. 15A, G, 16E). About 1.3 x longer than wide. Surface smooth. Anterior margin of notogaster broadly arched anteriad, lenticulus absent. Pteromorph small, curved ventrally. Ten pairs of notogastral setae thin, smooth, dp absent. Porose areas round to oval, their outer margins poorly developed; Aa and A1 larger than A2 and A3 (Fig. 15C). Lyrifissures ia, im, ih, ips, ip and opisthosomal gland opening small.</paragraph>
<paragraph id="5C4FEDB0DED5D7B7B9D7BC895577CD4E" pageId="27" pageNumber="28">Gnathosoma (Figs. 15D, 16F, G). Subcapitular mentum conspicuously wider than long, without microtubercles. Hypostomal setae a, h, m of medium length, thin, smooth. Palp and chelicerae typical for family as shown in Fig. 16F, G.</paragraph>
<paragraph id="961D7BB2048A55B7EDC79B0D950E147B" pageId="27" pageNumber="28">Epimeral region (Fig. 15D). Apodemes well developed, apo 2 and apo sj long, aligned obliquely, nearly parallel to each other; apo 3 short. Epimeral setae thin, smooth; setae 1c, 3b and 3c slightly longer than others, setal formula 3-1-3-3. Custodium short, just reaching anterior margin of pedotectum II; discidium markedly projecting laterally.</paragraph>
<paragraph id="84DE165E64E95D2E914C1066606503A6" pageId="27" pageNumber="28">Ano-genital region (Fig. 15D, F, G). Anal aperture slightly larger than genital aperture, anal and genital plates smooth. All ano-genital setae thin, smooth; adanal setae ad3 situated in a relatively posterior position; placement of genital setae somewhat variable as shown in Fig. 15D, F; setal formula same as in the previous species. Adanal lyrifissures lad short, situated at same level as anal seta an2, close to anterolateral margin of anal aperture. Postanal porose area narrowly elongate, slightly shorter than distance between bases of adanal setae ad2 (Fig. 15G).</paragraph>
<paragraph id="6FFAC1661CCAE47F188F486AD7410978" pageId="27" pageNumber="28">Legs (Fig. 17). Medial claw much thicker than lateral claws. Formula of leg setation (including famulus): I (1-5-3-4-20); II (1-5-3-4-15); III (2-3-1-3-15); IV (1-2-2-3-12); formula of solenidia: I (1-2-2); II (1-1-2); III (1-1-0); IV (0-1-0). Most leg setae distinctly barbed, except (p) and (u) on tarsi I-IV Setae l&quot; on tibiae and genua I-IV thicker than other setae, heavily barbed.</paragraph>
</subSubSection>
<subSubSection id="B431B9016F757B5D33DA914131A8898D" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="multiple">
<caption id="03039E3E48228D5BFBCA374DF840F5BE" pageId="28" pageNumber="29">
<paragraph id="91DEA2BF637EB62CC1BB68F0613F0989" pageId="28" pageNumber="29">
<pageBreakToken id="B21454D12A3DD87C1D7CE342FAD5F8A6" pageId="28" pageNumber="29" start="start">FIGURE</pageBreakToken>
15.
<taxonomicName id="8A90ECF485A3E877114D689E0C1CA1B8" class="Arachnida" family="Ceratozetidae" genus="Jugatala" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cribelliger" subGenus.completionEvidence="missing, matching species" subGenus.innerRound="1" subGenus.outerRound="1">Jugatala cribelliger</taxonomicName>
(Berlese, 1904).
<normalizedToken id="EC4F3DBB371C9AEB5D15421D55931627" originalValue="A—Dorsal" pageId="28" pageNumber="29">A-Dorsal</normalizedToken>
view;
<normalizedToken id="7A9FA7157BD26A1359E74CEF1A12081A" originalValue="B—Sensillus" pageId="28" pageNumber="29">B-Sensillus</normalizedToken>
and bothridium (lateral view);
<normalizedToken id="A024ADFF0547A106F6C6549A994CEF98" originalValue="C—" pageId="28" pageNumber="29">C-</normalizedToken>
Relative placement of porose areas, notogastral setae, lyrifissures and opisthosomal gland opening (dorso-lateral view);
<normalizedToken id="099D19918FF95ECAAFA9BB2064CBF7CE" originalValue="D—Ventral" pageId="28" pageNumber="29">D-Ventral</normalizedToken>
view;
<normalizedToken id="9F6DB670A101C20797D0B002387C0574" originalValue="E—Variation" pageId="28" pageNumber="29">E-Variation</normalizedToken>
of tutorium (right);
<normalizedToken id="36ED21B133BBAE62D04D772D3FB56758" originalValue="F—Genital" pageId="28" pageNumber="29">F-Genital</normalizedToken>
area;
<normalizedToken id="747A3C0B6076DC7F03DC80CBEFC30215" originalValue="G—Posterior" pageId="28" pageNumber="29">G-Posterior</normalizedToken>
view of opisthosoma. Scale bar is same for
<normalizedToken id="9751267F451F259EC0FB52F0E0FB07DF" originalValue="AD" pageId="28" pageNumber="29">A-D</normalizedToken>
;
<normalizedToken id="2824E95CB8F43BE2CB45863A4F518D80" originalValue="EG" pageId="28" pageNumber="29">E-G</normalizedToken>
, respectively.
</paragraph>
</caption>
<caption id="C37E26A49440403EBD1756432B728D72" pageId="29" pageNumber="30">
<paragraph id="78E078E09515E6FF92708707E55D2A1F" pageId="29" pageNumber="30">
<pageBreakToken id="77AD5F4B2A071C03F61BA022598B4048" pageId="29" pageNumber="30" start="start">FIGURE</pageBreakToken>
16.
<taxonomicName id="02D1D86AABF657896A1CBEFD89909A87" class="Arachnida" family="Ceratozetidae" genus="Jugatala" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cribelliger" subGenus.completionEvidence="missing, matching species" subGenus.innerRound="1" subGenus.outerRound="1">Jugatala cribelliger</taxonomicName>
(Berlese, 1904).
<normalizedToken id="E42F0C03CFD214B0A46080D109485703" originalValue="A—Prodorsum" pageId="29" pageNumber="30">A-Prodorsum</normalizedToken>
and anterior part of notogaster;
<normalizedToken id="D8D8A57E59681A94B8B3C59C49805038" originalValue="B-D—Lamellae" pageId="29" pageNumber="30">B-D-Lamellae</normalizedToken>
and lamellar setae, showing Variation of lamellar cusps;
<normalizedToken id="AEFCBFA25830890EE8F74CF5292C99F2" originalValue="E—Lateral" pageId="29" pageNumber="30">E-Lateral</normalizedToken>
view of notogaster;
<normalizedToken id="0EA982BB7723DA1C349F6DBE76196361" originalValue="F—Chelicera" pageId="29" pageNumber="30">F-Chelicera</normalizedToken>
(right, antiaxial view);
<normalizedToken id="2647D625288E48EA007469C2462BCF55" originalValue="G—Palp" pageId="29" pageNumber="30">G-Palp</normalizedToken>
(right, antiaxial view). Scale bar is same for A-D.
</paragraph>
</caption>
<caption id="B3DBC636B6F44F826F971E34521793A0" pageId="30" pageNumber="31">
<paragraph id="240F7F9383E82C1C46C81212BAA37099" pageId="30" pageNumber="31">
<pageBreakToken id="BFFB00E2FDF39F1372F7236BE171D1C4" pageId="30" pageNumber="31" start="start">FIGURE</pageBreakToken>
17.
<taxonomicName id="22B3E3E2167E52838C0E548D3E4428A0" class="Arachnida" family="Ceratozetidae" genus="Jugatala" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="cribelliger" subGenus.completionEvidence="missing, matching species" subGenus.innerRound="1" subGenus.outerRound="1">Jugatala cribelliger</taxonomicName>
(Berlese, 1904).
<normalizedToken id="6F9CDEEC40B69C85ED0D7A0F59D0D294" originalValue="A—Leg" pageId="30" pageNumber="31">A-Leg</normalizedToken>
I (right, antiaxial view);
<normalizedToken id="0281DFD242848D0F3A5CCACA570A9B03" originalValue="B—Leg" pageId="30" pageNumber="31">B-Leg</normalizedToken>
II (right, antiaxial view);
<normalizedToken id="926CE122607D787C6F6E92ABB3FF2A87" originalValue="C—Leg" pageId="30" pageNumber="31">C-Leg</normalizedToken>
III (left, antiaxial view);
<normalizedToken id="004ADAD5ADD76FB9CC9A89712B2DF44D" originalValue="D—Leg" pageId="30" pageNumber="31">D-Leg</normalizedToken>
IV (right, antiaxial view). Scale bar is same for all figures.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="4A456F55896A570D3E478E4BDE5F7291" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="5CF435F190AA8C8F164EC95C3410C345" pageId="31" pageNumber="32">
<pageBreakToken id="BB3DC3B491B04F3B30AE741F1053F5DC" pageId="31" pageNumber="32" start="start">Material</pageBreakToken>
examined.
<materialsCitation id="360AD39D453D57AEDC74C9B952389B2E" ID-GBIF-Occurrence="923893747" collectingDate="1988-07-17" collectorName="H. Schatz" country="Austria" latitude="47.0335" location="Kaiser Dorfertal - Daberklamm" longitude="12.6233" stateProvince="East Tyrol">
<collectingCountry id="2CB92F38D9C7538121F7E01EBEBB6891">Austria</collectingCountry>
,
<collectingRegion id="0F5238A7988B344B3F4B6B2DC238FF06">East Tyrol</collectingRegion>
,
<location id="1B8237002D9B834B7402AAA4B15D2795" LSID="urn:lsid:plazi:treatment:A6CAB06F4A14B5B4D8E5073259F92F46:1B8237002D9B834B7402AAA4B15D2795" country="Austria" latitude="47.0335" longitude="12.6233" name="Kaiser Dorfertal - Daberklamm" stateProvince="East Tyrol">Kalser Dorfertal - Daberklamm</location>
, in loose gravel with cushion plant Vegetation (1520 m a.s.l.,
<date id="8D5BC6185BFD2AAE752903F872A815C9" value="1988-07-17">
<collectingDate id="BCD30AB226C1657F22900919B95F814D" value="1988-07-17">17 July 1988</collectingDate>
</date>
, leg.
<collectorName id="E1108C0EB821C739564FED113B16A7B5">H. Schatz</collectorName>
, 4 ad.)
</materialsCitation>
;
<materialsCitation id="726CD3E18FA78BFCE794A78F88B38761" ID-GBIF-Occurrence="923893746" collectingDate="1988-09-01" collectorName="H. Schatz" country="Austria" latitude="47.0335" location="Kalser Dorfertal - Daberklamm" longitude="12.6233" stateProvince="East Tyrol">
<location id="644B35A8C053A26839B6484E73F7E48D" LSID="urn:lsid:plazi:treatment:A6CAB06F4A14B5B4D8E5073259F92F46:644B35A8C053A26839B6484E73F7E48D" country="Austria" latitude="47.0335" longitude="12.6233" name="Kalser Dorfertal - Daberklamm" stateProvince="East Tyrol">ibid.</location>
, (
<date id="AFA65E6D47C8201BF0853B672A8CE97D" value="1988-09-01">
<collectingDate id="E7678DE1C77711D3A2E67F836F9872D5" value="1988-09-01">01 September 1988</collectingDate>
</date>
, leg.
<collectorName id="CF8299E528DAD834A4AF0EF19CCF5910">H. Schatz</collectorName>
, 1 ad.), sub
<taxonomicName id="2FD21D73F5A6823240C6FCA45411481B" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="oxypterus">Trichoribates oxypterus</taxonomicName>
(Schatz 1989)
</materialsCitation>
,
<materialsCitation id="5BBD73E96D07D61FFC88060F0A86F615" ID-GBIF-Occurrence="923893754" collectingDate="2007-06-30" collectorName="B. M. Fischer" country="Italy" latitude="46.509144" location="Dolomites, Plattkofel / Sasso Piatto" longitude="11.698380" stateProvince="South Tyrol">
<collectingCountry id="279C61BC9EE2FA5CBD6E9AE9D63BCA34">Italy</collectingCountry>
,
<collectingRegion id="44E38B8C1616EEB2E9FE9BD5274F44CD">South Tyrol</collectingRegion>
,
<location id="3E0F1C9DDAC8474ADEF850ED5599B981" LSID="urn:lsid:plazi:treatment:A6CAB06F4A14B5B4D8E5073259F92F46:3E0F1C9DDAC8474ADEF850ED5599B981" country="Italy" latitude="46.509144" longitude="11.698380" name="Dolomites, Plattkofel / Sasso Piatto" stateProvince="South Tyrol">Dolomites, Plattkofel/Sasso Piatto</location>
, calcareous screen with grass, moss and herbs (2200 m a.s.l.,
<date id="8B5C3DB6D5A7DA4A51CC592F2BD42C89" value="2007-06-30">
<collectingDate id="793B3D5FDD42511A795493D2D953642F" value="2007-06-30">30 June 2007</collectingDate>
</date>
: 4 ad., leg.
<collectorName id="D9868A87F65ACE4815CC447FF3367F63">B. M. Fischer</collectorName>
; Fischer and Schatz 2007)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B88AE29090391912B119F3656E23A27A" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8EC053B8C60CFF94B046CA41B9848B0A" pageId="31" pageNumber="32">
Remarks. The main characters of the present material agree with those of the type specimen, examined by Mahunka and Mahunka-Papp (1995) and the material from Switzerland studied by Schweizer (1956). Since the original description of this species the generic Status has been changed several times. Schweizer (1956) considered it as a member of
<taxonomicName id="A856F4ECEE46E09D0F556F3B65481BF1" class="Arachnida" family="Ceratozetidae" genus="Ceratozetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Ceratozetes</taxonomicName>
, while Mahunka and Mahunka-Papp (1995) moved it to the genus
<taxonomicName id="BB7D73FC01018E20E0469EAEFB2A327B" class="Arachnida" family="Punctoribatidae" genus="Mycobates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Mycobates</taxonomicName>
. Recently,
<normalizedToken id="E674DB379E65F953E66604D6F47955B2" originalValue="Subías" pageId="31" pageNumber="32">Subias</normalizedToken>
(2000) combined it with
<taxonomicName id="8E5B2F65BE185969E6E9185BF56610CA" class="Arachnida" family="Ceratozetidae" genus="Jugatala" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Jugatala</taxonomicName>
. Later,
<normalizedToken id="D7CBC72750A8BDF787C0F6C7B6D609B4" originalValue="Subías" pageId="31" pageNumber="32">Subias</normalizedToken>
(2004, 2008) moved this species again to
<taxonomicName id="B2307CC15C003DFAD5B683DC47B724DD" class="Arachnida" family="Punctoribatidae" genus="Mycobates" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subGenus" subGenus="Calyptozetes">Mycobates (Calyptozetes)</taxonomicName>
.
</paragraph>
<paragraph id="AEE51B3F5BE5347DCACD4CD08ED94702" pageId="31" pageNumber="32">
We consider this species a member of
<taxonomicName id="1D1CA803EF673BEC6DE9F06A36508635" class="Arachnida" family="Ceratozetidae" genus="Jugatala" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Jugatala</taxonomicName>
(fam.
<taxonomicName id="237F69B61C6DD90564B30D06694CDB00" family="Ceratozetidae" pageId="31" pageNumber="32" rank="family">Ceratozetidae</taxonomicName>
) since it lacks the posterior notogastral tectum, which is one of the main characters of the representatives of adult
<taxonomicName id="D09ECD9FB22D066B10C9F57D0D8BA2AC" family="Mycobatidae" pageId="31" pageNumber="32" rank="family">Mycobatidae</taxonomicName>
(Behan-Pelletier 2000; Behan-Pelletier &amp; Eamer 2008). The setation of legs also confirms this opinion.
</paragraph>
</subSubSection>
<subSubSection id="5B1D8C5793BB11E5912DADF1613683AF" pageId="31" pageNumber="32" type="distribution">
<paragraph id="0AA5AEEDF3937C3640F9F177265A8539" pageId="31" pageNumber="32">
Distribution. This species is distributed in the Central and south-western Europe, and has been recorded in Northern Italy, Austria, Switzerland and in the Iberian Peninsula - Portugal and Spain (Schweizer 1956;
<normalizedToken id="829D5562C928CCA68C5B41B0C4D06AEB" originalValue="Subías" pageId="31" pageNumber="32">Subias</normalizedToken>
2000; Bernini et al. 1995; Mahunka &amp; Mahunka-Papp 1995, and present study).
</paragraph>
</subSubSection>
</treatment>
</document>