167 lines
16 KiB
XML
167 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="A6C9D7B96BDD570CBD050074E9685291" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Lithocharodes lituratus Irmler 2024, sp. nov." docType="treatment" docUuid="E1BE0BED-DBF9-4190-8CA4-AF15CFC96C41" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
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<mods:mods id="DB6FF0265E3BAD7B36D8465611E8E906" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="088EBA328FF57BA71C2FDFFDE470584E">
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<mods:title id="D77FD06676E98AFF5569725F576D2FBE">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="967E0B6B43F6E5474B8DFA92D43234CC" type="personal">
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<mods:role id="69CD2513591871401487D0A979FAFEE7">
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<mods:roleTerm id="C6111936DC5884571584FE17FC858A5F">Author</mods:roleTerm>
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<mods:namePart id="183C017C3DB8AD1E8A3A49507686EFA7">Irmler, Ulrich</mods:namePart>
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<mods:affiliation id="7EC5186A554F91FB0002BF97C51C17F9">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
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<mods:nameIdentifier id="6058F36E80E9F4C30005DD57A7E791D1" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
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<mods:typeOfResource id="5C398EC1F2D4894997DF74850F69D8B8">text</mods:typeOfResource>
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<mods:title id="6F358C7170E7C42458875E4C5C79F832">Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="4942779B542F5A89033416F9F15BE7F0">
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<mods:date id="A78923ABCB3B84EE58D855454B7EED5A">2024</mods:date>
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<mods:detail id="760F5ABE30897C4E1C555E2A15E084DA" type="pubDate">
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<mods:number id="8C55D4CEDD241FD6854BCE0D76833263">2024-01-31</mods:number>
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<mods:number id="1291F860B3F8C67E167EF2B85EF95838">74</mods:number>
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<mods:detail id="13C4029E0B3D0600371E23DD08041921" type="issue">
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<mods:number id="7DC7F59482299091930BA548FAA9843B">1</mods:number>
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<mods:extent id="54715F61768232882AC6D669001FFAEB" unit="page">
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<mods:start id="4C5C9B210B6793AB35424713AC700EB8">13</mods:start>
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<mods:url id="9B45CA5FD38F9A72CB3205D7052F34F7">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
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<mods:classification id="4EB1F7B627C3B1000C7AC8113677E2E8">journal article</mods:classification>
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<mods:identifier id="848AE6D61ED086A10597594CF9B7CCF5" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
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<treatment id="A6C9D7B96BDD570CBD050074E9685291" LSID="urn:lsid:zoobank.org:act:E1BE0BED-DBF9-4190-8CA4-AF15CFC96C41" httpUri="http://treatment.plazi.org/id/A6C9D7B96BDD570CBD050074E9685291" lastPageNumber="13" pageId="0" pageNumber="13">
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<subSubSection id="D44B2B09EF757C9EA22F504D3DA8FC51" pageId="0" pageNumber="13" type="nomenclature">
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<paragraph id="D710B4C51EE65DB835794BF7C2AD7131" pageId="0" pageNumber="13">
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<taxonomicName id="14D1E1704C35F8614A0A7745F10CF26D" LSID="https://zoobank.org/E1BE0BED-DBF9-4190-8CA4-AF15CFC96C41" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes lituratus" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="lituratus" status="sp. nov.">Lithocharodes lituratus</taxonomicName>
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<taxonomicNameLabel id="7DB9D3193950715EE5E2C867C694899B" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="EC807D32D874370F4FFE34034C6B8B2E" pageId="0" pageNumber="13" type="description">
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<paragraph id="AED0DABA477091F2ABA5CFE461C6D758" pageId="0" pageNumber="13">
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<figureCitation id="DCABADED5875461932737447C6C7A844" captionStart="Figures 5–8" captionStartId="F2" captionText="Figures 5 - 8. 5. Lithocharodes hansoni; 6. L. hibbsi; 7. L. lituratus; 8. L. montanus. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures5-8" httpUri="https://binary.pensoft.net/fig/975716" pageId="0" pageNumber="13">Figs 7a-d</figureCitation>
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<figureCitation id="E6E9499527E55F29CD638659BB6188CB" captionStart="Figure 20" captionStartId="F7" captionText="Figure 20. A. Fore-body of Lithocharodes lituratus; B. L. montanus; C. L. parallelus; D. L. zamorae; E. Somoleptus alajuelae; F. S. curtioculatus. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure20" httpUri="https://binary.pensoft.net/fig/975721" pageId="0" pageNumber="13">, 20A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6183794A79FB35099ADED50CA502D887" pageId="0" pageNumber="13" type="materials_examined">
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<paragraph id="B182E8382F1FF67314ACC5862618F3B0" pageId="0" pageNumber="13">Material examined.</paragraph>
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<paragraph id="6D5FA98C6C3DDABE88442FD5BEC59298" pageId="0" pageNumber="13">
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<materialsCitation id="2C3EDA6FA4D181DD7CBFB7BE0E69C3C9" collectingDate="1991-12-23" collectorName="C. Carlton, R. Leschen" country="ECUADOR" elevation="1300" location="Quito" specimenCount="1" stateProvince="Pichincha" typeStatus="Holotype">
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<emphasis id="5B1B3AD475601F989CCE96AA121D5B33" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="19CA60774625332DF67630239CCB6E93" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<collectingCountry id="6D113BD478FE86AA7CE6E745A9FE10D4" name="Ecuador">ECUADOR</collectingCountry>
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male;
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<collectingRegion id="A9605B49BBFD7F6555553140DE7F5074" country="Ecuador" name="Pichincha">Pichincha</collectingRegion>
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; Maquipucuna For. Res.;
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<quantity id="921E1972A8CF5927A9EDD4A6543F0257" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
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<location id="584BD71B7F91A661CBD345C2E8E0166D" LSID="urn:lsid:plazi:treatment:A6C9D7B96BDD570CBD050074E9685291:584BD71B7F91A661CBD345C2E8E0166D" country="ECUADOR" name="Quito" stateProvince="Pichincha">Quito</location>
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;
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<quantity id="A6A24E3F6DBDFE8A3D4F8C59135E7F5A" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
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<elevation id="7B160E1D42BF51647ED4C636FD7119A4" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
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</quantity>
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elevation; banana duff; berlesate;
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<collectingDate id="DF427F2F3A6007557C08C8F9BC986583" value="1991-12-23">23 Dec 1991</collectingDate>
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;
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<collectorName id="03CCD98A557F25098F591463B611AEE6">C. Carlton</collectorName>
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,
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<collectorName id="0EE36F68A24A804639A85E249D47FE0D">R. Leschen</collectorName>
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leg.; KNHM #72.
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</materialsCitation>
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</paragraph>
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<paragraph id="038F14393D373985DB911B417990FEE0" pageId="0" pageNumber="13">
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<materialsCitation id="C2549A5F3F08DCC06A997C6FEA6F6893" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="C. Carlton & R. Leschen" country="ECUADOR" elevation="1300" location="ECUADOR" specimenCount="1" specimenCount-female="1" stateProvince="Pichincha" typeStatus="Paratypes">
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<emphasis id="60BDFFA51FACBEB199D0437963251CC6" bold="true" italics="true" pageId="0" pageNumber="13">
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<typeStatus id="6006E96F79AE3BA3B400EA21BC196C3F" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<collectingCountry id="ED604445A858CD64540974B829D8C324" name="Ecuador">ECUADOR</collectingCountry>
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<specimenCount id="DCF4DB32E9C786E925F6B2DB858F10BA" count="1" type="female">1 female</specimenCount>
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; same data as holotype; KNHM ECU1A99
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</materialsCitation>
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;
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<materialsCitation id="E7DBD0F41F7D6EAC0836604BE7ADA331" collectingDate="1999-10-27" collectorName="R. Anderson" country="Ecuador" elevation="1200" latitude="0.11666667" location="Quito" longLatPrecision="21" longitude="-78.634995" specimenCount="1" specimenCount-male="1" stateProvince="Pichincha" typeStatus="paratype">
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<specimenCount id="509D5DCB1F344602938A54618F96D738" count="1" type="male">1 male</specimenCount>
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; same location as holotype but;
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<geoCoordinate id="A5C371BC14B96468725CC17077D4994A" degrees="0" direction="north" minutes="7" orientation="latitude" precision="15" seconds="0" value="0.11666667">0°7'0"N</geoCoordinate>
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,
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<geoCoordinate id="AD324D0A3C486A9E0B306E4AFF5734A2" degrees="78" direction="west" minutes="38" orientation="longitude" precision="15" seconds="6" value="-78.634995">78°38'6"W</geoCoordinate>
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;
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<quantity id="6D17D1896760F8B51501F26E85E10938" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
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<elevation id="98231ADD43B08D8742DE8E27A95D45C2" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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elevation; montane evergreen forest litter;
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<collectingDate id="B3A6359412E14DDD1890490C8527F9B3" value="1999-10-27">27 Oct 1999</collectingDate>
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;
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<collectorName id="4D2F114C22939873B94696C83FD89E50">R. Anderson</collectorName>
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leg.; KNHM ECU1A99 208D
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="E0239AF586CCCEF1D4414B346F97648B" pageId="0" pageNumber="13" type="diagnosis">
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<paragraph id="36C27EA8FE390CBEF9981E320EFF679E" pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph id="E32ACA20F139BCCD851F750FC1D203CB" pageId="0" pageNumber="13">
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According to the structure of the aedeagus,
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<taxonomicName id="5C018692514D0205AA0845EA9BAD56E2" lsidName="L. lituratus" pageId="0" pageNumber="13" rank="species" species="lituratus">
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<emphasis id="166004AD73C42C92503D9B7D76ECBC24" italics="true" pageId="0" pageNumber="13">L. lituratus</emphasis>
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</taxonomicName>
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belongs to the
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<taxonomicName id="45B14EF05ED508988C8A0F6FDA7BA508" lsidName="L. sordida" pageId="0" pageNumber="13" rank="species" species="sordida">
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<emphasis id="9F631D83159D2276657410DA4D15CF96" italics="true" pageId="0" pageNumber="13">L. sordida</emphasis>
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</taxonomicName>
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-group that is also characterised by short and thick parameres. Colouration of
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<taxonomicName id="31058C0ABA86ADF23166FD4D514BEA9B" lsidName="L. lituratus" pageId="0" pageNumber="13" rank="species" species="lituratus">
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<emphasis id="B6C8F7B9CE71B07C437E64091E6C74FF" italics="true" pageId="0" pageNumber="13">L. lituratus</emphasis>
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</taxonomicName>
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is similar as in
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<taxonomicName id="6B9BDD4B15D1BABC4EDC79830E764C3C" lsidName="L. bicornis" pageId="0" pageNumber="13" rank="species" species="bicornis">
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<emphasis id="7E3F3CB7A922BBD13386FFD10DE0F24D" italics="true" pageId="0" pageNumber="13">L. bicornis</emphasis>
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</taxonomicName>
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Irmler, 2021 by the lighter brown parts on elytra and pronotum. In contrast to
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<taxonomicName id="5A5E85E936D199E50F69CA84CAD9411F" lsidName="L. lituratus" pageId="0" pageNumber="13" rank="species" species="lituratus">
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<emphasis id="5E8DCEA2290951F727B4ADF0066AC964" italics="true" pageId="0" pageNumber="13">L. lituratus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="C0083E40F798BA8FFE6C602B4C44E98D" lsidName="L. sordida" pageId="0" pageNumber="13" rank="species" species="sordida">
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<emphasis id="47CF7077390D9F6F7D9166DB49442F65" italics="true" pageId="0" pageNumber="13">L. sordida</emphasis>
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</taxonomicName>
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Sharp, 1885 is nearly black. The species can be easily differentiated from these two species by the slenderer shape and the sparser punctation. The location, Ecuador, is between that of the two other species in Peru and Panama.
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</paragraph>
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</subSubSection>
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<subSubSection id="3ABA83240BB18362996B8673FD1BF0F9" pageId="0" pageNumber="13" type="description">
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<paragraph id="6CA6BA2D56802EF7EDC825B9E8EC7275" pageId="0" pageNumber="13">Description.</paragraph>
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<paragraph id="F160A89B5C2FEF339AC92E792F3FBA15" pageId="0" pageNumber="13">Length: 6.4 mm; Colouration: blackish with indistinct lighter spots on humeral angles of elytra. Legs and antennae light brown.</paragraph>
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<paragraph id="6AA9212918549E0C3A75A116F77FD293" pageId="0" pageNumber="13">Head: 0.93 mm long, 0.79 mm wide; eyes slightly prominent; PS:E ratio 5.2; without posterior angles; posteriorly, widely rounded; inter-antennal furrows short; between eyes with indistinct depression; setiferous punctation moderately fine and sparse; on average, interstices between punctures nearly twice as wide as diameter of punctures; at sides and at posterior margin, punctation deeper and denser than in centre; surface without microsculpture; polished; antennae with elongate first antennomere; longer than half-length of head; second and third antennomere each nearly twice as long as wide; combined half as long as first antennomere; following antennomeres transverse; approximately twice as wide as long; apically increasing in width. Pronotum: 1.24 mm long, 0.79 mm wide, widest in anterior third; in posterior part, slightly convergent, but nearly parallel; posterior margin nearly semi-circular; at posterior angle, 0.8 times as wide in anterior part; setiferous punctation deeper and denser than on head; on average, interstices half as wide as diameter of punctures; wide mid-line impunctate; surface without microsculpture; polished. Elytra: 1.05 mm long, 0.92 mm wide; humeral angles obtuse; posteriorly, sides approximately parallel; posterior angles sub-rectangular; posterior margin retreated to suture; setiferous punctation finer, but as dense as on pronotum; surface without microsculpture; polished. Abdomen with fine and sparse punctation; without microsculpture; polished; posterior margin of sternite VII of male sinuate with slightly prominent centre; posterior margin of tergite VII of male triangularly prominent; meso-tibia with 5, meta-tibia with 3 ctenidia. Aedeagus oval with sub-rectangular apical part; dorsal plate of central lobe covering nearly total central lobe; endophallus with several torsions, covered by long teeth; paramere short and thick; at apex, abruptly narrowed to acute top at inner edge; in basal part with few sensillae; at inner edge, near apex with few setae.</paragraph>
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<paragraph id="9ADE57A06C28404FDAB4E5C33E3F439E" pageId="0" pageNumber="13">
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<emphasis id="D61664641782085E548558CEABE005F0" bold="true" pageId="0" pageNumber="13">Etymology.</emphasis>
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The species name is derived from the same Latin word
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<emphasis id="D6302FDE1A53596707A962EA7577BCFF" italics="true" pageId="0" pageNumber="13">lituratus</emphasis>
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meaning
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<normalizedToken id="243FAAC91EA0391FF19AFB34C04625BC" originalValue="“spotted”">"spotted"</normalizedToken>
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and refers to the yellow spots on the elytra.
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</paragraph>
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</subSubSection>
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<subSubSection id="D5730B547D4B5B2418A9E2AB5DB85F19" pageId="0" pageNumber="13" type="geography">
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<paragraph id="C9E8960A0065C9AF696459B82FD503EC" pageId="0" pageNumber="13">Geography.</paragraph>
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<paragraph id="198ECECF7682EFF8A67565629A7AC8D1" pageId="0" pageNumber="13">Western Ecuador.</paragraph>
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</subSubSection>
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<subSubSection id="26025B41A54C26DDEE2487A22569994F" pageId="0" pageNumber="13" type="ecology">
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<paragraph id="9C4D1F73590CC5F8C7BB2AB8185931BB" pageId="0" pageNumber="13">Ecology.</paragraph>
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<paragraph id="8BE439BEFC322EA45C5E16DB8E0AE0E3" pageId="0" pageNumber="13">Montane forest and similar agricultural plantations at approximately 1600 m elevation from October to December.</paragraph>
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</subSubSection>
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</treatment>
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</document> |