480 lines
98 KiB
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480 lines
98 KiB
XML
<document ID-CLB-Dataset="121971" ID-DOI="http://dx.doi.org/10.3897/subtbiol.44.86914" ID-GBIF-Dataset="6dcce6d8-6a0c-4afd-bab2-ca640300ae07" ID-Pensoft-Pub="1314-2615-44-21" ID-Pensoft-UUID="27097103DFA85FF0954CAADE72F24005" ID-ZooBank="1865B08110724900BFC7563C0D29DA7D" ModsDocID="1314-2615-44-21" checkinTime="1660625097676" checkinUser="pensoft" docAuthor="Shintani, Aki, Lee, Chi-Woo & Tomikawa, Ko" docDate="2022" docId="A69D72015B56538EABEAA09A41282E03" docLanguage="en" docName="SubterBiol 44: 21-50" docOrigin="Subterranean Biology 44" docPubDate="2022-08-15" docSource="http://dx.doi.org/10.3897/subtbiol.44.86914" docTitle="Eoniphargus toriii Shintani & Lee & Tomikawa 2022, sp. nov." docType="treatment" docUuid="2A3DFB47-9FEC-46E4-A003-923CCD93E20D" docUuidSource="ZooBank" docVersion="5" id="27097103DFA85FF0954CAADE72F24005" lastPageNumber="21" masterDocId="27097103DFA85FF0954CAADE72F24005" masterDocTitle="Two new species add to the diversity of Eoniphargus in subterranean waters of Japan, with molecular phylogeny of the family Mesogammaridae (Crustacea, Amphipoda)" masterLastPageNumber="50" masterPageNumber="21" pageNumber="21" updateTime="1732798697937" updateUser="ExternalLinkService">
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<mods:titleInfo id="40F894ACCEE18E70EE2F9A8106EAD1AC">
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<mods:title id="724AE3ED0B2F830D266B89E39E41E134">Two new species add to the diversity of Eoniphargus in subterranean waters of Japan, with molecular phylogeny of the family Mesogammaridae (Crustacea, Amphipoda)</mods:title>
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<mods:name id="8000F9EBA8CA693B5B850B5BCCA24AC8" type="personal">
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<mods:roleTerm id="602ACB0F28133B22246E5E5745AB52A9">Author</mods:roleTerm>
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<mods:namePart id="1A6550A119705E9A72EFF5CBC91F526A">Shintani, Aki</mods:namePart>
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<mods:affiliation id="AB30FA578E12A7B2CFEED682384A1873">Graduate School of Humanities and Social Sciences, Hiroshima University, 1 - 1 - 1 Kagamiyama, Higashihiroshima, Hiroshima, 739 - 8524, Japan</mods:affiliation>
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<mods:namePart id="D77C9E22A594E0C3821CD9181E5188BD">Lee, Chi-Woo</mods:namePart>
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<mods:nameIdentifier id="A7B7180F9649FC7376052425EEAD0CB7" type="ORCID">https://orcid.org/0000-0002-0763-4271</mods:nameIdentifier>
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<mods:affiliation id="66F119AE155B2688DF4C6F55A8D5B87D">Nakdonggang National Institute of Biological Resources, 137, Donam 2 - gil, Sangju-si, Gyeongsangbuk-do, 37242, Republic of Korea</mods:affiliation>
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<mods:name id="DFED3BD38C1052540E756A4139A84EFC" type="personal">
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<mods:namePart id="EB3633E4FB99EF96D309343A25F2F95F">Tomikawa, Ko</mods:namePart>
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<mods:affiliation id="EC20FA489182FF7A0A173D77DAB8338A">Graduate School of Humanities and Social Sciences, Hiroshima University, 1 - 1 - 1 Kagamiyama, Higashihiroshima, Hiroshima, 739 - 8524, Japan</mods:affiliation>
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<mods:nameIdentifier id="882AF5DAD5986F7C521F92A873B026F7" type="email">tomikawa@hiroshima-u.ac.jp</mods:nameIdentifier>
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<mods:typeOfResource id="E372FAEA7E8662C95289811BF950931F">text</mods:typeOfResource>
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<mods:title id="1D73FF2589810413CCE0174E5C20D185">Subterranean Biology</mods:title>
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<mods:part id="7C7475310AC7CC0114E4F20C2D64779A">
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<mods:date id="C07A5B90E3DE64C8C917D581CEDA7C83">2022</mods:date>
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<mods:number id="098746A4D6F251554D66CC12E5E3A764">2022-08-15</mods:number>
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<mods:number id="A75699F933B2A0E2D1F5937CC6DA2ABD">44</mods:number>
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<treatment id="A69D72015B56538EABEAA09A41282E03" ID-GBIF-Taxon="199350556" LSID="urn:lsid:zoobank.org:act:2A3DFB47-9FEC-46E4-A003-923CCD93E20D" httpUri="http://treatment.plazi.org/id/A69D72015B56538EABEAA09A41282E03" lastPageNumber="21" pageId="0" pageNumber="21" scope_family="Mesogammaridae" scope_order="Amphipoda">
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<subSubSection id="468087D74B29916792214058DEF7DF8A" pageId="0" pageNumber="21" type="nomenclature">
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<paragraph id="C1ADCBA53F527DFB4625AE951A5AA1B5" pageId="0" pageNumber="21">
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<taxonomicName id="25A766A17F65A5D4F99CF88E891B2E98" ID-CoL="9GDMQ" LSID="https://zoobank.org/2A3DFB47-9FEC-46E4-A003-923CCD93E20D" authority="Shintani & Lee & Tomikawa, 2022" authorityName="Shintani & Lee & Tomikawa" authorityYear="2022" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eoniphargus toriii" order="Amphipoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="toriii" status="sp. nov.">Eoniphargus toriii</taxonomicName>
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<taxonomicNameLabel id="DA2BBBC5F1E3FFFC9A61A4B89FF2407E" pageId="0" pageNumber="21">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C9AF4C07CB4CF7A643F91E3A04224068" pageId="0" pageNumber="21" type="description">
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<paragraph id="DFFC61BED6D78C31D8E97348C7A51E0E" pageId="0" pageNumber="21">
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<figureCitation id="BC78D09C2E08C540E72A4BF3D3D4942E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">Figs 2</figureCitation>
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<figureCitation id="0D9DFDF6151304B7B6AB5303239DB050" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">, 3</figureCitation>
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<figureCitation id="87C37ECE1EC027F489FA269EB348F5B6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">, 4</figureCitation>
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<figureCitation id="7EF191BDFFB5D17C3EB821BB01765AAE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">, 5 New Japanese name: Torii-chikayokoebi</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="118864322623680953AEBEC559F638E9" pageId="0" pageNumber="21" type="reference_group">
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<paragraph id="38ABB5151DF9D2C151640AB1544E4840" pageId="0" pageNumber="21">
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<taxonomicName id="41C381A9BA5FDFE7599BED429F57443E" authority="Shintani & Lee & Tomikawa, 2022" authorityName="Shintani & Lee & Tomikawa" authorityYear="2022" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eoniphargus kojimai" order="Amphipoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="kojimai" status="sp. nov.">Eoniphargus kojimai</taxonomicName>
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:
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<bibRefCitation id="EDA9DE8B3FF8329A915424D0019A4A86" DOI="https://doi.org/10.1111/j.1096-3642.2007.00277.x" author="Tomikawa, K" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="21" pagination="643 - 670" refId="B37" refString="Tomikawa, K, Kobayashi, N, Morino, H, Mawatari, SF, 2007. New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda). Zoological Journal of the Linnean Society 149: 643 - 670, DOI: https://doi.org/10.1111/j.1096-3642.2007.00277.x" title="New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)." url="https://doi.org/10.1111/j.1096-3642.2007.00277.x" volume="149" year="2007">Tomikawa et al. 2007</bibRefCitation>
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, 647, figs 2-6.
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</paragraph>
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</subSubSection>
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<subSubSection id="B6225BFE3A269057EE101AC905A0025F" pageId="0" pageNumber="21" type="type material">
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<paragraph id="BA90171CE51900FAFCB88900F5BF31A0" pageId="0" pageNumber="21">Type material.</paragraph>
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<paragraph id="F9C10266FA4BC9CB7F3DE874180475BC" pageId="0" pageNumber="21">
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<emphasis id="941D143147479275C1ED20F71C6F6D7C" bold="true" italics="true" pageId="0" pageNumber="21">Holotype</emphasis>
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: ♂ 4.3 mm (NSMT-Cr 16652), Seto River (
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<geoCoordinate id="6143F686F78A1446E38C3D9603CD208C" degrees="34.880555" direction="north" orientation="latitude" precision="1" value="34.880554">34.880555°N</geoCoordinate>
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,
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<geoCoordinate id="4A3CE8C3BBCEBF02463BAD2AF58EA585" degrees="138.218888" direction="east" orientation="longitude" precision="1" value="138.21889">138.218888°E</geoCoordinate>
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), Terajima, Fujieda, Shizuoka Prefecture, Japan, collected by T. Torii on 3 June 2004.
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<emphasis id="8214F3AAEF28ED99382B8CD4708F4846" bold="true" italics="true" pageId="0" pageNumber="21">Paratypes</emphasis>
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:2 ♀♀, 4.6 mm and 4.3 mm (NSMT-Cr 16653 and 16654), data same as for the holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="A3401B584FDBF50F4B0B239D82B27813" pageId="0" pageNumber="21" type="diagnosis">
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<paragraph id="6B85CF4837BBBD97C4DC5A2BBDB59245" pageId="0" pageNumber="21">Diagnosis.</paragraph>
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<paragraph id="848500365F0BA3D50EC7059DCAA7A465" pageId="0" pageNumber="21">Urosomite 3 without dorsal setae. Epimeral plates 2-3 without ventral setae. Peduncular article 1 of antenna 1 with robust seta on posterodistal corner. Antenna 2 with peduncular article 2 gland cone not exceeding end of article 3; calceoli present in male. Mandible with 5-dentate left incisor; left lacinia mobilis 4-dentate, right one bifid with three or four teeth. Inner plate of maxilla 1 with six plumose setae. Inner plate of maxilla 2 with seven facial seta in oblique row. Peduncle of pleopod 3 with seta. Uropod 1 with peduncle bearing basofacial robust setae. Uropod 3 with inner ramus 0.3 times as long as outer ramus; outer ramus two-ariticulate, with plumose setae on medial margin. Telson length 1.1 times width, cleft for 67% of length.</paragraph>
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</subSubSection>
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<subSubSection id="800087FDFB3BB65BEC87E751EC7E6C6E" pageId="0" pageNumber="21" type="etymology">
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<paragraph id="74E118C9A1EBBA12BA04F88FAB6B458A" pageId="0" pageNumber="21">Etymology.</paragraph>
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<paragraph id="3328F3AC4D987F9AC9EC2B8D0BE2F43E" pageId="0" pageNumber="21">The new species is named after Dr. Takaaki Torii, who collected the species.</paragraph>
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</subSubSection>
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<subSubSection id="F37E8A738E2755C112F26ECFB099963E" pageId="0" pageNumber="21" type="description">
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<paragraph id="33CA6C0F275BE5CB58BEB5601E0CEAB2" pageId="0" pageNumber="21">Description.</paragraph>
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<paragraph id="835AB32FF23EF84FE2ABE295DD4F3DE5" pageId="0" pageNumber="21">
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<emphasis id="FA82CEC491FDDBEB1FEBDC466B3DE1A9" bold="true" pageId="0" pageNumber="21">Male holotype, NSMT-Cr 16652.</emphasis>
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Head (Fig.
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<figureCitation id="FAC6E49B6FD969174DB3B2B4B84241EB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2A</figureCitation>
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) as long as pereonites 1 and 2 combined; eyes absent; inferior antennal sinus distinct with rounded angle. Dorsal margins of pleonites 1-3 (Figs
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<figureCitation id="9DF43E527D13E8CB517DC1937C5C533A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2A</figureCitation>
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,
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<figureCitation id="EBB4293B6D0C75CA7E84E9DACA094F4A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5A-C</figureCitation>
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) each with four setae. Epimeral plates 1-3 (Figs
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<figureCitation id="867571FC74EDCAC5EB10885D1A3D71A9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2A</figureCitation>
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,
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|
<figureCitation id="01DC2D0A2193D1AF4F2E9956829CAD33" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5N-P</figureCitation>
|
|
) with weakly pointed posterodistal corners; ventral margins without setae; posterior margins of plates 1-3 with one, zero, one seta, respectively. Dorsal margins of urosomites 1 and 2 with robust seta, urosomite 3 dorsally bare.
|
|
</paragraph>
|
|
<caption id="73B9A247A4FA94D4DF0EB062BB0469D5" doi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21" start="Figure 2" startId="F2">
|
|
<paragraph id="7DEFCD447FA6B2D823E38D5BE6932100" pageId="0" pageNumber="21">
|
|
<emphasis id="D928B849A5B2B821B571A7D763A4CB4F" bold="true" pageId="0" pageNumber="21">Figure 2.</emphasis>
|
|
<taxonomicName id="B75BF1D5EAD364BECBACBC9D7FA3008A" authorityName="Shintani & Lee & Tomikawa" authorityYear="2022" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eoniphargus toriii" order="Amphipoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="toriii">
|
|
<emphasis id="E350BB8F22C41C1B419A78EF48459254" italics="true" pageId="0" pageNumber="21">Eoniphargus toriii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653)
|
|
<emphasis id="D91E9FC0C9C371510144075A7309BBC0" bold="true" pageId="0" pageNumber="21">A</emphasis>
|
|
habitus, lateral view
|
|
<emphasis id="AFA3C5F61B066278506DE3652F1821EE" bold="true" pageId="0" pageNumber="21">B</emphasis>
|
|
upper lip, anterior view
|
|
<emphasis id="C5EC7A153A46522611EE8B26A5607912" bold="true" pageId="0" pageNumber="21">C</emphasis>
|
|
left mandible, medial view
|
|
<emphasis id="F54E4F0A015FB238B2FF1D2338DE416F" bold="true" pageId="0" pageNumber="21">D</emphasis>
|
|
incisor and lacinia mobilis of left mandible, medial view
|
|
<emphasis id="9E08B04C75B81345B97FCC88C46E68E4" bold="true" pageId="0" pageNumber="21">E</emphasis>
|
|
and
|
|
<emphasis id="80968C2ED27F021247CA149A4EE2B3D0" bold="true" pageId="0" pageNumber="21">F</emphasis>
|
|
incisor and lacinia mobilis of right mandible, medial view
|
|
<emphasis id="81335C45E7B00FC2496E822004AF060B" bold="true" pageId="0" pageNumber="21">G</emphasis>
|
|
lower lip, anterior view
|
|
<emphasis id="DA45F8B0CF51DB0501453AABE5446235" bold="true" pageId="0" pageNumber="21">H</emphasis>
|
|
maxilla 1, dorsal view
|
|
<emphasis id="43EEAB49FD88D29286A7B5F0893E3BE3" bold="true" pageId="0" pageNumber="21">I</emphasis>
|
|
outer plate of maxilla 1, dorsal view
|
|
<emphasis id="8002C82ADB7B538EFCB4B1AA061D0768" bold="true" pageId="0" pageNumber="21">J</emphasis>
|
|
palp of maxilla 1, dorsal view
|
|
<emphasis id="8B0DA8FB7024B8D8F3E444BB3B2FC83A" bold="true" pageId="0" pageNumber="21">K</emphasis>
|
|
maxilla 2, dorsal view
|
|
<emphasis id="6C3848F21580EA3E736B1A69C2C8BD39" bold="true" pageId="0" pageNumber="21">L</emphasis>
|
|
maxilliped, dorsal view
|
|
<emphasis id="3F2BE78CDBBCFD159D144AFF5A0E305E" bold="true" pageId="0" pageNumber="21">M</emphasis>
|
|
palp article 4 of maxilliped, dorsal view
|
|
<emphasis id="9C4B0850075F81C4BBA0FB43D0E0DD81" bold="true" pageId="0" pageNumber="21">N</emphasis>
|
|
outer plate of maxilliped, dorsal view
|
|
<emphasis id="F4E4DE360A872C37D0D8E85D7AA05A56" bold="true" pageId="0" pageNumber="21">O, P</emphasis>
|
|
inner plate of maxilliped, dorsal view
|
|
<emphasis id="D222B17A74A49F4965D14A39A7F767C4" bold="true" pageId="0" pageNumber="21">F, P</emphasis>
|
|
female; others, male. Modified from
|
|
<bibRefCitation id="26ED16F6976D459575180291E8CB1B34" DOI="https://doi.org/10.1111/j.1096-3642.2007.00277.x" author="Tomikawa, K" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="21" pagination="643 - 670" refId="B37" refString="Tomikawa, K, Kobayashi, N, Morino, H, Mawatari, SF, 2007. New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda). Zoological Journal of the Linnean Society 149: 643 - 670, DOI: https://doi.org/10.1111/j.1096-3642.2007.00277.x" title="New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)." url="https://doi.org/10.1111/j.1096-3642.2007.00277.x" volume="149" year="2007">Tomikawa et al. (2007)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="0E08E0510AE54C43C3C3E301B4FF5C7F" pageId="0" pageNumber="21">
|
|
Antenna 1 (Figs
|
|
<figureCitation id="5F00848FCDA106C6E3A5F0EEA1BE4B96" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2A</figureCitation>
|
|
,
|
|
<figureCitation id="7DDD60E479C03C976611F2237863BF13" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3A</figureCitation>
|
|
) length 0.6 times as long as body; length ratio of peduncular articles 1-3 in 1.0: 0.8: 0.4; posterodistal corner of peduncular article 1 with robust seta; accessory flagellum three-articulate (Fig.
|
|
<figureCitation id="AA9199CAEA3124627BF739595B433343" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3B</figureCitation>
|
|
); primary flagellum with approximately 22 articles, each article with one aesthetasc (Fig.
|
|
<figureCitation id="DE9D94B8B7401A355D4C9E1EA3DC2F5B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3C</figureCitation>
|
|
). Antenna 2 (Figs
|
|
<figureCitation id="F867FB05248B70A656F7C7CC4D4B5CF1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2A</figureCitation>
|
|
,
|
|
<figureCitation id="8D3BF6D175C417044BBB77858CF04BDF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3D</figureCitation>
|
|
) length 0.6 times as long as antenna 1; peduncular article 4 length 0.9 times as long as article 5; right peduncular article 5 with calceoli, left one lacking calceoli; flagellum 16-articulate, with calceoli (Fig.
|
|
<figureCitation id="3BFAE2FA22FC30F8C9E3495BB2AD232B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="73B193D4C0152356A54393BE36188761" doi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21" start="Figure 3" startId="F3">
|
|
<paragraph id="C3D3F780104B2640949D792A892F4DAC" pageId="0" pageNumber="21">
|
|
<emphasis id="C024A8FA723F7E373610A14A091B8ED9" bold="true" pageId="0" pageNumber="21">Figure 3.</emphasis>
|
|
<taxonomicName id="D073F5A7BFB143691A836CBA86CDECA2" authorityName="Shintani & Lee & Tomikawa" authorityYear="2022" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eoniphargus toriii" order="Amphipoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="toriii">
|
|
<emphasis id="EEEC6D804614F4A15CF4425CC0392412" italics="true" pageId="0" pageNumber="21">Eoniphargus toriii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653)
|
|
<emphasis id="0D68C2A869F5B6155923C3512692A15F" bold="true" pageId="0" pageNumber="21">A</emphasis>
|
|
antenna 1, medial view
|
|
<emphasis id="79E155B96515BAF2C977F317DD76190D" bold="true" pageId="0" pageNumber="21">B</emphasis>
|
|
accessory flagellum of antenna 1, medial view
|
|
<emphasis id="DC551981F864D3BA216E0FA17E49C802" bold="true" pageId="0" pageNumber="21">C</emphasis>
|
|
aesthetasc on flagellar article of antenna 1, medial view
|
|
<emphasis id="92281CC8B4D2F8E53190459235FF3C1B" bold="true" pageId="0" pageNumber="21">D</emphasis>
|
|
antenna 2 (some flagellar articles are omitted), medial view
|
|
<emphasis id="E1041169D13EBE1FCD4474D5E86DE587" bold="true" pageId="0" pageNumber="21">E</emphasis>
|
|
calceolus on flagellar article of antenna 2, medial view
|
|
<emphasis id="72B527694343C3444FCE9EF5029E7A5E" bold="true" pageId="0" pageNumber="21">F</emphasis>
|
|
antenna 1 (female, some flagellar articles are omitted), medial view
|
|
<emphasis id="C272DC5FA61F706C0573A5A45FFA1E88" bold="true" pageId="0" pageNumber="21">G</emphasis>
|
|
accessory flagellum of antenna 1 (female), medial view
|
|
<emphasis id="A859B3F7D9079B245938896818526E44" bold="true" pageId="0" pageNumber="21">H</emphasis>
|
|
aesthetasc on flagellar article of antenna 1 (female), medial view
|
|
<emphasis id="3146D6B34198953D7CFFA49904144FC4" bold="true" pageId="0" pageNumber="21">I</emphasis>
|
|
antenna 2 (female, some flagellar articles are omitted), medial view
|
|
<emphasis id="DA536421CA4F0FC87EFDF3216336BEA7" bold="true" pageId="0" pageNumber="21">J</emphasis>
|
|
gnathopod 1, lateral view
|
|
<emphasis id="18ABDE3A083C2F2D64B453C05A9B9A50" bold="true" pageId="0" pageNumber="21">K</emphasis>
|
|
palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view
|
|
<emphasis id="CF73739B6E402779F0FBD3ABEB09DE63" bold="true" pageId="0" pageNumber="21">L</emphasis>
|
|
gnathopod 2, lateral view
|
|
<emphasis id="F6FD16C117A2BA432FCD81D49BEC92DF" bold="true" pageId="0" pageNumber="21">M</emphasis>
|
|
palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view
|
|
<emphasis id="418B40937ECCF9CEBFCCFB3853752F9F" bold="true" pageId="0" pageNumber="21">N</emphasis>
|
|
gnathopod 1 (female), lateral view
|
|
<emphasis id="FB47F4C4685CC74916BBDDEEBCE32540" bold="true" pageId="0" pageNumber="21">O</emphasis>
|
|
palmar margin and dactylus of gnathopod 1 (female), lateral view
|
|
<emphasis id="BC421EEF66079C4CAB7294F02CFCBBB0" bold="true" pageId="0" pageNumber="21">P</emphasis>
|
|
gnathopod 2 (female), lateral view
|
|
<emphasis id="C49CA76E97DD51FE04EE7ABC2C017CED" bold="true" pageId="0" pageNumber="21">Q</emphasis>
|
|
palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view
|
|
<emphasis id="042BAF4ABDE9C2712B922551ABA5F0A4" bold="true" pageId="0" pageNumber="21">F-H, N-Q</emphasis>
|
|
female; others, male. Modified from
|
|
<bibRefCitation id="E03C3AD53F575334DEBDCA3976D683CE" DOI="https://doi.org/10.1111/j.1096-3642.2007.00277.x" author="Tomikawa, K" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="21" pagination="643 - 670" refId="B37" refString="Tomikawa, K, Kobayashi, N, Morino, H, Mawatari, SF, 2007. New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda). Zoological Journal of the Linnean Society 149: 643 - 670, DOI: https://doi.org/10.1111/j.1096-3642.2007.00277.x" title="New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)." url="https://doi.org/10.1111/j.1096-3642.2007.00277.x" volume="149" year="2007">Tomikawa et al. (2007)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="C34032B58781B21F3E420D829859156E" pageId="0" pageNumber="21">
|
|
Upper lip (Fig.
|
|
<figureCitation id="63930B83FC904FD14C643AEE4C91C813" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2B</figureCitation>
|
|
) with rounded apical margin bearing fine setae. Mandibles (Fig.
|
|
<figureCitation id="77A0943FDEA56D72CA70DCB891ED65BC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2C-E</figureCitation>
|
|
) with left and right incisors comprising five and five-six teeth, respectively; left lacinia mobilis comprising four teeth, right lacinia mobilis bifid with three teeth; molar process triturative with plumose seta; length ratio of palp articles 1-3 in 1.0: 2.4: 1.9; palp article 1 without setae; palp article 2 with eight marginal setae; palp article 3 with pair of A-setae, many D-setae and E-setae, lateral face with fine setae. Lower lip (Fig.
|
|
<figureCitation id="3F0A643C91A2E5AD770CDFB192CEDA3F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2G</figureCitation>
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) lacking inner lobes; outer lobes broad, shoulder rounded, with fine setae. Maxilla 1 (Fig.
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|
<figureCitation id="E1B121422F0F32EFA2D6F897F2EE0668" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2H-J</figureCitation>
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) with triangular inner lobe, bearing six plumose setae on medial margin; outer plate rectangular, with 11 serrate robust setae apically; palp 2-articulate, article 1 without setae, article 2 with six robust setae apically. Maxilla 2 (Fig.
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|
<figureCitation id="26AB5F0CC0B8975E64F0CAEA8193744A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2K</figureCitation>
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) with inner plate bearing seven plumose setae in oblique row. Maxilliped (Fig.
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<figureCitation id="6E8B296B2FF3B521768A659AF1154397" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2L-O</figureCitation>
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) with inner plate reaching end of palp article 1, subquadrate, bearing three subapical robust setae and medial robust seta; outer plate ovate, reaching middle of palp article 2, with row of robust setae along apical to medial margins; palp 4-articulate, article 2 longest with medial setae, nail of article 4 distinct.
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</paragraph>
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<paragraph id="2D986296C8BB57DFE193065EFB83FAEE" pageId="0" pageNumber="21">
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Gnathopod 1 (Fig.
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<figureCitation id="5BBCFF932B03AA953C1AB880CB74C973" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3J, K</figureCitation>
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) with subquadrate coxa bearing setae along anterior to ventral margins; anterior and posterior margins with long setae; carpus length 0.9 times as long as length of propodus and 1.5 times width of carpus; propodus length 1.9 times width, palmar margin weakly serrate, oblique, with three medial and two lateral robust setae; dactylus not reaching posterodistal corner of propodus. Gnathopod 2 (Fig.
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<figureCitation id="715CD4AAA7F197231F09507C97452A1C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3L, M</figureCitation>
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) with coxa expanded proximally bearing setae along anterior to ventral margins; posterior margin of basis with long setae; carpus length 1.2 times as long as length of propodus and 2.8 times width of carpus; propodus length 2.4 times width, palmar margin weakly serrate, almost vertical, with two medial and two lateral robust setae; dactylus not reaching posterodistal corner of propodus.
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</paragraph>
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<paragraph id="915F145353E9973AE44279B91C58C891" pageId="0" pageNumber="21">
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Pereopod 3 (Fig.
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<figureCitation id="7209DB1C8993D0847D5308300F1BFD89" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">4A, B</figureCitation>
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) with subquadrate coxa, proximally expanded, bearing setae along anterior to ventral margins; posterior margin of basis with long setae. Pereopod 4 (Fig.
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|
<figureCitation id="6F85235F353EF02A613483DCA3DA36F3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">4D, E</figureCitation>
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) with coxa bearing setae along anterior to ventral margins, posteroproximally concave; posterior margin of basis with long setae. Pereopod 5 (Fig.
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<figureCitation id="F42D43E2035BDC1F24D6F0350BB3266A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">4G, H</figureCitation>
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) with bilobed coxa bearing seta on anterior and posterior lobes; anterior margin of basis with robust setae, posterodistal corner weakly lobate. Pereopod 6 (Fig.
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<figureCitation id="0BF7ACA477E5FACC4F72261D202A9EB5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">4J, K</figureCitation>
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) with bilobed coxa bearing seta on posterior lobe; basis ovate, with robust setae on anterior margin, posterodistal corner weakly lobate. Pereopod 7 (Fig.
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<figureCitation id="899BED7507F8840860DAB2F47F2B9F53" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">4M</figureCitation>
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) with subtriangular coxa bearing seta on posterior margin; basis elliptical, with robust setae on anterior margin, posterodistal corner weakly lobate.
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</paragraph>
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<caption id="1E074873B38A1E70E72A4FDA05432F2F" doi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21" start="Figure 4" startId="F4">
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<paragraph id="9D46322989E60869CD3F13C9088B2B5B" pageId="0" pageNumber="21">
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<emphasis id="1034F3BBE9BAEF164DA369C74002E6C0" bold="true" pageId="0" pageNumber="21">Figure 4.</emphasis>
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<taxonomicName id="046C21511B07E10F7402531914BB56AF" authorityName="Shintani & Lee & Tomikawa" authorityYear="2022" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eoniphargus toriii" order="Amphipoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="toriii">
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<emphasis id="247BCB8A56FF2DA779BDDB43FDCAEC6B" italics="true" pageId="0" pageNumber="21">Eoniphargus toriii</emphasis>
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</taxonomicName>
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sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653)
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<emphasis id="96A7A257366BC193F114C0753795B20A" bold="true" pageId="0" pageNumber="21">A</emphasis>
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pereopod 3, lateral view
|
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<emphasis id="55AEA725E13F653E3BB08E80E3B2F508" bold="true" pageId="0" pageNumber="21">B</emphasis>
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dactylus of pereopod 3, lateral view
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<emphasis id="F59C089A983DEE0A76E2E1AB0D542701" bold="true" pageId="0" pageNumber="21">C</emphasis>
|
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coxa of pereopod 3 (female), lateral view
|
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<emphasis id="7DCE88AB9A7738160A046BFE226A01F2" bold="true" pageId="0" pageNumber="21">D</emphasis>
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pereopod 4, lateral view
|
|
<emphasis id="BDC153BDDD1DD81554F151969ABA39C5" bold="true" pageId="0" pageNumber="21">E</emphasis>
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|
dactylus of pereopod 4, lateral view
|
|
<emphasis id="DA279CF7C96D6AE11B9BA5C76F12F012" bold="true" pageId="0" pageNumber="21">F</emphasis>
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coxa of pereopod 4 (female), lateral view
|
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<emphasis id="1B52D8A6C25AD2ECA20C49661336684A" bold="true" pageId="0" pageNumber="21">G</emphasis>
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pereopod 5, lateral view
|
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<emphasis id="A6698BEB6BC508371DCA04AB61150915" bold="true" pageId="0" pageNumber="21">H</emphasis>
|
|
dactylus of pereopod 5, lateral view
|
|
<emphasis id="DBEB8153407951AE06DFD3526B06CDD1" bold="true" pageId="0" pageNumber="21">I</emphasis>
|
|
coxa-merus of pereopod 5 (female), lateral view
|
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<emphasis id="0F73D22BBF54F9091FF3BE015A25E991" bold="true" pageId="0" pageNumber="21">J</emphasis>
|
|
pereopod 6, lateral view
|
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<emphasis id="2DBD9C9B5B45CC9BC23A4813550F8305" bold="true" pageId="0" pageNumber="21">K</emphasis>
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dactylus of pereopod 6, lateral view
|
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<emphasis id="AF5A6EE165090CAC172F6D44EA8B1F17" bold="true" pageId="0" pageNumber="21">L</emphasis>
|
|
coxa-merus of pereopod 6 (female), lateral view
|
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<emphasis id="E7A4683A58D3B43A6F5766EBA2E98643" bold="true" pageId="0" pageNumber="21">M</emphasis>
|
|
coxa-merus of pereopod 7, lateral view
|
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<emphasis id="58C6D04A554A6755D27FF4B92A26AEE4" bold="true" pageId="0" pageNumber="21">N</emphasis>
|
|
coxa-merus of pereopod 7 (female), lateral view
|
|
<emphasis id="37096E1C047FB35E3B3DB127FC187CFF" bold="true" pageId="0" pageNumber="21">C, F, I, L, N</emphasis>
|
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female; others, male. Modified from
|
|
<bibRefCitation id="AE9D3802EACFB7ECAC9B0E149B2BF643" DOI="https://doi.org/10.1111/j.1096-3642.2007.00277.x" author="Tomikawa, K" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="21" pagination="643 - 670" refId="B37" refString="Tomikawa, K, Kobayashi, N, Morino, H, Mawatari, SF, 2007. New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda). Zoological Journal of the Linnean Society 149: 643 - 670, DOI: https://doi.org/10.1111/j.1096-3642.2007.00277.x" title="New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)." url="https://doi.org/10.1111/j.1096-3642.2007.00277.x" volume="149" year="2007">Tomikawa et al. (2007)</bibRefCitation>
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.
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</paragraph>
|
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</caption>
|
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<paragraph id="D09D0DEFE2D6C112763AF22B2184DF53" pageId="0" pageNumber="21">
|
|
Coxal gills (Figs
|
|
<figureCitation id="4508101888E0D64F885494266B8A9584" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3L</figureCitation>
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,
|
|
<figureCitation id="00E4CBF37B79C369AE681CFC321D5EAB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">4A, D, G, J</figureCitation>
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) ovate with stalks on gnathopod 2 and pereopods 3-6.
|
|
</paragraph>
|
|
<paragraph id="48CBE9E2A216AAEEE482CE8B2A1356A3" pageId="0" pageNumber="21">
|
|
Pleopods 1-3 (Fig.
|
|
<figureCitation id="39973E191DF1C42DC050E856A5B4989E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5G-K</figureCitation>
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) with peduncles longer than rami, bearing distal seta on pleopod 2 and proximal seta on pleopod 3; retinacula paired (Fig.
|
|
<figureCitation id="E9ED1F9D248A80D21493485E44494088" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5H</figureCitation>
|
|
); mediobasal margin of inner ramus with bifid plumose setae (Fig.
|
|
<figureCitation id="F8D8B05CC67AF87E8A94D23CF30A66E2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5I</figureCitation>
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); rami well developed.
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|
</paragraph>
|
|
<caption id="273BD96207BF1C4331BD741916155D6A" doi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21" start="Figure 5" startId="F5">
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|
<paragraph id="06B951C690772308EA9FFAF0E1EA47FA" pageId="0" pageNumber="21">
|
|
<emphasis id="D41039413E4F66696021D586F32B4F26" bold="true" pageId="0" pageNumber="21">Figure 5.</emphasis>
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|
<taxonomicName id="8011873B994D00D20A30EB1386AA64D2" authorityName="Shintani & Lee & Tomikawa" authorityYear="2022" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eoniphargus toriii" order="Amphipoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="toriii">
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<emphasis id="49A4385F1E8CE292211050566B7447A4" italics="true" pageId="0" pageNumber="21">Eoniphargus toriii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653)
|
|
<emphasis id="A9985C1B1663736EBFA2FAEE6422EBDB" bold="true" pageId="0" pageNumber="21">A-C</emphasis>
|
|
dorsal margins of pleonites 1-3, dorsal views
|
|
<emphasis id="DAE89CA1D7E2E80B4D2CE9FEE86BD671" bold="true" pageId="0" pageNumber="21">D-F</emphasis>
|
|
dorsal margins of urosomites 1-3, dorsal views
|
|
<emphasis id="58D04CD3643C63CD49DF2EE6E7FCC15E" bold="true" pageId="0" pageNumber="21">G</emphasis>
|
|
pleopod 1 (plumose setae on rami are omitted), medial view
|
|
<emphasis id="B0A073B5810325C316EFCFB2C13B76B0" bold="true" pageId="0" pageNumber="21">H</emphasis>
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|
retinacula of pleopod 1, medial view
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|
<emphasis id="5DEC381C7E4A45514F8066513A99461C" bold="true" pageId="0" pageNumber="21">I</emphasis>
|
|
bifid setae of pleopod 1, medial view
|
|
<emphasis id="205E00C2569BF8744081DEA8A32E79E5" bold="true" pageId="0" pageNumber="21">J</emphasis>
|
|
pleopod 2 (plumose setae on rami are omitted), medial view
|
|
<emphasis id="1B4782A975BC55A651FD0FE79BEE39A3" bold="true" pageId="0" pageNumber="21">K</emphasis>
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|
pleopod 3 (plumose setae on rami are omitted), medial view
|
|
<emphasis id="A5B1E4BD59C2B1849C5AB312499A9B37" bold="true" pageId="0" pageNumber="21">L</emphasis>
|
|
pleopod 1 (female, rami are omitted), medial view
|
|
<emphasis id="F31C67AD850DDA90F947C4E8E6A5A86D" bold="true" pageId="0" pageNumber="21">M</emphasis>
|
|
pleopod 2 (female, rami are omitted), lateral view
|
|
<emphasis id="E6341163A31B9FB85D4D35A68BA2AE03" bold="true" pageId="0" pageNumber="21">N-P</emphasis>
|
|
epimeral plates 1-3, lateral views
|
|
<emphasis id="702CB39B556340872C36E5528BFD3F25" bold="true" pageId="0" pageNumber="21">Q</emphasis>
|
|
epimeral plate 2 (female), lateral view
|
|
<emphasis id="D87E4579D88FA01A87DA4F7AF413F692" bold="true" pageId="0" pageNumber="21">R</emphasis>
|
|
uropod 1, dorsal view
|
|
<emphasis id="2A41F02895DE563A4DF8F5A58CE22500" bold="true" pageId="0" pageNumber="21">S</emphasis>
|
|
uropod 2, dorsal view
|
|
<emphasis id="CD685ED5BF75044D97148BE048F44A66" bold="true" pageId="0" pageNumber="21">T</emphasis>
|
|
uropod 2 (female), dorsal view
|
|
<emphasis id="0379DA9949E1A5EA0DA9271C25346E48" bold="true" pageId="0" pageNumber="21">U</emphasis>
|
|
uropod 3, ventral view
|
|
<emphasis id="98AB7B26F031EF436212314D57CC2933" bold="true" pageId="0" pageNumber="21">V</emphasis>
|
|
terminal article of outer ramus of uropod 3, ventral view
|
|
<emphasis id="9B6E3D4973462DBB74EFB37B1E4877FC" bold="true" pageId="0" pageNumber="21">W</emphasis>
|
|
uropod 3 (female), ventral view
|
|
<emphasis id="55CB2087F04E89548F019A3F9B973F75" bold="true" pageId="0" pageNumber="21">X</emphasis>
|
|
telson, dorsal view
|
|
<emphasis id="152C857FFD91F7D422B1F551FD9BBDA3" bold="true" pageId="0" pageNumber="21">Y</emphasis>
|
|
telson (female), dorsal view
|
|
<emphasis id="8660E6CB37FBACBC20C2D728917503B4" bold="true" pageId="0" pageNumber="21">L, M, Q, T, W, Y</emphasis>
|
|
female; others, male. Modified from
|
|
<bibRefCitation id="8D04AE800F0B537BDEE165CC84B4FA39" DOI="https://doi.org/10.1111/j.1096-3642.2007.00277.x" author="Tomikawa, K" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="21" pagination="643 - 670" refId="B37" refString="Tomikawa, K, Kobayashi, N, Morino, H, Mawatari, SF, 2007. New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda). Zoological Journal of the Linnean Society 149: 643 - 670, DOI: https://doi.org/10.1111/j.1096-3642.2007.00277.x" title="New gammaroid family, genera and species from subterranean waters of Japan, and their phylogenetic relationships (Crustacea: Amphipoda)." url="https://doi.org/10.1111/j.1096-3642.2007.00277.x" volume="149" year="2007">Tomikawa et al. (2007)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="86A35A9AE80FEB7DEC1BE260D10A5F62" pageId="0" pageNumber="21">
|
|
Uropod 1 (Fig.
|
|
<figureCitation id="DB235AE724DEAFFAF45BE9063744A543" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5R</figureCitation>
|
|
) with peduncle bearing three basofacial robust setae and dorsal robust setae; inner ramus length 0.7 times as long as peduncle, with two medial and one lateral robust setae, and ventroproximal robust seta; outer ramus length 0.9 times as long as inner ramus, with two robust setae on medial margin. Uropod 2 (Fig.
|
|
<figureCitation id="9562134735730FA10184EF173D939461" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5S</figureCitation>
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|
) with peduncle bearing dorsal robust setae; inner ramus length 0.8 times as long as peduncle, bearing two medial robust setae and two ventroproximal slender setae; outer ramus length 0.6 times as long as inner ramus, without marginal setae. Uropod 3 (Fig.
|
|
<figureCitation id="801235449F195A82FDD311DDDC701D08" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5U, V</figureCitation>
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|
) with peduncle length 0.3 times as long as outer ramus, bearing bent robust setae on distal edge; inner ramus length 0.3 times as long as outer ramus, with two robust setae on medial margin; outer ramus 2-articulate, proximal article with plumose setae on medial margin and robust setae on medial and lateral margins, terminal article length 0.3 times as long as proximal article with subapical setae.
|
|
</paragraph>
|
|
<paragraph id="EC3B3470CF0A76E658809C4AE4AAD02F" pageId="0" pageNumber="21">
|
|
Telson (Fig.
|
|
<figureCitation id="D7F4F0F9A1B29FE012E1464B30560EEC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5X</figureCitation>
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|
) length 1.1 times width, each lobe with two lateral and three apical robust setae and two dorsolateral penicillate setae, cleft for 67%.
|
|
</paragraph>
|
|
<paragraph id="A52ED463C301EACC498A14C35FC81DB3" pageId="0" pageNumber="21">
|
|
<emphasis id="F1AC84636F70131C8215ED463AC2EEF0" bold="true" pageId="0" pageNumber="21">Female paratype, NSMT-Cr 16653.</emphasis>
|
|
Antenna 1 (Fig.
|
|
<figureCitation id="6A860040A8A4D603A39F6518CD74A6AE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3F-H</figureCitation>
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|
) with peduncles 1-3 of which length ratio in 1.0: 0.8: 0.5; accessory flagellum 4-articulate (Fig.
|
|
<figureCitation id="A6C7753EA3DBF9FE85270F1E00178F58" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3G</figureCitation>
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). Antenna 2 (Fig.
|
|
<figureCitation id="2E1156E81394F83F6A5090ABB72275D4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3I</figureCitation>
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) without calceoli.
|
|
</paragraph>
|
|
<paragraph id="D1465054F10B20749C3A116781702972" pageId="0" pageNumber="21">
|
|
Incisor of right mandible 6-dentate (Fig.
|
|
<figureCitation id="64B273A89581F97889732EB578D32A82" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2F</figureCitation>
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); right lacinia mobilis 4-dentate, weakly bifid. Apical robust setae of inner plate of maxilliped stiffer than those of male (Fig.
|
|
<figureCitation id="BC32A571467B93E07D31775C40F16880" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A habitus, lateral view B upper lip, anterior view C left mandible, medial view D incisor and lacinia mobilis of left mandible, medial view E and F incisor and lacinia mobilis of right mandible, medial view G lower lip, anterior view H maxilla 1, dorsal view I outer plate of maxilla 1, dorsal view J palp of maxilla 1, dorsal view K maxilla 2, dorsal view L maxilliped, dorsal view M palp article 4 of maxilliped, dorsal view N outer plate of maxilliped, dorsal view O, P inner plate of maxilliped, dorsal view F, P female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure2" httpUri="https://binary.pensoft.net/fig/729385" pageId="0" pageNumber="21">2P</figureCitation>
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|
).
|
|
</paragraph>
|
|
<paragraph id="8343A1AED34F7D7E0B5D984FFF5E4F21" pageId="0" pageNumber="21">
|
|
Gnathopod 1 (Fig.
|
|
<figureCitation id="2C0D208E85C90740C417CFD6BF4F47CB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3N, O</figureCitation>
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|
) with carpus length 1.8 times width; length of propodus 2.0 times width; dactylus reaching posterodistal corner of propodus. Gnathopod 2 (Fig.
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<figureCitation id="B9C8913D72AF99029EBCF085F26C4324" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3P, Q</figureCitation>
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) with dactylus reaching posterodistal corner of propodus.
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</paragraph>
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<paragraph id="9C1099532A6BE149D1A039B4A785D4D6" pageId="0" pageNumber="21">
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Brood plates (Figs
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<figureCitation id="2A0107ADE532C8F578FA8F659F35E9B0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A antenna 1, medial view B accessory flagellum of antenna 1, medial view C aesthetasc on flagellar article of antenna 1, medial view D antenna 2 (some flagellar articles are omitted), medial view E calceolus on flagellar article of antenna 2, medial view F antenna 1 (female, some flagellar articles are omitted), medial view G accessory flagellum of antenna 1 (female), medial view H aesthetasc on flagellar article of antenna 1 (female), medial view I antenna 2 (female, some flagellar articles are omitted), medial view J gnathopod 1, lateral view K palmar margin and dactylus of gnathopod 1 (some setae are omitted), lateral view L gnathopod 2, lateral view M palmar margin and dactylus of gnathopod 2 (some setae are omitted), lateral view N gnathopod 1 (female), lateral view O palmar margin and dactylus of gnathopod 1 (female), lateral view P gnathopod 2 (female), lateral view Q palmar margin and dactylus of gnathopod 2 (female, some setae are omitted), lateral view F-H, N-Q female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure3" httpUri="https://binary.pensoft.net/fig/729386" pageId="0" pageNumber="21">3P</figureCitation>
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,
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<figureCitation id="8D4131D1895B355984B827D59DC926B6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A pereopod 3, lateral view B dactylus of pereopod 3, lateral view C coxa of pereopod 3 (female), lateral view D pereopod 4, lateral view E dactylus of pereopod 4, lateral view F coxa of pereopod 4 (female), lateral view G pereopod 5, lateral view H dactylus of pereopod 5, lateral view I coxa-merus of pereopod 5 (female), lateral view J pereopod 6, lateral view K dactylus of pereopod 6, lateral view L coxa-merus of pereopod 6 (female), lateral view M coxa-merus of pereopod 7, lateral view N coxa-merus of pereopod 7 (female), lateral view C, F, I, L, N female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure4" httpUri="https://binary.pensoft.net/fig/729387" pageId="0" pageNumber="21">4I</figureCitation>
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) narrow, lacking setae, on gnathopod 2 and pereopods 3-5.
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</paragraph>
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<paragraph id="1642C32255CBCD55E23B991C38E28DE3" pageId="0" pageNumber="21">
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Peduncles of pleopods 1 and 3 with three and one setae, respectively (Fig.
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<figureCitation id="045DDD7E8DAD2855BC2B4D95E6AC1B8E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5L, M</figureCitation>
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).
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</paragraph>
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<paragraph id="C5A80B133DC60858F7ECCA0B43ABADC9" pageId="0" pageNumber="21">
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Uropod 2 (Fig.
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<figureCitation id="9391E980172A8264CD5CDCC35B6DAD2F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5T</figureCitation>
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) with inner ramus bearing two medial and one lateral robust setae; outer ramus with medial robust seta. Uropod 3 (Fig.
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<figureCitation id="1AA4A691663142FF72E7C0EE3FA69CE6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5W</figureCitation>
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) with terminal article of outer ramus length 0.2 times as long as proximal article.
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</paragraph>
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<paragraph id="8D105F315FCE6E995D177E286087EDDB" pageId="0" pageNumber="21">
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Telson (Fig.
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|
<figureCitation id="18CE31515AC1F8087E2D4FF9E84F2F72" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eoniphargus toriii sp. nov., male holotype (NSMT-Cr 16652), female paratype (NSMT-Cr 16653) A-C dorsal margins of pleonites 1 - 3, dorsal views D-F dorsal margins of urosomites 1 - 3, dorsal views G pleopod 1 (plumose setae on rami are omitted), medial view H retinacula of pleopod 1, medial view I bifid setae of pleopod 1, medial view J pleopod 2 (plumose setae on rami are omitted), medial view K pleopod 3 (plumose setae on rami are omitted), medial view L pleopod 1 (female, rami are omitted), medial view M pleopod 2 (female, rami are omitted), lateral view N-P epimeral plates 1 - 3, lateral views Q epimeral plate 2 (female), lateral view R uropod 1, dorsal view S uropod 2, dorsal view T uropod 2 (female), dorsal view U uropod 3, ventral view V terminal article of outer ramus of uropod 3, ventral view W uropod 3 (female), ventral view X telson, dorsal view Y telson (female), dorsal view L, M, Q, T, W, Y female; others, male. Modified from Tomikawa et al. (2007)." figureDoi="10.3897/subtbiol.44.86914.figure5" httpUri="https://binary.pensoft.net/fig/729388" pageId="0" pageNumber="21">5Y</figureCitation>
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) as long as wide, cleft for 71%.
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</paragraph>
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</subSubSection>
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|
<subSubSection id="3B7359EF0D44A972476FAD8BD36FFE31" pageId="0" pageNumber="21" type="distribution">
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<paragraph id="D71B08799321864BEB2D9E049DBDDF2F" pageId="0" pageNumber="21">Distribution and environment.</paragraph>
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<paragraph id="3CF5A929D3559F4983B5623B24868D34" pageId="0" pageNumber="21">The species is known only from its type locality in the Seto River, Shizuoka Prefecture, Japan. Specimens were collected at a depth of 20 cm, from the bank of the river.</paragraph>
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</subSubSection>
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<subSubSection id="9B5B4832D7B9EB7350A920B3CC8E1AEA" pageId="0" pageNumber="21" type="remarks">
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<paragraph id="5E7E57B7CC453BA031822D1A123DB37F" pageId="0" pageNumber="21">Remarks.</paragraph>
|
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<paragraph id="1C129303A6ECE821E126B8A9698BB19B" pageId="0" pageNumber="21">
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|
<taxonomicName id="25A49BE89FC6AE7305F778BAD5F88A2D" authorityName="Shintani & Lee & Tomikawa" authorityYear="2022" class="Malacostraca" family="Mesogammaridae" genus="Eoniphargus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eoniphargus toriii" order="Amphipoda" pageId="0" pageNumber="21" phylum="Arthropoda" rank="species" species="toriii">
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<emphasis id="9385F170EC2D6A563BD83533BA733A94" italics="true" pageId="0" pageNumber="21">Eoniphargus toriii</emphasis>
|
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</taxonomicName>
|
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sp. nov. is similar to
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<taxonomicName id="30879E223C31BFB86E1097BFDF8D4379" lsidName="E. kojimai" pageId="0" pageNumber="21" rank="species" species="kojimai">
|
|
<emphasis id="87C4782D8B0A5C92A05211053A7E6EBE" italics="true" pageId="0" pageNumber="21">E. kojimai</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="2A827E1CC01A6422DF0D87D5760360CB" lsidName="E. iwataorum" pageId="0" pageNumber="21" rank="species" species="iwataorum">
|
|
<emphasis id="FC16B77975DCB0D2726D092032425B7C" italics="true" pageId="0" pageNumber="21">E. iwataorum</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., with a head bearing deep antennal sinus, antenna 1 peduncular article 1 with a robust seta on the posterodistal corner, antenna 2 peduncular article 2 with a gland cone not exceeding peduncular article 3, uropod 1 peduncle with basofacial robust setae, and uropod 3 with 2-articulate outer ramus. However,
|
|
<taxonomicName id="61CC31B3D128132666DF65A48114E064" lsidName="E. toriii" pageId="0" pageNumber="21" rank="species" species="toriii">
|
|
<emphasis id="271CE430DD305A0FF664A663D6D235E5" italics="true" pageId="0" pageNumber="21">E. toriii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName id="03766F724D2ABC316209A99659C69E4A" lsidName="E. kojimai" pageId="0" pageNumber="21" rank="species" species="kojimai">
|
|
<emphasis id="71903FC4DE9EC68A94DF440E03C7C99C" italics="true" pageId="0" pageNumber="21">E. kojimai</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="AF7A19AC0CCC650AF75BF211CDB636CC" lsidName="E. iwataorum" pageId="0" pageNumber="21" rank="species" species="iwataorum">
|
|
<emphasis id="491BBB0E8433DBEAC16ACDB58741D512" italics="true" pageId="0" pageNumber="21">E. iwataorum</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in the following features: right mandible with lacinia mobilis bearing three or four teeth (more than four teeth in
|
|
<taxonomicName id="21E09A864336A133F8ABCEA28F615201" lsidName="E. kojimai" pageId="0" pageNumber="21" rank="species" species="kojimai">
|
|
<emphasis id="682F6A1DC0DCAC63A5F25E313AF8AECB" italics="true" pageId="0" pageNumber="21">E. kojimai</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="70D1B10FFABE6248A88E0C34936FD84B" lsidName="E. iwataorum" pageId="0" pageNumber="21" rank="species" species="iwataorum">
|
|
<emphasis id="12E4490FE9CE64D5133E3F916A45B39C" italics="true" pageId="0" pageNumber="21">E. iwataorum</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.) and maxilla 1 having six medial setae on the inner plate (eight medial setae in
|
|
<taxonomicName id="5FB32F4C2E8FA168E516023DDAFC8FDB" lsidName="E. kojimai" pageId="0" pageNumber="21" rank="species" species="kojimai">
|
|
<emphasis id="C8D2D1DC2CEFF3EE3669CC100529FDEF" italics="true" pageId="0" pageNumber="21">E. kojimai</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4744295201526192378760F62AE2A330" lsidName="E. iwataorum" pageId="0" pageNumber="21" rank="species" species="iwataorum">
|
|
<emphasis id="EBE8E32D8444EA4F935AB002205BE606" italics="true" pageId="0" pageNumber="21">E. iwataorum</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |