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<document id="79DB0A9DCB833B3305D9FC85C29AF179" ENCODING="UTF-8" ID-DOI="10.5281/zenodo.32035" ID-GBIF-Dataset="ef713f74-fc1f-47ea-8b87-ae6b959ab9fb" ID-HNS-Pub="20236" ModsDocAuthor="Fern<72>ndez, F." ModsDocID="20236" checkinTime="1244210911494" checkinUser="thomas" docAuthor="Fernández, F." docDate="2003" docId="A68114724760C8C582352DB3BA208E4C" docLanguage="en" docName="20236_gg2.xml" docOrigin="Zootaxa 361" docSource="http://www.mapress.com/zootaxa/2003/zt00361.pdf" docTitle="Adelomyrmex tristani" docType="treatment" docVersion="13" lastPageNumber="33" masterDocId="498CEE3BDC847973176576F046D5BAC9" masterDocTitle="Revision of the myrmicine ants of the Adelomyrmex genus-group (Hymenoptera: Formicidae)." masterLastPageNumber="52" masterPageNumber="1" pageNumber="30" updateTime="1701311136115" updateUser="plazi">
<mods:mods id="3EA1B4FAFFBCDDD24404D125FB24F9CD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B0E03F5C39B265B7136536CDB0B35F6F">
<mods:title id="FE3AE764EB3D037396E3C4B05B7D8CF0">Revision of the myrmicine ants of the Adelomyrmex genus-group (Hymenoptera: Formicidae).</mods:title>
</mods:titleInfo>
<mods:name id="5FACF4D078F96BE0E445D8FA9F8E7DF4" type="personal">
<mods:role id="CB97874F3029612687CEC834FE0E6AD3">
<mods:roleTerm id="9B64142640299567DF23B4EBD7C74E04">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D62E8D2BF359A4916212D7A1B6BF007A">Fernández, F.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CE3D11F6595A3C44700C1A8DEF1A576F">text</mods:typeOfResource>
<mods:relatedItem id="031F9B301305C9D3815A2FBEC99DC35E" type="host">
<mods:titleInfo id="C6B44770F6812FD0AB73B3A1946D6D1B">
<mods:title id="F789D00BD04D15A93FED80FF28FBB350">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3F4033C09C305D38EDF216F0B03837CE">
<mods:date id="A7EAD4382F672B693637D45AB06B6171">2003</mods:date>
<mods:detail id="07961AA57375BA3E31D3D7FABA5D5494" type="volume">
<mods:number id="B57A3715FF74DBD9A48B870D5DA7E943">361</mods:number>
</mods:detail>
<mods:extent id="386AB53813D9AF5D13561814665913B8" unit="page">
<mods:start id="229370A4BAF189CD20370253C85088CA">1</mods:start>
<mods:end id="1AAFE9EC16927E30BCC1DDF906DD2550">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="27151D04B1D441A5DCCD141759A3C4BE">
<mods:url id="AB69C0C403C461383338B8D649CDC059">http://www.mapress.com/zootaxa/2003/zt00361.pdf</mods:url>
</mods:location>
<mods:classification id="2DD2469B2E4DE0972AC00D91050045DC">journal article</mods:classification>
<mods:identifier id="E4F208B16AF961F5BD4B73C12DA01170" type="HNS-Pub">20236</mods:identifier>
<mods:identifier id="4401D3FD8522E8CF04330CDD8260DC3E" type="DOI">10.5281/zenodo.32035</mods:identifier>
</mods:mods>
<treatment id="A68114724760C8C582352DB3BA208E4C" ID-DOI="http://doi.org/10.5281/zenodo.6273934" ID-GBIF-Taxon="100126314" ID-Zenodo-Dep="6273934" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C" httpUri="http://treatment.plazi.org/id/A68114724760C8C582352DB3BA208E4C" lastPageNumber="33" pageNumber="30">
<subSubSection id="10652854D809C0FDDA0623DEA42E57B5" type="nomenclature">
<paragraph id="4C9032D6A5168C3279EC4B9140D264B5" pageNumber="30">
<taxonomicName id="6A847AFC1983BF3E55DF1CEA5B521F6B" ID-CoL="9XX7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24999" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex tristani (Menozzi)" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="tristani">Adelomyrmex tristani (Menozzi)</taxonomicName>
(Figs. 19-24, 33, 39-40, 73)
</paragraph>
</subSubSection>
<subSubSection id="797112884433C96BD08A8D092CF2E587" type="reference_group">
<paragraph id="A39507AC0255812FC00479AC6C614393" pageNumber="30">
<taxonomicName id="2D9B9DC8D907953761086A593ECBCB07" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:132668" class="Insecta" genus="Apsychomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Apsychomyrmex tristani Menozzi" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="tristani">Apsychomyrmex tristani Menozzi</taxonomicName>
, 1931:260 (w); Borgmeier, 1937:240 (w); M.R. Smith, 1947:472 (worker redescription). [Holotype not seen, presumably in Deutsches Entomologisches Institut].
</paragraph>
<paragraph id="3EE47A42B3249ECFBD9B341EDFAAE039" pageNumber="30">
<taxonomicName id="97CCF272F84FD2954C274CE414411D1E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24999" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex tristani (Menozzi)" order="Hymenoptera" pageNumber="30" phylum="Arthropoda" rank="species" species="tristani">Adelomyrmex tristani</taxonomicName>
: Kempf, 1972:18 (
<taxonomicNameLabel id="9ADAE79BB2288C84CC0ADB5ABFD825DD">New combination</taxonomicNameLabel>
); Bolton, 1995:58 (w.).
</paragraph>
</subSubSection>
<subSubSection id="4DF0DDFFB4BF3454A49AD06C2D9B0C38" type="description">
<paragraph id="38633C07FF34372F4B9D9758E53D9ECB" pageNumber="30">Worker measurements (n=40). HL 0.53 - 0.73 HW 0.45 - 0.65 SL 0.29 - 0.41 EL 0.05 - 0.08 WL 0.53 - 0.73 GL 0.63 - 0.92 TL 2.00 - 2.84 CI 90 - 91 SI 55 - 63.</paragraph>
</subSubSection>
<subSubSection id="430EDE78C98C13B27831294D1F47411A" type="diagnosis">
<paragraph id="FF596EC7F4C0B54E273191A62E1B2978" lastPageNumber="31" pageNumber="30">Worker diagnosis: Head with posterior margin weakly flat to feebly concave. Mandibles with 5 to 6 teeth decreasing in size from the apical teeth. Tooth of basal margin of mandible not protuberant, smaller than subapical masticatory tooth. Eyes small, with 7 to 22 facets. Hypostomal tooth small but visible. Promesonotum slightly convex, higher than propodeal dorsum; metanotal groove well marked. Lateral sides of pronotum angulated. Propodeal spiracle rounded. Propodeum sloping, with the spines variable in size and shape, from low, triangular to long and narrow. Petiole high, with anterior side sloping and posterior side more or less feebly convex; sometimes petiole nearly quadrate, with anterior and posterior faces nearly parallel. Postpetiole campaniform with a ventral transverse ridge toothlike as seen in profile. Sting well developed. Mandibles dorsally with moderate longitudinal rugulations. Clypeal plate smooth and shining. Head frons longitudinally rugulate, head sides irregularly rugulated and mixed with punctures. Promesonotal dorsum variously sculptured, from longitudinal fine rugulae to longitudinal to strong coarse costae, in some workers the rugae or striae mixed, anteriorly oblique to transverse, in a few workers only a very few central longitudinal rugae visible. Promesonotal rugae or striae mixed with punctures. Propodeal declivity transversely rugulated between propodeal spines. Declivitous face of propodeum from smooth to covered with transverse rugae. Petiole and postpetiole normally transversely rugulated, sometimes times dorsally smooth and shining. Legs and gaster smooth and shining.</paragraph>
<paragraph id="FA302874D1F69BF1B6022FB570AD21BF" pageNumber="31">Hairs yellowish, long and flexuous on the body, more short and appressed on antennae and legs. Body black, dark brown or brown, legs light brown, tarsi yellowish.</paragraph>
</subSubSection>
<subSubSection id="FFF2FA8675F78D0F98DF4323A3EF27DB" type="description">
<paragraph id="764F32110DD2CAF471928FE2A7F28B2E" pageNumber="31">Queen measurements: HL 0.75 HW 0.66 SL 0.45 EL 0.11 WL 0.84 GL 1.05 TL 3.22 CI 88 SI 68.</paragraph>
</subSubSection>
<subSubSection id="31231AD3E9BA0F708BA526B70DBA63C5" type="diagnosis">
<paragraph id="673942EC3313C6D703029F5602C6805B" pageNumber="31">As worker with the following differences: Mandibles with all teeth of masticatory border of similar size. Three ocelli present. 12 - 13 ommatidia in the largest diameter of eye. Pronotum laterally costulated. Promesonotum longitudinally costulated with areas smooth and shining variable, from anterior half to nearly all promesonotum. Metanotum longitudinally costulated. Mesopleurae with longitudinal costulation and areas smooth and shining. Propodeum costulated longitudinally in the sides, transversely in the declivitous face.</paragraph>
<paragraph id="AC0F4E9A82B248531A0F510F6F4C283D" pageNumber="31">Male: Unknown.</paragraph>
</subSubSection>
<subSubSection id="1381891ECC57088B843657165C2B4DF7" type="materials_examined">
<paragraph id="E4719FA05453039DA9E56D57F58C0FDB" pageNumber="31">
Material examined:
<materialsCitation id="7F623337A46CDB6F2B01F2E3BF40A355" ID-GBIF-Occurrence="923930699" collectingDate="1992-09-24" collectionCode="INBio" collectorName="R. S. Anderson" country="MEXICO" location="Bochi, 1800 m." locationDeviation="17.3 km NW" specimenCode="No. 92 - 115" stateProvince="Chiapas">
<collectingCountry id="5E032EFAB4EA674E24CF9EF2CFC6081F">MEXICO</collectingCountry>
: 3 w,
<collectingRegion id="50B10F31BEE1B1EA004DB572270E203B">Chiapas</collectingRegion>
,
<locationDeviation id="5EEB868B6CA7A29BCE03C83575F009CC">17.3 km NW</locationDeviation>
<location id="EFBE2E745D87CF05744751A8EB4ED7C0" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:EFBE2E745D87CF05744751A8EB4ED7C0" country="MEXICO" name="Bochi, 1800 m." stateProvince="Chiapas">Bochi, 1800 m.</location>
,
<date id="1F222F7B31A3320734F1B624DBF70646" day="24" month="09" value="1992-09-24" year="1992">
<collectingDate id="65480EC89305C5054F8978D45A334827" day="24" month="09" value="1992-09-24" year="1992">24.ix.1992</collectingDate>
</date>
,
<collectorName id="F4E20DB0F0EF3880C4E6B128120989E5">R.S. Anderson</collectorName>
<specimenCode id="755DC851E258490681410DE6FF3847B6">No. 92-115</specimenCode>
(
<collectionCode id="FE0B004C649D6D7571BDF2C8D1DF9353" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="774AC08C6B67DB924A607DAE16E56694" ID-GBIF-Occurrence="923930685" collectingDate="1992-09-21" collectionCode="INBio, IAvH" collectorName="R. S. Anderson" country="MEXICO" location="Motozintla de Mendoza, 2000 m." locationDeviation="7.4 km SSW" specimenCode="No. 92 - 110" stateProvince="Chiapas">
6 w, 10 q,
<collectingRegion id="C693472576C81FF30185806879061FE7">Chiapas</collectingRegion>
,
<locationDeviation id="6A8662462484C63F45BEBC12AF9E459D">7.4 km SSW</locationDeviation>
<location id="6BEA2D3EF19DCEBF732DEB00F028ECFE" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:6BEA2D3EF19DCEBF732DEB00F028ECFE" name="Motozintla de Mendoza, 2000 m." stateProvince="Chiapas">Motozintla de Mendoza, 2000 m.</location>
,
<date id="C8985A97A0FCEE47CD4B6649F7696242" day="21" month="09" value="1992-09-21" year="1992">
<collectingDate id="BD677842FC69D56E022D73D6FF59EE53" day="21" month="09" value="1992-09-21" year="1992">21.ix.1992</collectingDate>
</date>
,
<collectorName id="9401DADBE49EFC213C328A1636BAD762">R.S. Anderson</collectorName>
<specimenCode id="D9B21CFDE5A1AE81472F16388F60F95A">No. 92-110</specimenCode>
(
<collectionCode id="242A5FADC447581C8E079202F83A931E" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
,
<collectionCode id="C930B8D8B9EC4A7BF5BCF63757FF259D">IAvH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2E4C7CE353697BFBDAF2D694DD9881BB" ID-GBIF-Occurrence="923930679" collectingDate="1992-09-19" collectionCode="INBio" collectorName="R. S. Anderson" country="MEXICO" location="km N Union Juarez, Volcan Tacana, lower slopes, 2000 m" specimenCode="No. 92 - 110" stateProvince="Chiapas">
2 w,
<collectingRegion id="95675051282CFC03B6E4B45F7B3F6211">Chiapas</collectingRegion>
,
<location id="8B582E82DE98198A30D4548CA14B2D79" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:8B582E82DE98198A30D4548CA14B2D79" name="km N Union Juarez, Volcan Tacana, lower slopes, 2000 m" stateProvince="Chiapas">
km N
<normalizedToken id="5D6DD277328978F5A0E726748DB9BF76" originalValue="Unión">Union</normalizedToken>
Juarez,
<normalizedToken id="3AE42EBFF315963F49780820C561536B" originalValue="Volcán">Volcan</normalizedToken>
Tacana, lower slopes, 2000 m
</location>
,
<date id="E753E1622C7D7082CBE1F52A8BE2EFAA" day="19" month="09" value="1992-09-19" year="1992">
<collectingDate id="2312D0F4F3535BCC9DB56D306F46ED52" day="19" month="09" value="1992-09-19" year="1992">19.ix.1992</collectingDate>
</date>
,
<collectorName id="2E895FAD85D8364883729EDE3C527A07">R.S. Anderson</collectorName>
<specimenCode id="0D7EE19EB8857DC055A0A485F0903079">No. 92-110</specimenCode>
(
<collectionCode id="6F4F42F3E2D1026C956C5F527F0F24C0" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="61240B52C127341C95B0BE0A7CCB018E" ID-GBIF-Occurrence="923930695" collectingDate="1946-04-18" collectionCode="LACM, USNM" collectorName="F. Bonnet" country="MEXICO" location="Tezitlan" specimenCode="No. 1350" stateProvince="Puebla">
9 w,
<collectingRegion id="2F192B44010E46B33D777C6709D91334">Puebla</collectingRegion>
,
<normalizedToken id="194E40BB6C88AAF2A8EBA8038C3C5D7A" originalValue="Tezitlán">
<location id="C1FD3FA5E9A9989E2DC077D71172194C" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:C1FD3FA5E9A9989E2DC077D71172194C" name="Tezitlan" stateProvince="Puebla">Tezitlan</location>
</normalizedToken>
,
<collectingDate id="7977AB95C355A6F8880E434D6BB3373D">
<date id="CD1CCF84F041B08434A1A5EB5FE7C3D7" day="18" month="04">18.iv.</date>
46
</collectingDate>
,
<collectorName id="FE3FA0A14D1F7F2A3DBB1C90794F4AC8">F. Bonnet</collectorName>
<specimenCode id="460B3936FF00FDAAFF4CA93D57DF95DF">No. 1350</specimenCode>
(
<collectionCode id="0B4D18A572EBEB5EA79D4A808E4D64D7" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
,
<collectionCode id="6F3E9571AFA64F5D7971536964DDFB06" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D63C2E64EEB727E01497F11515CE9FCA" ID-GBIF-Occurrence="923930690" collectingDate="1969-08-05" collectionCode="BMNH" collectorName="S. Peck, J. Peck" country="MEXICO" location="Fortin Canyon, Metlac River" stateProvince="Veracruz">
6 w,
<collectingRegion id="5173A7C3B194130E49F06ACE58916EE0">Veracruz</collectingRegion>
,
<location id="0E842197C808EE0D3FECE9C150A1157F" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:0E842197C808EE0D3FECE9C150A1157F" name="Fortin Canyon, Metlac River" stateProvince="Veracruz">Fortin Canyon, Metlac River</location>
,
<collectingDate id="3217940EFCD2A76EEC7AD544351F594E">
<date id="C2665EE85D70C1EB296810B24612862E" day="05" month="08">5.viii.</date>
69
</collectingDate>
,
<collectorName id="86B8D9035220447FE841A7D9E46A1FD5">S. &amp; J. Peck</collectorName>
(
<collectionCode id="00D50CEDFB1A14C64A8EB702E1AD7440" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E5690F9A87F9F8BAA3A09B875C5CEBF9" ID-GBIF-Occurrence="923930701" collectingDate="1946-03-19" collectionCode="MZSP" country="GUATEMALA" location="San Francisco, in Odontoglossum sp." specimenCode="No. 20342">
<collectingCountry id="B85E9E1F7E56BCC5A8BCB9FBBE0D352C">GUATEMALA</collectingCountry>
: 1 w,
<location id="722CEF8D24A03B1F54C666F2B034A698" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:722CEF8D24A03B1F54C666F2B034A698" country="GUATEMALA" name="San Francisco">San Francisco</location>
,
<collectingDate id="5316E7D6861E418A445ABF4B1CA5774E">
<date id="496118D1C52B92568F5E7D1F8ECAA845" day="19" month="03">19.iii.</date>
46
</collectingDate>
<specimenCode id="8F7E3E45B0A57DC76502163D8A88BCC4">No. 20342</specimenCode>
,
<location id="67292214D18DD0C4EF3BB132ADDF9E9F" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:67292214D18DD0C4EF3BB132ADDF9E9F" country="GUATEMALA" name="in Odontoglossum sp.">
in
<taxonomicName id="1675157065DF15DC42AABDF7B2374B47" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:215700" class="Liliopsida" family="Orchidaceae" genus="Odontoglossum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName-HNS="Odontoglossum B." order="Asparagales" pageNumber="31" phylum="Tracheophyta" rank="genus">Odontoglossum</taxonomicName>
sp.
</location>
Lot. No. 46-3094 (
<collectionCode id="8BD76C2FFD9D65A760013F22A26426AE" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo">MZSP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="4E1BA727AF0B436E8BCF10B4DA641102" ID-GBIF-Occurrence="923930688" collectingDate="1985-08-22" collectionCode="INBio" collectorName="J. Longino" country="COSTA RICA" latitude="10.3" location="Monteverde, 950 m." locationDeviation="6.5 km E" longitude="-84.75" specimenCode="No. 861 - s" stateProvince="Alajuela">
<collectingCountry id="9D553AA81722433281CEB979A5C49C39">COSTA RICA</collectingCountry>
: 2 w,
<collectingRegion id="E9BE1E039B3A7602489B93FA91C5FE71">Alajuela</collectingRegion>
,
<locationDeviation id="05909129A0DFD9A4AAE75C09C82F9E26">6.5 km E</locationDeviation>
<location id="B6C6C55DB0E148969B176DD707D85539" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:B6C6C55DB0E148969B176DD707D85539" country="COSTA RICA" latitude="10.3" longitude="-84.75" name="Monteverde" stateProvince="Alajuela">Monteverde</location>
,
<geoCoordinate id="8DE155F9BA98A762066043AA0FE2C7CE" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="2698BE48EC594C3FDC4DFF1B92AD6BFE" direction="west" orientation="longitude" value="-84.75">84°45’W</geoCoordinate>
,
<location id="377A3975D621DE144FC50AF105751F5A" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:377A3975D621DE144FC50AF105751F5A" country="COSTA RICA" latitude="10.3" longitude="-84.75" name="950 m." stateProvince="Alajuela">950 m.</location>
,
<date id="158AB98CCB7227BE9676228FE1B26745" day="22" month="08" value="1985-08-22" year="1985">
<collectingDate id="90CBE28FB9669CBBABC54BEE2D20B91E" day="22" month="08" value="1985-08-22" year="1985">22.viii.1985</collectingDate>
</date>
,
<collectorName id="4EC436A1E5F68C1CDC2491355D7E6747">J. Longino</collectorName>
<specimenCode id="07F1E0444D733ED01DDF86E0ED080FF1">No. 861-s</specimenCode>
(
<collectionCode id="4D2FC7A893377E457C69EB7B3499AACD" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BA1BE015BA8340591B07EDF4608A9285" ID-GBIF-Occurrence="923930691" collectingDate="1990-04-26" collectionCode="INBio" collectorName="J. Longino" country="COSTA RICA" latitude="10.3" location="Monteverde, 1400 m." locationDeviation="3 km E" longitude="-84.78333" specimenCode="No. 2674 - s" stateProvince="Alajuela">
3 w, 1 q,
<collectingRegion id="B95FDA621C9975A09405E77B9544270D">Alajuela</collectingRegion>
,
<locationDeviation id="3FF9A98451053704B95195FD9882E5C8">3 km E</locationDeviation>
<location id="24788630FD1ACFACFB8F243094491480" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:24788630FD1ACFACFB8F243094491480" latitude="10.3" longitude="-84.78333" name="Monteverde" stateProvince="Alajuela">Monteverde</location>
,
<geoCoordinate id="F7245E8EBFA86045FC9F8C30C8975C36" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="8020DF08ADDD1895129C21B440BFF0BE" direction="west" orientation="longitude" value="-84.78333">84°47’W</geoCoordinate>
,
<location id="488481231664446AC04750D8E1832238" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:488481231664446AC04750D8E1832238" latitude="10.3" longitude="-84.78333" name="1400 m." stateProvince="Alajuela">1400 m.</location>
,
<date id="D3624D727FB3A8D467EDDE7764D87557" day="26" month="04" value="1990-04-26" year="1990">
<collectingDate id="6BC33D9F1688EEBBBDBD371A18629F7D" day="26" month="04" value="1990-04-26" year="1990">26.iv.1990</collectingDate>
</date>
,
<collectorName id="AC9C6B8101A717C5BE96997C73A703AE">J. Longino</collectorName>
<specimenCode id="27AE7A494FBE3A90304129A4DAF48D53">No. 2674-s</specimenCode>
(
<collectionCode id="D066E6CA74880342F029398F23F005D4" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0FA42BC78A1AB56A95825470B0EF4202" ID-GBIF-Occurrence="923930684" collectingDate="1989-02-11" collectionCode="INBio" collectorName="J. Longino" country="COSTA RICA" latitude="10.933333" location="Cerro Cacao, 1500 m." longitude="-85.45" specimenCode="No. 2339 - s" stateProvince="Guanacaste">
2 w,
<collectingRegion id="A580E764CB6FAC3C2BE2D98437ECC45D">Guanacaste</collectingRegion>
,
<location id="BDFF6E9DFD2096130B51337A8A43B124" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:BDFF6E9DFD2096130B51337A8A43B124" latitude="10.933333" longitude="-85.45" name="Cerro Cacao" stateProvince="Guanacaste">Cerro Cacao</location>
,
<geoCoordinate id="DDB97BDEB950E85812CEF96DF2163141" direction="north" orientation="latitude" value="10.933333">10°56’N</geoCoordinate>
<geoCoordinate id="8B7CE5A4D3D248B1CA438677FA68A103" direction="west" orientation="longitude" value="-85.45">85°27’W</geoCoordinate>
,
<location id="DBD88A3A4B7CE88730F6D4D4B6F427B1" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:DBD88A3A4B7CE88730F6D4D4B6F427B1" latitude="10.933333" longitude="-85.45" name="1500 m." stateProvince="Guanacaste">1500 m.</location>
, 9-
<date id="43A8DD847DB60FE33D60A997E0A794EE" day="11" month="02" value="1989-02-11" year="1989">
<collectingDate id="5689C5E59131549E53895DF95BE2A0F3" day="11" month="02" value="1989-02-11" year="1989">11.ii.1989</collectingDate>
</date>
,
<collectorName id="5649B5304CE33ADE36C9D337FD61EC98">J. Longino</collectorName>
<specimenCode id="B8472AB5D0233BF43C075E07A2DC5E3B">No. 2339-s</specimenCode>
(
<collectionCode id="05C80B5EE10A7081C166B69FF45DB86F" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC427CE0C6015AF9306A3A5263478787" ID-GBIF-Occurrence="923930689" collectingDate="1986-07-14" collectionCode="INBio" collectorName="J. Longino" country="COSTA RICA" latitude="10.283334" location="Volcan Barba, 800 m" locationDeviation="18 km N" longitude="-84.083336" specimenCode="No. 1383 - S" stateProvince="Heredia">
13 w,
<collectingRegion id="12993102DF808F740F8E0BBD3C11C2F5">Heredia</collectingRegion>
,
<locationDeviation id="C84E4DF7770735D3E850DE8FDFBABF77">18 km N</locationDeviation>
<location id="63A860C97848A33E06B17BDC79EB4F1D" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:63A860C97848A33E06B17BDC79EB4F1D" latitude="10.283334" longitude="-84.083336" name="Volcan Barba" stateProvince="Heredia">
<normalizedToken id="83CA06DF098387F357254C0DC8CD2C25" originalValue="Volcán">Volcan</normalizedToken>
Barba
</location>
,
<geoCoordinate id="2BC23B90381C6E64E801735ECB9A0BA4" direction="north" orientation="latitude" value="10.283334">10°17’N</geoCoordinate>
<geoCoordinate id="2B8976CE2E5E88B4D34F4DF26CDBF506" direction="west" orientation="longitude" value="-84.083336">84°05’W</geoCoordinate>
,
<location id="0BFD09E2A779097E962058797D5CBB68" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:0BFD09E2A779097E962058797D5CBB68" latitude="10.283334" longitude="-84.083336" name="800 m" stateProvince="Heredia">800 m</location>
,
<date id="22C1674096AB84F41A53D27294D4B5BD" day="14" month="07" value="1986-07-14" year="1986">
<collectingDate id="B325426D36A71753F7C5A6F1AD4312C9" day="14" month="07" value="1986-07-14" year="1986">14.vii.1986</collectingDate>
</date>
,
<collectorName id="40EFB4F6F62B4DAAB204690B8E0265F7">J. Longino</collectorName>
<specimenCode id="7A25D580F6C60DF4F76870CC5B8449FA">No. 1383-S</specimenCode>
(
<collectionCode id="5CC56DEC12DCFB8726AC65BFE0D43647" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7D71BD879FD314D69FEEC41467BCE462" ID-GBIF-Occurrence="923930683" collectingDate="1992-10" collectionCode="OET, INBio" country="COSTA RICA" latitude="10.433333" location="La Selva Biological Station, 50-150 m" longitude="-84.01666667" stateProvince="Heredia">
1 w,
<collectingRegion id="94127457617F8789A633294D34A9E1FE">Heredia</collectingRegion>
,
<location id="B36D547C63B7F7E8D650758BFB3E2CF4" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:B36D547C63B7F7E8D650758BFB3E2CF4" latitude="10.433333" name="La Selva Biological Station" stateProvince="Heredia">La Selva Biological Station</location>
,
<geoCoordinate id="D0AC269BAF27B899784E0E69E09AF607" direction="north" orientation="latitude" value="10.433333">10°26’N</geoCoordinate>
<geoCoordinate id="7FDDEE6B0E4FAFB23046D06AE05AE786">84°01W</geoCoordinate>
,
<location id="0A5E8C4857BDE624D3A47BFCA9438CBC" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:0A5E8C4857BDE624D3A47BFCA9438CBC" latitude="10.433333" name="50 - 150 m" stateProvince="Heredia">50-150 m</location>
,
<date id="444A38E7F275FE6E67F1AB1546ABB732" month="10" value="1992-10" year="1992">
<collectingDate id="C5CB8812898F0FB58FFB9C582B8DFEC8" month="10" value="1992-10" year="1992">x.1992</collectingDate>
</date>
,
<collectionCode id="6D6DDF84240BA220EC8660DB745CB9A6" collectionName="OET">OET</collectionCode>
(
<collectionCode id="505D8ECD15D6C99135613975747A08F9" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7D9DF5537F059BB10FF830ACE475D788" ID-GBIF-Occurrence="923930700" collectingDate="1984-07-15" collectionCode="LACM" collectorName="L. Longino" country="COSTA RICA" latitude="10.3" location="Monteverde, 1560 m" longitude="-84.8" stateProvince="Puntarenas">
14 w, 1 f.,
<collectingRegion id="392F63634E437E4CE8573E2B45EAE71F">Puntarenas</collectingRegion>
,
<location id="9432A359FB740B0851A080D5522549CA" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:9432A359FB740B0851A080D5522549CA" latitude="10.3" longitude="-84.8" name="Monteverde" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="86F357D56F898568B2F85C69886DE3E6" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="F1F39C0DCE8D202845E787B9310BD405" direction="west" orientation="longitude" value="-84.8">84°48’W</geoCoordinate>
,
<location id="2AE1932D18E48EECAB2958746A04DDAB" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:2AE1932D18E48EECAB2958746A04DDAB" latitude="10.3" longitude="-84.8" name="1560 m" stateProvince="Puntarenas">1560 m</location>
,
<date id="FA83D3CED8B8A1449AF4D98A128233D5" day="15" month="07" value="1984-07-15" year="1984">
<collectingDate id="309EDC7F50A6E40075BA791DA9F4405E" day="15" month="07" value="1984-07-15" year="1984">15.vii.1984</collectingDate>
</date>
,
<collectorName id="32C5FE61E09485DEBA515B0877E74531">L. Longino</collectorName>
(
<collectionCode id="2B276ADC95C0FC86A69B9CFD9188F454" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BC4951661B0526DF8799193391E65BC4" ID-GBIF-Occurrence="923930687" collectingDate="1991-05-14" collectionCode="INBio, IAvH" collectorName="J. Longino" country="COSTA RICA" latitude="10.3" location="Monteverde, 1300-1800 m" longitude="-84.8" specimenCode="Nos. 1572 - 2888 - s" stateProvince="Puntarenas">
24 w, 7 q,
<collectingRegion id="5EB93A557E4CE9B47DFB74AD19DFC226">Puntarenas</collectingRegion>
,
<location id="CDB50BD6803D83B0BC1E860D6CC5514A" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:CDB50BD6803D83B0BC1E860D6CC5514A" latitude="10.3" longitude="-84.8" name="Monteverde" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="27C065EB631615AE827B6203BEB7AB89" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="A832427E6DC61C565F969A4D09ACC8EA" direction="west" orientation="longitude" value="-84.8">84°48’W</geoCoordinate>
,
<location id="1EBA06909829D81E78537C497E34B92A" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:1EBA06909829D81E78537C497E34B92A" latitude="10.3" longitude="-84.8" name="1300 - 1800 m" stateProvince="Puntarenas">1300-1800 m</location>
,
<collectingDate id="89D27174158BE08BDDC65C1EE4EE278D">
<date id="79281720B2545FD5357B397985ACBD86" day="15" month="07" value="1984-07-15" year="1984">15.vii.1984</date>
-
<date id="1CFB529439E6B3D739284AD96955C138" day="14" month="05" value="1991-05-14" year="1991">14.v.1991</date>
</collectingDate>
,
<collectorName id="2771DD5659B9715BD2DB3C770B718CC3">J. Longino</collectorName>
<specimenCode id="E4D578FCB6A306ADDA992FBAD06DBB6A">Nos. 1572 - 2888-s</specimenCode>
(
<collectionCode id="2FD457CF29567A90BA6FC848D599298D" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
,
<collectionCode id="26AC3181571104807DFFD3A0EFCB2473">IAvH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C64B30FA1B8A7B4612745852EDCA66EF" ID-GBIF-Occurrence="923930697" collectingDate="1987-05" collectionCode="INBio, IAvH" collectorName="S. Little" country="COSTA RICA" latitude="10.3" location="Monteverde, 1400 m." longitude="-84.8" stateProvince="Puntarenas">
15 w,
<collectingRegion id="D063282F7907D6E5BBF719C37359BFCF">Puntarenas</collectingRegion>
,
<location id="A178B71A03A842D7058FDD8E02047F2B" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:A178B71A03A842D7058FDD8E02047F2B" latitude="10.3" longitude="-84.8" name="Monteverde" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="DBD752BCFCF07DA3B07CFF6F049E077E" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="0C8E593B947782C067FD20CAF26B1F98" direction="west" orientation="longitude" value="-84.8">84°48’W</geoCoordinate>
,
<location id="AF4DA00A02D7BBD43E9300C0DF0CE262" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:AF4DA00A02D7BBD43E9300C0DF0CE262" latitude="10.3" longitude="-84.8" name="1400 m." stateProvince="Puntarenas">1400 m.</location>
, iv.-
<date id="363F292F73C5363F39B5C9DD0C0F6977" month="05" value="1987-05" year="1987">
<collectingDate id="FADAEC99F46C99FF70F5625851458F93" month="05" value="1987-05" year="1987">v.1987</collectingDate>
</date>
,
<collectorName id="3C9EE287DAEBDEE8859B8BE570B867D1">S. Little</collectorName>
(
<collectionCode id="92CEBC085C14FD2D77B6E01DAB6946C0" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
,
<collectionCode id="7612F68B445916411DA0F67D2BB07A4E">IAvH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C71D378908EED308C2F340F21A5F0D84" ID-GBIF-Occurrence="923930698" collectingDate="1988-04-08" collectionCode="INBio" collectorName="J. Longino" country="COSTA RICA" latitude="10.3" location="Monteverde, 1550 m" longitude="-84.8" specimenCode="No. 1994 - s to 1973 - s" stateProvince="Puntarenas">
14 w, 5 q,
<collectingRegion id="77391D39A620F45EB3F56BFD738C3514">Puntarenas</collectingRegion>
,
<location id="B7CCF160FFDF52F3BA7A4F6EF01C41C2" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:B7CCF160FFDF52F3BA7A4F6EF01C41C2" latitude="10.3" longitude="-84.8" name="Monteverde" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="55D5EB37B97CCB78D131BCAD98F6359A" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="2656ED5BB99DD4239272B79AD8F258FF" direction="west" orientation="longitude" value="-84.8">84°48’W</geoCoordinate>
,
<location id="A8B6ECD8CF5FB8862778E70322786C88" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:A8B6ECD8CF5FB8862778E70322786C88" latitude="10.3" longitude="-84.8" name="1550 m" stateProvince="Puntarenas">1550 m</location>
,
<collectingDate id="44196E492DB7B8BABD0C49A97DD654BA">
<date id="C4A5C60AB1F988DDC2AC7CDD95455C8C" day="08" month="04" value="1988-04-08" year="1988">8.iv.1988</date>
-
<date id="FDC3087A2AA0EDE4792321B213AB6BC9" day="10" month="12" value="1987-12-10" year="1987">10.xii.1987</date>
</collectingDate>
,
<collectorName id="B64CD759134274C395912DBB6B05CE3A">J. Longino</collectorName>
<specimenCode id="43B479347ABA80E9FDEE346B578F68B1">No. 1994-s to 1973-s</specimenCode>
(
<collectionCode id="791EE7E608872F0DE3FAF5F7B49B78DF" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="CF6FC330565DC951C3A7C0E1781F3D01" ID-GBIF-Occurrence="923930692" collectingDate="1989-04-30" collectionCode="INBio" collectorName="L. Longino" country="COSTA RICA" latitude="10.3" location="Monteverde, 1600 m." longitude="-84.8" specimenCode="No. 2486 - s" stateProvince="Puntarenas">
6 w, 1 q,
<collectingRegion id="EB48546012F0C9B909F27301E9F42061">Puntarenas</collectingRegion>
,
<location id="E6031011A2E6A7F7693427356BC849AC" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:E6031011A2E6A7F7693427356BC849AC" latitude="10.3" longitude="-84.8" name="Monteverde" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="FE9542FE350DEF861FE5DC9866CCA817" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="9771D6E27EBA5B23B6D4A454340ADC60" direction="west" orientation="longitude" value="-84.8">84°48’W</geoCoordinate>
,
<location id="A6575C7B9560F28FFBCE2304147C9A9F" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:A6575C7B9560F28FFBCE2304147C9A9F" latitude="10.3" longitude="-84.8" name="1600 m." stateProvince="Puntarenas">1600 m.</location>
,
<date id="FB1BC78D9A5867AB3DC572B08DC861E7" day="30" month="04" value="1989-04-30" year="1989">
<collectingDate id="02598A0E2DE32FE60F5DB252C0C2A6C1" day="30" month="04" value="1989-04-30" year="1989">30.iv.1989</collectingDate>
</date>
,
<collectorName id="E929FB55DE5C170314F3CFFC8E8FCD78">L. Longino</collectorName>
<specimenCode id="46BF38670899482CA3205A89A18C913E">No. 2486-s</specimenCode>
(
<collectionCode id="B0FAEA52DE8C0A60329D67E6AEDFB519" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E0F69FA4890FAD2DEEB2DCEC85C01BB4" ID-GBIF-Occurrence="923930686" collectingDate="1991-05-01" collectionCode="INBio" collectorName="U. California EAP" country="COSTA RICA" latitude="10.3" location="Monteverde, 1600 m." longitude="-84.8" stateProvince="Puntarenas">
3 w, 1 f,
<collectingRegion id="4C85A5EA99E562CBE302A3D0C177A2F4">Puntarenas</collectingRegion>
,
<location id="599F904F42F004798DE19E2B330A6070" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:599F904F42F004798DE19E2B330A6070" latitude="10.3" longitude="-84.8" name="Monteverde" stateProvince="Puntarenas">Monteverde</location>
,
<geoCoordinate id="90CBD848079EFE40DC0A5F8B57A11BB2" direction="north" orientation="latitude" value="10.3">10°18’N</geoCoordinate>
<geoCoordinate id="73754557DD86DEFDD2D9FDA370A0234F" direction="west" orientation="longitude" value="-84.8">84°48’W</geoCoordinate>
,
<location id="DB3C08DDF13949C56DF0830D1275D9D7" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:DB3C08DDF13949C56DF0830D1275D9D7" latitude="10.3" longitude="-84.8" name="1600 m." stateProvince="Puntarenas">1600 m.</location>
,
<date id="0736BB09844A459F7550D1983F139792" day="01" month="05" value="1991-05-01" year="1991">
<collectingDate id="BC23A5D39988288750EC0133612BBE1A" day="01" month="05" value="1991-05-01" year="1991">1.v.1991</collectingDate>
</date>
,
<collectorName id="1C950B0D646C9AD71EAB886BBD131ABD">U. California EAP</collectorName>
(
<collectionCode id="1E11528BB0BB6FF08FDFBB742C61F4BF" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C01C4D5BB75C5149532F6ED6393D646B" ID-GBIF-Occurrence="923930702" collectingDate="1989-07-02" collectionCode="INBio" country="COSTA RICA" latitude="9.033334" location="Estacion Pittier, 1670 m." longitude="-82.916664" stateProvince="Puntarenas">
1 w,
<collectingRegion id="6A43FB15156578C40FF9CA9DEE64CFF3">Puntarenas</collectingRegion>
,
<location id="16177E3ADA56F75F38249B29F0E1AB2E" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:16177E3ADA56F75F38249B29F0E1AB2E" latitude="9.033334" longitude="-82.916664" name="Estacion Pittier" stateProvince="Puntarenas">
<normalizedToken id="2A122CACE28C45393566AC8CB87BD723" originalValue="Estación">Estacion</normalizedToken>
Pittier
</location>
,
<geoCoordinate id="2A82E936F2E917E4B703FD7EFBD23E68" direction="north" orientation="latitude" value="9.033334">9°02’N</geoCoordinate>
<geoCoordinate id="A557332D4AAEBC1424F8D6E8A5358B58" direction="west" orientation="longitude" value="-82.916664">82°55’W</geoCoordinate>
,
<location id="FDF1991EA0866F9B7DA014FAE52A151F" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:FDF1991EA0866F9B7DA014FAE52A151F" latitude="9.033334" longitude="-82.916664" name="1670 m." stateProvince="Puntarenas">1670 m.</location>
,
<date id="FDD97B96BFCD78BF22CE2EE037AB6092" day="02" month="07" value="1989-07-02" year="1989">
<collectingDate id="4F784CFA3B04F7DD5B54CA017117D1E1" day="02" month="07" value="1989-07-02" year="1989">2.vii.1989</collectingDate>
</date>
(
<collectionCode id="47B6BB79E87AB0678E04E5DC27D26C1F" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="A39872071E2E46317B24ED720FA4BB95" ID-GBIF-Occurrence="923930682" collectingDate="1995-06-29" collectionCode="INBio" collectorName="J. Longino" country="COSTA RICA" latitude="8.783334" location="Fila Cruces, 1200 m." longitude="-83.05" specimenCode="No. 3693 - s" stateProvince="Puntarenas">
2 w,
<collectingRegion id="1ECB5D7BE1DEAC13F9553B411BBF47F6">Puntarenas</collectingRegion>
,
<location id="0827A2B2824EC0E4FAEEDCE27C842B5A" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:0827A2B2824EC0E4FAEEDCE27C842B5A" latitude="8.783334" longitude="-83.05" name="Fila Cruces" stateProvince="Puntarenas">Fila Cruces</location>
,
<geoCoordinate id="3922B2D04EC357BFE0CCA325CDF5AB14" direction="north" orientation="latitude" value="8.783334">8°47’N</geoCoordinate>
<geoCoordinate id="153B2A9061B128A75F93CBD50BDDEF5E" direction="west" orientation="longitude" value="-83.05">83°03’W</geoCoordinate>
,
<location id="765869ADB39F66767C69D20820AD36D2" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:765869ADB39F66767C69D20820AD36D2" latitude="8.783334" longitude="-83.05" name="1200 m." stateProvince="Puntarenas">1200 m.</location>
,
<date id="AB140F59EB39F727661F31FB240D22BB" day="29" month="06" value="1995-06-29" year="1995">
<collectingDate id="EC09C8CB9F9E8B6E2FD02D929E1FF93F" day="29" month="06" value="1995-06-29" year="1995">29.vi.1995</collectingDate>
</date>
,
<collectorName id="ED051EE101DF12A7B1F6DE62647AFA51">J. Longino</collectorName>
<specimenCode id="6F3A2FD922AD481C16EF224DE5D12480">No. 3693-s</specimenCode>
(
<collectionCode id="F95F821549445699D7FA2FBBF959069A" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBio</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1AC35CFD1083CF572FA7DE5C86DA7B20" ID-GBIF-Occurrence="923930713" collectingDate="1957-05-08" collectingMethod="tunnel" collectionCode="USNM" country="COSTA RICA" location="Laredo, en Philodendron" stateProvince="San Jose">
3 w,
<collectingRegion id="DE5B54F03F38713B46B9F18A9C786A8E">
San
<normalizedToken id="52B9F74990A87D69A07CF134C731F175" originalValue="José">Jose</normalizedToken>
</collectingRegion>
,
<location id="4C03C37AE241962C8B935E66C3098773" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:4C03C37AE241962C8B935E66C3098773" name="Laredo, en Philodendron" stateProvince="San Jose">
Laredo, en
<taxonomicName id="C4D6DE387B86652F9C38505DD7F403F1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:215579" class="Liliopsida" family="Araceae" genus="Philodendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName-HNS="Philodendron Schott" order="Alismatales" pageNumber="32" phylum="Tracheophyta" rank="genus">Philodendron</taxonomicName>
</location>
,
<collectingMethod id="831E2BBF652B860D74EAC733786B9DBF">tunnel</collectingMethod>
,
<collectingDate id="E5E7BE26046C7232629A4122F6B74281">
<date id="0118C4CA6F8E6B1491445F5765F4E2ED" day="08" month="05">8.v.</date>
57
</collectingDate>
, (
<collectionCode id="6C3069CB309C9FBEDDFEA755FBC690E4" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="68EB4A9E6B570AAB8B7CADF987BD8C58" ID-GBIF-Occurrence="923930719" collectionCode="ICN" country="USA" location="From Mexico or Costa Rica, in orchid, intercepted in Brownsville" stateProvince="Texas">
1 w, &quot;
<location id="46DF78B59B55C4FE5E073903D5769C0A" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:46DF78B59B55C4FE5E073903D5769C0A" name="From Mexico or Costa Rica, in orchid, intercepted in Brownsville" stateProvince="Texas">
From
<normalizedToken id="290E2F335F58CC06E3E8243B0E694F18" originalValue="México">Mexico</normalizedToken>
or Costa Rica, in orchid, intercepted in Brownsville
</location>
,
<collectingRegion id="926DAAC106DA7618EF48172D706D0466">Texas</collectingRegion>
, 1 w, (
<collectionCode id="5E5AD06BD6CA13C18684586FEDF1D4D6" collectionName="Colombia, Bogota, Universidad Nacional de Colombia, Insituto de Ciencias Naturales de la Universidad Nacional">ICN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0A0C90983A7F7A45CFF24B868CE9BD50" ID-GBIF-Occurrence="923930704" collectingDate="1993-12-13" collectingMethod="winkler trap" collectionCode="ICN" collectorName="A. Vahos" country="COLOMBIA" latitude="11.85" location="El Retiro, Fca. El Barcino, 2100 m" longitude="-84.583336" stateProvince="Antioquia">
<collectingCountry id="73F4331155DE7478BE1D6657996DA5AC">COLOMBIA</collectingCountry>
: 2 w,
<collectingRegion id="7611F924ADB256E98178AB2D010F215B">Antioquia</collectingRegion>
,
<location id="003BC2D0DB413460047BF45B6BE006FC" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:003BC2D0DB413460047BF45B6BE006FC" country="COLOMBIA" latitude="11.85" longitude="-84.583336" name="El Retiro, Fca. El Barcino, 2100 m" stateProvince="Antioquia">El Retiro, Fca. El Barcino, 2100 m</location>
,
<geoCoordinate id="975FA40F80056FB0F73E9E63E8313BAF" direction="north" orientation="latitude" value="11.85">11°51’N</geoCoordinate>
<geoCoordinate id="5EB0972EEEABA7E22F875DC43DFE4F4B" direction="west" orientation="longitude" value="-84.583336">83°95’W</geoCoordinate>
,
<collectingMethod id="0FCFB93A3E383BA862424D483F19C663">winkler trap</collectingMethod>
,
<date id="86DFBD85BC68E1367B7C81F8A8FB2F1B" day="13" month="12" value="1993-12-13" year="1993">
<collectingDate id="C7998538F4420F5A19C4B1946D555DD8" day="13" month="12" value="1993-12-13" year="1993">13.xii.1993</collectingDate>
</date>
,
<collectorName id="A04807FFBDDD34FCF20BB8B076506EC2">A. Vahos</collectorName>
leg. (
<collectionCode id="4FC2FAC70C5CFBE320F00D533B7F6915" collectionName="Colombia, Bogota, Universidad Nacional de Colombia, Insituto de Ciencias Naturales de la Universidad Nacional">ICN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D34DE18E6F1A13D6AE876B37BC8AA14A" ID-GBIF-Occurrence="923930706" collectingDate="2002-06-01" collectionCode="IAvH" collectorName="J. Sossa" country="COLOMBIA" location="Filandia, 1870 m" stateProvince="Quindio">
1 w,
<normalizedToken id="C182794C7B0D430FC98C469BB6873719" originalValue="Quindío">
<collectingRegion id="D719A6E8915EB9017F10D2A0290F4C45">Quindio</collectingRegion>
</normalizedToken>
,
<location id="9AB2584BB830E074F39CA8497FFDD141" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:9AB2584BB830E074F39CA8497FFDD141" name="Filandia, 1870 m" stateProvince="Quindio">Filandia, 1870 m</location>
,
<date id="BC8B047F7AA32DF262BE5B00F6EC04AB" day="01" month="06" value="2002-06-01" year="2002">
<collectingDate id="1731B7E30501A78B6A1387BB0B5D0297" day="01" month="06" value="2002-06-01" year="2002">1.vi.2002</collectingDate>
</date>
,
<collectorName id="761FD97AF22822EC64C52BAC129C00B2">J. Sossa</collectorName>
leg. (
<collectionCode id="875216457F2476C9DC764F8A48551433">IAvH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9CD81FB519FC1C9E54A6546EEAD79EA8" ID-GBIF-Occurrence="923930725" collectingDate="1999-03-20" collectingMethod="winkler trap" collectionCode="IAvH, ICN" collectorName="E. L. Gonzalez" country="COLOMBIA" latitude="6.0666666" location="Encino, Cachalu Natural Reserve, 2000 m" longitude="-73.11667" stateProvince="Santander">
11 w,
<collectingRegion id="23462A3280B4FFBCF58B6F413A15BE91">Santander</collectingRegion>
,
<location id="F22D58B4E58676B9DB571FC26E5460D0" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:F22D58B4E58676B9DB571FC26E5460D0" latitude="6.0666666" longitude="-73.11667" name="Encino, Cachalu Natural Reserve" stateProvince="Santander">
Encino,
<normalizedToken id="DB818271CD48A4220EFFB01CFB91CFEE" originalValue="Cachalú">Cachalu</normalizedToken>
Natural Reserve
</location>
,
<geoCoordinate id="1EEFFF9F9D0FBB8F6D6A0A11B699DE36" direction="north" orientation="latitude" value="6.0666666">06°04’N</geoCoordinate>
<geoCoordinate id="A62C6A4B0BFFE80302166EBDB0D4079F" direction="west" orientation="longitude" value="-73.11667">73°07’W</geoCoordinate>
,
<location id="2F358040EE50D0A79A3ADDE7DFC6288F" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:2F358040EE50D0A79A3ADDE7DFC6288F" latitude="6.0666666" longitude="-73.11667" name="2000 m" stateProvince="Santander">2000 m</location>
,
<collectingMethod id="C98893060E8AF8CE954E39EAE543A66E">winkler trap</collectingMethod>
,
<date id="4B4732F609F5785C381475308D582710" day="20" month="03" value="1999-03-20" year="1999">
<collectingDate id="A35B025D6537C970070C5644D6E833F2" day="20" month="03" value="1999-03-20" year="1999">20.iii.1999</collectingDate>
</date>
,
<collectorName id="2D83DDB74E2EA874ED629F729BFB664A">E.L. Gonzalez</collectorName>
leg. (
<collectionCode id="6D5AC5FBBFF8436F3549A7032EC2F8EA">IAvH</collectionCode>
,
<collectionCode id="D5503A02FE3435BC96D4AD06804FDCFF" collectionName="Colombia, Bogota, Universidad Nacional de Colombia, Insituto de Ciencias Naturales de la Universidad Nacional">ICN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7E95EBF59C720D05E2377BB1A605AE96" ID-GBIF-Occurrence="923930707" collectingMethod="winkler trap" collectionCode="IAvH, ICN" collectorName="S. Sarria" country="COLOMBIA" location="Alto Anchicaya, Farallones de Cali National Park, 650 m" stateProvince="Valle">
4 w,
<collectingRegion id="F6FEFFDF86D854A10A891F52D2B42603">Valle</collectingRegion>
,
<location id="89DA4044C100643895EEC2DE186EE311" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:89DA4044C100643895EEC2DE186EE311" name="Alto Anchicaya, Farallones de Cali National Park, 650 m" stateProvince="Valle">Alto Anchicaya, Farallones de Cali National Park, 650 m</location>
,
<collectingMethod id="A8B8F5D6D0056EC8B83DFE048812AF10">winkler trap</collectingMethod>
,
<collectorName id="59274F3D04C48EF59D1521E3838692AD">S. Sarria</collectorName>
leg. (
<collectionCode id="CC6885EB1B54CDAFA66C514B57A75CA6">IAvH</collectionCode>
,
<collectionCode id="86102D0FE4AE602CA48C719572EE764C" collectionName="Colombia, Bogota, Universidad Nacional de Colombia, Insituto de Ciencias Naturales de la Universidad Nacional">ICN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B6A65795EF8D9A3C4C844110CDCC22DE" ID-GBIF-Occurrence="923930721" collectingDate="1993-12-08" collectionCode="ICN" collectorName="P. Chacon" country="COLOMBIA" location="Cali, El Ensueno, 27 km road Cali-Buenaventura, 1780 m" stateProvince="Valle">
1 w,
<collectingRegion id="AF2A823B658351E57216200159AD86B6">Valle</collectingRegion>
,
<location id="929E6EBC9139B161CA0A59338BC21349" LSID="urn:lsid:plazi:treatment:A68114724760C8C582352DB3BA208E4C:929E6EBC9139B161CA0A59338BC21349" name="Cali, El Ensueno, 27 km road Cali-Buenaventura, 1780 m" stateProvince="Valle">
Cali, El
<normalizedToken id="AA755D0CC2EDB4603A04F727587134FB" originalValue="Ensueño">Ensueno</normalizedToken>
, 27 km road Cali-Buenaventura, 1780 m
</location>
, in litt.,
<date id="02FE4FAE8823CDB1FAC372C4347D5FCA" day="08" month="12" value="1993-12-08" year="1993">
<collectingDate id="34C348146B81722B11DDC8B3F0E4D7C7" day="08" month="12" value="1993-12-08" year="1993">8.xii.1993</collectingDate>
</date>
,
<collectorName id="7FDAA81EB1EB85FBE7E5C23B866CD26B">
P.
<normalizedToken id="9E2925A5C9C9E93B000C7EB925ECCDBB" originalValue="Chacón">Chacon</normalizedToken>
</collectorName>
leg. (
<collectionCode id="F19EFD1C618AA0E416378AB222413A3F" collectionName="Colombia, Bogota, Universidad Nacional de Colombia, Insituto de Ciencias Naturales de la Universidad Nacional">ICN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F361504A5A51590C2883116B88FCD4CE" type="discussion">
<paragraph id="08F0DE0E0BDF3D823143F6237EC14D34" pageNumber="32">Comments. This species is highly variable in size, color, pilosity, configuration of the propodeal spines, and to a lesser extent, petiole configuration. Relative to other groups of monomorphic myrmicines, there is a great deal of size variation, with worker HW ranging from 0.45 to 0.65 mm. The eyes vary from 7-8 to 20-22 ommatidia, with this number correlated with size. In color, the majority of the specimens are black with brown appendages and yellowish extremities. The normal pilosity is moderately long, although very long in some small brown workers (Costa Rica, Barbas Volcano, 800 m). The petiole typically has a short peduncle, curving continuously into the dorsal rounded face; in some workers, the petiole is somewhat more square, with the anterior and posterior faces almost parallel. The characteristics with the greatest variability are the sculpturing of the promesonotum and the size and form of the propodeal spines. The most typical tendency in worker sculpturing is the formation of rugulae or longitudinal carinae, either moderately or strongly marked, which form 14-15 coarse costulae over the dorsal area. Most of the time these costae or rugulae do not touch the anterior border of the pronotum, instead being replaced by oblique or transverse rugae. In a few cases these rugae reach the anterior margin of the pronotum, and in some workers they are concentric around a few central longitudinal rugae. In a few other workers, they are mostly oblique, almost transverse. Figures 23, 39 and 40 show some examples of the sculpturing. The teeth or propodeal spines are variable as well, from low and triangular (shorter than their bases), to elongated and slender in shape.</paragraph>
<paragraph id="243980428EAC65B5966DE1D7B45B16A0" lastPageNumber="33" pageNumber="32">
With such wide variation it would be easy to interpret the endpoints (in isolation) as different species, and in fact at the beginning of this revision some of the extreme examples seemed to represent new species, based especially on the size and sculpturing of the promesonotum. Nevertheless, critical examination of wider samples, from
<normalizedToken id="D3A01CAA013750F4937E929F89877EF2" originalValue="México">Mexico</normalizedToken>
to Colombia, reveals a continuum of size, sculpturing, and color mixtures. There are samples from the same locality and even the same nest in which the workers are different in size and promesonotum sculpturing. Under these circumstances, it is a challenge to identify characteristics by which species can be unambiguously separated. It is possible that
<taxonomicName id="A6FDF5FAFD84300322D2F99EB26E69D6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24999" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex tristani (Menozzi)" order="Hymenoptera" pageNumber="33" phylum="Arthropoda" rank="species" species="tristani">A. tristani</taxonomicName>
is actually a complex of closely-related species that have very recently undergone (or are undergoing) speciation. Populations in valleys or isolated mountains might be in the process of becoming discontinuous from neighboring populations, with accompanying differences in size, color, pilosity, or habits. Males associated with workers (lacking to date) could provide important information for evaluating these alternatives. Bigger samples, males, and genetic study should elucidate whether
<taxonomicName id="0E8F89F2B3D9F8FE2B6EF48674D44CB4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24999" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex tristani (Menozzi)" order="Hymenoptera" pageNumber="33" phylum="Arthropoda" rank="species" species="tristani">A. tristani</taxonomicName>
is an extremely variable species or a set of closely-related species.
</paragraph>
<paragraph id="18DE69885FA08296F843AFAFD2024876" pageNumber="33">
Some of the biggest specimens of
<taxonomicName id="B499B82E0B8D23A03DDBFDC532577E21" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24999" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex tristani (Menozzi)" order="Hymenoptera" pageNumber="33" phylum="Arthropoda" rank="species" species="tristani">A. tristani</taxonomicName>
look like
<taxonomicName id="5B5F3F22D9ED93F8338632046798FB40" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182913" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex robustus Fernández" order="Hymenoptera" pageNumber="33" phylum="Arthropoda" rank="species" species="robustus">A. robustus</taxonomicName>
. Nevertheless, the latter has a typical HW equal or greater than 0.83 mm, and most are around 0.85 mm. The rugulae of the head and promesonotum in
<taxonomicName id="4A8141E1462B0BB584F34A0F1AF3DB27" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182913" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex robustus Fernández" order="Hymenoptera" pageNumber="33" phylum="Arthropoda" rank="species" species="robustus">A. robustus</taxonomicName>
are more irregular, never straight, with more abundant and marked piliferous punctures. The tooth of the basal margin is large and robust, as big as or bigger than the subapical tooth of the masticatory margin; the clypeal teeth are also conspicuous. The propodeal spines are always low and black.
</paragraph>
</subSubSection>
<subSubSection id="AEBB809B148DDDB8213E7457FBAD7A06" type="distribution">
<paragraph id="67D915CBE51F0A882C764015438FD6D3" pageNumber="33">
<taxonomicName id="49561BDB9EDB8885989303DE66028108" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:182913" class="Insecta" family="Formicidae" genus="Adelomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Adelomyrmex robustus Fernández" order="Hymenoptera" pageNumber="33" phylum="Arthropoda" rank="species" species="robustus">A. robustus</taxonomicName>
is known only from Mexico.
</paragraph>
</subSubSection>
</treatment>
</document>
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