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<document id="517FAA0C68188841A77683D42B6694FD" ID-CLB-Dataset="45463" ID-DOI="10.5281/zenodo.210568" ID-GBIF-Dataset="d418cb1f-e1fd-420d-9483-cbcbb7d4f84e" ID-ISSN="1175-5326" ID-Zenodo-Dep="210568" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460438124540" checkinUser="plazi" docAuthor="Naglis, Stefan &amp; Bickel, Daniel J." docDate="2012" docId="A67E87A6845BFF81F5D5FC1C2B9AFE39" docLanguage="en" docName="zt03188p063.pdf" docOrigin="Zootaxa 3188" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Medetera nuwarensis Naglis &amp; Bickel, 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="60" masterDocId="5A47FFDE845FFF84F542FF8C2E01FFF6" masterDocTitle="Medetera (Diptera, Dolichopodidae) of Sri Lanka" masterLastPageNumber="63" masterPageNumber="55" pageNumber="59" updateTime="1698332490315" updateUser="ExternalLinkService">
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<mods:title id="E9D0DB0E9601DC00CC2D6745985D60F9">Medetera (Diptera, Dolichopodidae) of Sri Lanka</mods:title>
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<mods:namePart id="D9D0452F757CAE7A69AD5B2667EC7153">Naglis, Stefan</mods:namePart>
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<mods:namePart id="7A0D38820D319695316BE1E1C39A7BEA">Bickel, Daniel J.</mods:namePart>
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<mods:date id="A9E49C6574B8844799BF9AB671166458">2012</mods:date>
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<treatment id="A67E87A6845BFF81F5D5FC1C2B9AFE39" ID-DOI="http://doi.org/10.5281/zenodo.6170185" ID-GBIF-Taxon="119507113" ID-Zenodo-Dep="6170185" LSID="urn:lsid:plazi:treatment:A67E87A6845BFF81F5D5FC1C2B9AFE39" httpUri="http://treatment.plazi.org/id/A67E87A6845BFF81F5D5FC1C2B9AFE39" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59">
<subSubSection id="66CD653B845BFF80F5D5FC1C2F2AFC3D" pageId="4" pageNumber="59" type="nomenclature">
<paragraph id="2E6836B0845BFF80F5D5FC1C2FFEFC5C" blockId="4.[151,511,912,971]" box="[151,511,912,938]" pageId="4" pageNumber="59">
<heading id="752081DC845BFF80F5D5FC1C2FFEFC5C" bold="true" box="[151,511,912,938]" fontSize="11" level="1" pageId="4" pageNumber="59" reason="1">
<emphasis id="1CA3EAA2845BFF80F5D5FC1C2FFEFC5C" bold="true" box="[151,511,912,938]" pageId="4" pageNumber="59">
<taxonomicName id="E9D74D33845BFF80F5D5FC1C2F9AFC5C" box="[151,411,912,938]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="nuwarensis" status="sp. nov.">
<emphasis id="1CA3EAA2845BFF80F5D5FC1C2F9AFC5C" bold="true" box="[151,411,912,938]" italics="true" pageId="4" pageNumber="59">Medetera nuwarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="079057D9845BFF80F4E3FC1C2FFEFC5C" box="[417,511,912,938]" pageId="4" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2E6836B0845BFF80F5D5FC3F2F2AFC3D" blockId="4.[151,511,912,971]" box="[151,299,947,971]" pageId="4" pageNumber="59">
(
<figureCitation id="B6EC2A35845BFF80F5DDFC3F2EECFC3D" box="[159,237,947,971]" captionStart="FIGURE 4" captionStartId="5.[151,250,1153,1176]" captionTargetBox="[156,1420,526,1120]" captionTargetId="figure@5.[151,1436,515,1131]" captionTargetPageId="5" captionText="FIGURE 4. Medetera nuwarensis sp. nov. male hypopygium: (A) left lateral view; (B) hypandrium and aedeagus, ventral view (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/210572/files/figure.png" pageId="4" pageNumber="59">Figs. 4</figureCitation>
A, B)
</paragraph>
</subSubSection>
<subSubSection id="66CD653B845BFF80F5D5FC702A9EFBAD" pageId="4" pageNumber="59" type="materials_examined">
<paragraph id="2E6836B0845BFF80F5D5FC702A9EFBAD" blockId="4.[151,1437,1019,2018]" pageId="4" pageNumber="59">
<emphasis id="1CA3EAA2845BFF80F5D5FC702C68FBE2" bold="true" box="[151,617,1019,1044]" pageId="4" pageNumber="59">
<typeStatus id="F16C8812845BFF80F5D5FC702ED3FBE2" box="[151,210,1020,1044]" pageId="4" pageNumber="59">Type</typeStatus>
material examined.
<typeStatus id="F16C8812845BFF80F488FC772C68FBE2" box="[458,617,1019,1044]" pageId="4" pageNumber="59" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
:
<emphasis id="1CA3EAA2845BFF80F738FC772D17FBE2" bold="true" box="[634,790,1019,1044]" pageId="4" pageNumber="59">
<collectingCountry id="56C07620845BFF80F738FC772D17FBE2" box="[634,790,1019,1044]" name="Sri Lanka" pageId="4" pageNumber="59">
<collectionCode id="48C6AE75845BFF80F738FC772CA8FBE2" box="[634,681,1019,1044]" httpUri="http://grbio.org/cool/z5p4-dpar" name="Serengetti Research Institute" pageId="4" pageNumber="59">SRI</collectionCode>
<collectionCode id="48C6AE75845BFF80F7F3FC772D17FBE2" box="[689,790,1019,1044]" pageId="4" pageNumber="59">LANKA</collectionCode>
</collectingCountry>
</emphasis>
. 3: “
<collectingCountry id="56C07620845BFF80F614FC772DC6FBE5" box="[854,967,1019,1043]" name="Sri Lanka" pageId="4" pageNumber="59">Sri Lanka</collectingCountry>
N. E. Dist. Nuwara Eliya blacklight trap
<date id="5A691070845BFF80F5D5FB932F41FBC1" box="[151,320,1055,1079]" pageId="4" pageNumber="59" value="1975-09-26" valueMax="1975-09-27" valueMin="1975-09-26">26-27.IX.1975</date>
”; “collected by D.M. Davies, S. Karunaratne, D.W. Balasooriya” (
<collectionCode id="48C6AE75845BFF80F115FB932AAFFBC1" LSID="urn:lsid:biocol.org:col:34871" box="[1111,1198,1055,1079]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="59">USNM</collectionCode>
).
<emphasis id="1CA3EAA2845BFF80F187FB932B58FBCE" bold="true" box="[1221,1369,1055,1080]" pageId="4" pageNumber="59">
<typeStatus id="F16C8812845BFF80F187FB932B58FBCE" box="[1221,1369,1055,1080]" pageId="4" pageNumber="59" type="paratype">PARATYPE</typeStatus>
</emphasis>
:
<emphasis id="1CA3EAA2845BFF80F02FFB932EFDFBAA" bold="true" pageId="4" pageNumber="59">
<collectingCountry id="56C07620845BFF80F02FFB932EFDFBAA" name="Sri Lanka" pageId="4" pageNumber="59">
<collectionCode id="48C6AE75845BFF80F02FFB932B9DFBCE" box="[1389,1436,1055,1080]" httpUri="http://grbio.org/cool/z5p4-dpar" name="Serengetti Research Institute" pageId="4" pageNumber="59">SRI</collectionCode>
<collectionCode id="48C6AE75845BFF80F5D5FBCF2EFDFBAA" box="[151,252,1091,1116]" pageId="4" pageNumber="59">LANKA</collectionCode>
</collectingCountry>
</emphasis>
. 1 Ƥ: Bad. Dist. Kande-ela Reservoir,
<quantity id="E92F9B55845BFF80F7ECFBCF2D02FBAD" box="[686,771,1091,1115]" metricMagnitude="3" metricUnit="m" metricValue="1.8897599999999997" pageId="4" pageNumber="59" unit="ft" value="6200.0">6200 ft</quantity>
.,
<date id="5A691070845BFF80F651FBCF2D92FBAD" box="[787,915,1091,1115]" pageId="4" pageNumber="59" value="1970-10-01" valueMax="1970-10-05" valueMin="1970-10-01">15.x.1970</date>
, O.S. Flint Jr. (
<collectionCode id="48C6AE75845BFF80F17EFBCF2A92FBAD" LSID="urn:lsid:biocol.org:col:34871" box="[1084,1171,1091,1115]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="59">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="66CD653B845BFF80F585FBEB2C90F8A4" pageId="4" pageNumber="59" type="description">
<paragraph id="2E6836B0845BFF80F585FBEB2C90F8A4" blockId="4.[151,1437,1019,2018]" pageId="4" pageNumber="59">
<emphasis id="1CA3EAA2845BFF80F585FBEB2F9DFB76" bold="true" box="[199,412,1127,1152]" pageId="4" pageNumber="59">Description. Male.</emphasis>
Length (
<typeStatus id="F16C8812845BFF80F4BFFBEB2C62FB89" box="[509,611,1127,1151]" pageId="4" pageNumber="59" type="holotype">holotype</typeStatus>
): Body
<quantity id="E92F9B55845BFF80F7F4FBEB2D0DFB89" box="[694,780,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="59" unit="mm" value="2.4">2.4 mm</quantity>
, wing
<quantity id="E92F9B55845BFF80F611FBEB2DA8FB89" box="[851,937,1127,1151]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="59" unit="mm" value="2.7">2.7 mm</quantity>
.
<emphasis id="1CA3EAA2845BFF80F6F1FBE42DFDFB76" bold="true" box="[947,1020,1128,1152]" pageId="4" pageNumber="59">Head:</emphasis>
Face and clypeus dark metallic blueviolet with little grey pruinosity; frons dark metallic green with dense grey pruinosity; palpus and proboscis dark brown; scape and pedicel yellow, first flagellomere black; lower postoculars white.
<emphasis id="1CA3EAA2845BFF80F17DFB3C2AA0FB3E" bold="true" box="[1087,1185,1200,1224]" pageId="4" pageNumber="59">Thorax:</emphasis>
Dorsum dark metallic bronze-green with dusting of grayish-brown pruinosity; all thoracic setae black; 5 strong dc, and some hair-like anterior setae; ac strong and long, posteriormost ac as long as anteriormost dc; 2 strong scutellars and 2 lateral setae two-thirds length of medians; pleura dark metallic bronze-green, proepisternum with 2 strong setae below.
<emphasis id="1CA3EAA2845BFF80F019FA902B9DFAC2" bold="true" box="[1371,1436,1308,1332]" pageId="4" pageNumber="59">Legs:</emphasis>
Entirely black, tip of femora pale; all setae black, except as noted. Fore leg: bare of major setae; relative podomere ratios: 49/50/28/
<date id="5A691070845BFF80F41BFAEF2FADFA8D" box="[345,428,1379,1403]" pageId="4" pageNumber="59" value="2010-08-17">10/17/8</date>
/5. Mid leg: coxa with 3 strong anterolateral setae; femur with av row of setae, length nearly subequal to diameter of femur; tibia with strong ad/pd setal pair at 1/4, and 1 apicoventral seta; relative podomere ratios: 56/60/42/
<date id="5A691070845BFF80F490FA272C24FA35" box="[466,549,1451,1475]" pageId="4" pageNumber="59" value="1919-08-16">19/16/8</date>
/5.
<collectingCountry id="56C07620845BFF80F705FA272C7EFA35" box="[583,639,1451,1475]" name="India" pageId="4" pageNumber="59">Hind</collectingCountry>
leg: coxa with strong lateral seta; femur with anterior and av row of strong setae; tibia lacking major setae, with black pv apical tooth-like projection (MSSC); basitarsus with small basal tubercle opposite of tibial tooth (MSSC); relative podomere ratios: 64/87/19/42/
<date id="5A691070845BFF80F12DFA7F2AC9F9FD" box="[1135,1224,1523,1547]" pageId="4" pageNumber="59" value="1923-09-14">23/14/9.</date>
<emphasis id="1CA3EAA2845BFF80F192FA7F2B18F9FA" bold="true" box="[1232,1305,1523,1548]" pageId="4" pageNumber="59">Wing:</emphasis>
Membrane slightly darkened; veins black; R4+5 slightly arching posteriad distally, R4+5 and M converging and becoming subparallel before wing margin; CuAx ratio: 0.6; RMx ratio: 4.8; lower calypter yellowish-brown with fan of yellowish setae; halter yellowish with dark brown stem.
<emphasis id="1CA3EAA2845BFF80F783F9EE2D3EF98D" bold="true" box="[705,831,1634,1659]" pageId="4" pageNumber="59">Abdomen:</emphasis>
Dark metallic green; setae black Male genitalia (
<figureCitation id="B6EC2A35845BFF80F026F9EE2EA7F968" captionStart="FIGURE 4" captionStartId="5.[151,250,1153,1176]" captionTargetBox="[156,1420,526,1120]" captionTargetId="figure@5.[151,1436,515,1131]" captionTargetPageId="5" captionText="FIGURE 4. Medetera nuwarensis sp. nov. male hypopygium: (A) left lateral view; (B) hypandrium and aedeagus, ventral view (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/210572/files/figure.png" pageId="4" pageNumber="59">Figs. 4</figureCitation>
A, B): hypandrium with rounded apex, with narrow lobe on either side; epandrial seta absent; epandrial lobes well separated each bearing strong curved seta; surstylus trifurcate, dorsal lobe narrow and acute, with apical and subapical seta, ventral lobe rounded, with blade-like subapical and 3 normal setae; cercus elongate, with strong, hornlike apical projection and 2 blade-like and 1 normal setae ventrally; aedeagus with dorsal bifurcate process projecting laterad of cercus (= “bottle-brush-like” aedeagal projection
<emphasis id="1CA3EAA2845BFF80F61CF89B2D9DF8D8" box="[862,924,1815,1838]" italics="true" pageId="4" pageNumber="59">sensu</emphasis>
<bibRefCitation id="4A464B41845BFF80F6E6F89A2A4AF8D8" author="Bickel" box="[932,1099,1814,1838]" pageId="4" pageNumber="59" refString="Bickel, D. J. (1985) A revision of the Nearctic Medetera (Diptera: Dolichopodidae). United States Department of Agriculture Technical Bulletin, 1692, 1 - 109." type="journal article" year="1985">Bickel (1985))</bibRefCitation>
.
<emphasis id="1CA3EAA2845BFF80F114F89A2AB7F8D8" bold="true" box="[1110,1206,1814,1838]" pageId="4" pageNumber="59">Female:</emphasis>
Similar to male, but tibiae yellowish-brown, wings less darkened.
</paragraph>
</subSubSection>
<subSubSection id="66CD653B845BFF80F585F8D22D3EF880" box="[199,831,1886,1911]" pageId="4" pageNumber="59" type="etymology">
<paragraph id="2E6836B0845BFF80F585F8D22D3EF880" blockId="4.[151,1437,1019,2018]" box="[199,831,1886,1911]" pageId="4" pageNumber="59">
<emphasis id="1CA3EAA2845BFF80F585F8D22F4FF880" bold="true" box="[199,334,1886,1910]" pageId="4" pageNumber="59">Etymology.</emphasis>
The species is named after the
<typeStatus id="F16C8812845BFF80F7EFF8D32CDCF881" box="[685,733,1887,1911]" pageId="4" pageNumber="59">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="66CD653B845BFF81F585F80F2B9AFE39" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59" type="discussion">
<paragraph id="2E6836B0845BFF81F585F80F2A51FF01" blockId="4.[151,1437,1019,2018]" lastBlockId="5.[151,1436,151,247]" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59">
<emphasis id="1CA3EAA2845BFF80F585F80F2F3AF86D" bold="true" box="[199,315,1923,1947]" pageId="4" pageNumber="59">Remarks.</emphasis>
<emphasis id="1CA3EAA2845BFF80F404F80E2C2EF86C" box="[326,559,1922,1946]" italics="true" pageId="4" pageNumber="59">
Medtera
<taxonomicName id="E9D74D33845BFF80F4F2F80F2C2EF86C" box="[432,559,1923,1946]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="nuwarensis">nuwarensis</taxonomicName>
</emphasis>
probably represents a new species-group. It is similar to
<taxonomicName id="E9D74D33845BFF80F19AF80F2F27F848" authority="Bickel, 1987" authorityName="Bickel" authorityYear="1987" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="1CA3EAA2845BFF80F19AF80F2B9DF86C" box="[1240,1436,1922,1946]" italics="true" pageId="4" pageNumber="59">M. kinabaluensis</emphasis>
<bibRefCitation id="4A464B41845BFF80F5D5F82A2F27F848" author="Bickel" box="[151,294,1958,1982]" pageId="4" pageNumber="59" refString="Bickel, D. J. (1987) A revision of the Oriental and Australasian Medetera (Diptera: Dolichopodidae). Records of the Australian Museum, 39, 195 - 259." type="journal article" year="1987">Bickel, 1987</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="56C07620845BFF80F42EF82A2FD7F848" box="[364,470,1958,1982]" name="Malaysia" pageId="4" pageNumber="59">Malaysia</collectingCountry>
, the single species of the
<emphasis id="1CA3EAA2845BFF80F7B5F82A2DEFF848" box="[759,1006,1958,1982]" italics="true" pageId="4" pageNumber="59">signaticornis-pinicola</emphasis>
species-group in the Oriental Region. Both species can be separated by the characters given below. The species bears a long, narrow, apically bifurcate dorsal projection, the “bottle-brush-like” aedeagal projection according to
<bibRefCitation id="4A464B41845AFF81F69AFF1B2A6AFF59" author="Bickel" box="[984,1131,151,175]" pageId="5" pageNumber="60" refString="Bickel, D. J. (1985) A revision of the Nearctic Medetera (Diptera: Dolichopodidae). United States Department of Agriculture Technical Bulletin, 1692, 1 - 109." type="journal article" year="1985">Bickel (1985</bibRefCitation>
,
<bibRefCitation id="4A464B41845AFF81F135FF1B2AB2FF59" author="Bickel" box="[1143,1203,151,175]" pageId="5" pageNumber="60" refString="Bickel, D. J. (1987) A revision of the Oriental and Australasian Medetera (Diptera: Dolichopodidae). Records of the Australian Museum, 39, 195 - 259." type="journal article" year="1987">1987</bibRefCitation>
), but in
<taxonomicName id="E9D74D33845AFF81F04EFF152EB4FF22" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="nuwarensis">
<emphasis id="1CA3EAA2845AFF81F04EFF152EB4FF22" italics="true" pageId="5" pageNumber="60">M. nuwarensis</emphasis>
</taxonomicName>
the apex of this projection is not hairy as in other species of this group. Moreover,
<taxonomicName id="E9D74D33845AFF81F122FF312B04FF22" box="[1120,1285,189,212]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="nuwarensis">
<emphasis id="1CA3EAA2845AFF81F122FF312B04FF22" box="[1120,1285,189,212]" italics="true" pageId="5" pageNumber="60">M. nuwarensis</emphasis>
</taxonomicName>
has a narrow lobe on either side of the hypandrium, a unique character within the Oriental species.
</paragraph>
<paragraph id="2E6836B0845AFF81F5D5FEAB2B9DFE81" blockId="5.[151,1436,295,463]" pageId="5" pageNumber="60">
1 Body length more than
<quantity id="E92F9B55845AFF81F487FEAB2FFEFECA" box="[453,511,295,316]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="60" unit="mm" value="3.0">3 mm</quantity>
; lower calypter with pale setae; cercus distally bulbous; hypandrium with pointed apex, without lateral lobes; aedeagus without dorsal process projecting near base of cercus (see
<bibRefCitation id="4A464B41845AFF81F6AFFEC92A65FEAC" author="Bickel" box="[1005,1124,325,346]" pageId="5" pageNumber="60" refString="Bickel, D. J. (1987) A revision of the Oriental and Australasian Medetera (Diptera: Dolichopodidae). Records of the Australian Museum, 39, 195 - 259." type="journal article" year="1987">Bickel 1987</bibRefCitation>
, figs. 19a, b)...................................................................................................
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<emphasis id="1CA3EAA2845AFF81F191FEEE2B56FE81" box="[1235,1367,354,375]" italics="true" pageId="5" pageNumber="60">kinabaluensis</emphasis>
Bickel
</taxonomicName>
</paragraph>
<paragraph id="2E6836B0845AFF81F5D5FEF32B9AFE39" blockId="5.[151,1436,295,463]" pageId="5" pageNumber="60">
- Body length less than
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; lower calypter with dark setae; cercus distally tapering; hypandrium with rounded apex, with lateral lobes on either side; aedeagus with long, narrow, bifurcate dorsal process projecting near base of cercus (
<figureCitation id="B6EC2A35845AFF81F1BBFE112B3DFE44" box="[1273,1340,413,434]" captionStart="FIGURE 4" captionStartId="5.[151,250,1153,1176]" captionTargetBox="[156,1420,526,1120]" captionTargetId="figure@5.[151,1436,515,1131]" captionTargetPageId="5" captionText="FIGURE 4. Medetera nuwarensis sp. nov. male hypopygium: (A) left lateral view; (B) hypandrium and aedeagus, ventral view (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/210572/files/figure.png" pageId="5" pageNumber="60">Figs. 4</figureCitation>
A, B).......................................................................................
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<emphasis id="1CA3EAA2845AFF81F19FFE362B48FE39" box="[1245,1353,442,463]" italics="true" pageId="5" pageNumber="60">nuwarensis</emphasis>
</taxonomicName>
<emphasis id="1CA3EAA2845AFF81F00DFE362B9AFE39" bold="true" box="[1359,1435,442,463]" pageId="5" pageNumber="60">
<taxonomicNameLabel id="079057D9845AFF81F00DFE362B9AFE39" box="[1359,1435,442,463]" pageId="5" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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