treatments-xml/data/A6/7E/52/A67E52E71A76CD44CAFA44CE53C0720F.xml

327 lines
44 KiB
XML

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.745.23685" ID-PMC="PMC5904534" ID-Pensoft-Pub="1313-2970-745-101" ID-Pensoft-UUID="047DFFCAFFA5F32EA97C873F4708943F" ID-PubMed="29670449" ID-Zenodo-Dep="1222435" ID-ZooBank="8785DC6BC2A244FD94B6243EB07C717F" ModsDocID="1313-2970-745-101" checkinTime="1521799180220" checkinUser="pensoft" docAuthor="Moore, Matthew R., Cave, Ronald D. &amp; Branham, Marc A." docDate="2018" docId="A67E52E71A76CD44CAFA44CE53C0720F" docLanguage="en" docName="ZooKeys 745: 101-378" docOrigin="ZooKeys 745" docPubDate="2018-03-22" docSource="http://dx.doi.org/10.3897/zookeys.745.23685" docTitle="Ancognatha vulgaris Arrow 1911" docType="treatment" docVersion="7" id="047DFFCAFFA5F32EA97C873F4708943F" lastPageNumber="118" masterDocId="047DFFCAFFA5F32EA97C873F4708943F" masterDocTitle="Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)" masterLastPageNumber="378" masterPageNumber="101" pageNumber="117" updateTime="1740595812083" updateUser="felipe">
<mods:mods id="295489AB7CE92C986B124A97667E54FA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A6F43999F39C2C5E4957390D8B409CB3">
<mods:title id="38987176AB86F571111B50F76EC11D0F">Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)</mods:title>
</mods:titleInfo>
<mods:name id="4150218933B5F67D7AED872117B8B095" type="personal">
<mods:role id="37849EA50A4CE0C132F38697E5766E91">
<mods:roleTerm id="813A7510B9667F5EFE5DAA420691507E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31263564F40E9728B440B21F3A97E84D">Moore, Matthew R.</mods:namePart>
<mods:affiliation id="E7AC903FE38473B015B5B51898A6254C">Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
<mods:nameIdentifier id="96410FEFE248EA7A80432EB8B5FB4DD6" type="email">cyclocephala@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="86CAC17957B0098D0CD5EE4610E44403" type="personal">
<mods:role id="D3EF289543EF5AD1ECD61B061595E400">
<mods:roleTerm id="728FEC8C51D8A1EEBE2A779C6BD5CAD0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="07A9C618F0D73E9E1E228CD77D881C9A">Cave, Ronald D.</mods:namePart>
<mods:affiliation id="098058A20EC7E9A1FA9F6A6F4F7CF1C2">Department of Entomology and Nematology, University of Florida, Indian River Research and Education Center, 2199 South Rock Road, Fort Pierce, FL 34945, USA</mods:affiliation>
</mods:name>
<mods:name id="15D0CB7F86B756C600063A65D711EA70" type="personal">
<mods:role id="FA442A1F3ECA62C009BAFA19F08B084E">
<mods:roleTerm id="A7A0CC383CAED7B4AA344A4AFB43568E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2B344961AD6A7FB0B6FA6D93E9E81896">Branham, Marc A.</mods:namePart>
<mods:affiliation id="D59EC5B3FA71308244043B0E46992FCC">Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E8E6B7608B3EB8A2A15A013D43872974">text</mods:typeOfResource>
<mods:relatedItem id="83799F51BFF91918BCF113607259ED95" type="host">
<mods:titleInfo id="00A0913906C7449BC21301C8259B5849">
<mods:title id="B259D1BEAAD8B054EEDA4BEDD0D11EFE">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="ABC9439521675EFA5F8EB13BF518438C">
<mods:date id="4EA7D6B22614B336FEE823E2495CFF16">2018</mods:date>
<mods:detail id="714AAEFBED6F4286E18AC72AD6897F26" type="pubDate">
<mods:number id="3E4C420C1198160E1232510AB14C077F">2018-03-22</mods:number>
</mods:detail>
<mods:detail id="61E33E02A23843043B769D07D43D0B87" type="volume">
<mods:number id="B45EA55BEE4488B4AB0F3C985E979A4D">745</mods:number>
</mods:detail>
<mods:extent id="54732E673357AE47E1FF579A24C2088A" unit="page">
<mods:start id="0F419EC75B82AEBA441F3D8C11651C24">101</mods:start>
<mods:end id="07378E0A57E35DCB80D49E9E4EBC1D12">378</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C643B45C737614ACFE0881C8916A179F">
<mods:url id="4AFBACCE2ECE21D5E59BB38127084C76">http://dx.doi.org/10.3897/zookeys.745.23685</mods:url>
</mods:location>
<mods:classification id="2BA3DCBED2C72B4B74E696BC9C1942F6">journal article</mods:classification>
<mods:identifier id="9D3E0456ED536F13D519F846E63B32A8" type="DOI">http://dx.doi.org/10.3897/zookeys.745.23685</mods:identifier>
<mods:identifier id="0814D63958F2D1D51270C2BC00A780EC" type="Pensoft-Pub">1313-2970-745-101</mods:identifier>
<mods:identifier id="D64FD6B54478F7C2BF983792D26CE024" type="ZooBank">8785DC6BC2A244FD94B6243EB07C717F</mods:identifier>
<mods:identifier id="54ADAD40EE37384C71CDA00106E435FF" type="Pensoft-UUID">047DFFCAFFA5F32EA97C873F4708943F</mods:identifier>
<mods:identifier id="D05404F26AA41954896F3ED13A5BCEA4" type="Zenodo-Dep">1222435</mods:identifier>
</mods:mods>
<treatment id="A67E52E71A76CD44CAFA44CE53C0720F" ID-GBIF-Taxon="142480105" LSID="urn:lsid:plazi:treatment:A67E52E71A76CD44CAFA44CE53C0720F" httpUri="http://treatment.plazi.org/id/A67E52E71A76CD44CAFA44CE53C0720F" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117">
<subSubSection id="F97B762C74F6438C0ADF23037FA39252" pageId="16" pageNumber="117" type="nomenclature">
<paragraph id="4DDD6E0B4B607160DCC489AB7111421E" pageId="16" pageNumber="117">
<taxonomicName id="6176B714EC9BD5722BEAD47014D73F00" ID-CoL="DN9B" authority="Arrow, 1911" authorityName="Arrow" authorityYear="1911" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="vulgaris">Ancognatha vulgaris Arrow, 1911</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0BB47963AA66CCBC576C355251D0A496" pageId="16" pageNumber="117" type="reference_group">
<paragraph id="8CD3E502F6A6BB9A1A655FC4ECE7FC6A" pageId="16" pageNumber="117">
<taxonomicName id="E2AC17337325CAC73F898ECF3FCFC313" ID-CoL="DN9B" authority="Arrow, 1911: 169 - 170" authorityName="Arrow" authorityPageNumber="169 - 170" authorityYear="1911" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="vulgaris">Ancognatha vulgaris Arrow, 1911: 169-170</taxonomicName>
[original combination].
</paragraph>
<paragraph id="562A6376225D64ABD4EFE3A1B1A7EEBC" pageId="16" pageNumber="117">
<emphasis id="4EC2879E4A6DC3BA6365A7C9E8023DAA" bold="true" pageId="16" pageNumber="117">syn.</emphasis>
<taxonomicName id="D5BA6380C070FD97C6D37C4C9B8FCB26" ID-CoL="DN8F" authority="Otoya, 1945: 275 - 282" authorityName="Otoya" authorityPageNumber="275 - 282" authorityYear="1945" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nigriventris" subGenus="Pseudoancognatha">Ancognatha (Pseudoancognatha) nigriventris Otoya, 1945: 275-282</taxonomicName>
[original combination].
</paragraph>
<paragraph id="AC7162F63A693986C9AD316C6177D38B" pageId="16" pageNumber="117">
<taxonomicName id="ED599BD9DBF3F34CA8096D48C6AA4245" ID-CoL="DN9B" authority="Arrow" authorityName="Arrow" authorityYear="1911" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="vulgaris">Ancognatha vulgaris Arrow</taxonomicName>
[synonymy by
<bibRefCitation id="C7A0AD8AF2EC4AA4C214C90274CBC130" author="Martinez, A" pageId="205" pageNumber="306" refId="B744" refString="Martinez, A, 1965a. Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae). Anales de la Sociedad Cientifica Argentina 179: 63-74." title="Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae). Anales de la Sociedad Cientifica Argentina 179: 63 - 74." year="1965 a">
<normalizedToken id="04425F55D071DCF68CDFB991CA9244C7" originalValue="Martínez">Martinez</normalizedToken>
1965a
</bibRefCitation>
: 64].
</paragraph>
</subSubSection>
<subSubSection id="9BEFE04647DBF91C743C412EB14D2FB9" pageId="16" pageNumber="117" type="types">
<paragraph id="518D2746B52060F8A6C52B273B235163" pageId="16" pageNumber="117">Types.</paragraph>
<paragraph id="B45F7496EDE67CC9FBD6DEB704E9F289" pageId="16" pageNumber="117">
Type of
<taxonomicName id="DA74EBE247DF83C2AF14810CE3F8A9E2" authorityName="Arrow" authorityYear="1911" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="7C0EFF2EE0FDDDCB6D3838712399B080" italics="true" pageId="16" pageNumber="117">A. vulgaris</emphasis>
</taxonomicName>
at BMNH (
<bibRefCitation id="AF3F1A6827A1372C30BF379A2C76F86E" author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken id="30980D7EB61D0EAA7A0107D64F5F36AF" originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
). Holotype ♂ of
<taxonomicName id="24A29A892981FE7A507C4269DE62D291" authorityName="Otoya" authorityYear="1945" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="AD6BD8809D38C95768847E291114A810" italics="true" pageId="16" pageNumber="117">A. nigriventris</emphasis>
</taxonomicName>
at ICN (
<bibRefCitation id="C972164B1DD575DB09F21444B8C2E6F3" author="Otoya, IA" pageId="211" pageNumber="312" refId="B861" refString="Otoya, IA, 1945. Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273-292." title="Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273 - 292." year="1945">Otoya 1945</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A48D9A0F294410BE16082020B36D5DF2" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117" type="distribution">
<paragraph id="96D18917456E3D8E452B87E5E1483FD1" pageId="16" pageNumber="117">Distribution.</paragraph>
<paragraph id="4A78F0AA0973C3D3AF6470A4D0438BDD" lastPageId="17" lastPageNumber="118" pageId="16" pageNumber="117">
BOLIVIA. BRAZIL: Amazonas. COLOMBIA: Antioquia,
<normalizedToken id="D521F9A347410D019E43376B29E11A70" originalValue="Boyacá">Boyaca</normalizedToken>
,
<taxonomicName id="C3E2E3AD0985C7CF1085366050A9B4B4" class="Insecta" family="Cerambycidae" genus="Cauca" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">Cauca</taxonomicName>
, Cundinamarca, Huila, Magdalena, Meta,
<normalizedToken id="B7BDC4A402C17FFBE35A906964D1C4D9" originalValue="Nariño">Narino</normalizedToken>
, Norte de Santander,
<normalizedToken id="88EF287E233DF211C59D18E8D9887C6E" originalValue="Quindío">Quindio</normalizedToken>
, Risaralda, Santander, Tolima. COSTA RICA: Alajuela, Cartago, Guanacaste, Heredia,
<normalizedToken id="EB3B90106CD266FF553C64C3A6969E9E" originalValue="Limón">Limon</normalizedToken>
, Puntarenas, San
<normalizedToken id="44944BD182364B1E6EE7216E7B15827C" originalValue="José">Jose</normalizedToken>
. ECUADOR. PANAMA: Bocas del Toro,
<pageBreakToken id="D2890851A29AB09775EF4878B176150C" pageId="17" pageNumber="118" start="start">
<normalizedToken id="FA99569D3877CE8FA9EB379A201083C1" originalValue="Chiriquí">Chiriqui</normalizedToken>
</pageBreakToken>
,
<normalizedToken id="270734C446D748706A1B2789B3AEFE02" originalValue="Coclé">Cocle</normalizedToken>
,
<normalizedToken id="85A6B7081E93AFC23C1F38351BF34561" originalValue="Panamá">Panama</normalizedToken>
, Veraguas. PERU: Ayacucho, Cajamarca, Cuzco,
<normalizedToken id="59815232AC8F90E922EB2CC2513F5734" originalValue="Huánuco">Huanuco</normalizedToken>
, La Libertad, Lima, Loreto, Pasco, Piura, San
<normalizedToken id="4B4C4B63452827BDC57F76D942FC1248" originalValue="Martín">Martin</normalizedToken>
, Ucayali. VENEZUELA:
<normalizedToken id="CF30305900B16FA87C3E47DE42FD43D6" originalValue="Mérida">Merida</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="A5E8F6700216D88D6781C88ED3BCCCB9" pageId="17" pageNumber="118" type="references">
<paragraph id="FA0240F2C284BED35E77DDC60B3D514F" pageId="17" pageNumber="118">References.</paragraph>
<paragraph id="E5E069D2AA2E2E3E5AD9EA52F913D25D" pageId="17" pageNumber="118">
<bibRefCitation id="5A79A6958D4AF4470230F5F0A86650C9" author="Arrow, GJ" pageId="163" pageNumber="264" refId="B46" refString="Arrow, GJ, 1911. Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. The Annals and Magazine of Natural History 8 (8th series): 151-176." title="Notes on the coleopterous subfamily Dynastinae, with descriptions of new genera and species. The Annals and Magazine of Natural History 8 (8 th series): 151 - 176." year="1911">Arrow 1911</bibRefCitation>
,
<bibRefCitation id="E985B8ECB9BC35E69F8EADBF80A69F22" author="Arrow, GJ" pageId="164" pageNumber="265" refId="B51" refString="Arrow, GJ, 1937b. Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1-124." title="Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1 - 124." year="1937 b">1937b</bibRefCitation>
,
<bibRefCitation id="883AAE1B0422C806F65287C51EC04B4A" author="Blackwelder, RE" pageId="167" pageNumber="268" refId="B101" refString="Blackwelder, RE, 1944. Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189-341." title="Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189 - 341." year="1944">Blackwelder 1944</bibRefCitation>
,
<bibRefCitation id="F7AEB1E3041ACE613A82F9A44178C7EB" author="Otoya, IA" pageId="211" pageNumber="312" refId="B861" refString="Otoya, IA, 1945. Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273-292." title="Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273 - 292." year="1945">Otoya 1945</bibRefCitation>
,
<bibRefCitation id="40C930FE0C21A4E149C0750E973AC178" author="Gutierrez, AR" pageId="186" pageNumber="287" refId="B445" refString="Gutierrez, AR, 1950. Scarabaeidae del Norte de Chile. Anales de la Sociedad Cientifica Argentina 149 (2): 52-75." title="Scarabaeidae del Norte de Chile. Anales de la Sociedad Cientifica Argentina 149 (2): 52 - 75." year="1950">
<normalizedToken id="746AF23D5D620C5B156276251690BB51" originalValue="Gutiérrez">Gutierrez</normalizedToken>
1950
</bibRefCitation>
,
<bibRefCitation id="BDFA3E74EF6EC0E3D03BC7FDE61D87BB" author="Martinez, A" pageId="205" pageNumber="306" refId="B744" refString="Martinez, A, 1965a. Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae). Anales de la Sociedad Cientifica Argentina 179: 63-74." title="Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae). Anales de la Sociedad Cientifica Argentina 179: 63 - 74." year="1965 a">
<normalizedToken id="9D7580BE11ADE6F4C875E6950653E488" originalValue="Martínez">Martinez</normalizedToken>
1965a
</bibRefCitation>
,
<bibRefCitation id="81228E59B6A4C8F05D2491C8B8DDDA11" author="Howden, HF" pageId="194" pageNumber="295" refId="B575" refString="Howden, HF, Campbell, JM, 1974. Observations on some Scarabaeoidea in the Colombian Sierra Nevada de Santa Marta. The Coleopterists Bulletin 28: 109-114." title="Observations on some Scarabaeoidea in the Colombian Sierra Nevada de Santa Marta. The Coleopterists Bulletin 28: 109 - 114." year="1974">Howden and Campbell 1974</bibRefCitation>
,
<bibRefCitation id="806A5FB4FA4B9ACC8A7DB983760731EC" author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken id="83ADF28198483CB238D00E6188341CE3" originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
,
<bibRefCitation id="D8EF5EA4B85E58F14455355788ABDC45" author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
,
<bibRefCitation id="A27EDF06206DE1D1EBB9FDB90C36CFF0" author="Ruiz, BN" pageId="222" pageNumber="323" refId="B1047" refString="Ruiz, BN, Pamalpa, CN, 1990. Observaciones sobre las chisas (Coleoptera: Scarabaeidae) en Narino. Revista ICA: Publicación Científica del Instituto Colombiano Agropecuario 25: 275-282." title="Observaciones sobre las chisas (Coleoptera: Scarabaeidae) en Narino. Revista ICA: Publicacio ́ n Cienti ́ fica del Instituto Colombiano Agropecuario 25: 275 - 282." year="1990">Ruiz and Pamalpa 1990</bibRefCitation>
,
<bibRefCitation id="43CBCF25000A44788A02BAAA3F623427" author="Montoya, GC" pageId="207" pageNumber="308" refId="B791" refString="Montoya, GC, Madrigal-C, A, Ramirez, CA, 1994. Evaluacion de trampas de luz para el control de adultos de Scarabaeidae (Coleoptera) en cultivos de papa en La Union (Antioquia). Revista Colombiana de Entomologia 20: 130-136." title="Evaluacion de trampas de luz para el control de adultos de Scarabaeidae (Coleoptera) en cultivos de papa en La Union (Antioquia). Revista Colombiana de Entomologia 20: 130 - 136." year="1994">Montoya et al. 1994</bibRefCitation>
, Restrepo et al. 2003,
<bibRefCitation id="F2D969472F9EDF718933946EDC4CF7A6" DOI="https://doi.org/10.11646/zootaxa.32.1.1" author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B972" refString="Ratcliffe, BC, 2002a. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1-48. https://doi.org/10.11646/zootaxa.32.1.1" title="A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1 - 48." url="https://doi.org/10.11646/zootaxa.32.1.1" year="2002 a">Ratcliffe 2002a</bibRefCitation>
,
<bibRefCitation id="1184FA159DC1D7F36E9B94595624AE2D" author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B974" refString="Ratcliffe, BC, 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1-506." title="The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." year="2003">2003</bibRefCitation>
,
<bibRefCitation id="6261092E5DC180CBE4F02DAEE709274B" DOI="https://doi.org/10.1080/03670244.1997.9991520" author="Onore, G" pageId="210" pageNumber="311" refId="B858" refString="Onore, G, 1997. A brief note on edible insects in Ecuador. Ecology of Food and Nutrition 36: 277-285. https://doi.org/10.1080/03670244.1997.9991520" title="A brief note on edible insects in Ecuador. Ecology of Food and Nutrition 36: 277 - 285." url="https://doi.org/10.1080/03670244.1997.9991520" year="1997">Onore 1997</bibRefCitation>
,
<bibRefCitation id="A661D0756BF41984FB85D9AB4B02D6C4" author="Onore, G" pageId="211" pageNumber="312" refId="B859" refString="Onore, G, 2005. Edible insects in Ecuador. In: Paoletti MG (Ed.) Ecological Implications of Minilivestock: potential of insects, rodents, frogs, and snails. Science publishers INC., Enfield, 343-352." title="Edible insects in Ecuador. In: Paoletti MG (Ed.) Ecological Implications of Minilivestock: potential of insects, rodents, frogs, and snails. Science publishers INC., Enfield, 343 - 352." year="2005">2005</bibRefCitation>
,
<bibRefCitation id="0FEF35D9B0FA07BBE06C2D2BFF2633F5" DOI="https://doi.org/10.1653/0015-4040(2005)88[355:SACOTW]2.0.CO;2" author="Pardo-Locarno, LC" pageId="211" pageNumber="312" refId="B869" refString="Pardo-Locarno, LC, Montoya-Lerma, J, Bellotti, AC, van Schoonhoven, A, 2005a. Structure and composition of the white grub complex (Coleoptera: Scarabaeidae) in agroecological systems of northern Cauca, Colombia. Florida Entomologist 88: 355-363. https://doi.org/10.1653/0015-4040(2005)88[355:SACOTW]2.0.CO;2" title="Structure and composition of the white grub complex (Coleoptera: Scarabaeidae) in agroecological systems of northern Cauca, Colombia. Florida Entomologist 88: 355 - 363." url="https://doi.org/10.1653/0015-4040(2005)88[355:SACOTW]2.0.CO;2" year="2005 a">Pardo-Locarno et al. 2005a</bibRefCitation>
,
<bibRefCitation id="1C736AA12B0C4DAC838D719BDE68AAFD" author="Utima, OA" pageId="229" pageNumber="330" refId="B1157" refString="Utima, OA, Vallejo-E, LF, 2008. Escarabajos Melolonthidae (Scarabaeidae-Pleurosticti) de la Montana Cafetera, departamento de Risaralda, Colombia. Agronomica 16 (2): 31-44." title="Escarabajos Melolonthidae (Scarabaeidae-Pleurosticti) de la Montana Cafetera, departamento de Risaralda, Colombia. Agronomica 16 (2): 31 - 44." year="2008">
<normalizedToken id="159D1F3F256EA58D97732BBE693715A8" originalValue="Útima">Utima</normalizedToken>
and Vallejo 2008
</bibRefCitation>
,
<bibRefCitation id="D179EA980B008A56810F9614FF9697A9" DOI="https://doi.org/10.3897/zookeys.34.309" author="Gasca-Alvarez, HJ" pageId="182" pageNumber="283" refId="B381" refString="Gasca-Alvarez, HJ, Amat-Garcia, G, 2010. Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia. ZooKeys 34: 153-192. https://doi.org/10.3897/zookeys.34.309" title="Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia. ZooKeys 34: 153 - 192." url="https://doi.org/10.3897/zookeys.34.309" year="2010">
<normalizedToken id="336E0F03630A4A59ECD207C8AEF51A81" originalValue="Gasca-Álvarez">Gasca-Alvarez</normalizedToken>
and
<normalizedToken id="0924C96487E31B3B53626EF2C2DD00CE" originalValue="Amat-García">Amat-Garcia</normalizedToken>
2010
</bibRefCitation>
,
<bibRefCitation id="0EDED48900A740CF699E40D0CA76074A" author="Krajcik, M" pageId="200" pageNumber="301" refId="B658" refString="Krajcik, M, 2005. Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma.X, Supplement No. 2, Pilsen, 122 pp." title="Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma. X, Supplement No. 2, Pilsen, 122 pp." year="2005">Krajcik 2005</bibRefCitation>
,
<bibRefCitation id="A3C37CE7E8937DADFF64E46A542BAC06" author="Krajcik, M" pageId="200" pageNumber="301" refId="B659" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma.X Supplement 5, Pilsen, 278 pp." title="Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Pilsen, 278 pp." year="2012">2012</bibRefCitation>
,
<bibRefCitation id="1830AAA88B86328646C2FF823531814E" DOI="https://doi.org/10.1649/072.066.0303" author="Moore, MR" pageId="208" pageNumber="309" refId="B794" refString="Moore, MR, 2012. A new female elytral character for the tribe Cyclocephalini (Coleoptera: Scarabaeidae: Dynastinae) and an observation of its possible function. The Coleopterists Bulletin 66: 200-2002. https://doi.org/10.1649/072.066.0303" title="A new female elytral character for the tribe Cyclocephalini (Coleoptera: Scarabaeidae: Dynastinae) and an observation of its possible function. The Coleopterists Bulletin 66: 200 - 2002." url="https://doi.org/10.1649/072.066.0303" year="2012">Moore 2012</bibRefCitation>
,
<bibRefCitation id="A9972598260FC8F65A1F18C035A21E79" DOI="https://doi.org/10.1603/EN13123" author="Breeschoten, T" pageId="167" pageNumber="268" refId="B123" refString="Breeschoten, T, Clark, DR, Schilthuizen, M, 2013. Evolutionary patterns of asymmetric genitalia in the beetle tribe Cyclocephalini (Coleoptera: Scarabaeidae: Dynastinae). Contributions to Zoology 82: 95-106. https://doi.org/10.1603/EN13123" title="Evolutionary patterns of asymmetric genitalia in the beetle tribe Cyclocephalini (Coleoptera: Scarabaeidae: Dynastinae). Contributions to Zoology 82: 95 - 106." url="https://doi.org/10.1603/EN13123" year="2013">Breeschoten et al. 2013</bibRefCitation>
,
<bibRefCitation id="CCB81CAFFD0F7FC7C8619C585EA199C3" author="Lopez-Garcia, MM" pageId="203" pageNumber="304" refId="B695" refString="Lopez-Garcia, MM, Garcia-Atencia, S, Amat-Garcia, G, 2015. Escarabajos fitofagos (Coleoptera: Scarabaeidae &quot;Pleurosticti&quot;) de los Andes orientales de Colombia (Departamentos de Santander, Boyaca y Cundinamarca). Boletin Cientifico Centro de Museos: Museo de Historia Natural 19: 322-358." title="Escarabajos fitofagos (Coleoptera: Scarabaeidae &quot; Pleurosticti &quot;) de los Andes orientales de Colombia (Departamentos de Santander, Boyaca y Cundinamarca). Boletin Cientifico Centro de Museos: Museo de Historia Natural 19: 322 - 358." year="2015">
<normalizedToken id="CA9BDEC41BF8589673F710BEDBAFD966" originalValue="López-García">Lopez-Garcia</normalizedToken>
et al. 2015
</bibRefCitation>
,
<bibRefCitation id="A37E723B5A2D1DF7E5D97B743A0B2B16" DOI="https://doi.org/10.2317/kent-88-02-186-207.1" author="Ratcliffe, BC" pageId="219" pageNumber="320" refId="B990" refString="Ratcliffe, BC, Jameson, ML, Figueroa, L, Cave, RD, Paulsen, MJ, Cano, EB, Beza-Beza, C, Jimenez-Ferbans, L, Reyes-Castillo, P, 2015. Beetles (Coleoptera) of Peru: A survey of families. Scarabaeoidea. The Journal of the Kansas Entomogical Society 88: 186-207. https://doi.org/10.2317/kent-88-02-186-207.1" title="Beetles (Coleoptera) of Peru: A survey of families. Scarabaeoidea. The Journal of the Kansas Entomogical Society 88: 186 - 207." url="https://doi.org/10.2317/kent-88-02-186-207.1" year="2015">Ratcliffe et al. 2015</bibRefCitation>
,
<bibRefCitation id="39E32ADDE98068E5DFC3513C4BD07376" author="Dossey, AT" pageId="176" pageNumber="277" refId="B276" refString="Dossey, AT, Morales-Ramos, JA, Guadalupe, Rojas M, 2016. Documented information for 1555 species of insects and spiders. In: Insects as sustainable food ingredients: production, processing and food applications. Elsevier Academic Press, San Diego, 311-376." title="Documented information for 1555 species of insects and spiders. In: Insects as sustainable food ingredients: production, processing and food applications. Elsevier Academic Press, San Diego, 311 - 376." year="2016">Dossey et al. 2016</bibRefCitation>
,
<bibRefCitation id="08FA13B097E193CF9299DC581A9E6AF2" DOI="https://doi.org/10.1649/072.070.0107" author="Figueroa, L" pageId="180" pageNumber="281" refId="B348" refString="Figueroa, L, Ratcliffe, BC, 2016. A new species of Ancognatha Erichson (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) from Peru, with distributions of Peruvian Ancognatha species. The Coleopterists Bulletin 70: 65-72. https://doi.org/10.1649/072.070.0107" title="A new species of Ancognatha Erichson (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) from Peru, with distributions of Peruvian Ancognatha species. The Coleopterists Bulletin 70: 65 - 72." url="https://doi.org/10.1649/072.070.0107" year="2016">Figueroa and Ratcliffe 2016</bibRefCitation>
,
<bibRefCitation id="33983313BFBF90611BDDCF1E6FAF7C55" DOI="https://doi.org/10.1201/9781315367927" author="Mitasuhashi, J" pageId="207" pageNumber="308" refId="B782" refString="Mitasuhashi, J, 2016. Edible insects of the world. CRC Press, Boca Raton, 296 pp. https://doi.org/10.1201/9781315367927" title="Edible insects of the world. CRC Press, Boca Raton, 296 pp." url="https://doi.org/10.1201/9781315367927" year="2016">Mitasuhashi 2016</bibRefCitation>
,
<bibRefCitation id="2C4DBEF4A85DC113E01F53D4CFF04325" author="Villalobos-Moreno, A" pageId="229" pageNumber="330" refId="B1167" refString="Villalobos-Moreno, A, Pardo-Locarno, LC, Cabrero-Sanudo, FJ, Ospina-Torres, R, Gomez-Murillo, IJ, 2016. Inventario preliminar de los escarabajos de la familia Melolonthidae (Coleoptera: Scarabaeoidea) en un robledal del nororiente de los Andes Colombianos. Boletin de la Sociedad Entomologica Aragonesa 58: 159-167." title="Inventario preliminar de los escarabajos de la familia Melolonthidae (Coleoptera: Scarabaeoidea) en un robledal del nororiente de los Andes Colombianos. Boletin de la Sociedad Entomologica Aragonesa 58: 159 - 167." year="2016">Villalobos-Moreno et al. 2016</bibRefCitation>
,
<bibRefCitation id="F8A0229419D20F89465A25AD92D6B58E" author="Villalobos-Moreno, A" pageId="229" pageNumber="330" refId="B1168" refString="Villalobos-Moreno, A, Pardo-Locarno, LC, Cabrero-Sanudo, FJ, Ospina-Torres, R, Gomez-Murillo, IJ, 2017. Escarabajos fitofagos (Coleoptera: Scarabaeoidea) de un robledal Andino del nororiente Colombiano. Boletin de la Sociedad Entomologica Aragonesa 61: 115-136." title="Escarabajos fitofagos (Coleoptera: Scarabaeoidea) de un robledal Andino del nororiente Colombiano. Boletin de la Sociedad Entomologica Aragonesa 61: 115 - 136." year="2017">2017</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="11BC151E0E3BD238B9F1CA766588414D" pageId="17" pageNumber="118" type="remarks">
<paragraph id="F709E18456F53DF99CEC4C067B3987C5" pageId="17" pageNumber="118">Remarks.</paragraph>
<paragraph id="E357C79162C9595DB431CBFD25E2F2B5" pageId="17" pageNumber="118">
Some authors attributed the name
<taxonomicName id="3945030AFD2E783A8F2BE6691B32B17F" baseAuthorityName="Burmeister" baseAuthorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="B996CD0E305F88E13FE46F5514AF5061" italics="true" pageId="17" pageNumber="118">A. humeralis</emphasis>
</taxonomicName>
to
<bibRefCitation id="7D4313BC7D9B37686DA5193B8947096D" author="Bates, HW" pageId="164" pageNumber="265" refId="B65" refString="Bates, HW, 1888. Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296-342." title="Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296 - 342." year="1888">Bates (1888)</bibRefCitation>
and subsequently treated this taxon as a synonym of
<taxonomicName id="3167488F0B9779F7522DECC27C0BFD61" authorityName="Arrow" authorityYear="1911" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="5ED22CD9714480C4E6779B742B23DC02" italics="true" pageId="17" pageNumber="118">A. vulgaris</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="63B6A723B7695710E2A9A5D2407C14EB" author="Arrow, GJ" pageId="164" pageNumber="265" refId="B51" refString="Arrow, GJ, 1937b. Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1-124." title="Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1 - 124." year="1937 b">Arrow 1937b</bibRefCitation>
,
<bibRefCitation id="B01644993D551512632B6EAA4AEC7BD9" author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken id="E17C7CD402A129C8E548229803CD3BD5" originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
,
<bibRefCitation id="03A45B348384A160364A3B7F19587F46" author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
,
<bibRefCitation id="2D647B49FCB11B4603495D06D2AE2E77" author="Krajcik, M" pageId="200" pageNumber="301" refId="B658" refString="Krajcik, M, 2005. Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma.X, Supplement No. 2, Pilsen, 122 pp." title="Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma. X, Supplement No. 2, Pilsen, 122 pp." year="2005">Krajcik 2005</bibRefCitation>
,
<bibRefCitation id="B900F6D32786C770DBE7310487D80A49" author="Krajcik, M" pageId="200" pageNumber="301" refId="B659" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma.X Supplement 5, Pilsen, 278 pp." title="Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Pilsen, 278 pp." year="2012">2012</bibRefCitation>
).
<bibRefCitation id="2848C3F516AA0DE1B3996331FB8A8F03" author="Bates, HW" pageId="164" pageNumber="265" refId="B65" refString="Bates, HW, 1888. Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296-342." title="Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296 - 342." year="1888">Bates (1888)</bibRefCitation>
clearly attributed the name
<taxonomicName id="202DD0457FE0083BB3E84AF019642C2F" baseAuthorityName="Burmeister" baseAuthorityYear="1847" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="DC774F3D3BE9414F8FE9A8FFF67CB862" italics="true" pageId="17" pageNumber="118">A. humeralis</emphasis>
</taxonomicName>
to
<bibRefCitation id="B8D591A63BEFF217D184F998F430237A" author="Burmeister, H" pageId="168" pageNumber="269" refId="B140" refString="Burmeister, H, 1847. Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." title="Handbuch der Entomologie, volume 5. TCF Enslin, Berlin, 584 pp." year="1847">Burmeister (1847)</bibRefCitation>
, and his notes on this species should not be considered a description of a new species.
</paragraph>
<paragraph id="236B5AEC4DF850408E89F1A4ED71B886" pageId="17" pageNumber="118">
The identity and species status of
<taxonomicName id="952045196E8D52751BF972117134759C" authorityName="Otoya" authorityYear="1945" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="0AD1D345AEFE4AF1B2E4F25136534E87" italics="true" pageId="17" pageNumber="118">A. nigriventris</emphasis>
</taxonomicName>
is ambiguous and needs clarification.
<taxonomicName id="76C81793AB9955789C067AE611BAC82E" authorityName="Otoya" authorityYear="1945" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="0EACCD6CBAAEABFBC46CE42CCDCB7122" italics="true" pageId="17" pageNumber="118">Ancognatha nigriventris</emphasis>
</taxonomicName>
was described from male and female specimens collected in the Colombian departments of Meta and Santander (
<bibRefCitation id="9F77D9D69386451FE2B7BDAC56BE6919" author="Otoya, IA" pageId="211" pageNumber="312" refId="B861" refString="Otoya, IA, 1945. Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273-292." title="Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273 - 292." year="1945">Otoya 1945</bibRefCitation>
). The species was placed in a new subgenus based on its relatively well-developed maxillary teeth (
<bibRefCitation id="3D3181126C9803809F7FAA375C3F2504" author="Otoya, IA" pageId="211" pageNumber="312" refId="B861" refString="Otoya, IA, 1945. Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273-292." title="Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273 - 292." year="1945">Otoya 1945</bibRefCitation>
).
<bibRefCitation id="D2F3F112CF8A74D278DC659D1B4DA310" author="Gutierrez, AR" pageId="186" pageNumber="287" refId="B445" refString="Gutierrez, AR, 1950. Scarabaeidae del Norte de Chile. Anales de la Sociedad Cientifica Argentina 149 (2): 52-75." title="Scarabaeidae del Norte de Chile. Anales de la Sociedad Cientifica Argentina 149 (2): 52 - 75." year="1950">
<normalizedToken id="B198FEEF311E4E50282AF4FF8D372741" originalValue="Gutiérrez">Gutierrez</normalizedToken>
(1950)
</bibRefCitation>
discussed the subgenus and
<bibRefCitation id="DDFA56CE231C5E95E4FECDA01CDBAE76" author="Martinez, A" pageId="205" pageNumber="306" refId="B744" refString="Martinez, A, 1965a. Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae). Anales de la Sociedad Cientifica Argentina 179: 63-74." title="Notas Coleopterologicas X. Dos nuevas especies de Cyclocephalini Neotropicales (Dynastinae). Anales de la Sociedad Cientifica Argentina 179: 63 - 74." year="1965 a">
<normalizedToken id="7456B212CEB52F3BDA55A5723E67027D" originalValue="Martínez">Martinez</normalizedToken>
(1965a)
</bibRefCitation>
implied that the species was a synonym of
<taxonomicName id="E9B69E52674A001E29656650DE2A10F2" authorityName="Arrow" authorityYear="1911" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="CE267195E6C6FAAFB4D76176D5D101F8" italics="true" pageId="17" pageNumber="118">A. vulgaris</emphasis>
</taxonomicName>
.
<bibRefCitation id="0103FADD1715A835C391DDC75B096F8F" author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken id="5031F3815981B7E261C33ABD056639FB" originalValue="Endrődi">Endrodi</normalizedToken>
(1966)
</bibRefCitation>
remarked that he had not seen the type series, or any specimens, of
<taxonomicName id="C2C3EFF1EDB396381A7C236FA7EB9C42" authorityName="Otoya" authorityYear="1945" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="798CD0170B06A8F94F7636D0F09B9F17" italics="true" pageId="17" pageNumber="118">A. nigriventris</emphasis>
</taxonomicName>
and treated the species as valid.
<bibRefCitation id="A670849D3B3EB8E08CE75A393661F9F5" author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">
<normalizedToken id="BD6EE85BD1916672C2FCC49788BC579C" originalValue="Endrődi">Endrodi</normalizedToken>
(1985a)
</bibRefCitation>
did not further treat
<taxonomicName id="DF2FB8D3C6C11621A1C45F51250835A2" authorityName="Otoya" authorityYear="1945" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="9584C748D0CD1A025076C851A9DD98CE" italics="true" pageId="17" pageNumber="118">A. nigriventris</emphasis>
</taxonomicName>
. Some subsequent papers have cited the species from Colombia (
<bibRefCitation id="C1B2ED3ACFC1A83C07D30A73E7D22D2C" author="Ruiz, BN" pageId="222" pageNumber="323" refId="B1047" refString="Ruiz, BN, Pamalpa, CN, 1990. Observaciones sobre las chisas (Coleoptera: Scarabaeidae) en Narino. Revista ICA: Publicación Científica del Instituto Colombiano Agropecuario 25: 275-282." title="Observaciones sobre las chisas (Coleoptera: Scarabaeidae) en Narino. Revista ICA: Publicacio ́ n Cienti ́ fica del Instituto Colombiano Agropecuario 25: 275 - 282." year="1990">Ruiz and Pamalpa 1990</bibRefCitation>
,
<bibRefCitation id="58F59727D7AEFBF49F8AA0C95842D31C" author="Montoya, GC" pageId="207" pageNumber="308" refId="B791" refString="Montoya, GC, Madrigal-C, A, Ramirez, CA, 1994. Evaluacion de trampas de luz para el control de adultos de Scarabaeidae (Coleoptera) en cultivos de papa en La Union (Antioquia). Revista Colombiana de Entomologia 20: 130-136." title="Evaluacion de trampas de luz para el control de adultos de Scarabaeidae (Coleoptera) en cultivos de papa en La Union (Antioquia). Revista Colombiana de Entomologia 20: 130 - 136." year="1994">Montoya et al. 1994</bibRefCitation>
), while others have ignored it (e.g., Restrepo et al. 2003 and
<bibRefCitation id="F3AD3EF60CBD61FCBA73D1D099E0E0C8" DOI="https://doi.org/10.3897/zookeys.34.309" author="Gasca-Alvarez, HJ" pageId="182" pageNumber="283" refId="B381" refString="Gasca-Alvarez, HJ, Amat-Garcia, G, 2010. Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia. ZooKeys 34: 153-192. https://doi.org/10.3897/zookeys.34.309" title="Synopsis and key to the genera of Dynastinae (Coleoptera, Scarabaeoidea, Scarabaeidae) of Colombia. ZooKeys 34: 153 - 192." url="https://doi.org/10.3897/zookeys.34.309" year="2010">
<normalizedToken id="7D85CBAADA00F5B58CE18ABB80163595" originalValue="Gasca-Álvarez">Gasca-Alvarez</normalizedToken>
and
<normalizedToken id="E6DF37C2361FDB0EC4CBF2E258DA1AE7" originalValue="Amat-García">Amat-Garcia</normalizedToken>
2010
</bibRefCitation>
).
<taxonomicName id="AFDB3CDD34FCA8C2742A731FA1BD8276" authorityName="Otoya" authorityYear="1945" class="Insecta" family="Scarabaeidae" genus="Ancognatha" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="8E608714D4E5AD55844AF1A9A97F9A89" italics="true" pageId="17" pageNumber="118">Ancognatha nigriventris</emphasis>
</taxonomicName>
is reported from the Colombian states of Meta,
<normalizedToken id="0AF9DEE306DC94BB860888514CEF251D" originalValue="Nariño">Narino</normalizedToken>
, and Santander (
<bibRefCitation id="B8B5FCC03251781D991F8C8CE5888065" author="Otoya, IA" pageId="211" pageNumber="312" refId="B861" refString="Otoya, IA, 1945. Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273-292." title="Anotaciones sobre el genero Ancognatha y descripcion de una nueva especie (Scarabaeidae). Caldasia 3: 273 - 292." year="1945">Otoya 1945</bibRefCitation>
,
<bibRefCitation id="D46F12B74FA5359A675DC836E3C63A9B" author="Ruiz, BN" pageId="222" pageNumber="323" refId="B1047" refString="Ruiz, BN, Pamalpa, CN, 1990. Observaciones sobre las chisas (Coleoptera: Scarabaeidae) en Narino. Revista ICA: Publicación Científica del Instituto Colombiano Agropecuario 25: 275-282." title="Observaciones sobre las chisas (Coleoptera: Scarabaeidae) en Narino. Revista ICA: Publicacio ́ n Cienti ́ fica del Instituto Colombiano Agropecuario 25: 275 - 282." year="1990">Ruiz and Pamalpa 1990</bibRefCitation>
,
<bibRefCitation id="5781037136D07AAD316ECC6E0ED58BCF" author="Montoya, GC" pageId="207" pageNumber="308" refId="B791" refString="Montoya, GC, Madrigal-C, A, Ramirez, CA, 1994. Evaluacion de trampas de luz para el control de adultos de Scarabaeidae (Coleoptera) en cultivos de papa en La Union (Antioquia). Revista Colombiana de Entomologia 20: 130-136." title="Evaluacion de trampas de luz para el control de adultos de Scarabaeidae (Coleoptera) en cultivos de papa en La Union (Antioquia). Revista Colombiana de Entomologia 20: 130 - 136." year="1994">Montoya et al. 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>