141 lines
15 KiB
XML
141 lines
15 KiB
XML
<document id="BD43CACAB85554FFC991C00782EE9999" ID-DOI="http://dx.doi.org/10.3897/zookeys.102.1148" ID-GBIF-Dataset="7fa288c5-61ce-4f13-b08f-56c96f19d9c0" ID-PMC="PMC3131081" ID-Pensoft-Pub="1313-2970-102-65" ID-PubMed="21747675" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-102-65" ModsDocOrigin="ZooKeys 102" ModsDocTitle="Review of the subgenus Polyphylla (Granida) from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae)" checkinTime="1451250337947" checkinUser="pensoft" docAuthor="Sehnal, Richard & Bezdek, Ales" docDate="2011" docId="A67DCC3BD9A91914D710A03CD26D30D5" docLanguage="en" docName="ZooKeys 102: 65-76" docOrigin="ZooKeys 102" docSource="http://dx.doi.org/10.3897/zookeys.102.1148" docTitle="Polyphylla ((Granida)) phongsali Zidek 2006" docType="treatment" docVersion="4" lastPageNumber="73" masterDocId="FF87FFE2FFB7FFC1BD11FFBF6C6D4F56" masterDocTitle="Review of the subgenus Polyphylla (Granida) from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae)" masterLastPageNumber="76" masterPageNumber="65" pageNumber="72" updateTime="1732710903437" updateUser="ExternalLinkService">
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<mods:titleInfo id="7C9F0B2BB5BC60EC20C5FFAAC9713890">
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<mods:title id="FB8583628A090239E81C8A17004D6C59">Review of the subgenus Polyphylla (Granida) from continental Asia (Coleoptera, Scarabaeidae, Melolonthinae)</mods:title>
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<mods:namePart id="0FA9AF2EE111ACE760EEB4DC96F99B22">Sehnal, Richard</mods:namePart>
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<mods:namePart id="22B662E2F42599D5321386F00215EE59">Bezdek, Ales</mods:namePart>
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<mods:typeOfResource id="37491DEF0FF88FB4F910B06FD738BEF3">text</mods:typeOfResource>
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<mods:title id="D51602E9C377FD506B5BD6D59FD3284C">ZooKeys</mods:title>
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<mods:part id="E1794455E44B116E14416B49082A1464">
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<mods:date id="9D36A7435CFE33C1B2395E09E28C5CCE">2011</mods:date>
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<mods:detail id="37064646D455D5F746CC1DB73CEDB9D9" type="volume">
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<mods:number id="DFA75BC73D25516B44D284E807F24AAA">102</mods:number>
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<mods:start id="F6413528BD5FE0959E584A1309AC6B90">65</mods:start>
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<mods:end id="2B9E6537D30CA1318C571A43CBC32652">76</mods:end>
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<mods:url id="2FE728B29E7EF5C9C75BE63C1879152B">http://dx.doi.org/10.3897/zookeys.102.1148</mods:url>
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<mods:classification id="74773C89281305A23A5398A0BCABFC80">journal article</mods:classification>
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<mods:identifier id="5542342B59704C1973D1BB35F7D52F82" type="DOI">http://dx.doi.org/10.3897/zookeys.102.1148</mods:identifier>
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<mods:identifier id="022E2867D69CF2EDEF8283E403C6F41E" type="Pensoft-Pub">1313-2970-102-65</mods:identifier>
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<treatment id="A67DCC3BD9A91914D710A03CD26D30D5" ID-GBIF-Taxon="152029561" LSID="urn:lsid:plazi:treatment:A67DCC3BD9A91914D710A03CD26D30D5" httpUri="http://treatment.plazi.org/id/A67DCC3BD9A91914D710A03CD26D30D5" lastPageId="8" lastPageNumber="73" pageId="7" pageNumber="72" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_subFamily="Melolonthinae" scope_subGenus="Granida">
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<subSubSection id="1025065639D8F967691754A7E8A4516E" pageId="7" pageNumber="72" type="nomenclature">
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<paragraph id="B6FB0A364B56495C4BFC5FF788B01355" pageId="7" pageNumber="72">
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<taxonomicName id="1745877BFED24C666AE2CF3C12FA5BDA" ID-CoL="4LDXR" LSID="http://species-id.net/wiki/Polyphylla_(Granida)_phongsali" authority="Zidek, 2006" authorityName="Zidek" authorityYear="2006" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla ((Granida)) phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali" subGenus="(Granida)">
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<pageBreakToken id="B630704E127CB1C24C5A6AF1BC5110E6" pageId="7" pageNumber="72" start="start">Polyphylla</pageBreakToken>
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(Granida) phongsali
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<normalizedToken id="49335A22DA59B140A40336A18E4FCEF4" originalValue="Zídek">Zidek</normalizedToken>
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, 2006
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</taxonomicName>
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Figs 16-19
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</paragraph>
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</subSubSection>
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<subSubSection id="EA237D91DACCA4E45BC4F815E6EC7481" pageId="7" pageNumber="72" type="reference_group">
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<paragraph id="82B873500ADDF4EE2D7F509858E787ED" pageId="7" pageNumber="72">
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<taxonomicName id="CE07F29C7D478E505888A887EE291425" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali">Polyphylla phongsali</taxonomicName>
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<normalizedToken id="6F0A4ECF521EDCDF8091EF890AAB15F3" originalValue="Zídek">Zidek</normalizedToken>
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, 2006: 10, pl. 3.
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</paragraph>
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</subSubSection>
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<subSubSection id="B05BC05579BB7B6216BD6F6ABAC889C4" pageId="7" pageNumber="72" type="type locality">
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<paragraph id="E2A1658DC3BC77AFC4006BCEBF0ABF28" pageId="7" pageNumber="72">Type locality.</paragraph>
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<paragraph id="F84A66135B47734C20F06B3A9882D9D4" pageId="7" pageNumber="72">"N. Laos, Phongsali, Gnoi-ou".</paragraph>
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</subSubSection>
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<subSubSection id="E01E6D16F3A7D983FB1F16B9FBE791A2" pageId="7" pageNumber="72" type="type material examined">
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<paragraph id="850A1CCF158A17D07FAE38F28C9BEAD2" pageId="7" pageNumber="72">Type material examined.</paragraph>
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<paragraph id="73D86E4B21432236BD6902DBE6E6C3E0" pageId="7" pageNumber="72">
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Holotype (male), labeled: "N. Laos, Phongsali, Gnoi-ou, Li Jingke VI-2003 [h] // BMNH (E) 2006-162 [h] // J. ZIDEK det. 2006 [p] Polyphylla phongsali
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<normalizedToken id="EF6145AB25BA121E49AF69D02DFD3464" originalValue="Zídek">Zidek</normalizedToken>
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[h] HOLOTYPE [p, red label]", in BMNH.
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</paragraph>
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</subSubSection>
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<subSubSection id="AAE3CE46A639F1E41B501BFCF1EAD43A" pageId="7" pageNumber="72" type="additional material examined">
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<paragraph id="F8BD6416C49B3B32C89A8B46CE44F8C0" pageId="7" pageNumber="72">Additional material examined.</paragraph>
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<paragraph id="8EC72FAD8E38D7EE6B21C57FF665EB21" pageId="7" pageNumber="72">
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LAO-NE, Hua Phan prov., ~20˚12′N, 104˚01′E, PHU PHAN Mt. 1500-1900m, 17.v.-31.vi. 2007, M. Brancucci leg., 1 male in ABCC; LAOS-NE, Houa Phan prov., 20˚13′09-19″N, 103˚59′54″-104˚00′03″E, 1480-1510m, PHOU PHANE Mt., 22.iv.-14.v.2008,
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<normalizedToken id="C700C76E980882EE5D0203206C9FF188" originalValue="Vít">Vit</normalizedToken>
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<normalizedToken id="60E2B1CF0B402FEB5E6A410FC0A6621F" originalValue="Kubáň">Kubaň</normalizedToken>
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leg., 1 male and 1 female in NMPC; LAOS-NE, Houa Phan prov., 20˚13′N 103˚59′E, Ban SALUEI village, 16.vi.2009, 1350 m, at light,
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<normalizedToken id="59629F698C32755B7E6B1E8C39C33948" originalValue="Vít">Vit</normalizedToken>
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.
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<normalizedToken id="716AFBB3C6FF6A8B8505A0B96BFE2EBA" originalValue="Kubáň">Kubaň</normalizedToken>
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leg., 1 male in NMPC; LAOS-NE, Houa Phan prov., 20˚12-13.5′N 103˚59.5′-104˚01′E, Ban Saluei → Phou Pane Mt., 1340-1870 m, 1.v.-16.vi. 2009, Lao collectors leg., 1 male in NMPC; Laos, Houaphan prov., 38 km S of Sam Neua, Saluei 9.-22.5.2009,
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<normalizedToken id="0112EECCC605704652D4ED6BCE8F1127" originalValue="Martinů">Martinu</normalizedToken>
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lgt. 1350-1900 m, 1 male in JZCP and 1 male in RSCV; Laos, Houaphan prov., 38 km S of Sam Neua, Saluei 9.-22.5.2009,
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<normalizedToken id="64C5DDEB90DADC29B1C78E44A6AC6E6C" originalValue="Bednařík">Bednarik</normalizedToken>
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lgt. 1350-1900 m, 1 male in RSCV.
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</paragraph>
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</subSubSection>
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<subSubSection id="ACBFEBB39B92A2FE91516E4365A1C107" pageId="7" pageNumber="72" type="diagnosis of female">
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<paragraph id="8A40E0A84A29C259AC809896DAEBD015" pageId="7" pageNumber="72">Diagnosis of female</paragraph>
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<paragraph id="E339162A58EE9C5398BB940D8C4CD3BF" pageId="7" pageNumber="72">(Fig. 17). Similar to male (Fig. 16), with the following exceptions. The length of the only known female specimen is 28.0 mm (except of pygidium), while the length of males varies between 21.5-24.5 mm. Anterior margin of clypeus only very feebly upturned, nearly flat. Antennal club pentamerous. Outer margin of anterior tibia distinctly tridentate. Upper apical spur of metatibia broad, flattened, blunt apically. Tarsal claws of all pairs of legs equal in length.</paragraph>
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<caption id="CC568F41D7F8F536F710938BE00A71F3" pageId="7" pageNumber="72">
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<paragraph id="B45C7EC8A060F23CDF3CBA8E00877B92" pageId="7" pageNumber="72">
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Figures 16-19.
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<taxonomicName id="B77BA0E7C95DFECFADE05E145FCD85C4" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali" subGenus="Granida">Polyphylla (Granida) phongsali</taxonomicName>
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16 Habitus of male (Hua Phan, Laos, length 22.5 mm), dorsal view 17 Habitus of female (Hua Phan, Laos, length 28.0 mm), dorsal view 18 Male genitalia, dorsal view 19 The same, right lateral view, shaded area indicates overlapping part of left paramere. Scale bar: 5 mm for Figs 18-19.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="A995269CCF8DD12BF41986147FD6B91C" pageId="7" pageNumber="72" type="collecting methods">
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<paragraph id="7C75E065636F0B46CD96FC889483F393" pageId="7" pageNumber="72">Collecting methods.</paragraph>
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<paragraph id="E6FD94ED141D35172221770ED4489A51" pageId="7" pageNumber="72">The female specimen was collected at light.</paragraph>
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</subSubSection>
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<subSubSection id="7247B0DD4B290D2B9798A81B8C8CB84E" pageId="7" pageNumber="72" type="diagnosis">
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<paragraph id="51BDBCB64EC06C92395B44C44EBF46CB" pageId="7" pageNumber="72">Diagnosis.</paragraph>
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<paragraph id="AB1D45C33EF33B1B1A354C1ACB20BC96" pageId="7" pageNumber="72">
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An easily recognizable
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<taxonomicName id="337D2720433B30E875AB1D162E3C412C" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida)" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="subGenus" subGenus="Granida">Polyphylla (Granida)</taxonomicName>
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species because of its maculate elytra. Antennomere 3 long and slender, more than three times longer than antennomere 2. Basal margin of pronotum is almost straight against the scutellum, while convex in other
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<taxonomicName id="46345FD45D30625515D02822161C20A4" class="Insecta" family="Scarabaeidae" genus="Granida" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Granida" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="genus">Granida</taxonomicName>
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species. It is most likely to be confused only with some
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<taxonomicName id="DAC9E2FC9571D332027F492E74295D78" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) jessopi" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="jessopi" subGenus="Granida">Polyphylla (Granida) jessopi</taxonomicName>
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specimens bearing the same maculate elytral pattern.
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<taxonomicName id="FEC50F1277C68E55BF0D175DF0C97D1A" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) phongsali" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="phongsali" subGenus="Granida">Polyphylla (Granida) phongsali</taxonomicName>
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in average larger than
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<taxonomicName id="B881B93D136DFE789B77B742A5A3EC68" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) jessopi" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="jessopi" subGenus="Granida">Polyphylla (Granida) jessopi</taxonomicName>
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(the length of males varies between 21.5-24.5 mm versus 18.5-20.0 mm in
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<taxonomicName id="77EDF9BFF4A68572DDF6FCB843A898E2" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) jessopi" order="Coleoptera" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="jessopi" subGenus="Granida">Polyphylla (Granida) jessopi</taxonomicName>
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). Male genitalia as in Figs 18-19.
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</paragraph>
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</subSubSection>
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<subSubSection id="7A3E2672CC6A8EC4CA12E947F9C19BCB" pageId="7" pageNumber="72" type="distribution">
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<paragraph id="C59BA856BB5A9A74CA27D420B13C1406" pageId="7" pageNumber="72">Distribution.</paragraph>
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<paragraph id="A1BC993E74E1FD424056C8B074E38BDD" pageId="7" pageNumber="72">Northern Laos.</paragraph>
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</subSubSection>
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<subSubSection id="B2775683AF8C532EC3F9E8BE17D3878B" lastPageId="8" lastPageNumber="73" pageId="7" pageNumber="72" type="remarks">
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<paragraph id="194975D69791E154D079DF2770F919A3" pageId="7" pageNumber="72">Remarks.</paragraph>
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<paragraph id="42CCC9203891641A2AAAB02AC0509A4A" pageId="7" pageNumber="72">The species was originally described from a single male. Here we recorded eight additional specimens from northern Laos (Fig. 20). The specimens with altitude data were collected between 1350-1900 m a.s.l.</paragraph>
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<caption id="45295E78138806B1818D744612CBDE42" pageId="8" pageNumber="73">
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<paragraph id="F474A92E55F480F10F8BCAADB9F68ACC" pageId="8" pageNumber="73">
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<pageBreakToken id="120440E7672F987E4128104F2C4A635B" pageId="8" pageNumber="73" start="start">Figure</pageBreakToken>
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20. Distribution of
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<taxonomicName id="3BBEE3CFC0FE8249A624A875E332AE20" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida)" order="Coleoptera" pageId="8" pageNumber="73" phylum="Arthropoda" rank="subGenus" subGenus="Granida">Polyphylla (Granida)</taxonomicName>
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species in continental Asia. Because of imprecise locality data, distributional mark of
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<taxonomicName id="804FA8E14851E164B212E3EC0BD00C64" class="Insecta" family="Scarabaeidae" genus="Polyphylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphylla (Granida) nikodymi" order="Coleoptera" pageId="8" pageNumber="73" phylum="Arthropoda" rank="species" species="nikodymi" subGenus="Granida">Polyphylla (Granida) nikodymi</taxonomicName>
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in Myanmar is omitted.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |