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<mods:title id="2D6C24A78C1018A626096AEDC183D591">Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)</mods:title>
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<mods:namePart id="1FF7F2F70AD153D7B567D13ACB7BDEDB">Dankittipakul, Pakawin</mods:namePart>
<mods:affiliation id="98604E2F87CBE89CF4D7B17A148F3D99">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand; &amp; Manukau Institute of Technology, Manukau, New Zealand;</mods:affiliation>
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<mods:namePart id="DFCC790C45BBAC8E693586C7B7936B21">Tavano, Maria</mods:namePart>
<mods:affiliation id="23E525C64AC480DC372052BBEE00A3F3">Museo Civico di Storia Naturale ‘ Giacomo Doria’, Genova, Italy</mods:affiliation>
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<mods:namePart id="F296FAC7036A31DF4AD4747D431EDA7B">Singtripop, Tippawan</mods:namePart>
<mods:affiliation id="1CA0C89D1E250095B6CA04BE19283C36">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand;</mods:affiliation>
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Genus 
<taxonomicName id="E9DF2D87FFDBFF8C057FF9AC11688A1B" ID-CoL="63866" authority="Thorell, 1897" authorityName="Thorell" authorityYear="1897" box="[608,853,1625,1649]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1543" phylum="Arthropoda" rank="genus">
<emphasis id="1CAB8A16FFDBFF8C057FF9AC10848A1B" bold="true" box="[608,697,1626,1649]" italics="true" pageId="6" pageNumber="1543">Jacaena</emphasis>
<bibRefCitation id="4A4E2BF5FFDBFF8C05DFF9AF11688A1B" author="Thorell T" box="[704,853,1625,1649]" pageId="6" pageNumber="1543" pagination="161 - 267" refId="ref11552" refString="Thorell T. 1897. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXIII. Secondo saggio sui Ragni birmani. I. Parallelodontes. Tubitelariae. Annali Mus Civ St Nat Genova. 37: 161 - 267." type="journal article" year="1897">Thorell, 1897</bibRefCitation>
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<heading id="7528E168FFDBFF8C07CCF96C115A8ADB" box="[211,871,1689,1713]" fontSize="10" level="3" pageId="6" pageNumber="1543" reason="5">
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<taxonomicName id="E9DF2D87FFDBFF8C07CCF96C103C8ADB" ID-CoL="63866" authority="Thorell, 1897: 230" authorityName="Thorell" authorityPageNumber="230" authorityYear="1897" box="[211,513,1689,1713]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="1543" phylum="Arthropoda" rank="genus">
<emphasis id="1CAB8A16FFDBFF8C07CCF96C13168ADB" box="[211,299,1690,1713]" italics="true" pageId="6" pageNumber="1543">Jacaena</emphasis>
<bibRefCitation id="4A4E2BF5FFDBFF8C062DF96F103C8ADB" author="Thorell T" box="[306,513,1689,1713]" pageId="6" pageNumber="1543" pagination="161 - 267" refId="ref11552" refString="Thorell T. 1897. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXIII. Secondo saggio sui Ragni birmani. I. Parallelodontes. Tubitelariae. Annali Mus Civ St Nat Genova. 37: 161 - 267." type="journal article" year="1897">Thorell, 1897: 230</bibRefCitation>
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; 
<treatmentCitation id="AF7E7015FFDBFF8C0513F96F115A8ADB" author="Deeleman-Reinhold CL" box="[524,871,1689,1713]" page="465" pageId="6" pageNumber="1543" year="2001">
<bibRefCitation id="4A4E2BF5FFDBFF8C0513F96F115A8ADB" author="Deeleman-Reinhold CL" box="[524,871,1689,1713]" pageId="6" pageNumber="1543" refId="ref11331" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." type="book" year="2001">Deeleman-Reinhold, 2001: 465</bibRefCitation>
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Figure 3. 
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<emphasis id="1CAB8A16FFDAFF8D07E7FB61137A88C6" box="[248,327,1175,1196]" italics="true" pageId="7" pageNumber="1544">Jacaena</emphasis>
spp.
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Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). 
<taxonomicName id="E9DF2D87FFDAFF8D06F4FB2410AF888D" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[491,658,1233,1255]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="1544" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis id="1CAB8A16FFDAFF8D06F4FB2410AF888D" box="[491,658,1233,1255]" italics="true" pageId="7" pageNumber="1544">Jacaena lunulata</emphasis>
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<taxonomicNameLabel id="0798376DFFDAFF8D0587FB2410DA888D" box="[664,743,1234,1255]" pageId="7" pageNumber="1544" rank="species">sp. nov.</taxonomicNameLabel>
, male, paratype (A–C, E); 
<taxonomicName id="E9DF2D87FFDAFF8D04EAFB241674888D" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Dankittipakul &amp; Tavano &amp; Singtripop" baseAuthorityYear="2013" box="[1013,1097,1233,1255]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="1544" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFDAFF8D04EAFB241674888D" box="[1013,1097,1233,1255]" italics="true" pageId="7" pageNumber="1544">J. mihun</emphasis>
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, male, new material (D).
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species
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<taxonomicName id="E9DF2D87FFDAFF8D0790FA8213D589E1" ID-CoL="3QHZC" authority="Thorell, 1897" authorityName="Thorell" authorityYear="1897" box="[143,488,1395,1419]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="1544" phylum="Arthropoda" rank="species" species="distincta">
<emphasis id="1CAB8A16FFDAFF8D0790FA82137189E1" box="[143,332,1395,1419]" italics="true" pageId="7" pageNumber="1544">Jacaena distincta</emphasis>
<bibRefCitation id="4A4E2BF5FFDAFF8D064CFA8513D589E1" author="Thorell T" box="[339,488,1395,1419]" pageId="7" pageNumber="1544" pagination="161 - 267" refId="ref11552" refString="Thorell T. 1897. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXIII. Secondo saggio sui Ragni birmani. I. Parallelodontes. Tubitelariae. Annali Mus Civ St Nat Genova. 37: 161 - 267." type="journal article" year="1897">Thorell, 1897</bibRefCitation>
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<heading id="7528E168FFDAFF8D0790FA2D12C68998" box="[143,251,1499,1522]" fontSize="10" level="3" pageId="7" pageNumber="1544" reason="5">
<emphasis id="1CAB8A16FFDAFF8D0790FA2D12C68998" box="[143,251,1499,1522]" italics="true" pageId="7" pageNumber="1544">Diagnosis</emphasis>
</heading>
</paragraph>
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Representatives of 
<taxonomicName id="E9DF2D87FFDAFF8D0678F9F513828A70" authorityName="Thorell" authorityYear="1897" box="[359,447,1539,1562]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="1544" phylum="Arthropoda" rank="genus">
<emphasis id="1CAB8A16FFDAFF8D0678F9F513828A70" box="[359,447,1539,1562]" italics="true" pageId="7" pageNumber="1544">Jacaena</emphasis>
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can be recognized by a combined presence of characters: the integument of carapace is strongly granulate or punctate, forming radiating striae (
<figureCitation id="B6E44A81FFDAFF8D0788F9B4131C8A30" box="[151,289,1602,1626]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="5.[143,210,1100,1122]" captionStartId-1="6.[211,278,1132,1154]" captionTargetBox-0="[145,1102,168,1066]" captionTargetBox-1="[213,1170,168,1099]" captionTargetId-0="figure-5@5.[145,1102,168,1066]" captionTargetId-1="figure-5@6.[213,1170,168,1099]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." captionText-1="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi-0="http://doi.org/10.5281/zenodo.6517638" figureDoi-1="http://doi.org/10.5281/zenodo.6517642" httpUri-0="https://zenodo.org/record/6517638/files/figure.png" httpUri-1="https://zenodo.org/record/6517642/files/figure.png" pageId="7" pageNumber="1544">Figures 1–2</figureCitation>
); the fovea is deep and circular (
<figureCitation id="B6E44A81FFDAFF8D05B7F9B4111D8A30" box="[680,800,1602,1626]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="7" pageNumber="1544">Figure 3B</figureCitation>
); the dorsal opisthosomal scutum occupying approximately half the length of the opisthosoma or much less in males (
<figureCitation id="B6E44A81FFDAFF8D07FEF974137F8AF0" box="[225,322,1666,1690]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="7" pageNumber="1544">Figure 1</figureCitation>
), completely absent in females (
<figureCitation id="B6E44A81FFDAFF8D05A8F97411208AF0" box="[695,797,1666,1690]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="7" pageNumber="1544">Figure 2</figureCitation>
); the dorsal pattern of the opisthosoma consists of paired, oval patches, followed by three to five pale chevrons (
<figureCitation id="B6E44A81FFD5FF8207C4F9B4135C8A30" box="[219,353,1602,1626]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="5.[143,210,1100,1122]" captionStartId-1="6.[211,278,1132,1154]" captionTargetBox-0="[145,1102,168,1066]" captionTargetBox-1="[213,1170,168,1099]" captionTargetId-0="figure-5@5.[145,1102,168,1066]" captionTargetId-1="figure-5@6.[213,1170,168,1099]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." captionText-1="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi-0="http://doi.org/10.5281/zenodo.6517638" figureDoi-1="http://doi.org/10.5281/zenodo.6517642" httpUri-0="https://zenodo.org/record/6517638/files/figure.png" httpUri-1="https://zenodo.org/record/6517642/files/figure.png" pageId="8" pageNumber="1545">Figures 1–2</figureCitation>
) on a dark greenish background; the male palp is provided with an apically excavated proximal RTA and a truncated distal RTA (
<figureCitation id="B6E44A81FFD5FF820473F99411CC8A10" box="[876,1009,1634,1658]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="8" pageNumber="1545">Figure 12A</figureCitation>
); the pyriform tegulum bears a distal conductor and a hyaline tegular apophysis (
<figureCitation id="B6E44A81FFD5FF8204C2F97416648AF0" box="[989,1113,1666,1690]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A–C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="8" pageNumber="1545">Figure 9D</figureCitation>
); the embolic base originates disto-prolaterally, its filiform embolus wrapping behind the tegulum (
<figureCitation id="B6E44A81FFD4FF8307E7FF3113B28CB5" box="[248,399,199,223]" captionStart="Figure 4" captionStartId="8.[211,278,1428,1450]" captionTargetBox="[213,1170,168,1394]" captionTargetId="figure-5@8.[213,1170,168,1394]" captionTargetPageId="8" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (A–C); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (D–F); J. mihun, holotype (G–I)." figureDoi="http://doi.org/10.5281/zenodo.6517648" httpUri="https://zenodo.org/record/6517648/files/figure.png" pageId="9" pageNumber="1546">Figure 4C, D</figureCitation>
); the elongated female internal ducts are coiled, moving peripherally before leading to poorly defined or elliptical spermathecae, which are posteriorly situated.
</paragraph>
</subSubSection>
<caption id="7AA0068CFFD5FF8207CCFA6216BA898E" ID-DOI="http://doi.org/10.5281/zenodo.6517648" ID-Zenodo-Dep="6517648" httpUri="https://zenodo.org/record/6517648/files/figure.png" pageId="8" pageNumber="1545" startId="8.[211,278,1428,1450]" targetBox="[213,1170,168,1394]" targetPageId="8">
<paragraph id="2E605604FFD5FF8207CCFA6216BA898E" blockId="8.[211,1171,1428,1508]" pageId="8" pageNumber="1545">
Figure 4. 
<taxonomicName id="E9DF2D87FFD5FF82065FFA6313F889C3" box="[320,453,1428,1450]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="1545" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="1CAB8A16FFD5FF82065FFA6313B289C0" box="[320,399,1429,1450]" italics="true" pageId="8" pageNumber="1545">Jacaena</emphasis>
spp.
</taxonomicName>
Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). 
<taxonomicName id="E9DF2D87FFD5FF8206AEFA4410AE89AD" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" box="[433,659,1457,1479]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="1545" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis id="1CAB8A16FFD5FF8206AEFA4410AE89AD" box="[433,659,1457,1479]" italics="true" pageId="8" pageNumber="1545">Jacaena schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD5FF820584FA47113289AD" box="[667,783,1457,1479]" pageId="8" pageNumber="1545" rank="species">comb. nov.</taxonomicNameLabel>
, holotype (A–C); 
<taxonomicName id="E9DF2D87FFD5FF8204D3FA44160B89AD" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[972,1078,1457,1479]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="1545" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis id="1CAB8A16FFD5FF8204D3FA44160B89AD" box="[972,1078,1457,1479]" italics="true" pageId="8" pageNumber="1545">J. lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD5FF820320FA4416AD89AD" box="[1087,1168,1458,1479]" pageId="8" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (originally misidentified as 
<taxonomicName id="E9DF2D87FFD5FF82055BFA381113898E" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" box="[580,814,1486,1508]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="1545" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis id="1CAB8A16FFD5FF82055BFA381113898E" box="[580,814,1486,1508]" italics="true" pageId="8" pageNumber="1545">Sesieutes schwendingeri</emphasis>
</taxonomicName>
) (D–F); 
<taxonomicName id="E9DF2D87FFD5FF820497FA3911E6898E" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Dankittipakul &amp; Tavano &amp; Singtripop" baseAuthorityYear="2013" box="[904,987,1486,1508]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="1545" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFD5FF820497FA3911E6898E" box="[904,987,1486,1508]" italics="true" pageId="8" pageNumber="1545">J. mihun</emphasis>
</taxonomicName>
, holotype (G–I).
</paragraph>
</caption>
<subSubSection id="66C5058FFFD4FF800790FE9610D08856" lastPageId="10" lastPageNumber="1547" pageId="9" pageNumber="1546" type="description">
<paragraph id="2E605604FFD4FF830790FE9613338D1D" blockId="9.[143,1104,352,1690]" box="[143,270,352,375]" pageId="9" pageNumber="1546">
<heading id="7528E168FFD4FF830790FE9613338D1D" box="[143,270,352,375]" fontSize="10" level="3" pageId="9" pageNumber="1546" reason="5">
<emphasis id="1CAB8A16FFD4FF830790FE9613338D1D" box="[143,270,352,375]" italics="true" pageId="9" pageNumber="1546">Description</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFD4FF830790FE7013FA8911" blockId="9.[143,1104,352,1690]" pageId="9" pageNumber="1546">
Carapace (
<figureCitation id="B6E44A81FFD4FF830615FE70134B8DF4" box="[266,374,390,414]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="9" pageNumber="1546">Figures 1</figureCitation>
, 
<figureCitation id="B6E44A81FFD4FF83069AFE7013AE8DF4" box="[389,403,390,414]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="9" pageNumber="1546">2</figureCitation>
) ovoid in dorsal view, attenuated in front, narrowed opposite palpal insertion, widest between coxae II and III, in profile highest between PME and fovea; pars thoracica rounded, marked with deep, circular fovea (
<figureCitation id="B6E44A81FFD4FF830464FE3011CC8DB4" box="[891,1009,454,478]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3B</figureCitation>
); ocular area and clypeus with few stiff setae; pars thoracica with fine recumbent setae; integument of carapace usually with numerous granules (
<figureCitation id="B6E44A81FFD4FF8305D4FDF0118E8E74" box="[715,947,518,542]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="9" pageNumber="1546">Figures 1A–B, D–H</figureCitation>
, 
<figureCitation id="B6E44A81FFD4FF8304A1FDF0167A8E74" box="[958,1095,518,542]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="9" pageNumber="1546">2A–E, G–H</figureCitation>
) or punctures (
<figureCitation id="B6E44A81FFD4FF83062DFDD013898E54" box="[306,436,550,574]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="9" pageNumber="1546">Figures 1C</figureCitation>
, 
<figureCitation id="B6E44A81FFD4FF8306DCFDD013DE8E54" box="[451,483,550,574]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="9" pageNumber="1546">2F</figureCitation>
) forming radiating striae; colour from dark chestnutbrown to brownish orange. Both eye rows straight in dorsal view (
<figureCitation id="B6E44A81FFD4FF83049BFDB011CD8E34" box="[900,1008,582,606]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="9" pageNumber="1546">Figures 1</figureCitation>
, 
<figureCitation id="B6E44A81FFD4FF8304E0FDB016328E34" box="[1023,1039,582,606]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="9" pageNumber="1546">2</figureCitation>
, 
<figureCitation id="B6E44A81FFD4FF830303FDB0167F8E34" box="[1052,1090,582,606]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">3D</figureCitation>
); in frontal view, AER very slightly procurved, PER slightly procurved (
<figureCitation id="B6E44A81FFD4FF8304D5FD90167C8E14" box="[970,1089,614,638]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3E</figureCitation>
); AME circular, dark; PME oval, pearly white; ALE and PLE subcircular, light; anterior eyes separated by their diameter or less; posterior eyes evenly spaced, separated by 1.5–2 their diameter (except in 
<taxonomicName id="E9DF2D87FFD4FF8306E9FD3010688EB4" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Dankittipakul &amp; Tavano &amp; Singtripop" baseAuthorityYear="2013" box="[502,597,710,734]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="1546" phylum="Arthropoda" rank="species" species="mihun" status="comb. nov.">
<emphasis id="1CAB8A16FFD4FF8306E9FD3010688EB4" box="[502,597,710,734]" italics="true" pageId="9" pageNumber="1546">J. mihun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD4FF830542FD3310E78EB7" box="[605,730,709,733]" pageId="9" pageNumber="1546" rank="species">comb. nov.</taxonomicNameLabel>
and 
<taxonomicName id="E9DF2D87FFD4FF830409FD3011418EB4" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Jager" baseAuthorityYear="2007" box="[790,892,710,734]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="1546" phylum="Arthropoda" rank="species" species="thakek" status="comb. nov.">
<emphasis id="1CAB8A16FFD4FF830409FD3011418EB4" box="[790,892,710,734]" italics="true" pageId="9" pageNumber="1546">J. thakek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD4FF83049AFD33163C8EB7" box="[901,1025,709,733]" pageId="9" pageNumber="1546" rank="species">comb. nov.</taxonomicNameLabel>
whose PME are distinctly enlarged); ALE and PLE separated by 1–1.5 times their diameter. MOQ longer than wide, slightly wider in front than behind (wider behind than in front in 
<taxonomicName id="E9DF2D87FFD4FF8307B1FCD013318F57" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Dankittipakul &amp; Tavano &amp; Singtripop" baseAuthorityYear="2013" box="[174,268,805,829]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="1546" phylum="Arthropoda" rank="species" species="mihun" status="comb. nov.">
<emphasis id="1CAB8A16FFD4FF8307B1FCD013318F57" box="[174,268,805,829]" italics="true" pageId="9" pageNumber="1546">J. mihun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD4FF83060BFCD313AD8F57" box="[276,400,805,829]" pageId="9" pageNumber="1546" rank="species">comb. nov.</taxonomicNameLabel>
and 
<taxonomicName id="E9DF2D87FFD4FF8306D4FCD0100D8F57" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Jager" baseAuthorityYear="2007" box="[459,560,805,829]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="1546" phylum="Arthropoda" rank="species" species="thakek" status="comb. nov.">
<emphasis id="1CAB8A16FFD4FF8306D4FCD0100D8F57" box="[459,560,805,829]" italics="true" pageId="9" pageNumber="1546">J. thakek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD4FF830526FCD310888F57" box="[569,693,805,829]" pageId="9" pageNumber="1546" rank="species">comb. nov.</taxonomicNameLabel>
because of enlarged PME). Clypeal height 1.5 times diameter of AME. Chelicerae convex anteriorly, with conspicuous condyles, fang furrow with three promarginal teeth and one to three retromarginal denticles. Chilum heavily sclerotised, trapezoidal (
<figureCitation id="B6E44A81FFD4FF8305BEFC7311248FF7" box="[673,793,901,925]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3E</figureCitation>
). Mouthparts and sternum brownish orange, darker along sternal margin. Labium (
<figureCitation id="B6E44A81FFD4FF83043AFC5211A28FD6" box="[805,927,932,956]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3A</figureCitation>
) subtriangular, longer than wide, invaginated at posterolateral corners, with few strong setae on anterior surface. Gnathocoxae (
<figureCitation id="B6E44A81FFD4FF8306EFFC12105E8F96" box="[496,611,996,1020]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3A</figureCitation>
) rectangular, with very faint oblique depressions, each with anterolateral serrula and anteromedian scopula arising from pale apical area. Sternum (
<figureCitation id="B6E44A81FFD4FF83068EFBD210378856" box="[401,522,1060,1084]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3A</figureCitation>
) scutiform, strongly rebordered, posterior margin truncated, not protruded between coxae IV; anterior margin medially excavated to accommodate labium; lateral margin with triangular extensions fitting to and between coxae. Leg formula 4123; legs brownish orange; femora I always with two elongated disto-prolateral spines; patellae unmodified; anterior tibiae ventrally not flattened, with five to nine pairs of ventral spines and single prolateral spine; anterior metatarsi with four to six pairs of ventral spines; posterior metatarsi with brush of distoventral hairs, anterior without such brush; anterior tarsi devoid of spines; posterior tarsi with few spines; two pectinate claws carrying two to three denticles; claw tufts indistinct; trichobothria long, in two dorsal rows on tarsi and metatarsi, one row on tibiae increasing in length distally.
</paragraph>
<paragraph id="2E605604FFD4FF8007A5FA75130A8D15" blockId="9.[143,1104,352,1690]" lastBlockId="10.[211,1172,167,383]" lastPageId="10" lastPageNumber="1547" pageId="9" pageNumber="1546">
Opisthosoma (
<figureCitation id="B6E44A81FFD4FF83067DFA7513ED89F1" box="[354,464,1411,1435]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="9" pageNumber="1546">Figures 1</figureCitation>
, 
<figureCitation id="B6E44A81FFD4FF8306FFFA7513D389F1" box="[480,494,1411,1435]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="9" pageNumber="1546">2</figureCitation>
) ovoid in dorsal view, dark greenish dorsally, dorsal pattern consisting of two pale oval patches situated medially, followed by three to five chevrons. Dorsal scutum with poorly diffused edge, covering less than half the length of the opisthosoma in male, completely absent in female. Two pairs of muscle apodemes, their surface covered by minute, circular, heavily sclerotised spots; first pair of spots locating just behind dorsal scutum (
<figureCitation id="B6E44A81FFD4FF8305CFF9D411038A50" box="[720,830,1570,1594]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="9" pageNumber="1546">Figures 1</figureCitation>
, 
<figureCitation id="B6E44A81FFD4FF830451F9D411638A50" box="[846,862,1570,1594]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (A–H). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="9" pageNumber="1546">2</figureCitation>
), second pair always located at centre of second pale oval patches. Epigastric region entirely covered with scutum (
<figureCitation id="B6E44A81FFD4FF8307F0F99413598A10" box="[239,356,1634,1658]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3C</figureCitation>
), slightly extending anteriorly to form short collar. Ventral scutum absent. Postgenital scuta (
<figureCitation id="B6E44A81FFD4FF8306A2F97410098AF0" box="[445,564,1666,1690]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="9" pageNumber="1546">Figure 3C</figureCitation>
) triangular or semi-circular, situated posterior laterally, surrounding book lung openings but clearly separated from epigastric scutum. Colour of venter uniform in all species (
<figureCitation id="B6E44A81FFD7FF8005CCFF3111758CB5" box="[723,840,199,223]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (A–C, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="10" pageNumber="1547">Figure 3C</figureCitation>
), dark greenish, marked with dark trident on pale rectangular area, with numerous tiny sclerotised spots arranged into four longitudinal lines running between epigastric furrow and transverse tracheal spiracle. Spinnerets two-segmented; ALS conical, contiguous, distal segment short; PMS small, slender in males, flattened and elongated in females; PLS with short distal segment.
</paragraph>
<paragraph id="2E605604FFD7FF8007CCFE5216BB8EB1" blockId="10.[211,1172,420,731]" pageId="10" pageNumber="1547">
<emphasis id="1CAB8A16FFD7FF8007CCFE5213708DD6" box="[211,333,420,444]" italics="true" pageId="10" pageNumber="1547">Male palp.</emphasis>
Palpal femur without modification. Retrolateral tibial apophysis consisting of basal and distal parts (
<figureCitation id="B6E44A81FFD7FF800501FE32109E8DB6" box="[542,675,452,476]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="10" pageNumber="1547">Figure 12A</figureCitation>
); basal RTA triangular or digitiform, apical surface usually with deep excavation; distal RTA short and membranous, digitiform. Cymbium dorsally with mid-longitudinal patch of chemosensitive hairs (
<figureCitation id="B6E44A81FFD7FF8004E3FDF512D18E51" captionStart="Figure 4" captionStartId="8.[211,278,1428,1450]" captionTargetBox="[213,1170,168,1394]" captionTargetId="figure-5@8.[213,1170,168,1394]" captionTargetPageId="8" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (A–C); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (D–F); J. mihun, holotype (G–I)." figureDoi="http://doi.org/10.5281/zenodo.6517648" httpUri="https://zenodo.org/record/6517648/files/figure.png" pageId="10" pageNumber="1547">Figure 4B, E, H</figureCitation>
). Tegulum elongate-ovoid, with distinct apico-prolateral depression accommodating base of embolus. Embolus filiform, elongated, curving behind tegulum (
<figureCitation id="B6E44A81FFD7FF800306FDB512D58E11" captionStart="Figure 4" captionStartId="8.[211,278,1428,1450]" captionTargetBox="[213,1170,168,1394]" captionTargetId="figure-5@8.[213,1170,168,1394]" captionTargetPageId="8" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (A–C); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (D–F); J. mihun, holotype (G–I)." figureDoi="http://doi.org/10.5281/zenodo.6517648" httpUri="https://zenodo.org/record/6517648/files/figure.png" pageId="10" pageNumber="1547">Figure 4A, B</figureCitation>
). Tegular apophysis hyaline, gradually tapered, originating near base of conductor (
<figureCitation id="B6E44A81FFD7FF8007C4FD7513638EF1" box="[219,350,643,667]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A–C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="10" pageNumber="1547">Figures 6A</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF800672FD7513AE8EF1" box="[365,403,643,667]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A–C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="10" pageNumber="1547">9D</figureCitation>
). Conductor originating on disto-prolateral surface of tegulum; its shape from simple, triangular projection (
<figureCitation id="B6E44A81FFD7FF8005B1FD55116A8ED1" box="[686,855,675,699]" captionStart="Figure 10" captionStartId="18.[211,278,1446,1468]" captionTargetBox="[213,1170,167,1412]" captionTargetId="figure-5@18.[213,1170,167,1413]" captionTargetPageId="18" captionText="Figure 10. Jacaena mihun, holotype (A–C); J. thakek comb. nov., holotype (D–F). Male palp, ventral (A, D). Ditto, prolateral (B, E). Ditto, retrolateral (C, F)." figureDoi="http://doi.org/10.5281/zenodo.6517670" httpUri="https://zenodo.org/record/6517670/files/figure.png" pageId="10" pageNumber="1547">Figure 10A, D</figureCitation>
) to elongated cylinder with median groove (
<figureCitation id="B6E44A81FFD7FF800698FD3510348EB1" box="[391,521,707,731]" captionStart="Figure 5" captionStartId="11.[143,210,1565,1587]" captionTargetBox="[145,1102,161,1532]" captionTargetId="figure-5@11.[145,1102,161,1532]" captionTargetPageId="11" captionText="Figure 5. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena erawan comb. nov., holotype (A–C); J. punctata sp. nov., holotype (D–F); J. thakek comb. nov., holotype (G–I)." figureDoi="http://doi.org/10.5281/zenodo.6517654" httpUri="https://zenodo.org/record/6517654/files/figure.png" pageId="10" pageNumber="1547">Figures 5D</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF80050AFD3510788EB1" box="[533,581,707,731]" captionStart="Figure 13" captionStartId="23.[143,210,1158,1180]" captionTargetBox="[146,1102,171,1126]" captionTargetId="figure-5@23.[145,1102,168,1126]" captionTargetPageId="23" captionText="Figure 13. Jacaena punctata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal genitalia, dorsal (E). CO, copulatory orifice; P, secretory plug." figureDoi="http://doi.org/10.5281/zenodo.6517676" httpUri="https://zenodo.org/record/6517676/files/figure.png" pageId="10" pageNumber="1547">13A</figureCitation>
) or large crescent-shaped lamina (
<figureCitation id="B6E44A81FFD7FF8004D9FD35167A8EB1" box="[966,1095,707,731]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A–C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="10" pageNumber="1547">Figures 6A</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF80034DFD3516458EB1" box="[1106,1144,707,731]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (A–C); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="10" pageNumber="1547">9D</figureCitation>
).
</paragraph>
<paragraph id="2E605604FFD7FF8007CCFCF310D08856" blockId="10.[211,1172,773,1084]" pageId="10" pageNumber="1547">
<emphasis id="1CAB8A16FFD7FF8007CCFCF313CB8F77" box="[211,502,773,797]" italics="true" pageId="10" pageNumber="1547">Female copulatory organ.</emphasis>
Epigynal region (
<figureCitation id="B6E44A81FFD7FF8005D1FCF311098F77" box="[718,820,773,797]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A–F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="10" pageNumber="1547">Figure 7</figureCitation>
) heavily sclerotised, with two copulatory orifices usually filled with dark secretory substance (
<figureCitation id="B6E44A81FFD7FF80048DFCD3161B8F57" box="[914,1062,805,829]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A–F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="10" pageNumber="1547">Figure 7A, B</figureCitation>
). Internal duct system consisting of semi-transparent proximal duct (
<figureCitation id="B6E44A81FFD7FF800483FCB3161C8F37" box="[924,1057,837,861]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A–C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="10" pageNumber="1547">Figures 6F</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF80032AFCB3165E8F37" box="[1077,1123,837,861]" captionStart="Figure 14" captionStartId="26.[211,278,1060,1082]" captionTargetBox="[213,1167,169,1027]" captionTargetId="figure-5@26.[213,1170,169,1027]" captionTargetPageId="26" captionText="Figure 14. Jacaena angoonae sp. nov., holotype, female (A–C); J. peculiaris sp. nov., holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C)." figureDoi="http://doi.org/10.5281/zenodo.6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="10" pageNumber="1547">14C</figureCitation>
) or enlarged chamber (
<figureCitation id="B6E44A81FFD7FF8006B0FC93100E8F17" box="[431,563,869,893]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A–C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="10" pageNumber="1547">Figures 6D</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF80055EFC9310538F17" box="[577,622,869,893]" captionStart="Figure 13" captionStartId="23.[143,210,1158,1180]" captionTargetBox="[146,1102,171,1126]" captionTargetId="figure-5@23.[145,1102,168,1126]" captionTargetPageId="23" captionText="Figure 13. Jacaena punctata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal genitalia, dorsal (E). CO, copulatory orifice; P, secretory plug." figureDoi="http://doi.org/10.5281/zenodo.6517676" httpUri="https://zenodo.org/record/6517676/files/figure.png" pageId="10" pageNumber="1547">13E</figureCitation>
) connected to loosely coiled insemination ducts (
<figureCitation id="B6E44A81FFD7FF8007C4FC7313628FF7" box="[219,351,901,925]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="10" pageNumber="1547">Figure 12E</figureCitation>
: ID) and spermathecae. Spermathecae slightly enlarged, poorly defined (
<figureCitation id="B6E44A81FFD7FF8007C4FC5213AB8FD6" box="[219,406,932,956]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="10" pageNumber="1547">Figure 8A–D, G</figureCitation>
) or with elliptical and spherical enlargement (
<figureCitation id="B6E44A81FFD7FF800484FC5216108FD6" box="[923,1069,932,956]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="10" pageNumber="1547">Figure 8E, F</figureCitation>
); surface of spermathecae usually provided with glandular pores (
<figureCitation id="B6E44A81FFD7FF800457FC3211F78FB6" box="[840,970,964,988]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (A–G). Jacaena mihun (A–D); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="10" pageNumber="1547">Figures 8G</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF8004C9FC3216388FB6" box="[982,1029,964,988]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="10" pageNumber="1547">12E</figureCitation>
). Small, perforated ampulla (
<figureCitation id="B6E44A81FFD7FF80068DFC12100C8F96" box="[402,561,996,1020]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A–C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="10" pageNumber="1547">Figures 6D, F</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF800524FC1210578F96" box="[571,618,996,1020]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="10" pageNumber="1547">12E</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF80056BFC12109F8F96" box="[628,674,996,1020]" captionStart="Figure 13" captionStartId="23.[143,210,1158,1180]" captionTargetBox="[146,1102,171,1126]" captionTargetId="figure-5@23.[145,1102,168,1126]" captionTargetPageId="23" captionText="Figure 13. Jacaena punctata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal genitalia, dorsal (E). CO, copulatory orifice; P, secretory plug." figureDoi="http://doi.org/10.5281/zenodo.6517676" httpUri="https://zenodo.org/record/6517676/files/figure.png" pageId="10" pageNumber="1547">13E</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF8005B2FC1210C78F96" box="[685,762,996,1020]" captionStart="Figure 14" captionStartId="26.[211,278,1060,1082]" captionTargetBox="[213,1167,169,1027]" captionTargetId="figure-5@26.[213,1170,169,1027]" captionTargetPageId="26" captionText="Figure 14. Jacaena angoonae sp. nov., holotype, female (A–C); J. peculiaris sp. nov., holotype, female (D, E). Epigyne, ventral (A, D). Internal copulatory organ, dorsal (B, E). Proximal part of insemination duct (C)." figureDoi="http://doi.org/10.5281/zenodo.6517678" httpUri="https://zenodo.org/record/6517678/files/figure.png" pageId="10" pageNumber="1547">14B, E</figureCitation>
: AM) connected to proximal portion of insemination ducts or chambers (
<figureCitation id="B6E44A81FFD7FF80057CFBF210D88876" box="[611,741,1028,1052]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (A–C); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="10" pageNumber="1547">Figures 6D</figureCitation>
, 
<figureCitation id="B6E44A81FFD7FF8005F0FBF211228876" box="[751,799,1028,1052]" captionStart="Figure 13" captionStartId="23.[143,210,1158,1180]" captionTargetBox="[146,1102,171,1126]" captionTargetId="figure-5@23.[145,1102,168,1126]" captionTargetPageId="23" captionText="Figure 13. Jacaena punctata sp. nov., holotype, male (A–C) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal genitalia, dorsal (E). CO, copulatory orifice; P, secretory plug." figureDoi="http://doi.org/10.5281/zenodo.6517676" httpUri="https://zenodo.org/record/6517676/files/figure.png" pageId="10" pageNumber="1547">13E</figureCitation>
), usually with hardened secretion filling proximal portion of ducts (
<figureCitation id="B6E44A81FFD7FF800555FBD210DD8856" box="[586,736,1060,1084]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (A–F). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="10" pageNumber="1547">Figure 7A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFD7FF8007CCFB8A13C189F1" pageId="10" pageNumber="1547" type="biology_ecology">
<paragraph id="2E605604FFD7FF8007CCFB8A134188FE" blockId="10.[211,1172,1148,1435]" box="[211,380,1148,1172]" pageId="10" pageNumber="1547">
<heading id="7528E168FFD7FF8007CCFB8A134188FE" box="[211,380,1148,1172]" fontSize="10" level="3" pageId="10" pageNumber="1547" reason="5">
<emphasis id="1CAB8A16FFD7FF8007CCFB8A134188FE" box="[211,380,1148,1172]" italics="true" pageId="10" pageNumber="1547">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFD7FF8007CCFB5213C189F1" blockId="10.[211,1172,1148,1435]" pageId="10" pageNumber="1547">
Most 
<taxonomicName id="E9DF2D87FFD7FF800606FB5313F588D1" box="[281,456,1187,1212]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="1CAB8A16FFD7FF800606FB53134C88D6" box="[281,369,1189,1212]" italics="true" pageId="10" pageNumber="1547">Jacaena</emphasis>
species
</taxonomicName>
were collected by sifting decomposing organic litter in moist, primary evergreen hill forests at relatively high elevation. These forests are generally covered with a thick layer of humus on the forest floor and the forest canopy is often dense, allowing limited sunlight to reach the ground. A species from 
<collectingCountry id="56C81694FFD7FF800332FAF516A98971" box="[1069,1172,1283,1307]" name="Thailand" pageId="10" pageNumber="1547">Thailand</collectingCountry>
(
<taxonomicName id="E9DF2D87FFD7FF8007C3FAD213048951" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Dankittipakul &amp; Tavano &amp; Singtripop" baseAuthorityYear="2013" box="[220,313,1315,1339]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFD7FF8007C3FAD213048951" box="[220,313,1315,1339]" italics="true" pageId="10" pageNumber="1547">J. mihun</emphasis>
</taxonomicName>
) inhabits deciduous forests and open plantations. 
<taxonomicName id="E9DF2D87FFD7FF80046DFAD216208951" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Jager" baseAuthorityYear="2007" box="[882,1053,1315,1339]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="thakek">
<emphasis id="1CAB8A16FFD7FF80046DFAD216208951" box="[882,1053,1315,1339]" italics="true" pageId="10" pageNumber="1547">Jacaena thakek</emphasis>
</taxonomicName>
was sifted from relatively dry litter in a narrow belt of trees, bamboo or Rattan (disturbed secondary habitat); the 
<typeStatus id="F164E8A6FFD7FF8006A1FA9213D38916" box="[446,494,1380,1404]" pageId="10" pageNumber="1547">type</typeStatus>
locality of this species is almost of the same altitude of the Mekong basin (
<quantity id="E927FBE1FFD7FF80069DFA7513F589F1" box="[386,456,1411,1435]" metricMagnitude="2" metricUnit="m" metricValue="1.59" pageId="10" pageNumber="1547" unit="m" value="159.0">159 m</quantity>
alt.)
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFD7FF8007CCFA2D10328AD0" pageId="10" pageNumber="1547" type="discussion">
<paragraph id="2E605604FFD7FF8007CCFA2D13BE8998" blockId="10.[211,1172,1499,1722]" box="[211,387,1499,1523]" pageId="10" pageNumber="1547">
<heading id="7528E168FFD7FF8007CCFA2D13BE8998" box="[211,387,1499,1523]" fontSize="10" level="3" pageId="10" pageNumber="1547" reason="5">
<emphasis id="1CAB8A16FFD7FF8007CCFA2D13BE8998" box="[211,387,1499,1523]" italics="true" pageId="10" pageNumber="1547">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFD7FF8007CCF9F510328AD0" blockId="10.[211,1172,1499,1722]" pageId="10" pageNumber="1547">
<taxonomicName id="E9DF2D87FFD7FF8007CCF9F513A18A70" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[211,412,1539,1562]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="angoonae" status="sp. nov.">
<emphasis id="1CAB8A16FFD7FF8007CCF9F513A18A70" box="[211,412,1539,1562]" italics="true" pageId="10" pageNumber="1547">Jacaena angoonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF8006BBF9F513C18A70" box="[420,508,1539,1562]" pageId="10" pageNumber="1547" rank="species">sp. nov.</taxonomicNameLabel>
; 
<taxonomicName id="E9DF2D87FFD7FF800516F9F511198A70" authority="Thorell, 1897" authorityName="Thorell" authorityYear="1897" box="[521,804,1538,1562]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="distincta">
<emphasis id="1CAB8A16FFD7FF800516F9F510BC8A70" box="[521,641,1538,1562]" italics="true" pageId="10" pageNumber="1547">J. distincta</emphasis>
<bibRefCitation id="4A4E2BF5FFD7FF800596F9F411198A70" author="Thorell T" box="[649,804,1538,1562]" pageId="10" pageNumber="1547" pagination="161 - 267" refId="ref11552" refString="Thorell T. 1897. Viaggio di Leonardo Fea in Birmania e regioni vicine. LXXIII. Secondo saggio sui Ragni birmani. I. Parallelodontes. Tubitelariae. Annali Mus Civ St Nat Genova. 37: 161 - 267." type="journal article" year="1897">Thorell, 1897</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="E9DF2D87FFD7FF80042FF9F5132C8A50" authority="(Deeleman-Reinhold, 2001)" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="erawan" status="comb. nov.">
<emphasis id="1CAB8A16FFD7FF80042FF9F511A48A70" box="[816,921,1539,1562]" italics="true" pageId="10" pageNumber="1547">J. erawan</emphasis>
(
<bibRefCitation id="4A4E2BF5FFD7FF8004B7F9F413348A50" author="Deeleman-Reinhold CL" pageId="10" pageNumber="1547" refId="ref11331" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF800606F9D413AB8A50" box="[281,406,1570,1594]" pageId="10" pageNumber="1547" rank="species">comb. nov.</taxonomicNameLabel>
; 
<taxonomicName id="E9DF2D87FFD7FF8006BDF9D5102B8A50" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[418,534,1570,1594]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis id="1CAB8A16FFD7FF8006BDF9D5102B8A50" box="[418,534,1570,1594]" italics="true" pageId="10" pageNumber="1547">J. lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF800502F9D510498A50" box="[541,628,1571,1594]" pageId="10" pageNumber="1547" rank="species">sp. nov.</taxonomicNameLabel>
; 
<taxonomicName id="E9DF2D87FFD7FF80059FF9D516338A50" authority="Deeleman-Reinhold, 2001" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[640,1038,1570,1594]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFD7FF80059FF9D510E38A50" box="[640,734,1570,1594]" italics="true" pageId="10" pageNumber="1547">J. mihun</emphasis>
<bibRefCitation id="4A4E2BF5FFD7FF8005FAF9D416338A50" author="Deeleman-Reinhold CL" box="[741,1038,1570,1594]" pageId="10" pageNumber="1547" refId="ref11331" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="E9DF2D87FFD7FF800305F9D516A98A50" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[1050,1172,1571,1594]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="punctata" status="sp. nov.">
<emphasis id="1CAB8A16FFD7FF800305F9D516A98A50" box="[1050,1172,1571,1594]" italics="true" pageId="10" pageNumber="1547">J. punctata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF8007CCF9B513178A30" box="[211,298,1603,1626]" pageId="10" pageNumber="1547" rank="species">sp. nov.</taxonomicNameLabel>
; 
<taxonomicName id="E9DF2D87FFD7FF80062BF9B5138B8A30" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[308,438,1602,1626]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="peculiaris" status="sp. nov.">
<emphasis id="1CAB8A16FFD7FF80062BF9B5138B8A30" box="[308,438,1602,1626]" italics="true" pageId="10" pageNumber="1547">J. peculiaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF8006A2F9B510298A30" box="[445,532,1603,1626]" pageId="10" pageNumber="1547" rank="species">sp. nov.</taxonomicNameLabel>
; 
<taxonomicName id="E9DF2D87FFD7FF800501F9B516318A30" authority="(Deeleman-Reinhold, 2001)" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" box="[542,1036,1602,1626]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis id="1CAB8A16FFD7FF800501F9B510F28A30" box="[542,719,1602,1626]" italics="true" pageId="10" pageNumber="1547">J. schwendingeri</emphasis>
(
<bibRefCitation id="4A4E2BF5FFD7FF8005C2F9B416398A30" author="Deeleman-Reinhold CL" box="[733,1028,1602,1626]" pageId="10" pageNumber="1547" refId="ref11331" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." type="book" year="2001">Deeleman-Reinhold, 2001</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF80030CF9B416B28A30" box="[1043,1167,1602,1626]" pageId="10" pageNumber="1547" rank="species">comb. nov.</taxonomicNameLabel>
; 
<taxonomicName id="E9DF2D87FFD7FF8007CCF99513FA8A10" authority="(Jager, 2007)" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Jager" baseAuthorityYear="2007" box="[211,455,1634,1658]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="thakek" status="comb. nov.">
<emphasis id="1CAB8A16FFD7FF8007CCF995130B8A10" box="[211,310,1634,1658]" italics="true" pageId="10" pageNumber="1547">J. thakek</emphasis>
(
<bibRefCitation id="4A4E2BF5FFD7FF80065DF99413FD8A10" author="Jager P" box="[322,448,1634,1658]" pageId="10" pageNumber="1547" pagination="29 - 58" refId="ref11377" refString="Jager P. 2007. Spiders from Laos with descriptions of new species (Arachnida: Araneae). Acta arachn Tokyo. 56: 29 - 58." type="journal article" year="2007">Jäger, 2007</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF8006D2F99410758A10" box="[461,584,1634,1658]" pageId="10" pageNumber="1547" rank="species">comb. nov.</taxonomicNameLabel>
; 
<taxonomicName id="E9DF2D87FFD7FF80054EF99511B18A10" authority="(Zhang and Fu, 2011)" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Zhang and Fu" baseAuthorityYear="2011" box="[593,908,1634,1658]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="1547" phylum="Arthropoda" rank="species" species="zhui" status="comb. nov.">
<emphasis id="1CAB8A16FFD7FF80054EF99510A88A10" box="[593,661,1634,1658]" italics="true" pageId="10" pageNumber="1547">J. zhui</emphasis>
(
<bibRefCitation id="4A4E2BF5FFD7FF8005BCF99411B88A10" author="Zhang F &amp; Fu JY" box="[675,901,1634,1658]" pageId="10" pageNumber="1547" pagination="69 - 74" refId="ref11594" refString="Zhang F, Fu JY. 2011. First report of the genus Sesieutes Simon (Araneae: Liocranidae) from China, with description of one new species. Ent News. 121: 69 - 74." type="journal article" year="2011">Zhang and Fu, 2011</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD7FF80048DF99416378A10" box="[914,1034,1634,1658]" pageId="10" pageNumber="1547" rank="species">comb. nov.</taxonomicNameLabel>
Two further species were also collected from 
<collectingCountry id="56C81694FFD7FF80055FF974109D8AF0" box="[576,672,1666,1690]" name="Vietnam" pageId="10" pageNumber="1547">Vietnam</collectingCountry>
(male and female) and 
<collectingCountry id="56C81694FFD7FF8004B3F97411CC8AF0" box="[940,1009,1666,1690]" name="China" pageId="10" pageNumber="1547">China</collectingCountry>
(female); they have not yet been described.
</paragraph>
</subSubSection>
<caption id="7AA0068CFFD6FF810790F9EB10698A07" ID-DOI="http://doi.org/10.5281/zenodo.6517654" ID-Zenodo-Dep="6517654" httpUri="https://zenodo.org/record/6517654/files/figure.png" pageId="11" pageNumber="1548" startId="11.[143,210,1565,1587]" targetBox="[145,1102,161,1532]" targetPageId="11">
<paragraph id="2E605604FFD6FF810790F9EB10698A07" blockId="11.[143,1104,1565,1646]" pageId="11" pageNumber="1548">
Figure 5. 
<taxonomicName id="E9DF2D87FFD6FF8107E3F9E813BC8A59" box="[252,385,1566,1587]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="1548" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="1CAB8A16FFD6FF8107E3F9E813768A59" box="[252,331,1566,1587]" italics="true" pageId="11" pageNumber="1548">Jacaena</emphasis>
spp.
</taxonomicName>
Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). 
<taxonomicName id="E9DF2D87FFD6FF81067CF9CD13C38A3A" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" box="[355,510,1595,1616]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="1548" phylum="Arthropoda" rank="species" species="erawan" status="comb. nov.">
<emphasis id="1CAB8A16FFD6FF81067CF9CD13C38A3A" box="[355,510,1595,1616]" italics="true" pageId="11" pageNumber="1548">Jacaena erawan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD6FF81051BF9CC10488A3A" box="[516,629,1594,1616]" pageId="11" pageNumber="1548" rank="species">comb. nov.</taxonomicNameLabel>
, holotype (A–C); 
<taxonomicName id="E9DF2D87FFD6FF810434F9CD11AA8A3A" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[811,919,1595,1616]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="1548" phylum="Arthropoda" rank="species" species="punctata" status="sp. nov.">
<emphasis id="1CAB8A16FFD6FF810434F9CD11AA8A3A" box="[811,919,1595,1616]" italics="true" pageId="11" pageNumber="1548">J. punctata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD6FF810482F9CD11D18A3A" box="[925,1004,1595,1616]" pageId="11" pageNumber="1548" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (D–F); 
<taxonomicName id="E9DF2D87FFD6FF8107C4F9AE13088A04" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Jager" baseAuthorityYear="2007" box="[219,309,1624,1646]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="1548" phylum="Arthropoda" rank="species" species="thakek" status="comb. nov.">
<emphasis id="1CAB8A16FFD6FF8107C4F9AE13088A04" box="[219,309,1624,1646]" italics="true" pageId="11" pageNumber="1548">J. thakek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD6FF810624F9A113918A07" box="[315,428,1623,1645]" pageId="11" pageNumber="1548" rank="species">comb. nov.</taxonomicNameLabel>
, holotype (G–I).
</paragraph>
</caption>
<caption id="7AA0068CFFD1FF8607CCFB8010678956" ID-DOI="http://doi.org/10.5281/zenodo.6517660" ID-Zenodo-Dep="6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="12" pageNumber="1549" startId="12.[211,278,1142,1164]" targetBox="[213,1170,171,1109]" targetPageId="12">
<paragraph id="2E605604FFD1FF8607CCFB8010678956" blockId="12.[211,1172,1142,1340]" pageId="12" pageNumber="1549">
Figure 6. 
<taxonomicName id="E9DF2D87FFD1FF860621FB8113FD88E6" box="[318,448,1143,1164]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="1549" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="1CAB8A16FFD1FF860621FB8113B088E6" box="[318,397,1143,1164]" italics="true" pageId="12" pageNumber="1549">Jacaena</emphasis>
spp.
</taxonomicName>
Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). 
<taxonomicName id="E9DF2D87FFD1FF8607E0FB1A13988968" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[255,421,1260,1282]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="1549" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis id="1CAB8A16FFD1FF8607E0FB1A13988968" box="[255,421,1260,1282]" italics="true" pageId="12" pageNumber="1549">Jacaena lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD1FF8606B6FB1A13CA896B" box="[425,503,1260,1281]" pageId="12" pageNumber="1549" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (A–C); 
<taxonomicName id="E9DF2D87FFD1FF8605B5FB1A11238968" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[682,798,1260,1282]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="1549" phylum="Arthropoda" rank="species" species="peculiaris" status="sp. nov.">
<emphasis id="1CAB8A16FFD1FF8605B5FB1A11238968" box="[682,798,1260,1282]" italics="true" pageId="12" pageNumber="1549">J. peculiaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD1FF86043CFB1A114C896B" box="[803,881,1260,1281]" pageId="12" pageNumber="1549" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (D); 
<taxonomicName id="E9DF2D87FFD1FF860318FB1A164E896B" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[1031,1139,1260,1281]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="1549" phylum="Arthropoda" rank="species" species="punctata" status="sp. nov.">
<emphasis id="1CAB8A16FFD1FF860318FB1A164E896B" box="[1031,1139,1260,1281]" italics="true" pageId="12" pageNumber="1549">J. punctata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFD1FF860367FB1A133C8974" pageId="12" pageNumber="1549" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (E); 
<taxonomicName id="E9DF2D87FFD1FF860685FAFF13D28975" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[410,495,1289,1311]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="1549" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFD1FF860685FAFF13D28975" box="[410,495,1289,1311]" italics="true" pageId="12" pageNumber="1549">J. mihun</emphasis>
</taxonomicName>
(F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct.
</paragraph>
</caption>
<subSubSection id="66C5058FFFD1FF8607CCFA8D167289B0" pageId="12" pageNumber="1549" type="distribution">
<paragraph id="2E605604FFD1FF8607CCFA8D136589F9" blockId="12.[211,1171,1403,1499]" box="[211,344,1403,1427]" pageId="12" pageNumber="1549">
<heading id="7528E168FFD1FF8607CCFA8D136589F9" box="[211,344,1403,1427]" fontSize="10" level="3" pageId="12" pageNumber="1549" reason="5">
<emphasis id="1CAB8A16FFD1FF8607CCFA8D136589F9" box="[211,344,1403,1427]" italics="true" pageId="12" pageNumber="1549">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFD1FF8607CCFA55167289B0" blockId="12.[211,1171,1403,1499]" pageId="12" pageNumber="1549">
Predominantly Indo-Burmese sub-region of the Oriental Region (
<collectingCountry id="56C81694FFD1FF8604B5FA55162189D1" box="[938,1052,1443,1467]" name="Myanmar" pageId="12" pageNumber="1549">Myanmar</collectingCountry>
, 
<collectingCountry id="56C81694FFD1FF86033AFA5516B289D1" box="[1061,1167,1443,1467]" name="Thailand" pageId="12" pageNumber="1549">Thailand</collectingCountry>
, 
<collectingCountry id="56C81694FFD1FF8607CCFA35133189B0" box="[211,268,1475,1498]" name="Laos" pageId="12" pageNumber="1549">Laos</collectingCountry>
, 
<collectingCountry id="56C81694FFD1FF860607FA34134189B0" box="[280,380,1474,1498]" name="Vietnam" pageId="12" pageNumber="1549">Vietnam</collectingCountry>
), also in evergreen forests of southern 
<collectingCountry id="56C81694FFD1FF860437FA34115089B0" box="[808,877,1474,1498]" name="China" pageId="12" pageNumber="1549">China</collectingCountry>
(
<collectingRegion id="EC1B98E6FFD1FF860464FA34167989B0" box="[891,1092,1474,1498]" country="China" name="Yunnan" pageId="12" pageNumber="1549">Yunnan Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>