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<document id="A7C0A71CCA549ADDA5E7960BBDC74B29" ID-CLB-Dataset="55811" ID-DOI="10.1080/00222933.2012.763059" ID-GBIF-Dataset="dfc793f4-7033-4586-9849-718e7e526360" ID-ISSN="1464-5262" ID-Zenodo-Dep="556127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628879433880" checkinUser="felipe" docAuthor="Dankittipakul, Pakawin, Tavano, Maria &amp; Singtripop, Tippawan" docDate="2013" docId="A676E712FFCEFF9C06FAF9841095891A" docLanguage="en" docName="JNatHist.47.23-24.1539-1567.pdf" docOrigin="Journal of Natural History 47 (23 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2012.763059" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Jacaena lunulata Dankittipakul &amp; Tavano &amp; Singtripop 2013, sp. nov." docType="treatment" docVersion="11" lastPageNumber="1559" masterDocId="5A4F9F6AFFDDFF8A071FFFF6123D8C6A" masterDocTitle="Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)" masterLastPageNumber="1567" masterPageNumber="1539" pageNumber="1556" updateTime="1699079059524" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F0EFC163B52D18DEC3F37926A6E35A6E">Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)</mods:title>
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<mods:namePart id="2A729D1BBB9AA579E754B38F829AD1CF">Dankittipakul, Pakawin</mods:namePart>
<mods:affiliation id="EB39359EA97969F856E6F08C32FA440C">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand; &amp; Manukau Institute of Technology, Manukau, New Zealand;</mods:affiliation>
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<mods:namePart id="D07748BF868A40606647C79D9512BA24">Tavano, Maria</mods:namePart>
<mods:affiliation id="D84D74E13BFF3836CB85B5B9789D7FD9">Museo Civico di Storia Naturale Giacomo Doria, Genova, Italy</mods:affiliation>
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<mods:namePart id="A355A0DBDE03D6BBA5522EBC45B8E3AA">Singtripop, Tippawan</mods:namePart>
<mods:affiliation id="F8E45E7FFCDF94EB5F52F97B52418CD4">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand;</mods:affiliation>
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<mods:title id="D54B4FCE034ADC445551583E2F840D40">Journal of Natural History</mods:title>
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<mods:date id="BCC8FE72D39043085508921BB887112D">2013</mods:date>
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<mods:number id="F88C00D183D6F119E6AF9E96A01306E7">23 - 24</mods:number>
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<treatment id="A676E712FFCEFF9C06FAF9841095891A" ID-DOI="http://doi.org/10.5281/zenodo.5196930" ID-GBIF-Taxon="194854858" ID-Zenodo-Dep="5196930" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A" httpUri="http://treatment.plazi.org/id/A676E712FFCEFF9C06FAF9841095891A" lastPageId="22" lastPageNumber="1559" pageId="19" pageNumber="1556">
<subSubSection id="66C5058FFFCEFF9906FAF98410C78AE0" box="[485,762,1649,1674]" pageId="19" pageNumber="1556" type="nomenclature">
<paragraph id="2E605604FFCEFF9906FAF98410C78AE0" blockId="19.[258,989,1649,1706]" box="[485,762,1649,1674]" pageId="19" pageNumber="1556">
<heading id="7528E168FFCEFF9906FAF98410C78AE0" box="[485,762,1649,1674]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="1556" reason="2">
<taxonomicName id="E9DF2D87FFCEFF9906FAF98410A28AE0" ID-CoL="6NDFH" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[485,671,1649,1674]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="1556" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis id="1CAB8A16FFCEFF9906FAF98410A28AE0" bold="true" box="[485,671,1649,1674]" italics="true" pageId="19" pageNumber="1556">Jacaena lunulata</emphasis>
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<taxonomicNameLabel id="0798376DFFCEFF9905BAF98510C78AE0" box="[677,762,1651,1674]" pageId="19" pageNumber="1556" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="66C5058FFFCEFF99061DF96411E08AC0" box="[258,989,1682,1706]" pageId="19" pageNumber="1556" type="description">
<paragraph id="2E605604FFCEFF99061DF96411E08AC0" blockId="19.[258,989,1649,1706]" box="[258,989,1682,1706]" pageId="19" pageNumber="1556">
(
<figureCitation id="B6E44A81FFCEFF990616F96413978AC3" box="[265,426,1682,1706]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (AH). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="19" pageNumber="1556">Figures 1B, H</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF9906A9F96413CA8AC0" box="[438,503,1682,1706]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (AH). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="19" pageNumber="1556">2B, C</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF99051DF96410588AC3" box="[514,613,1682,1706]" captionStart="Figure 3" captionStartId="7.[143,210,1174,1196]" captionTargetBox="[145,1102,169,1141]" captionTargetId="figure-5@7.[145,1102,169,1141]" captionTargetPageId="7" captionText="Figure 3. Jacaena spp. Prosoma and anterior part of opisthosoma showing epigastric scutum, ventral (A). Fovea, dorsal (B). Opisthosoma, ventral (C). Carapace, dorsal, showing ocular region (D). Carapace, frontal (E). Jacaena lunulata sp. nov., male, paratype (AC, E); J. mihun, male, new material (D)." figureDoi="http://doi.org/10.5281/zenodo.6517646" httpUri="https://zenodo.org/record/6517646/files/figure.png" pageId="19" pageNumber="1556">3AC, E</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF99056EF964108F8AC0" box="[625,690,1682,1706]" captionStart="Figure 4" captionStartId="8.[211,278,1428,1450]" captionTargetBox="[213,1170,168,1394]" captionTargetId="figure-5@8.[213,1170,168,1394]" captionTargetPageId="8" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (DF); J. mihun, holotype (GI)." figureDoi="http://doi.org/10.5281/zenodo.6517648" httpUri="https://zenodo.org/record/6517648/files/figure.png" pageId="19" pageNumber="1556">4DF</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF9905A2F964113F8AC0" box="[701,770,1682,1706]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (AC); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="19" pageNumber="1556">6AC</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF990412F96411128AC0" box="[781,815,1682,1706]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (AF). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="19" pageNumber="1556">7C</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF990425F96411638AC0" box="[826,862,1682,1706]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (AG). Jacaena mihun (AD); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="19" pageNumber="1556">8G</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF990475F96411938AC3" box="[874,942,1682,1706]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="19" pageNumber="1556">9D, E</figureCitation>
,
<figureCitation id="B6E44A81FFCEFF9904A6F96411E98AC0" box="[953,980,1682,1706]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="19" pageNumber="1556">12</figureCitation>
)
</paragraph>
</subSubSection>
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<treatmentCitationGroup id="0ECF712AFFC9FF9E07CCFF5E13A68CB5" pageId="20" pageNumber="1557">
<treatmentCitation id="AF7E7015FFC9FF9E07CCFF5E11028CD5" author="Deeleman-Reinhold CL" box="[211,831,167,192]" page="464" pageId="20" pageNumber="1557" year="2001">
<taxonomicName id="E9DF2D87FFC9FF9E07CCFF5E11028CD5" authority="Deeleman-Reinhold, 2001: 464" authorityName="Deeleman-Reinhold" authorityPageNumber="464" authorityYear="2001" box="[211,831,167,192]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="erawan">
<emphasis id="1CAB8A16FFC9FF9E07CCFF5E139D8CD5" box="[211,416,168,192]" italics="true" pageId="20" pageNumber="1557">Sesieutes erawan</emphasis>
<bibRefCitation id="4A4E2BF5FFC9FF9E06A5FF5111028CD5" author="Deeleman-Reinhold CL" box="[442,831,167,191]" pageId="20" pageNumber="1557" refId="ref11331" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." type="book" year="2001">Deeleman-Reinhold, 2001: 464</bibRefCitation>
</taxonomicName>
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,
<specimenCount id="38D99D8DFFC9FF9E0442FF5E114F8CAA" box="[861,882,168,192]" pageId="20" pageNumber="1557" type="male"></specimenCount>
from Doi Inthanon. Misidentification.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2E605604FFC9FF9E07CCFF0110148D45" blockId="20.[211,1172,247,303]" pageId="20" pageNumber="1557">
<treatmentCitationGroup id="0ECF712AFFC9FF9E07CCFF0110148D45" pageId="20" pageNumber="1557">
<treatmentCitation id="AF7E7015FFC9FF9E07CCFF01117E8D65" author="Deeleman-Reinhold CL" box="[211,835,247,271]" page="461" pageId="20" pageNumber="1557" year="2001">
<taxonomicName id="E9DF2D87FFC9FF9E07CCFF01117E8D65" authority="Deeleman-Reinhold, 2001: 461" authorityName="Deeleman-Reinhold" authorityPageNumber="461" authorityYear="2001" box="[211,835,247,271]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis id="1CAB8A16FFC9FF9E07CCFF0113E48D65" box="[211,473,247,271]" italics="true" pageId="20" pageNumber="1557">Sesieutes schwendingeri</emphasis>
<bibRefCitation id="4A4E2BF5FFC9FF9E06FEFF01117E8D65" author="Deeleman-Reinhold CL" box="[481,835,247,271]" pageId="20" pageNumber="1557" refId="ref11331" refString="Deeleman-Reinhold CL. 2001. Forest spiders of South East Asia - with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]. Leiden, the Netherland: Brill." type="book" year="2001">Deeleman-Reinhold, 2001: 461</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="38D99D8DFFC9FF9E044FFF0111478D65" box="[848,890,247,271]" pageId="20" pageNumber="1557" type="male">♂♂</specimenCount>
from Doi Inthanon and Doi Suthep. Misidentification.
</treatmentCitationGroup>
</paragraph>
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<heading id="7528E168FFC9FF9E07CCFE9313518D16" box="[211,364,356,380]" fontSize="10" level="3" pageId="20" pageNumber="1557" reason="5">
<emphasis id="1CAB8A16FFC9FF9E07CCFE9313518D16" box="[211,364,356,380]" italics="true" pageId="20" pageNumber="1557">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC9FF9E07CCFE7A10C08D8E" blockId="20.[211,1171,356,484]" pageId="20" pageNumber="1557">
<materialsCitation id="9EB75C59FFC9FF9E07CCFE7A10C08D8E" ID-GBIF-Occurrence="3757642334" collectingDate="2006-01-11" collectingMethod="extraction of soil sample" collectionCode="MHNG" collectorName="P. Dankittipakul" country="Thailand" county="Chomthong District" elevation="1640" location="Doi Inthanon NP" municipality="Chomthong District" pageId="20" pageNumber="1557" specimenCode="MHNG, PDC 5746" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai Province" typeStatus="holotype">
<typeStatus id="F164E8A6FFC9FF9E07CCFE7A137F8DCE" box="[211,322,396,420]" pageId="20" pageNumber="1557" type="holotype">Holotype</typeStatus>
:
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,
<collectingCountry id="56C81694FFC9FF9E0672FE7A13EA8DCE" box="[365,471,396,420]" name="Thailand" pageId="20" pageNumber="1557">Thailand</collectingCountry>
:
<collectingRegion id="EC1B98E6FFC9FF9E06FFFE7A10F28DCE" box="[480,719,396,420]" country="Thailand" name="Chiang Mai" pageId="20" pageNumber="1557">Chiang Mai Province</collectingRegion>
,
<collectingMunicipality id="CE04CC7EFFC9FF9E05C6FE7A11828DCE" box="[729,959,396,420]" pageId="20" pageNumber="1557">Chomthong District</collectingMunicipality>
,
<location id="2B0000DFFFC9FF9E04D6FE7A16B28DCE" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E04D6FE7A16B28DCE" box="[969,1167,396,420]" country="Thailand" county="Chomthong District" municipality="Chomthong District" name="Doi Inthanon NP" pageId="20" pageNumber="1557" stateProvince="Chiang Mai Province">Doi Inthanon NP</location>
,
<location id="2B0000DFFFC9FF9E07CCFE5A134C8DAE" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E07CCFE5A134C8DAE" box="[211,369,428,452]" country="Thailand" county="Chomthong District" municipality="Chomthong District" name="Doi Inthanon" pageId="20" pageNumber="1557" stateProvince="Chiang Mai Province">Doi Inthanon</location>
,
<location id="2B0000DFFFC9FF9E0664FE5B106B8DAE" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E0664FE5B106B8DAE" box="[379,598,428,453]" country="Thailand" county="Chomthong District" municipality="Chomthong District" name="evergreen hill forest" pageId="20" pageNumber="1557" stateProvince="Chiang Mai Province">evergreen hill forest</location>
,
<quantity id="E927FBE1FFC9FF9E057FFE5A10CB8DAE" box="[608,758,428,452]" metricMagnitude="3" metricUnit="m" metricValue="1.64" metricValueMax="1.68" metricValueMin="1.6" pageId="20" pageNumber="1557" unit="m" value="1640.0" valueMax="1680.0" valueMin="1600.0">
<elevation id="A5F2B137FFC9FF9E057FFE5A10CB8DAE" box="[608,758,428,452]" metricMagnitude="3" metricUnit="m" metricValue="1.64" metricValueMax="1.68" metricValueMin="1.6" pageId="20" pageNumber="1557" unit="m" value="1640.0" valueMax="1680.0" valueMin="1600.0">16001680 m</elevation>
</quantity>
,
<collectingMethod id="F79E2E13FFC9FF9E05E0FE5A162C8DAE" box="[767,1041,428,452]" pageId="20" pageNumber="1557">extraction of soil sample</collectingMethod>
,
<date id="5A6170C4FFC9FF9E0305FE5A13368D8E" pageId="20" pageNumber="1557" value="2006-01-11">
<collectingDate id="4A25892CFFC9FF9E0305FE5A13368D8E" pageId="20" pageNumber="1557" value="2006-01-11">11 January 2006</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E0609FE3A13EC8D8E" box="[278,465,460,484]" pageId="20" pageNumber="1557">P. Dankittipakul</collectorName>
leg. (
<collectionCode id="48CECEC1FFC9FF9E0511FE3A10518D8E" box="[526,620,460,484]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="20" pageNumber="1557">MHNG</collectionCode>
, PDC 5746).
</materialsCitation>
</paragraph>
<paragraph id="2E605604FFC9FF9E07CCFE0C11D18F05" blockId="20.[211,1172,505,880]" pageId="20" pageNumber="1557">
<materialsCitation id="9EB75C59FFC9FF9E07CCFE0C10098E5B" ID-GBIF-Occurrence="3757642317" collectingDate="2000-03-25" collectingDateMax="2000-04-23" collectingDateMin="2000-03-25" collectingMethod="pitfall trap" collectionCode="MHNG" collectorName="P. Dankittipakul" country="Thailand" county="Chomthong District" elevation="1640" location="Doi Inthanon" municipality="Doi Inthanon" pageId="20" pageNumber="1557" specimenCode="MHNG, PDC 5747" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus id="F164E8A6FFC9FF9E07CCFE0C13788E78" box="[211,325,506,530]" pageId="20" pageNumber="1557" type="paratype">Paratypes</typeStatus>
: From type locality,
<collectingMethod id="F79E2E13FFC9FF9E0531FE0F10958E7B" box="[558,680,505,529]" pageId="20" pageNumber="1557">pitfall trap</collectingMethod>
,
<date id="5A6170C4FFC9FF9E05ABFE0F11F18E7B" box="[692,972,505,529]" pageId="20" pageNumber="1557" value="2000-03-25" valueMax="2000-04-23" valueMin="2000-03-25">
<collectingDate id="4A25892CFFC9FF9E05ABFE0F11F18E7B" box="[692,972,505,529]" pageId="20" pageNumber="1557" value="2000-03-25" valueMax="2000-04-23" valueMin="2000-03-25">25 March23 April 2000</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E04C7FE0C16AE8E7B" box="[984,1171,505,529]" pageId="20" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E0612FDEF13148E5B" box="[269,297,537,561]" pageId="20" pageNumber="1557" type="female">1♀</specimenCount>
(
<collectionCode id="48CECEC1FFC9FF9E0622FDEF13A68E5B" box="[317,411,537,561]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="20" pageNumber="1557">MHNG</collectionCode>
, PDC 5747);
</materialsCitation>
<materialsCitation id="9EB75C59FFC9FF9E055FFDEF13E68E3A" ID-GBIF-Occurrence="3757642321" collectingDate="2005-09-10" collectingMethod="litter sample" collectionCode="TNHM" collectorName="P. Dankittipakul" county="Chomthong District" elevation="1640" location="Doi Inthanon" municipality="Doi Inthanon" pageId="20" pageNumber="1557" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiang Mai" typeStatus="paratype">
<collectingMethod id="F79E2E13FFC9FF9E055FFDEF10EE8E5B" box="[576,723,537,561]" pageId="20" pageNumber="1557">litter sample</collectingMethod>
,
<date id="5A6170C4FFC9FF9E05FDFDEF11F98E5B" box="[738,964,537,561]" pageId="20" pageNumber="1557" value="2005-09-10">
<collectingDate id="4A25892CFFC9FF9E05FDFDEF11F98E5B" box="[738,964,537,561]" pageId="20" pageNumber="1557" value="2005-09-10">10 September 2005</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E04CCFDEF16AE8E5B" box="[979,1171,537,561]" pageId="20" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E0614FDCE13138E3B" box="[267,302,568,593]" pageId="20" pageNumber="1557" type="male">2♂</specimenCount>
,
<specimenCount id="38D99D8DFFC9FF9E0620FDCE13678E3B" box="[319,346,568,593]" pageId="20" pageNumber="1557" type="female">1♀</specimenCount>
, (
<collectionCode id="48CECEC1FFC9FF9E066CFDCE13F28E3A" box="[371,463,568,592]" country="USA" name="University of Texas" pageId="20" pageNumber="1557">TNHM</collectionCode>
);
</materialsCitation>
<materialsCitation id="9EB75C59FFC9FF9E06FAFDCF10C88E1A" ID-GBIF-Occurrence="3757642306" collectingDate="1999-10-09" collectingMethod="litter sample" collectionCode="THNM" collectorName="P. Dankittipakul" county="Chomthong District" elevation="1650" location="evergreen hill forest" municipality="Doi Inthanon" pageId="20" pageNumber="1557" specimenCode="THNM, PDC 5748" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<location id="2B0000DFFFC9FF9E06FAFDCF10F48E3A" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E06FAFDCF10F48E3A" box="[485,713,568,593]" county="Chomthong District" municipality="Doi Inthanon" name="evergreen hill forest" pageId="20" pageNumber="1557" stateProvince="Chiang Mai">evergreen hill forest</location>
,
<quantity id="E927FBE1FFC9FF9E05C8FDCE11128E3A" box="[727,815,568,592]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="20" pageNumber="1557" unit="m" value="1650.0">
<elevation id="A5F2B137FFC9FF9E05C8FDCE11128E3A" box="[727,815,568,592]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="20" pageNumber="1557" unit="m" value="1650.0">1650 m</elevation>
</quantity>
,
<collectingMethod id="F79E2E13FFC9FF9E0423FDCE11F08E3A" box="[828,973,568,592]" pageId="20" pageNumber="1557">litter sample</collectingMethod>
,
<date id="5A6170C4FFC9FF9E04C5FDCE16B28E3A" box="[986,1167,568,592]" pageId="20" pageNumber="1557" value="1999-10-09">
<collectingDate id="4A25892CFFC9FF9E04C5FDCE16B28E3A" box="[986,1167,568,592]" pageId="20" pageNumber="1557" value="1999-10-09">9 October 1999</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E07CCFDAF13AC8E1A" box="[211,401,600,624]" pageId="20" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E06CEFDAE13CE8E1B" box="[465,499,600,625]" pageId="20" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode id="48CECEC1FFC9FF9E051AFDAE105D8E1A" box="[517,608,600,624]" pageId="20" pageNumber="1557">THNM</collectionCode>
, PDC 5748).
</materialsCitation>
<materialsCitation id="9EB75C59FFC9FF9E05E1FDAE13248EDA" ID-GBIF-Occurrence="3757642329" collectedFrom="litter sample" collectingDate="2000-01-15" collectionCode="THNM" collectorName="P. Dankittipakul" county="Chomthong District" elevation="1750" location="Doi Inthanon" municipality="Doi Inthanon" pageId="20" pageNumber="1557" specimenCode="THNM, PDC 5749" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<location id="2B0000DFFFC9FF9E05E1FDAE11A28E1A" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E05E1FDAE11A28E1A" box="[766,927,600,624]" county="Chomthong District" municipality="Doi Inthanon" name="Doi Inthanon" pageId="20" pageNumber="1557" stateProvince="Chiang Mai">Doi Inthanon</location>
,
<location id="2B0000DFFFC9FF9E04B3FDAF16B28E1A" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E04B3FDAF16B28E1A" box="[940,1167,600,625]" county="Chomthong District" municipality="Doi Inthanon" name="evergreen hill forest" pageId="20" pageNumber="1557" stateProvince="Chiang Mai">evergreen hill forest</location>
,
<quantity id="E927FBE1FFC9FF9E07CCFD8E13108EFA" box="[211,301,632,656]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="20" pageNumber="1557" unit="m" value="1750.0">
<elevation id="A5F2B137FFC9FF9E07CCFD8E13108EFA" box="[211,301,632,656]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="20" pageNumber="1557" unit="m" value="1750.0">1750 m</elevation>
</quantity>
,
<collectedFrom id="9DC315F9FFC9FF9E0622FD8E13ED8EFA" box="[317,464,632,656]" pageId="20" pageNumber="1557">litter sample</collectedFrom>
,
<date id="5A6170C4FFC9FF9E06C0FD8E109F8EFA" box="[479,674,632,657]" pageId="20" pageNumber="1557" value="2000-01-15">
<collectingDate id="4A25892CFFC9FF9E06C0FD8E109F8EFA" box="[479,674,632,657]" pageId="20" pageNumber="1557" value="2000-01-15">15 January 2000</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E05ADFD8F114F8EFA" box="[690,882,632,656]" pageId="20" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E04A7FD8E11E68EFB" box="[952,987,632,657]" pageId="20" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode id="48CECEC1FFC9FF9E04F0FD8E16778EFA" box="[1007,1098,632,656]" pageId="20" pageNumber="1557">THNM</collectionCode>
, PDC 5749).
</materialsCitation>
<materialsCitation id="9EB75C59FFC9FF9E063BFD6E11058EBA" ID-GBIF-Occurrence="3757642304" collectingDate="1999-10-09" collectionCode="THNM, PDC" collectorName="P. Dankittipakul" county="Chomthong District" elevation="1500" location="Doi Inthanon" municipality="Doi Inthanon" pageId="20" pageNumber="1557" specimenCode="THNM, PDC 5750" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<collectingMunicipality id="CE04CC7EFFC9FF9E063BFD6E13F58EDA" box="[292,456,664,688]" pageId="20" pageNumber="1557">Doi Inthanon</collectingMunicipality>
, evergreen hill forest,
<quantity id="E927FBE1FFC9FF9E05D2FD6E111A8EDA" box="[717,807,664,688]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="1557" unit="m" value="1500.0">
<elevation id="A5F2B137FFC9FF9E05D2FD6E111A8EDA" box="[717,807,664,688]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="1557" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, litter sample,
<date id="5A6170C4FFC9FF9E04C8FD6E16B28EDA" box="[983,1167,664,688]" pageId="20" pageNumber="1557" value="1999-10-09">
<collectingDate id="4A25892CFFC9FF9E04C8FD6E16B28EDA" box="[983,1167,664,688]" pageId="20" pageNumber="1557" value="1999-10-09">9 October 1999</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E07CCFD4F13A98EBA" box="[211,404,696,720]" pageId="20" pageNumber="1557">P. Dankittipakul</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E06C4FD4E13C08EBA" box="[475,509,696,720]" pageId="20" pageNumber="1557" type="male">1♂</specimenCount>
,
<specimenCount id="38D99D8DFFC9FF9E050EFD4E10118EBA" box="[529,556,696,720]" pageId="20" pageNumber="1557" type="female">2♀</specimenCount>
(
<collectionCode id="48CECEC1FFC9FF9E055EFD4E10A18EBA" box="[577,668,696,720]" pageId="20" pageNumber="1557">THNM</collectionCode>
,
<collectionCode id="48CECEC1FFC9FF9E05B2FD4E10DB8EBA" box="[685,742,696,720]" pageId="20" pageNumber="1557">PDC</collectionCode>
5750).
</materialsCitation>
<materialsCitation id="9EB75C59FFC9FF9E045CFD4E13808F7A" ID-GBIF-Occurrence="3757642302" collectingDate="1985-11-06" collectionCode="MHNG" collectorName="P. J. Schwendinger" county="Chomthong District" elevation="1250" location="Doi Inthanon" municipality="Doi Inthanon" pageId="20" pageNumber="1557" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<location id="2B0000DFFFC9FF9E045CFD4E11D58EBA" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E045CFD4E11D58EBA" box="[835,1000,696,720]" county="Chomthong District" municipality="Doi Inthanon" name="Doi Inthanon" pageId="20" pageNumber="1557" stateProvince="Chiang Mai">Doi Inthanon</location>
,
<location id="2B0000DFFFC9FF9E04E7FD4F13288E9A" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E04E7FD4F13288E9A" county="Chomthong District" municipality="Doi Inthanon" name="evergreen hill forest" pageId="20" pageNumber="1557" stateProvince="Chiang Mai">evergreen hill forest</location>
,
<quantity id="E927FBE1FFC9FF9E0639FD2E13BC8E9A" box="[294,385,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="20" pageNumber="1557" unit="m" value="1250.0">
<elevation id="A5F2B137FFC9FF9E0639FD2E13BC8E9A" box="[294,385,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="20" pageNumber="1557" unit="m" value="1250.0">1250 m</elevation>
</quantity>
,
<date id="5A6170C4FFC9FF9E068EFD2E105B8E9A" box="[401,614,728,752]" pageId="20" pageNumber="1557" value="1985-11-06">
<collectingDate id="4A25892CFFC9FF9E068EFD2E105B8E9A" box="[401,614,728,752]" pageId="20" pageNumber="1557" value="1985-11-06">6 November 1985</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E0568FD2F11788E9A" box="[631,837,728,752]" pageId="20" pageNumber="1557">P.J. Schwendinger</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E0492FD2E11928E9A" box="[909,943,728,752]" pageId="20" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode id="48CECEC1FFC9FF9E04DBFD2E161F8E9A" box="[964,1058,728,752]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="20" pageNumber="1557">MHNG</collectionCode>
, misidentified as
<taxonomicName id="E9DF2D87FFC9FF9E0623FD0E13908F7A" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[316,429,760,784]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="erawan">
<emphasis id="1CAB8A16FFC9FF9E0623FD0E13908F7A" box="[316,429,760,784]" italics="true" pageId="20" pageNumber="1557">S. erawan</emphasis>
</taxonomicName>
).
</materialsCitation>
<materialsCitation id="9EB75C59FFC9FF9E06D6FD0E13A48F25" ID-GBIF-Occurrence="3757642333" collectingDate="1987-06-18" collectionCode="MHNG" collectorName="P. J. Schwendinger" county="Chiang Mai" elevation="1200" location="Doi Suthep-Pui NP" municipality="Doi Suthep" pageId="20" pageNumber="1557" specimenCount="1" specimenCount-female="1" stateProvince="Chiang Mai Province" typeStatus="paratype">
<collectingRegion id="EC1B98E6FFC9FF9E06D6FD0E10F98F7A" box="[457,708,760,784]" country="Thailand" name="Chiang Mai" pageId="20" pageNumber="1557">Chiang Mai Province</collectingRegion>
and District,
<location id="2B0000DFFFC9FF9E046FFD0E166A8F7A" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E046FFD0E166A8F7A" box="[880,1111,760,784]" county="Chiang Mai" municipality="Doi Suthep" name="Doi Suthep-Pui NP" pageId="20" pageNumber="1557" stateProvince="Chiang Mai Province">Doi Suthep-Pui NP</location>
,
<location id="2B0000DFFFC9FF9E0377FD0E13198F5A" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E0377FD0E13198F5A" county="Chiang Mai" municipality="Doi Suthep" name="Doi Suthep" pageId="20" pageNumber="1557" stateProvince="Chiang Mai Province">Doi Suthep</location>
,
<quantity id="E927FBE1FFC9FF9E062DFCEE13B48F45" box="[306,393,792,816]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="1557" unit="m" value="1200.0">
<elevation id="A5F2B137FFC9FF9E062DFCEE13B48F45" box="[306,393,792,816]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="1557" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="5A6170C4FFC9FF9E0688FCEE100D8F5A" box="[407,560,792,816]" pageId="20" pageNumber="1557" value="1987-06-18">
<collectingDate id="4A25892CFFC9FF9E0688FCEE100D8F5A" box="[407,560,792,816]" pageId="20" pageNumber="1557" value="1987-06-18">18 June 1987</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E0521FCEE11358F5A" box="[574,776,792,816]" pageId="20" pageNumber="1557">P.J. Schwendinger</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E0456FCEE11598F5A" box="[841,868,792,816]" pageId="20" pageNumber="1557" type="female">1♀</specimenCount>
(
<collectionCode id="48CECEC1FFC9FF9E0469FCEE11E98F5A" box="[886,980,792,816]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="20" pageNumber="1557">MHNG</collectionCode>
, misidentified as
<taxonomicName id="E9DF2D87FFC9FF9E07CCFCCE13B48F3A" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[211,393,824,848]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis id="1CAB8A16FFC9FF9E07CCFCCE13B48F3A" box="[211,393,824,848]" italics="true" pageId="20" pageNumber="1557">S. schwendingeri</emphasis>
</taxonomicName>
).
</materialsCitation>
<materialsCitation id="9EB75C59FFC9FF9E0680FCC111DA8F05" ID-GBIF-Occurrence="3757642325" collectingDate="1986-11-31" collectingDateMax="1987-01-02" collectingDateMin="1986-11-31" collectingMethod="pitfall trap" collectionCode="RMNH" collectorName="P. J. Schwendinger" county="Chiang Mai" elevation="960" httpUri="https://data.biodiversitydata.nl/naturalis/specimen/RMNH.ARA.15205" location="Doi Suthep" pageId="20" pageNumber="1557" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<location id="2B0000DFFFC9FF9E0680FCC1101F8F25" LSID="urn:lsid:plazi:treatment:A676E712FFCEFF9C06FAF9841095891A:2B0000DFFFC9FF9E0680FCC1101F8F25" box="[415,546,823,847]" county="Chiang Mai" municipality="Doi Inthanon" name="Doi Suthep" pageId="20" pageNumber="1557" stateProvince="Chiang Mai">Doi Suthep</location>
,
<quantity id="E927FBE1FFC9FF9E0532FCC110498F25" box="[557,628,823,847]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="20" pageNumber="1557" unit="m" value="960.0">
<elevation id="A5F2B137FFC9FF9E0532FCC110498F25" box="[557,628,823,847]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="20" pageNumber="1557" unit="m" value="960.0">960 m</elevation>
</quantity>
,
<collectingMethod id="F79E2E13FFC9FF9E0560FCC110C48F25" box="[639,761,823,847]" pageId="20" pageNumber="1557">pitfall trap</collectingMethod>
,
<date id="5A6170C4FFC9FF9E041BFCC111EB8F25" box="[772,982,823,847]" pageId="20" pageNumber="1557" value="1986-11-31">
<collectingDate id="4A25892CFFC9FF9E041BFCC111EB8F25" box="[772,982,823,847]" pageId="20" pageNumber="1557" value="1986-11-31">31 November 1986</collectingDate>
</date>
<date id="5A6170C4FFC9FF9E04FAFCC116B28F25" box="[997,1167,823,848]" pageId="20" pageNumber="1557" value="1987-01-02">
<collectingDate id="4A25892CFFC9FF9E04FAFCC116B28F25" box="[997,1167,823,848]" pageId="20" pageNumber="1557" value="1987-01-02">2 January 1987</collectingDate>
</date>
,
<collectorName id="832A33D2FFC9FF9E07CCFCAE13A68F05" box="[211,411,855,879]" pageId="20" pageNumber="1557">P.J. Schwendinger</collectorName>
leg.,
<specimenCount id="38D99D8DFFC9FF9E06C8FCA113C48F1A" box="[471,505,855,880]" pageId="20" pageNumber="1557" type="male">1♂</specimenCount>
(
<collectionCode id="48CECEC1FFC9FF9E0517FCA1105B8F05" box="[520,614,855,879]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="20" pageNumber="1557">RMNH</collectionCode>
, misidentified as
<taxonomicName id="E9DF2D87FFC9FF9E0439FCAE11E18F1A" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[806,988,856,880]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri">
<emphasis id="1CAB8A16FFC9FF9E0439FCAE11E18F1A" box="[806,988,856,880]" italics="true" pageId="20" pageNumber="1557">S. schwendingeri</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC9FF9E07CCFC55107A888B" pageId="20" pageNumber="1557" type="diagnosis">
<paragraph id="2E605604FFC9FF9E07CCFC5513028FD0" blockId="20.[211,1172,931,1249]" box="[211,319,931,954]" pageId="20" pageNumber="1557">
<heading id="7528E168FFC9FF9E07CCFC5513028FD0" box="[211,319,931,954]" fontSize="10" level="3" pageId="20" pageNumber="1557" reason="5">
<emphasis id="1CAB8A16FFC9FF9E07CCFC5513028FD0" box="[211,319,931,954]" italics="true" pageId="20" pageNumber="1557">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC9FF9E07CCFC3C107A888B" blockId="20.[211,1172,931,1249]" pageId="20" pageNumber="1557">
Males of
<taxonomicName id="E9DF2D87FFC9FF9E065DFC3D13858F88" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[322,440,970,994]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis id="1CAB8A16FFC9FF9E065DFC3D13858F88" box="[322,440,970,994]" italics="true" pageId="20" pageNumber="1557">J. lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC9FF9E06DEFC3D10258F88" box="[449,536,971,994]" pageId="20" pageNumber="1557" rank="species">sp. nov.</taxonomicNameLabel>
are similar to those of
<taxonomicName id="E9DF2D87FFC9FF9E0434FC3D11E38F88" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" box="[811,990,970,994]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis id="1CAB8A16FFC9FF9E0434FC3D11E38F88" box="[811,990,970,994]" italics="true" pageId="20" pageNumber="1557">J. schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC9FF9E04F7FC3C16588F88" box="[1000,1125,970,994]" pageId="20" pageNumber="1557" rank="species">comb. nov.</taxonomicNameLabel>
but differ by having a crescent-shaped conductor that is distinctly broader and strongly curved backward (
<figureCitation id="B6E44A81FFC9FF9E06AFFBFF1005884B" box="[432,568,1033,1057]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (AC); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="20" pageNumber="1557">Figures 6A</figureCitation>
,
<figureCitation id="B6E44A81FFC9FF9E0555FBFF10AB884B" box="[586,662,1033,1057]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="20" pageNumber="1557">9D, E</figureCitation>
), a tegulum that is greatly expanded, and the distal RTA of the male palp that is digitiform (
<figureCitation id="B6E44A81FFC9FF9E042EFBDF1196882B" box="[817,939,1065,1089]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (AC); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="20" pageNumber="1557">Figure 6C</figureCitation>
). In comparison, in
<taxonomicName id="E9DF2D87FFC9FF9E07CCFBBC13B8880B" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Deeleman-Reinhold" baseAuthorityYear="2001" box="[211,389,1097,1121]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="1557" phylum="Arthropoda" rank="species" species="schwendingeri" status="comb. nov.">
<emphasis id="1CAB8A16FFC9FF9E07CCFBBC13B8880B" box="[211,389,1097,1121]" italics="true" pageId="20" pageNumber="1557">J. schwendingeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC9FF9E0692FBBF1035880B" box="[397,520,1097,1121]" pageId="20" pageNumber="1557" rank="species">comb. nov.</taxonomicNameLabel>
the conductor is narrower and its sharply pointed apex is abruptly bent (
<figureCitation id="B6E44A81FFC9FF9E0667FB9F103088EB" box="[376,525,1129,1153]" captionStart="Figure 4" captionStartId="8.[211,278,1428,1450]" captionTargetBox="[213,1170,168,1394]" captionTargetId="figure-5@8.[213,1170,168,1394]" captionTargetPageId="8" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (DF); J. mihun, holotype (GI)." figureDoi="http://doi.org/10.5281/zenodo.6517648" httpUri="https://zenodo.org/record/6517648/files/figure.png" pageId="20" pageNumber="1557">Figure 4A, C</figureCitation>
), the tegulum and the distal RTA are significantly smaller. Females can be distinguished by the rather simple internal genitalia, the ID of which consists of circular proximal ducts, the more convoluted ID, and the poorly defined spermathecae (
<figureCitation id="B6E44A81FFC9FF9E0664FB3F13CD888B" box="[379,496,1225,1249]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (AG). Jacaena mihun (AD); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="20" pageNumber="1557">Figure 8G</figureCitation>
cf. 8F).
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC9FF9E07CCFAE811518917" pageId="20" pageNumber="1557" type="etymology">
<paragraph id="2E605604FFC9FF9E07CCFAE81376895C" blockId="20.[211,1171,1310,1405]" box="[211,331,1310,1334]" pageId="20" pageNumber="1557">
<heading id="7528E168FFC9FF9E07CCFAE81376895C" box="[211,331,1310,1334]" fontSize="10" level="3" pageId="20" pageNumber="1557" reason="5">
<emphasis id="1CAB8A16FFC9FF9E07CCFAE81376895C" box="[211,331,1310,1334]" italics="true" pageId="20" pageNumber="1557">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC9FF9E07CCFAB011518917" blockId="20.[211,1171,1310,1405]" pageId="20" pageNumber="1557">
The specific epithet is derived from Latin adjective (
<emphasis id="1CAB8A16FFC9FF9E045EFAB011988934" box="[833,933,1350,1374]" italics="true" pageId="20" pageNumber="1557">lunulatus</emphasis>
= crescent-shaped), referring to the lamina-shaped conductor on the male palp.
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFC9FF9C07CCFA4A10028856" lastPageId="22" lastPageNumber="1559" pageId="20" pageNumber="1557" type="description">
<paragraph id="2E605604FFC9FF9E07CCFA4A136F89B9" blockId="20.[211,1172,1468,1691]" box="[211,338,1468,1491]" pageId="20" pageNumber="1557">
<heading id="7528E168FFC9FF9E07CCFA4A136F89B9" box="[211,338,1468,1491]" fontSize="10" level="3" pageId="20" pageNumber="1557" reason="5">
<emphasis id="1CAB8A16FFC9FF9E07CCFA4A136F89B9" box="[211,338,1468,1491]" italics="true" pageId="20" pageNumber="1557">Description</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFC9FF9F07CCFA1511668CB5" blockId="20.[211,1172,1468,1691]" lastBlockId="21.[143,1104,167,542]" lastPageId="21" lastPageNumber="1558" pageId="20" pageNumber="1557">
<emphasis id="1CAB8A16FFC9FF9E07CCFA1513308991" box="[211,269,1507,1531]" italics="true" pageId="20" pageNumber="1557">Male</emphasis>
(
<typeStatus id="F164E8A6FFC9FF9E0603FA1413B88990" box="[284,389,1506,1530]" pageId="20" pageNumber="1557" type="holotype">holotype</typeStatus>
). Total length 6.6. Carapace 3.2 long, 2.5 wide. Opisthosoma 3.4 long, 2.0 wide. Eye sizes and interdistances: AME 0.15; ALE 0.13; PME 0.10; PLE 0.13; AMEAME 0.06; AMEALE 0.05; PMEPME 0.15; PMEPLE 0.16. MOQ 0.33 long; front width 0.33; back width 0.35.
<collectorName id="832A33D2FFC9FF9E05DEF9B510D18A31" box="[705,748,1603,1627]" pageId="20" pageNumber="1557">Leg</collectorName>
formula 4123.
<collectorName id="832A33D2FFC9FF9E048EF9B4167A8A30" box="[913,1095,1602,1627]" pageId="20" pageNumber="1557">Spination. Leg I</collectorName>
: femur 11pl; tibia 8pl 7rl; metatarsus 4pl 4rl.
<collectorName id="832A33D2FFC9FF9E059BF99510F18A10" box="[644,716,1635,1659]" pageId="20" pageNumber="1557">Leg II</collectorName>
: tibia 7pl 6rl; metatarsus 4pl 4rl.
<collectorName id="832A33D2FFC9FF9E0323F995165A8A11" box="[1084,1127,1635,1659]" pageId="20" pageNumber="1557">Leg</collectorName>
III: tibia 11v 1pl; metatarsus 11v 1pl.
<collectorName id="832A33D2FFC9FF9E0570F97510FC8AF0" box="[623,705,1667,1691]" pageId="20" pageNumber="1557">Leg IV</collectorName>
: tibia 122v 11rl; metatarsus 1111v 1pl 1rl.
<collectorName id="832A33D2FFC8FF9F07F6FF5E13298CAA" box="[233,276,168,192]" pageId="21" pageNumber="1558">Leg</collectorName>
measurements:
<collectorName id="832A33D2FFC8FF9F06D4FF5E10358CD5" box="[459,520,168,192]" pageId="21" pageNumber="1558">Leg I</collectorName>
8.1 (2.3, 0.9, 2.2, 1.8, 0.9); II 7.5 (2.1, 0.9, 1.9, 1.7, 0.9); III 6.7 (1.9, 0.9, 1.4, 1.6, 0.9); IV 9.4 (2.6, 1.0, 2.1, 2.5, 1.2).
</paragraph>
<paragraph id="2E605604FFC8FF9F07A5FF1111428DB4" blockId="21.[143,1104,167,542]" pageId="21" pageNumber="1558">
Colouration and pattern (
<figureCitation id="B6E44A81FFC8FF9F06C2FF11106C8C95" box="[477,593,231,255]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (AH). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="21" pageNumber="1558">Figure 1B</figureCitation>
): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown. Coxae, trochanters and femora dark brown; patellae and tibiae brown, distal part of tibiae orange; metatarsi III brown, IIIIV orange; tarsi orange. Opisthosoma ovoid; dorsum of opisthosoma dark grey, with pattern consisting of one pair of pale patches, followed by four transverse chevrons and pale area above spinnerets. Dorsal scutum occupying approximately one-third of opisthosoma length, posterior margin rounded, with oval extensions fusing with dorsal muscle apodemes.
</paragraph>
<paragraph id="2E605604FFC8FF9C07A5FE1011618D55" blockId="21.[143,1104,167,542]" lastBlockId="22.[211,1172,167,319]" lastPageId="22" lastPageNumber="1559" pageId="21" pageNumber="1558">
Palp (
<figureCitation id="B6E44A81FFC8FF9F07E0FE1013988D94" box="[255,421,486,510]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (AC); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="21" pageNumber="1558">Figures 6AC</figureCitation>
,
<figureCitation id="B6E44A81FFC8FF9F06ACFE1013C68D94" box="[435,507,486,510]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="21" pageNumber="1558">9D, E</figureCitation>
,
<figureCitation id="B6E44A81FFC8FF9F0514FE1010638D94" box="[523,606,486,510]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="21" pageNumber="1558">12AC</figureCitation>
): Proximal RTA triangular in ventral view (
<figureCitation id="B6E44A81FFC8FF9F0788FDF013278E74" box="[151,282,518,542]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="21" pageNumber="1558">Figure 12A</figureCitation>
); apical surface partially membranous, slightly excavated, forming shallow concavity (
<figureCitation id="B6E44A81FFCBFF9C064FFF5113CD8CD5" box="[336,496,167,191]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="22" pageNumber="1559">Figure 9D, E</figureCitation>
). Distal RTA digitiform, its apex blunt (
<figureCitation id="B6E44A81FFCBFF9C04D4FF5116738CD5" box="[971,1102,167,191]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (AC); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="22" pageNumber="1559">Figures 6C</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C0342FF5112D48CB5" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="22" pageNumber="1559">12A, C</figureCitation>
). Conductor (
<figureCitation id="B6E44A81FFCBFF9C068FFF3110288CB5" box="[400,533,199,223]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (AC); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="22" pageNumber="1559">Figures 6A</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C0539FF31104D8CB5" box="[550,624,199,223]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="22" pageNumber="1559">9D, E</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C059FFF3110ED8CB5" box="[640,720,199,223]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="22" pageNumber="1559">12AC</figureCitation>
) crescent-shaped, enlarged and broad, semi-transparent, with heavily pigmented outer margin, sharply pointed apex curving backwards. Tegular apophysis hyaline, lanceolate, originating behind base of conductor, aligning with tip of embolus (
<figureCitation id="B6E44A81FFCBFF9C0582FED111208D55" box="[669,797,295,319]" captionStart="Figure 6" captionStartId="12.[211,278,1142,1164]" captionTargetBox="[213,1170,171,1109]" captionTargetId="figure-5@12.[213,1170,167,1109]" captionTargetPageId="12" captionText="Figure 6. Jacaena spp. Male palps, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Internal copulatory organ, anterior part, dorsal (D). Ditto, showing proximal chamber and secretory ampulla (E). Internal copulatory organ (middle portion of insemination duct removed on right spermathecae, showing connection between copulatory orifice and insemination duct) (F). Jacaena lunulata sp. nov., holotype (AC); J. peculiaris sp. nov., paratype (D); J. punctata sp. nov., paratype (E); J. mihun (F). AM, secretory ampulla. CH, semi-transparent chamber. CO, copulatory orifice. F, fertilisation duct." figureDoi="http://doi.org/10.5281/zenodo.6517660" httpUri="https://zenodo.org/record/6517660/files/figure.png" pageId="22" pageNumber="1559">Figures 6A</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C0436FED111728D55" box="[809,847,295,319]" captionStart="Figure 9" captionStartId="17.[143,210,924,946]" captionTargetBox="[146,1102,170,891]" captionTargetId="figure-5@17.[145,1102,169,891]" captionTargetPageId="17" captionText="Figure 9. Jacaena spp. Distal part of male palp, ventral, showing conductor, tip of embolus and tegular apophysis (A, D). Ditto, retrolateral (B). Ditto, prolateral (C, E). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., holotype (D, E). C, conductor. E, embolus. TA, tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6517666" httpUri="https://zenodo.org/record/6517666/files/figure.png" pageId="22" pageNumber="1559">9D</figureCitation>
).
</paragraph>
<caption id="7AA0068CFFC8FF9F0790F9FC115C8AFF" ID-DOI="http://doi.org/10.5281/zenodo.6517674" ID-Zenodo-Dep="6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="21" pageNumber="1558" startId="21.[143,210,1546,1568]" targetBox="[145,1102,609,1513]" targetPageId="21">
<paragraph id="2E605604FFC8FF9F0790F9FC115C8AFF" blockId="21.[143,1104,1546,1685]" pageId="21" pageNumber="1558">
Figure 12.
<taxonomicName id="E9DF2D87FFC8FF9F061BF9FD13968A4A" authority="Dankittipakul &amp; Tavano &amp; Singtripop, 2013" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" box="[260,427,1546,1568]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="1558" phylum="Arthropoda" rank="species" species="lunulata" status="sp. nov.">
<emphasis id="1CAB8A16FFC8FF9F061BF9FD13968A4A" box="[260,427,1546,1568]" italics="true" pageId="21" pageNumber="1558">Jacaena lunulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFC8FF9F06ADF9FD103F8A4A" box="[434,514,1547,1568]" pageId="21" pageNumber="1558" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca.
</paragraph>
</caption>
<paragraph id="2E605604FFCBFF9C07CCFE9113388E14" blockId="22.[211,1172,359,1084]" pageId="22" pageNumber="1559">
<emphasis id="1CAB8A16FFCBFF9C07CCFE91131F8D15" box="[211,290,359,383]" italics="true" pageId="22" pageNumber="1559">Female</emphasis>
(
<typeStatus id="F164E8A6FFCBFF9C062DFE9113A58D15" box="[306,408,359,383]" pageId="22" pageNumber="1559" type="paratype">paratype</typeStatus>
, NHML, PDC 5747). Total length 8.1. Carapace 3.3 long, 2.5 wide. Opisthosoma 4.5 long, 2.7 wide. Eye sizes and interdistances: AME 0.11; ALE 0.13; PME 0.08; PLE 0.13; AMEAME 0.08; AMEALE 0.05; PMEPME 0.15; PME PLE 0.16. MOQ 0.38 long; front width 0.31; back width 0.35. Leg formula 1423. Spination. Leg I: femur 11pl; tibia 8pl 7rl; metatarsus 4pl 4rl. Leg II: tibia 7pl 6rl; metatarsus 4pl 4rl. Leg III: tibia 21v 1pl; metatarsus 11v 1pl. Leg IV: tibia 112v 1rl; metatarsus 1111v 1pl 1rl. Leg measurements: Leg I 8.6 (2.3, 1.0, 2.3, 2.0, 1.0); II 7.9 (2.2, 1.0, 1.9, 1.8, 1.0); III 6.6 (1.8, 2.8, 1.5, 1.5, 1.0); IV 8.3 (2.3, 0.8, 2.0, 2.2, 1.0).
</paragraph>
<paragraph id="2E605604FFCBFF9C07E1FD7013E08F37" blockId="22.[211,1172,359,1084]" pageId="22" pageNumber="1559">
Colouration and pattern (
<figureCitation id="B6E44A81FFCBFF9C053EFD7010AB8EF4" box="[545,662,646,670]" captionStart="Figure 2" captionStartId="6.[211,278,1132,1154]" captionTargetBox="[213,1170,168,1099]" captionTargetId="figure-5@6.[213,1170,168,1099]" captionTargetPageId="6" captionText="Figure 2. Females of Jacaena spp., habitus, dorsal (AH). Jacaena schwendingeri comb. nov., paratype (A); J. lunulata sp. nov., paratypes (B, C); J. angoonae sp. nov., holotype (D); J. punctata sp. nov., paratypes (E,F); J. mihun (G); J. peculiaris sp. nov., holotype (H)." figureDoi="http://doi.org/10.5281/zenodo.6517642" httpUri="https://zenodo.org/record/6517642/files/figure.png" pageId="22" pageNumber="1559">Figure 2B</figureCitation>
): Carapace dark chestnut-brown; integument coarsely granulated. Chelicerae, sternum, labium and gnathocoxae dark reddishbrown. Legs III brown, femora slightly darker than other segments, tibiae orangebrown distally; legs IIIIV orange-brown except femora dark brown. Opisthosoma elongate-ovoid; dorsum grey, with pair of pale patches connected anteriorly, followed by four chevrons and pale area above spinnerets (first pair very faint and narrowed, disconnected medially).
</paragraph>
<paragraph id="2E605604FFCBFF9C07E1FC9310028856" blockId="22.[211,1172,359,1084]" pageId="22" pageNumber="1559">
Copulatory organ (
<figureCitation id="B6E44A81FFCBFF9C06C9FC9310688F17" box="[470,597,869,893]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (AF). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="22" pageNumber="1559">Figures 7C</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C057EFC9310BE8F17" box="[609,643,869,893]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (AG). Jacaena mihun (AD); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="22" pageNumber="1559">8F</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C0590FC9310DC8F17" box="[655,737,869,893]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="22" pageNumber="1559">12D, E</figureCitation>
): Copulatory orifices situated medially on heavily sclerotized epigynal region (
<figureCitation id="B6E44A81FFCBFF9C0592FC7311308FF7" box="[653,781,901,925]" captionStart="Figure 7" captionStartId="13.[143,210,790,812]" captionTargetBox="[145,1102,169,757]" captionTargetId="figure-5@13.[145,1102,169,758]" captionTargetPageId="13" captionText="Figure 7. Jacaena spp. Epigynal region, ventral (AF). Jacaena schwendingeri comb. nov., paratype (A); J. punctata sp. nov., paratype (B); J. lunulata sp. nov., paratype (C); J. mihun (D); J. angoonae sp. nov., holotype (E); J. peculiaris sp. nov., holotype (F)." figureDoi="http://doi.org/10.5281/zenodo.6517662" httpUri="https://zenodo.org/record/6517662/files/figure.png" pageId="22" pageNumber="1559">Figures 7C</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C0404FC7311728FF7" box="[795,847,901,925]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="22" pageNumber="1559">12D</figureCitation>
). Proximal half of insemination ducts forming two loops encircling copulatory ducts (
<figureCitation id="B6E44A81FFCBFF9C0462FC52163E8FD6" box="[893,1027,932,956]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (AG). Jacaena mihun (AD); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="22" pageNumber="1559">Figures 8G</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C030DFC52167C8FD6" box="[1042,1089,932,956]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="22" pageNumber="1559">12E</figureCitation>
); inner loop distinctly enlarged, filled with dark substance (
<figureCitation id="B6E44A81FFCBFF9C0436FC3211998FB6" box="[809,932,964,988]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (AG). Jacaena mihun (AD); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="22" pageNumber="1559">Figure 8G</figureCitation>
); distal half of ducts ascending anteriorly then traversing and descending to connect with poorly defined spermathecae (
<figureCitation id="B6E44A81FFCBFF9C069DFBF210368876" box="[386,523,1028,1052]" captionStart="Figure 8" captionStartId="14.[211,278,950,972]" captionTargetBox="[213,1170,169,916]" captionTargetId="figure-5@14.[213,1170,169,917]" captionTargetPageId="14" captionText="Figure 8. Jacaena spp. Internal copulatory organ, dorsal (AG). Jacaena mihun (AD); J. angoonae sp. nov., holotype (E); J. schwendingeri comb. nov., paratype (F); J. lunulata sp. nov., paratype (G)." figureDoi="http://doi.org/10.5281/zenodo.6517664" httpUri="https://zenodo.org/record/6517664/files/figure.png" pageId="22" pageNumber="1559">Figures 8G</figureCitation>
,
<figureCitation id="B6E44A81FFCBFF9C0501FBF210708876" box="[542,589,1028,1052]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="22" pageNumber="1559">12E</figureCitation>
); glandular pores situated at anterior margin of spermathecae (
<figureCitation id="B6E44A81FFCBFF9C0664FBD213C18856" box="[379,508,1060,1084]" captionStart="Figure 12" captionStartId="21.[143,210,1546,1568]" captionTargetBox="[145,1102,609,1513]" captionTargetId="figure-146@21.[145,1102,605,1513]" captionTargetPageId="21" captionText="Figure 12. Jacaena lunulata sp. nov., holotype, male (AC) and paratype, female (D, E). Male palp, ventral (A). Ditto, prolateral (B). Ditto, retrolateral (C). Epigyne, ventral (D). Internal copulatory organ, dorsal (E). AM, secretory ampulla; dRTA, distal process of retrolateral tibial apophysis; EB, embolic base; GP, glandular pore on spermathecae; ID, insemination duct. pRTA, proximal process of retrolateral tibia apophysis; S, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6517674" httpUri="https://zenodo.org/record/6517674/files/figure.png" pageId="22" pageNumber="1559">Figure 12E</figureCitation>
, GF).
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFCBFF9C07CCFB711051888C" pageId="22" pageNumber="1559" type="biology_ecology">
<paragraph id="2E605604FFCBFF9C07CCFB71134188F5" blockId="22.[211,1172,1159,1254]" box="[211,380,1159,1183]" pageId="22" pageNumber="1559">
<heading id="7528E168FFCBFF9C07CCFB71134188F5" box="[211,380,1159,1183]" fontSize="10" level="3" pageId="22" pageNumber="1559" reason="5">
<emphasis id="1CAB8A16FFCBFF9C07CCFB71134188F5" box="[211,380,1159,1183]" italics="true" pageId="22" pageNumber="1559">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFCBFF9C07CCFB591051888C" blockId="22.[211,1172,1159,1254]" pageId="22" pageNumber="1559">
Types of this new species were collected in evergreen hill forests of Doi Suthep and Doi Inthanon between 1250 and
<quantity id="E927FBE1FFCBFF9C050CFB381055888C" box="[531,616,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="22" pageNumber="1559" unit="m" value="1750.0">1750 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFCBFF9C07CCFAC71095891A" box="[211,680,1329,1392]" pageId="22" pageNumber="1559" type="distribution">
<paragraph id="2E605604FFCBFF9C07CCFAC713658923" blockId="22.[211,680,1329,1392]" box="[211,344,1329,1353]" pageId="22" pageNumber="1559">
<heading id="7528E168FFCBFF9C07CCFAC713658923" box="[211,344,1329,1353]" fontSize="10" level="3" pageId="22" pageNumber="1559" reason="5">
<emphasis id="1CAB8A16FFCBFF9C07CCFAC713658923" box="[211,344,1329,1353]" italics="true" pageId="22" pageNumber="1559">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFCBFF9C07CCFAAE1095891A" blockId="22.[211,680,1329,1392]" box="[211,680,1368,1392]" pageId="22" pageNumber="1559">
<collectingRegion id="EC1B98E6FFCBFF9C07CCFAAE13F8891A" box="[211,453,1368,1392]" country="Thailand" name="Chiang Mai" pageId="22" pageNumber="1559">Chiang Mai Province</collectingRegion>
, northern
<collectingCountry id="56C81694FFCBFF9C0525FAAE1099891A" box="[570,676,1368,1392]" name="Thailand" pageId="22" pageNumber="1559">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>