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<document id="22BB106A77C4EEC548D59CD268C40D62" ID-CLB-Dataset="55811" ID-DOI="10.1080/00222933.2012.763059" ID-GBIF-Dataset="dfc793f4-7033-4586-9849-718e7e526360" ID-ISSN="1464-5262" ID-Zenodo-Dep="556127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628879433880" checkinUser="felipe" docAuthor="Dankittipakul, Pakawin, Tavano, Maria &amp; Singtripop, Tippawan" docDate="2013" docId="A676E712FFCDFF9A06CCFDF3119089B0" docLanguage="en" docName="JNatHist.47.23-24.1539-1567.pdf" docOrigin="Journal of Natural History 47 (23 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2012.763059" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Jacaena thakek Dankittipakul &amp; Tavano &amp; Singtripop 2013, comb. nov." docType="treatment" docVersion="11" lastPageNumber="1553" masterDocId="5A4F9F6AFFDDFF8A071FFFF6123D8C6A" masterDocTitle="Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)" masterLastPageNumber="1567" masterPageNumber="1539" pageNumber="1553" updateTime="1699079059524" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="13B676B9CA03F16C4C6333B81EA06C86">Revision of the spider genus Jacaena Thorell, 1897, with descriptions of four new species from Thailand (Araneae: Corinnidae)</mods:title>
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<mods:namePart id="94366F75CE6F439383E2F505CEA48780">Dankittipakul, Pakawin</mods:namePart>
<mods:affiliation id="C5BFD5246E0EE4AF3DF28B31A9905EF7">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand; &amp; Manukau Institute of Technology, Manukau, New Zealand;</mods:affiliation>
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<mods:namePart id="E957CD574BD3E31D3923335662534111">Tavano, Maria</mods:namePart>
<mods:affiliation id="3A00779A1D621A1FB89D485EB1C30E32">Museo Civico di Storia Naturale Giacomo Doria, Genova, Italy</mods:affiliation>
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<mods:namePart id="3E2E0815D4B66175EE5D9AC1ED088D4F">Singtripop, Tippawan</mods:namePart>
<mods:affiliation id="79FAD659D8E4FD6A5DC758224AAE1C02">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai, Thailand;</mods:affiliation>
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<mods:title id="1F4D1E15D487E9FE4EC2756A2A9A0796">Journal of Natural History</mods:title>
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<treatment id="A676E712FFCDFF9A06CCFDF3119089B0" ID-DOI="http://doi.org/10.5281/zenodo.6502524" ID-GBIF-Taxon="194854860" ID-Zenodo-Dep="6502524" LSID="urn:lsid:plazi:treatment:A676E712FFCDFF9A06CCFDF3119089B0" httpUri="http://treatment.plazi.org/id/A676E712FFCDFF9A06CCFDF3119089B0" lastPageNumber="1553" pageId="16" pageNumber="1553">
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<heading id="7528E168FFCDFF9A06CCFDF311A98E76" box="[467,916,516,541]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="1553" reason="2">
<taxonomicName id="E9DF2D87FFCDFF9A06CCFDF3112E8E76" ID-CoL="3QHZL" authority="(Jager, 2007)" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Jager" baseAuthorityYear="2007" box="[467,787,516,541]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="thakek" status="comb. nov.">
<emphasis id="1CAB8A16FFCDFF9A06CCFDF310428E77" bold="true" box="[467,639,516,541]" italics="true" pageId="16" pageNumber="1553">Jacaena thakek</emphasis>
(
<bibRefCitation id="4A4E2BF5FFCDFF9A0592FDF211318E76" author="Jager P" box="[653,780,516,540]" pageId="16" pageNumber="1553" pagination="29 - 58" refId="ref11377" refString="Jager P. 2007. Spiders from Laos with descriptions of new species (Arachnida: Araneae). Acta arachn Tokyo. 56: 29 - 58." type="journal article" year="2007">Jäger, 2007</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0798376DFFCDFF9A0405FDF211A98E76" box="[794,916,516,540]" pageId="16" pageNumber="1553" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="66C5058FFFCDFF9A0503FDD211768E56" box="[540,843,548,572]" pageId="16" pageNumber="1553" type="description">
<paragraph id="2E605604FFCDFF9A0503FDD211768E56" blockId="16.[467,916,516,572]" box="[540,843,548,572]" pageId="16" pageNumber="1553">
(
<figureCitation id="B6E44A81FFCDFF9A053BFDD210A38E56" box="[548,670,548,572]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (AH). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="16" pageNumber="1553">Figures 1F</figureCitation>
,
<figureCitation id="B6E44A81FFCDFF9A05B5FDD210DA8E56" box="[682,743,548,572]" captionStart="Figure 5" captionStartId="11.[143,210,1565,1587]" captionTargetBox="[145,1102,161,1532]" captionTargetId="figure-5@11.[145,1102,161,1532]" captionTargetPageId="11" captionText="Figure 5. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena erawan comb. nov., holotype (AC); J. punctata sp. nov., holotype (DF); J. thakek comb. nov., holotype (GI)." figureDoi="http://doi.org/10.5281/zenodo.6517654" httpUri="https://zenodo.org/record/6517654/files/figure.png" pageId="16" pageNumber="1553">5GI</figureCitation>
,
<figureCitation id="B6E44A81FFCDFF9A05EDFDD2117E8E56" box="[754,835,548,572]" captionStart="Figure 10" captionStartId="18.[211,278,1446,1468]" captionTargetBox="[213,1170,167,1412]" captionTargetId="figure-5@18.[213,1170,167,1413]" captionTargetPageId="18" captionText="Figure 10. Jacaena mihun, holotype (AC); J. thakek comb. nov., holotype (DF). Male palp, ventral (A, D). Ditto, prolateral (B, E). Ditto, retrolateral (C, F)." figureDoi="http://doi.org/10.5281/zenodo.6517670" httpUri="https://zenodo.org/record/6517670/files/figure.png" pageId="16" pageNumber="1553">10DF</figureCitation>
)
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<treatmentCitation id="AF7E7015FFCDFF9A07CCFD9210058E16" ID-DOI="http://doi.org/10.5281/zenodo.6250833" ID-Zenodo-Dep="6250833" author="Jager P" box="[211,568,612,637]" httpUri="http://treatment.plazi.org/id/D27487BFFF913B10FF763EAD7D93F79D" page="48" pageId="16" pageNumber="1553" year="2007">
<taxonomicName id="E9DF2D87FFCDFF9A07CCFD9210058E16" authority="Jager, 2007: 48" authorityName="Jager" authorityPageNumber="48" authorityYear="2007" box="[211,568,612,637]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="thakek">
<emphasis id="1CAB8A16FFCDFF9A07CCFD9213B78E16" box="[211,394,612,636]" italics="true" pageId="16" pageNumber="1553">Sesieutes thakek</emphasis>
<bibRefCitation id="4A4E2BF5FFCDFF9A068FFD9310058E16" author="Jager P" box="[400,568,612,637]" pageId="16" pageNumber="1553" pagination="29 - 58" refId="ref11377" refString="Jager P. 2007. Spiders from Laos with descriptions of new species (Arachnida: Araneae). Acta arachn Tokyo. 56: 29 - 58." type="journal article" year="2007">Jäger, 2007: 48</bibRefCitation>
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, figs 7382, description of
<specimenCount id="38D99D8DFFCDFF9A047CFD9211458E16" box="[867,888,612,636]" pageId="16" pageNumber="1553" type="male"></specimenCount>
.
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<emphasis id="1CAB8A16FFCDFF9A07CCFD2413518E80" box="[211,364,722,746]" italics="true" pageId="16" pageNumber="1553">Type material</emphasis>
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<materialsCitation id="9EB75C59FFCDFF9A07CCFD0F16018F3B" ID-GBIF-Occurrence="3757642322" collectingDate="2004-10-30" collectingMethod="sieving leaf litter" collectionCode="SMF" collectorName="P. Jager &amp; V. Vedel" country="Laos" elevation="159" latitude="17.448933" location="in front of cave" longLatPrecision="1" longitude="104.874985" pageId="16" pageNumber="1553" specimenCode="SMF-56635" specimenCount="1" specimenCount-male="1" stateProvince="Khammouan Province" typeStatus="holotype">
<typeStatus id="F164E8A6FFCDFF9A07CCFD0F137F8F7B" box="[211,322,761,785]" pageId="16" pageNumber="1553" type="holotype">Holotype</typeStatus>
:
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,
<collectingCountry id="56C81694FFCDFF9A0664FD0C13898F7B" box="[379,436,762,785]" name="Laos" pageId="16" pageNumber="1553">Laos</collectingCountry>
,
<collectingRegion id="EC1B98E6FFCDFF9A06DAFD0F10F18F7B" box="[453,716,761,785]" country="Laos" name="Khammouan" pageId="16" pageNumber="1553">Khammouan Province</collectingRegion>
,
<locationDeviation id="3FD24F26FFCDFF9A05C2FD0F160C8F7B" box="[733,1073,761,785]" pageId="16" pageNumber="1553">
<quantity id="E927FBE1FFCDFF9A05C2FD0F110D8F7B" box="[733,816,761,785]" metricMagnitude="3" metricUnit="m" metricValue="9.5" pageId="16" pageNumber="1553" unit="km" value="9.5">9.5 km</quantity>
northeast of Thakek
</locationDeviation>
,
<quantity id="E927FBE1FFCDFF9A035DFD0F16B28F7B" box="[1090,1167,761,785]" metricMagnitude="2" metricUnit="m" metricValue="1.59" pageId="16" pageNumber="1553" unit="m" value="159.0">
<elevation id="A5F2B137FFCDFF9A035DFD0F16B28F7B" box="[1090,1167,761,785]" metricMagnitude="2" metricUnit="m" metricValue="1.59" pageId="16" pageNumber="1553" unit="m" value="159.0">159 m</elevation>
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,
<geoCoordinate id="4BEB30C3FFCDFF9A07CCFCEF13568F5B" box="[211,363,789,817]" degrees="17" direction="north" minutes="26.936" orientation="latitude" pageId="16" pageNumber="1553" precision="1" value="17.448933">
17
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26.936
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N
</geoCoordinate>
,
<geoCoordinate id="4BEB30C3FFCDFF9A0663FCEF10238F5B" box="[380,542,789,817]" degrees="104" direction="east" minutes="52.499" orientation="longitude" pageId="16" pageNumber="1553" precision="1" value="104.874985">
104
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52.499
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E
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,
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<location id="2B0000DFFFCDFF9A05E1FCEF11858F5B" LSID="urn:lsid:plazi:treatment:A676E712FFCDFF9A06CCFDF3119089B0:2B0000DFFFCDFF9A05E1FCEF11858F5B" box="[766,952,793,817]" country="Laos" latitude="17.448933" longLatPrecision="1" longitude="104.874985" name="in front of cave" pageId="16" pageNumber="1553" stateProvince="Khammouan Province">in front of cave</location>
,
<date id="5A6170C4FFCDFF9A04D6FCEF16B28F5B" box="[969,1167,793,817]" pageId="16" pageNumber="1553" value="2004-10-30">
<collectingDate id="4A25892CFFCDFF9A04D6FCEF16B28F5B" box="[969,1167,793,817]" pageId="16" pageNumber="1553" value="2004-10-30">30 October 2004</collectingDate>
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,
<collectorName id="832A33D2FFCDFF9A07CCFCCC13148F38" box="[211,297,826,850]" pageId="16" pageNumber="1553">P. Jäger</collectorName>
and
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leg. (
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, examined,
<figureCitation id="B6E44A81FFCDFF9A0414FCCF11BA8F3B" box="[779,903,825,849]" captionStart="Figure 1" captionStartId="5.[143,210,1100,1122]" captionTargetBox="[145,1102,168,1066]" captionTargetId="figure-5@5.[145,1102,168,1066]" captionTargetPageId="5" captionText="Figure 1. Males of Jacaena spp., habitus, dorsal (AH). Jacaena schwendingeri comb. nov., holotype (A); J. lunulata sp. nov., holotype (B); J. punctata sp. nov., holotype (C); J. mihun (D); J. erawan comb. nov. (E); J. thakek comb. nov., holotype (F); J. schwendingeri comb. nov. (G); J. lunulata sp. nov., paratype (H). DS, dorsal scutum." figureDoi="http://doi.org/10.5281/zenodo.6517638" httpUri="https://zenodo.org/record/6517638/files/figure.png" pageId="16" pageNumber="1553">Figures 1F</figureCitation>
,
<figureCitation id="B6E44A81FFCDFF9A048CFCCF11EC8F3B" box="[915,977,825,849]" captionStart="Figure 5" captionStartId="11.[143,210,1565,1587]" captionTargetBox="[145,1102,161,1532]" captionTargetId="figure-5@11.[145,1102,161,1532]" captionTargetPageId="11" captionText="Figure 5. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena erawan comb. nov., holotype (AC); J. punctata sp. nov., holotype (DF); J. thakek comb. nov., holotype (GI)." figureDoi="http://doi.org/10.5281/zenodo.6517654" httpUri="https://zenodo.org/record/6517654/files/figure.png" pageId="16" pageNumber="1553">5GI</figureCitation>
,
<figureCitation id="B6E44A81FFCDFF9A04C3FCCF16128F3B" box="[988,1071,825,849]" captionStart="Figure 10" captionStartId="18.[211,278,1446,1468]" captionTargetBox="[213,1170,167,1412]" captionTargetId="figure-5@18.[213,1170,167,1413]" captionTargetPageId="18" captionText="Figure 10. Jacaena mihun, holotype (AC); J. thakek comb. nov., holotype (DF). Male palp, ventral (A, D). Ditto, prolateral (B, E). Ditto, retrolateral (C, F)." figureDoi="http://doi.org/10.5281/zenodo.6517670" httpUri="https://zenodo.org/record/6517670/files/figure.png" pageId="16" pageNumber="1553">10DF</figureCitation>
).
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</paragraph>
</subSubSection>
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<emphasis id="1CAB8A16FFCDFF9A07CCFC5E13028FD5" box="[211,319,936,959]" italics="true" pageId="16" pageNumber="1553">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFCDFF9A07CCFC381140892F" blockId="16.[211,1172,936,1349]" pageId="16" pageNumber="1553">
This species seems to be closest to
<taxonomicName id="E9DF2D87FFCDFF9A0542FC3910878F8D" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[605,698,975,999]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFCDFF9A0542FC3910878F8D" box="[605,698,975,999]" italics="true" pageId="16" pageNumber="1553">J. mihun</emphasis>
</taxonomicName>
, because both species have very distinctly enlarged PME; their palps can be distinguished from other members of
<taxonomicName id="E9DF2D87FFCDFF9A04EFFC191675886C" authorityName="Thorell" authorityYear="1897" box="[1008,1096,1007,1030]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="genus">
<emphasis id="1CAB8A16FFCDFF9A04EFFC191675886C" box="[1008,1096,1007,1030]" italics="true" pageId="16" pageNumber="1553">Jacaena</emphasis>
</taxonomicName>
by the pear-shaped bulb anteriorly narrowed, and the apex of the short, triangular conductor is pointing apically (
<figureCitation id="B6E44A81FFCDFF9A06A9FBD81073882C" box="[438,590,1070,1094]" captionStart="Figure 4" captionStartId="8.[211,278,1428,1450]" captionTargetBox="[213,1170,168,1394]" captionTargetId="figure-5@8.[213,1170,168,1394]" captionTargetPageId="8" captionText="Figure 4. Jacaena spp. Male palps, ventral (A, D, G). Ditto, prolateral (B, E, H). Ditto, retrolateral (C, F, I). Jacaena schwendingeri comb. nov., holotype (AC); J. lunulata sp. nov., paratype (originally misidentified as Sesieutes schwendingeri) (DF); J. mihun, holotype (GI)." figureDoi="http://doi.org/10.5281/zenodo.6517648" httpUri="https://zenodo.org/record/6517648/files/figure.png" pageId="16" pageNumber="1553">Figures 4GI</figureCitation>
cf. 5GI). The male of this species can be separated from that of
<taxonomicName id="E9DF2D87FFCDFF9A067AFBB913FF880C" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[357,450,1102,1126]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFCDFF9A067AFBB913FF880C" box="[357,450,1102,1126]" italics="true" pageId="16" pageNumber="1553">J. mihun</emphasis>
</taxonomicName>
by the absence of a small, triangular denticle on the dorsal surface of the conductor (
<figureCitation id="B6E44A81FFCDFF9A06C5FB98106088EC" box="[474,605,1134,1158]" captionStart="Figure 10" captionStartId="18.[211,278,1446,1468]" captionTargetBox="[213,1170,167,1412]" captionTargetId="figure-5@18.[213,1170,167,1413]" captionTargetPageId="18" captionText="Figure 10. Jacaena mihun, holotype (AC); J. thakek comb. nov., holotype (DF). Male palp, ventral (A, D). Ditto, prolateral (B, E). Ditto, retrolateral (C, F)." figureDoi="http://doi.org/10.5281/zenodo.6517670" httpUri="https://zenodo.org/record/6517670/files/figure.png" pageId="16" pageNumber="1553">Figure 10A</figureCitation>
cf. 10D). However, we have a private notion that the boundary between these two species is vague and
<taxonomicName id="E9DF2D87FFCDFF9A042EFB7911AB88CC" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Dankittipakul &amp; Tavano &amp; Singtripop" baseAuthorityYear="2013" box="[817,918,1166,1190]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="thakek" status="comb. nov.">
<emphasis id="1CAB8A16FFCDFF9A042EFB7911AB88CC" box="[817,918,1166,1190]" italics="true" pageId="16" pageNumber="1553">J. thakek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFCDFF9A0482FB78162588CF" box="[925,1048,1166,1190]" pageId="16" pageNumber="1553" rank="species">comb. nov.</taxonomicNameLabel>
is possibly a junior synonym of
<taxonomicName id="E9DF2D87FFCDFF9A06DCFB59101C88AC" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[451,545,1198,1222]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFCDFF9A06DCFB59101C88AC" box="[451,545,1198,1222]" italics="true" pageId="16" pageNumber="1553">J. mihun</emphasis>
</taxonomicName>
. Much larger samples would be needed to corroborate these slight differences as species-specific rather than intraspecific variation because more material recently obtained suggests that
<taxonomicName id="E9DF2D87FFCDFF9A05FBFB18117E896C" authorityName="Deeleman-Reinhold" authorityYear="2001" box="[740,835,1262,1286]" class="Arachnida" family="Liocranidae" genus="Jacaena" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="mihun">
<emphasis id="1CAB8A16FFCDFF9A05FBFB18117E896C" box="[740,835,1262,1286]" italics="true" pageId="16" pageNumber="1553">J. mihun</emphasis>
</taxonomicName>
has relatively broad distribution covering northern and north-eastern parts of
<collectingCountry id="56C81694FFCDFF9A041EFAFB1155894F" box="[769,872,1293,1317]" name="Thailand" pageId="16" pageNumber="1553">Thailand</collectingCountry>
(and most likely extending eastwards to
<collectingCountry id="56C81694FFCDFF9A0678FAD813A2892F" box="[359,415,1326,1349]" name="Laos" pageId="16" pageNumber="1553">Laos</collectingCountry>
where
<taxonomicName id="E9DF2D87FFCDFF9A06F1FAD8106F892F" authorityName="Dankittipakul &amp; Tavano &amp; Singtripop" authorityYear="2013" baseAuthorityName="Dankittipakul &amp; Tavano &amp; Singtripop" baseAuthorityYear="2013" box="[494,594,1325,1349]" class="Arachnida" family="Liocranidae" genus="Sesieutes" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="1553" phylum="Arthropoda" rank="species" species="thakek" status="comb. nov.">
<emphasis id="1CAB8A16FFCDFF9A06F1FAD8106F892F" box="[494,594,1325,1349]" italics="true" pageId="16" pageNumber="1553">J. thakek</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0798376DFFCDFF9A0546FADB10EE892F" box="[601,723,1325,1349]" pageId="16" pageNumber="1553" rank="species">comb. nov.</taxonomicNameLabel>
was collected).
</paragraph>
</subSubSection>
<subSubSection id="66C5058FFFCDFF9A07CCFA6D119089B0" box="[211,941,1435,1499]" pageId="16" pageNumber="1553" type="distribution">
<paragraph id="2E605604FFCDFF9A07CCFA6D136589D9" blockId="16.[211,941,1435,1499]" box="[211,344,1435,1459]" pageId="16" pageNumber="1553">
<heading id="7528E168FFCDFF9A07CCFA6D136589D9" box="[211,344,1435,1459]" fontSize="10" level="3" pageId="16" pageNumber="1553" reason="5">
<emphasis id="1CAB8A16FFCDFF9A07CCFA6D136589D9" box="[211,344,1435,1459]" italics="true" pageId="16" pageNumber="1553">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="2E605604FFCDFF9A07CCFA35119089B0" blockId="16.[211,941,1435,1499]" box="[211,941,1474,1499]" pageId="16" pageNumber="1553">
Known only from the
<typeStatus id="F164E8A6FFCDFF9A06D2FA3513C089B1" box="[461,509,1475,1499]" pageId="16" pageNumber="1553">type</typeStatus>
locality,
<collectingRegion id="EC1B98E6FFCDFF9A057CFA34115989B0" box="[611,868,1474,1498]" country="Laos" name="Khammouan" pageId="16" pageNumber="1553">Khammouan Province</collectingRegion>
,
<collectingCountry id="56C81694FFCDFF9A0470FA35119489B0" box="[879,937,1475,1498]" name="Laos" pageId="16" pageNumber="1553">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>