treatments-xml/data/A6/6B/1A/A66B1A7CEF54D32EFF0FFB4609E0FD48.xml
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<document id="F017939B76A1713BCE14CE84B8FDA8E0" ID-CLB-Dataset="7563" ID-DOI="10.11646/zootaxa.4948.2.1" ID-GBIF-Dataset="82bdee28-6a14-4f2d-b7b1-b4b103383775" ID-ISSN="1175-5326" ID-Zenodo-Dep="4620704" ID-ZooBank="6E073009-5EB5-4C00-89C6-0728E7FA544B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616141388973" checkinUser="plazi" docAuthor="Joya, Daniel Chirivi" docDate="2021" docId="A66B1A7CEF54D32EFF0FFB4609E0FD48" docLanguage="en" docName="zootaxa.4928.2.1.pdf" docOrigin="Zootaxa 4948 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrynus alfonsoi Joya 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="163" masterDocId="5A526204EF53D322FF98FFAA0D6FFF86" masterDocTitle="Four new species of Phrynus, Lamarck (Arachnida: Amblypygi) from Mexico" masterLastPageNumber="183" masterPageNumber="151" pageNumber="158" updateTime="1701998102619" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="44B107E13DCE949DFFEDA12D3DD51FEC">Four new species of Phrynus, Lamarck (Arachnida: Amblypygi) from Mexico</mods:title>
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<mods:name id="AC6235A8BCCDA718A4F8FBC00228EEBC" type="personal">
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<mods:namePart id="3F779A1B4C3EBE1C89BA046818B46C33">Joya, Daniel Chirivi</mods:namePart>
<mods:affiliation id="1A9BDE4F42433E8A743F3A92D97FD5A7">Unidad de Ecología y Sistemática (UNESIS), Laboratorio de Entomología, Departamento de Biología, Facultad de Ciencias, Pontificia Universidad Javeriana. Bogotá, Colombia. Carrera 7 N °. 40 - 62, Bogotá.</mods:affiliation>
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<mods:title id="42294B1E2ED82CFDD4F2FE67F5313D34">Zootaxa</mods:title>
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<mods:date id="5278BFB6A8E30C6A22AC42C4AFB79276">2021</mods:date>
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<mods:number id="96D29A337B30FC0EAF56E13763D6D1C2">2021-03-19</mods:number>
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<treatment id="A66B1A7CEF54D32EFF0FFB4609E0FD48" ID-DOI="http://doi.org/10.5281/zenodo.4651562" ID-GBIF-Taxon="180781950" ID-Zenodo-Dep="4651562" LSID="urn:lsid:plazi:treatment:A66B1A7CEF54D32EFF0FFB4609E0FD48" httpUri="http://treatment.plazi.org/id/A66B1A7CEF54D32EFF0FFB4609E0FD48" lastPageId="12" lastPageNumber="163" pageId="7" pageNumber="158">
<subSubSection id="66D8F8E1EF54D325FF0FFB460CA3FA81" box="[151,460,1260,1287]" pageId="7" pageNumber="158" type="nomenclature">
<paragraph id="2E7DAB6AEF54D325FF0FFB460CA3FA81" blockId="7.[151,513,1260,1323]" box="[151,460,1260,1287]" pageId="7" pageNumber="158">
<heading id="75351C06EF54D325FF0FFB460CA3FA81" bold="true" box="[151,460,1260,1287]" fontSize="11" level="1" pageId="7" pageNumber="158" reason="1">
<emphasis id="1CB67778EF54D325FF0FFB460CA3FA81" bold="true" box="[151,460,1260,1287]" pageId="7" pageNumber="158">
<taxonomicName id="E9C2D0E9EF54D325FF0FFB460C08FA81" ID-CoL="8VMY7" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[151,359,1260,1287]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF54D325FF0FFB460C08FA81" bold="true" box="[151,359,1260,1287]" italics="true" pageId="7" pageNumber="158">Phrynus alfonsoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0785CA03EF54D325FEF6FB470CA3FA81" box="[366,460,1261,1287]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF54D325FF0FFABA0F6EFAAC" box="[151,513,1296,1323]" pageId="7" pageNumber="158" type="description">
<paragraph id="2E7DAB6AEF54D325FF0FFABA0F6EFAAC" blockId="7.[151,513,1260,1323]" box="[151,513,1296,1323]" pageId="7" pageNumber="158">
<figureCitation id="B6F9B7EFEF54D325FF0FFABA0C49FAAC" box="[151,294,1296,1322]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="8.[151,250,1964,1988]" captionStartId-1="10.[151,250,1650,1674]" captionStartId-2="11.[151,250,1779,1803]" captionTargetBox-0="[174,1415,459,1933]" captionTargetBox-1="[162,1425,246,1606]" captionTargetBox-2="[155,1432,855,1752]" captionTargetId-0="figure-192@8.[161,1427,453,1940]" captionTargetId-1="figure-54@10.[151,1437,237,1627]" captionTargetId-2="figure-323@11.[151,1437,849,1755]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." captionText-1="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." captionText-2="FIGURE 6. Male genitalia: Phrynus alfonsoi sp. nov. (CNAN-T01445): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi-0="http://doi.org/10.5281/zenodo.4620720" figureDoi-1="http://doi.org/10.5281/zenodo.4620722" figureDoi-2="http://doi.org/10.5281/zenodo.4620724" httpUri-0="https://zenodo.org/record/4620720/files/figure.png" httpUri-1="https://zenodo.org/record/4620722/files/figure.png" httpUri-2="https://zenodo.org/record/4620724/files/figure.png" pageId="7" pageNumber="158">Figures. 46</figureCitation>
,
<figureCitation id="B6F9B7EFEF54D325FEA9FABB0C15FAAD" box="[305,378,1297,1323]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStartId-0="27.[151,250,1596,1620]" captionStartId-1="28.[151,250,1479,1503]" captionTargetBox-0="[158,1428,188,1562]" captionTargetBox-1="[151,1436,813,1456]" captionTargetId-0="figure-15@27.[151,1437,181,1572]" captionTargetId-1="figure-330@28.[151,1436,813,1456]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." captionText-1="FIGURE 18. Distribution: Phrynus alfonsoi sp. nov. (blue circle), Phrynus aliciae sp. nov. (yellow circle), Phrynus giseae sp. nov. (purple circle), Phrynus jalisco (light blue circle), Phrynus operculatus (red circle), Phrynus tresmarias sp. nov. (orange circle). Circles with black rings: type localities." figureDoi-0="http://doi.org/10.5281/zenodo.4620748" figureDoi-1="http://doi.org/10.5281/zenodo.4620750" httpUri-0="https://zenodo.org/record/4620748/files/figure.png" httpUri-1="https://zenodo.org/record/4620750/files/figure.png" pageId="7" pageNumber="158">1718</figureCitation>
.
<tableCitation id="63409ED1EF54D325FE1DFABA0C8FFAAD" box="[389,480,1296,1323]" captionStart="TABLE 2" captionStartId="9.[151,239,548,572]" captionText="TABLE 2. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus alfonsoi sp. nov. When the segment is missing, the cell is empty." pageId="7" pageNumber="158">Table. 2</tableCitation>
,
<tableCitation id="63409ED1EF54D325FE74FABA0C92FAAC" box="[492,509,1296,1322]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="7" pageNumber="158">6</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF54D325FF0FFAF30E36F9A1" pageId="7" pageNumber="158" type="materials_examined">
<paragraph id="2E7DAB6AEF54D325FF0FFAF3090FFA11" blockId="7.[151,1437,1368,2043]" pageId="7" pageNumber="158">
<materialsCitation id="9EAAA137EF54D325FF0FFAF3090FFA11" ID-GBIF-Occurrence="3064396319" collectingDate="2007-06-22" collectionCode="CNAN-T" collectorName="O. Francke &amp; A. Ballesteros &amp; L. Escalante &amp; H. Montano" country="Mexico" county="Municipio Ayutla" elevation="15317" latitude="16.902433" location="Loc" longLatPrecision="1" longitude="-99.09485" municipality="Tepango" pageId="7" pageNumber="158" specimenCode="CNAN-T01383" specimenCount="1" stateProvince="Guerrero" typeStatus="holotype">
<emphasis id="1CB67778EF54D325FF0FFAF30C12FAF4" bold="true" box="[151,381,1368,1395]" pageId="7" pageNumber="158">
<typeStatus id="F17915C8EF54D325FF0FFAF30C6BFAF5" box="[151,260,1369,1395]" pageId="7" pageNumber="158" type="holotype">Holotype</typeStatus>
(Female):
</emphasis>
<collectingCountry id="56D5EBFAEF54D325FE1CFAF30C99FAF5" box="[388,502,1369,1395]" name="Mexico" pageId="7" pageNumber="158">MEXICO</collectingCountry>
.
<emphasis id="1CB67778EF54D325FD98FAF30F07FAF5" box="[512,616,1369,1395]" italics="true" pageId="7" pageNumber="158">
<collectingRegion id="EC066588EF54D325FD98FAF30F07FAF5" box="[512,616,1369,1395]" country="Mexico" name="Guerrero" pageId="7" pageNumber="158">Guerrero</collectingRegion>
</emphasis>
:
<collectingCounty id="C71CD3E6EF54D325FDE0FAF20E51FAF4" box="[632,830,1368,1394]" pageId="7" pageNumber="158">Municipio Ayutla</collectingCounty>
,
<collectingMunicipality id="CE193110EF54D325FCD0FAF30EC7FAF5" box="[840,936,1369,1395]" pageId="7" pageNumber="158">Tepango</collectingMunicipality>
(
<geoCoordinate id="4BF6CDADEF54D325FC2EFAF2095EFAF4" box="[950,1073,1368,1394]" degrees="16.902433" direction="north" orientation="latitude" pageId="7" pageNumber="158" precision="1" value="16.902433">16.902433</geoCoordinate>
,
<geoCoordinate id="4BF6CDADEF54D325FBA4FAF209ACFAF5" box="[1084,1219,1368,1395]" degrees="99.094850" direction="west" orientation="longitude" pageId="7" pageNumber="158" precision="1" value="-99.09485">-99.094850</geoCoordinate>
):
<location id="2B1DFDB1EF54D325FB4CFAF3086FFAF5" LSID="urn:lsid:plazi:treatment:A66B1A7CEF54D32EFF0FFB4609E0FD48:2B1DFDB1EF54D325FB4CFAF3086FFAF5" box="[1236,1280,1369,1395]" country="Mexico" county="Municipio Ayutla" latitude="16.902433" longLatPrecision="1" longitude="-99.09485" municipality="Tepango" name="Loc" pageId="7" pageNumber="158" stateProvince="Guerrero">Loc</location>
<quantity id="E93A068FEF54D325FA9EFAF208F7FAF4" box="[1286,1432,1368,1395]" metricMagnitude="4" metricUnit="m" metricValue="1.5317" pageId="7" pageNumber="158" unit="m" value="15317.0">
<elevation id="A5EF4C59EF54D325FA9EFAF208F7FAF4" box="[1286,1432,1368,1395]" metricMagnitude="4" metricUnit="m" metricValue="1.5317" pageId="7" pageNumber="158" unit="m" value="15317.0">15, 317 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF54D325FF0FFAD70C5FFA11" box="[151,304,1405,1431]" pageId="7" pageNumber="158" value="2007-06-22">
<collectingDate id="4A387442EF54D325FF0FFAD70C5FFA11" box="[151,304,1405,1431]" pageId="7" pageNumber="158" value="2007-06-22">22 June 2007</collectingDate>
</date>
,
<collectorName id="8337CEBCEF54D325FEA3FAD70CD5FA10" box="[315,442,1404,1431]" pageId="7" pageNumber="158">O. Francke</collectorName>
,
<collectorName id="8337CEBCEF54D325FE5BFAD70F0AFA10" box="[451,613,1404,1431]" pageId="7" pageNumber="158">A. Ballesteros</collectorName>
,
<collectorName id="8337CEBCEF54D325FDE8FAD70F92FA10" box="[624,765,1404,1431]" pageId="7" pageNumber="158">L. Escalante</collectorName>
,
<collectorName id="8337CEBCEF54D325FC90FAD70EFFFA11" box="[776,912,1405,1431]" pageId="7" pageNumber="158">H. Montaño</collectorName>
(
<specimenCode id="7E640311EF54D325FC38FAD7093FFA11" box="[928,1104,1405,1431]" collectionCode="CNAN-T" pageId="7" pageNumber="158">CNAN-T01383</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="2E7DAB6AEF54D325FF5FFA0B0E36F9A1" blockId="7.[151,1437,1368,2043]" pageId="7" pageNumber="158">
<materialsCitation id="9EAAA137EF54D325FF5FFA0B0873FA59" ID-GBIF-Occurrence="3064396308" collectingDate="2007-06-22" collectionCode="CNAN-T" collectorName="O. Francke &amp; A. Ballesteros &amp; L. Escalante &amp; H. Montano" country="Mexico" county="Municipio Ayutla" elevation="15317" location="Loc" municipality="Tepango" pageId="7" pageNumber="158" specimenCode="CNAN-T01445" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Guerrero" typeStatus="paratype">
<emphasis id="1CB67778EF54D325FF5FFA0B0F56FA3C" bold="true" box="[199,569,1440,1467]" pageId="7" pageNumber="158">
<typeStatus id="F17915C8EF54D325FF5FFA0B0C50FA3D" box="[199,319,1441,1467]" pageId="7" pageNumber="158" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="38C460E3EF54D325FED7FA0A0CAFFA3C" box="[335,448,1440,1466]" pageId="7" pageNumber="158" type="female">5 females</specimenCount>
,
<specimenCount id="38C460E3EF54D325FE55FA0B0F43FA3C" box="[461,556,1440,1467]" pageId="7" pageNumber="158" type="male">5 males</specimenCount>
):
</emphasis>
<collectingCountry id="56D5EBFAEF54D325FDD8FA0B0FDDFA3D" box="[576,690,1441,1467]" name="Mexico" pageId="7" pageNumber="158">MEXICO</collectingCountry>
.
<emphasis id="1CB67778EF54D325FD25FA0B0E4AFA3D" box="[701,805,1441,1467]" italics="true" pageId="7" pageNumber="158">
<collectingRegion id="EC066588EF54D325FD25FA0B0E4AFA3D" box="[701,805,1441,1467]" country="Mexico" name="Guerrero" pageId="7" pageNumber="158">Guerrero</collectingRegion>
</emphasis>
:
<collectingCounty id="C71CD3E6EF54D325FCADFA0A0E93FA3C" box="[821,1020,1440,1466]" pageId="7" pageNumber="158">Municipio Ayutla</collectingCounty>
,
<collectingMunicipality id="CE193110EF54D325FB9EFA0B0904FA3D" box="[1030,1131,1441,1467]" pageId="7" pageNumber="158">Tepango</collectingMunicipality>
:
<location id="2B1DFDB1EF54D325FBEDFA0B09CEFA3D" LSID="urn:lsid:plazi:treatment:A66B1A7CEF54D32EFF0FFB4609E0FD48:2B1DFDB1EF54D325FBEDFA0B09CEFA3D" box="[1141,1185,1441,1467]" country="Mexico" county="Municipio Ayutla" municipality="Tepango" name="Loc" pageId="7" pageNumber="158" stateProvince="Guerrero">Loc</location>
<quantity id="E93A068FEF54D325FB30FA0A0854FA3C" box="[1192,1339,1440,1467]" metricMagnitude="4" metricUnit="m" metricValue="1.5317" pageId="7" pageNumber="158" unit="m" value="15317.0">
<elevation id="A5EF4C59EF54D325FB30FA0A0854FA3C" box="[1192,1339,1440,1467]" metricMagnitude="4" metricUnit="m" metricValue="1.5317" pageId="7" pageNumber="158" unit="m" value="15317.0">15, 317 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF54D325FADEFA0B0DBDFA59" pageId="7" pageNumber="158" value="2007-06-22">
<collectingDate id="4A387442EF54D325FADEFA0B0DBDFA59" pageId="7" pageNumber="158" value="2007-06-22">22 June 2007</collectingDate>
</date>
,
<collectorName id="8337CEBCEF54D325FF45FA6F0C34FA58" box="[221,347,1476,1503]" pageId="7" pageNumber="158">O. Francke</collectorName>
,
<collectorName id="8337CEBCEF54D325FEFDFA6F0F6AFA58" box="[357,517,1476,1503]" pageId="7" pageNumber="158">A. Ballesteros</collectorName>
,
<collectorName id="8337CEBCEF54D325FD88FA6F0FF2FA58" box="[528,669,1476,1503]" pageId="7" pageNumber="158">L. Escalante</collectorName>
,
<collectorName id="8337CEBCEF54D325FD30FA6F0E5EFA59" box="[680,817,1477,1503]" pageId="7" pageNumber="158">H. Montaño</collectorName>
, two females, three males (
<specimenCode id="7E640311EF54D325FBF9FA6E087EFA59" box="[1121,1297,1476,1503]" collectionCode="CNAN-T" pageId="7" pageNumber="158">CNAN-T01445</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="9EAAA137EF54D325FAB0FA6E0E3BF9A1" ID-GBIF-Occurrence="3064396303" collectingDate="2008-08-01" collectionCode="CNAN-T" collectorName="O. Francke &amp; H. Montano &amp; J. Ponce &amp; A. Quijano" country="Mexico" elevation="72" latitude="17.64491" location="Colonia Agua de Correa" longLatPrecision="1" longitude="-101.51822" municipality="Municipio Jose Asueto" pageId="7" pageNumber="158" specimenCode="CNAN-T01446" specimenCount="1" stateProvince="Guerrero" typeStatus="paratype">
<collectingMunicipality id="CE193110EF54D325FAB0FA6E0C71F985" pageId="7" pageNumber="158">Municipio José Asueto</collectingMunicipality>
,
<location id="2B1DFDB1EF54D325FEB1FA420F57F985" LSID="urn:lsid:plazi:treatment:A66B1A7CEF54D32EFF0FFB4609E0FD48:2B1DFDB1EF54D325FEB1FA420F57F985" box="[297,568,1512,1539]" country="Mexico" latitude="17.64491" longLatPrecision="1" longitude="-101.51822" municipality="Municipio Jose Asueto" name="Colonia Agua de Correa" pageId="7" pageNumber="158" stateProvince="Guerrero">Colonia Agua de Correa</location>
(
<geoCoordinate id="4BF6CDADEF54D325FDDEFA420FDCF984" box="[582,691,1512,1538]" degrees="17.64491" direction="north" orientation="latitude" pageId="7" pageNumber="158" precision="1" value="17.64491">17.64491</geoCoordinate>
,
<geoCoordinate id="4BF6CDADEF54D325FD26FA420E2AF985" box="[702,837,1512,1539]" degrees="101.51822" direction="west" orientation="longitude" pageId="7" pageNumber="158" precision="1" value="-101.51822">-101.51822</geoCoordinate>
):
<quantity id="E93A068FEF54D325FCCEFA430EDEF984" box="[854,945,1512,1539]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="7" pageNumber="158" unit="m" value="72.0">
<elevation id="A5EF4C59EF54D325FCCEFA430EDEF984" box="[854,945,1512,1539]" metricMagnitude="1" metricUnit="m" metricValue="7.2" pageId="7" pageNumber="158" unit="m" value="72.0">72 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF54D325FC24FA43090EF985" box="[956,1121,1513,1539]" pageId="7" pageNumber="158" value="2008-08-01">
<collectingDate id="4A387442EF54D325FC24FA43090EF985" box="[956,1121,1513,1539]" pageId="7" pageNumber="158" value="2008-08-01">1 August 2008</collectingDate>
</date>
,
<collectorName id="8337CEBCEF54D325FBF4FA430984F984" box="[1132,1259,1512,1539]" pageId="7" pageNumber="158">O. Francke</collectorName>
,
<collectorName id="8337CEBCEF54D325FB6EFA430810F985" box="[1270,1407,1513,1539]" pageId="7" pageNumber="158">H. Montaño</collectorName>
,
<collectorName id="8337CEBCEF54D325FA12FA430DB1F9A1" pageId="7" pageNumber="158">J. Ponce</collectorName>
,
<collectorName id="8337CEBCEF54D325FF70F9A70C08F9A0" box="[232,359,1548,1575]" pageId="7" pageNumber="158">A. Quijano</collectorName>
, three females, two males (
<specimenCode id="7E640311EF54D325FD01F9A60E26F9A0" box="[665,841,1548,1574]" collectionCode="CNAN-T" pageId="7" pageNumber="158">CNAN-T01446</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF54D325FF5FF99A0E71F9CD" box="[199,798,1584,1611]" pageId="7" pageNumber="158" type="distribution">
<paragraph id="2E7DAB6AEF54D325FF5FF99A0E71F9CD" blockId="7.[151,1437,1368,2043]" box="[199,798,1584,1611]" pageId="7" pageNumber="158">
<emphasis id="1CB67778EF54D325FF5FF99A0C0EF9CC" bold="true" box="[199,353,1584,1610]" pageId="7" pageNumber="158">Distribution.</emphasis>
<collectingCountry id="56D5EBFAEF54D325FEFFF99B0CB5F9CD" box="[359,474,1585,1611]" name="Mexico" pageId="7" pageNumber="158">MEXICO</collectingCountry>
: State of
<collectingRegion id="EC066588EF54D325FDDAF99B0FC4F9CD" box="[578,683,1585,1611]" country="Mexico" name="Guerrero" pageId="7" pageNumber="158">Guerrero</collectingRegion>
. (
<figureCitation id="B6F9B7EFEF54D325FD25F99A0E7EF9CD" box="[701,785,1584,1611]" captionStart="FIGURE 18" captionStartId="28.[151,250,1479,1503]" captionTargetBox="[151,1436,813,1456]" captionTargetId="figure-330@28.[151,1436,813,1456]" captionTargetPageId="28" captionText="FIGURE 18. Distribution: Phrynus alfonsoi sp. nov. (blue circle), Phrynus aliciae sp. nov. (yellow circle), Phrynus giseae sp. nov. (purple circle), Phrynus jalisco (light blue circle), Phrynus operculatus (red circle), Phrynus tresmarias sp. nov. (orange circle). Circles with black rings: type localities." figureDoi="http://doi.org/10.5281/zenodo.4620750" httpUri="https://zenodo.org/record/4620750/files/figure.png" pageId="7" pageNumber="158">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF54D325FF5FF9FF0CB5F915" pageId="7" pageNumber="158" type="etymology">
<paragraph id="2E7DAB6AEF54D325FF5FF9FF0CB5F915" blockId="7.[151,1437,1368,2043]" pageId="7" pageNumber="158">
<emphasis id="1CB67778EF54D325FF5FF9FF0C21F9E9" bold="true" box="[199,334,1621,1647]" pageId="7" pageNumber="158">Etymology.</emphasis>
The species is dedicated to my father, Alfonso Chirivi. My scientific contributions were not being possible without his support.
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF54D32BFF5FF9360EEDFF7F" lastPageId="9" lastPageNumber="160" pageId="7" pageNumber="158" type="diagnosis">
<paragraph id="2E7DAB6AEF54D325FF5FF9360975F809" blockId="7.[151,1437,1368,2043]" pageId="7" pageNumber="158">
<emphasis id="1CB67778EF54D325FF5FF9360C2DF930" bold="true" box="[199,322,1692,1718]" pageId="7" pageNumber="158">Diagnosis.</emphasis>
<taxonomicName id="E9C2D0E9EF54D325FED1F9360F66F930" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[329,521,1692,1718]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF54D325FED1F9360F66F930" box="[329,521,1692,1718]" italics="true" pageId="7" pageNumber="158">Phrynus alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF54D325FD89F9370F03F931" bold="true" box="[529,620,1693,1719]" pageId="7" pageNumber="158">
<taxonomicNameLabel id="0785CA03EF54D325FD89F9370F03F931" box="[529,620,1693,1719]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from most of
<taxonomicName id="E9C2D0E9EF54D325FC51F9360949F930" authorityName=", Lamarck" authorityYear="1801" box="[969,1062,1692,1718]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="1CB67778EF54D325FC51F9360949F930" box="[969,1062,1692,1718]" italics="true" pageId="7" pageNumber="158">Phrynus</emphasis>
</taxonomicName>
species by having only one conspicuous tooth in the ectal row on the base of chelicerae, with an acuminated apex (
<figureCitation id="B6F9B7EFEF54D325FBCAF96A09C6F95D" box="[1106,1193,1728,1755]" captionStart="FIGURE 4" captionStartId="8.[151,250,1964,1988]" captionTargetBox="[174,1415,459,1933]" captionTargetId="figure-192@8.[161,1427,453,1940]" captionTargetPageId="8" captionText="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="7" pageNumber="158">Fig. 4F</figureCitation>
). Does not exhibit an inconspicuous spine in the dorsomedial area of the pedipalp tarsus, Pd2 similar in size to Pd4, Fv3 shorter than Fv5, Td1 is very small with the same size as Td4, Tv3 shorter than Tv1 (
<figureCitation id="B6F9B7EFEF54D325FC0EF8A20EB4F8A5" box="[918,987,1800,1827]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="7" pageNumber="158">Fig. 5</figureCitation>
). The female genitalia have tubular and elongated gonopod bases, with sclerites of the gonopods claw-like, sticking out of the gonopod bases, and curved at the apical region, the sclerites are placed on the medial portion of the gonopod bases (
<figureCitation id="B6F9B7EFEF54D325FBCAF8FA09D8F8ED" box="[1106,1207,1872,1899]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="7" pageNumber="158">Fig. 17B</figureCitation>
). In the male genitalia, the LoL 2 present projections with widened bases, and the LoD thin (
<figureCitation id="B6F9B7EFEF54D325FC50F8DE0963F809" box="[968,1036,1908,1935]" captionStart="FIGURE 6" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[155,1432,855,1752]" captionTargetId="figure-323@11.[151,1437,849,1755]" captionTargetPageId="11" captionText="FIGURE 6. Male genitalia: Phrynus alfonsoi sp. nov. (CNAN-T01445): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620724" httpUri="https://zenodo.org/record/4620724/files/figure.png" pageId="7" pageNumber="158">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF54D32AFF5FF8320ED2FE28" blockId="7.[151,1437,1368,2043]" lastBlockId="8.[151,1437,151,430]" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158">
Species comparisons: The species
<taxonomicName id="E9C2D0E9EF54D325FDCCF8330FA3F834" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[596,716,1944,1970]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF54D325FDCCF8330FA3F834" box="[596,716,1944,1970]" italics="true" pageId="7" pageNumber="158">P. alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF54D325FD4EF8330E5DF835" bold="true" box="[726,818,1945,1971]" pageId="7" pageNumber="158">
<taxonomicNameLabel id="0785CA03EF54D325FD4EF8330E5DF835" box="[726,818,1945,1971]" pageId="7" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically similar to
<taxonomicName id="E9C2D0E9EF54D325FB09F8330995F834" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[1169,1274,1944,1970]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="jalisco">
<emphasis id="1CB67778EF54D325FB09F8330995F834" box="[1169,1274,1944,1970]" italics="true" pageId="7" pageNumber="158">P. jalisco</emphasis>
</taxonomicName>
, however, the female genitalia are different, in
<taxonomicName id="E9C2D0E9EF54D325FD9EF8170F03F850" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[518,620,1980,2006]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="jalisco">
<emphasis id="1CB67778EF54D325FD9EF8170F03F850" box="[518,620,1980,2006]" italics="true" pageId="7" pageNumber="158">P. jalisco</emphasis>
</taxonomicName>
the gonopod bases are more elongated, the sclerites are thicker and have a pronounced inward curvature, the sclerites are placed on the apex of the gonopod bases (
<figureCitation id="B6F9B7EFEF54D325FB0CF84A0998F87D" box="[1172,1271,2016,2043]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="7" pageNumber="158">Fig. 17F</figureCitation>
). These differences are also visible in the female genitalia of specimens of
<taxonomicName id="E9C2D0E9EF5BD32AFCDDFF3D0EC4FF37" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[837,939,151,177]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="jalisco">
<emphasis id="1CB67778EF5BD32AFCDDFF3D0EC4FF37" box="[837,939,151,177]" italics="true" pageId="8" pageNumber="159">P. jalisco</emphasis>
</taxonomicName>
from different localities (
<bibRefCitation id="4A53D69BEF5BD32AFB52FF3D08F7FF37" author="Armas, L. F. &amp; Guzman, A. &amp; Francke, O." box="[1226,1432,151,177]" pageId="8" pageNumber="159" pagination="3 - 7" refId="ref18743" refString="Armas, L. F., Guzman, A. &amp; Francke, O. (2014) Una especie nueva de Phrynus (Amblypygi: Phrynidae) de Mexico. Revista Iberica de Aracnologia, 25, 3 - 7." type="journal article" year="2014">
Armas
<emphasis id="1CB67778EF5BD32AFA85FF3D0839FF37" box="[1309,1366,151,177]" italics="true" pageId="8" pageNumber="159">et al.</emphasis>
2014
</bibRefCitation>
: fig. 3AC) in which the inward curvature is always present.
<taxonomicName id="E9C2D0E9EF5BD32AFCDEFF11096AFF53" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[838,1029,187,213]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF5BD32AFCDEFF11096AFF53" box="[838,1029,187,213]" italics="true" pageId="8" pageNumber="159">Phrynus alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF5BD32AFB96FF110907FF53" bold="true" box="[1038,1128,187,213]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="0785CA03EF5BD32AFB96FF110907FF53" box="[1038,1128,187,213]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also similar to
<taxonomicName id="E9C2D0E9EF5BD32AFAACFF1108F3FF53" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[1332,1436,187,213]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="1CB67778EF5BD32AFAACFF1108F3FF53" box="[1332,1436,187,213]" italics="true" pageId="8" pageNumber="159">P. aliciae</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF5BD32AFF0FFF750D9DFF7F" bold="true" box="[151,242,223,249]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="0785CA03EF5BD32AFF0FFF750D9DFF7F" box="[151,242,223,249]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, however, in this species, the base of sclerite of gonopod of the female genitalia is more straight and surrounding the lateral margin of the base (
<figureCitation id="B6F9B7EFEF5BD32AFDF8FEA90FAAFE9B" box="[608,709,259,285]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="8" pageNumber="159">Fig. 17C</figureCitation>
) in
<taxonomicName id="E9C2D0E9EF5BD32AFD6BFEA90E05FE9B" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[755,874,259,285]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF5BD32AFD6BFEA90E05FE9B" box="[755,874,259,285]" italics="true" pageId="8" pageNumber="159">P. alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF5BD32AFCEAFEA90EA2FE9B" bold="true" box="[882,973,259,285]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="0785CA03EF5BD32AFCEAFEA90EA2FE9B" box="[882,973,259,285]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the base is smaller.
<taxonomicName id="E9C2D0E9EF5BD32AFB2CFEA908F3FE9B" authorityName="Pocock" authorityYear="1902" box="[1204,1436,259,285]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF5BD32AFB2CFEA908F3FE9B" box="[1204,1436,259,285]" italics="true" pageId="8" pageNumber="159">Phrynus operculatus</emphasis>
</taxonomicName>
has a similar female genital but in this species, the sclerite of the female gonopod is smaller with a less pronounced curvature at the apical region (
<figureCitation id="B6F9B7EFEF5BD32AFE68FEE10F36FEE3" box="[496,601,331,357]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="8" pageNumber="159">Fig. 17A</figureCitation>
). In the male genitalia of
<emphasis id="1CB67778EF5BD32AFCE2FEE109D5FEE3" box="[890,1210,331,357]" italics="true" pageId="8" pageNumber="159">
<taxonomicName id="E9C2D0E9EF5BD32AFCE2FEE10976FEE3" authorityName="Pocock" authorityYear="1902" box="[890,1049,331,357]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="operculatus">P. operculatus</taxonomicName>
and
<taxonomicName id="E9C2D0E9EF5BD32AFBCAFEE109D5FEE3" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[1106,1210,331,357]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">P. aliciae</taxonomicName>
</emphasis>
<emphasis id="1CB67778EF5BD32AFB5AFEE10873FEE3" bold="true" box="[1218,1308,331,357]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="0785CA03EF5BD32AFB5AFEE10873FEE3" box="[1218,1308,331,357]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the projections on LoL 2 are tubular (
<figureCitation id="B6F9B7EFEF5BD32AFE53FEC50F4CFE0F" box="[459,547,367,393]" captionStart="FIGURE 3" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[156,1432,852,1747]" captionTargetId="figure-368@6.[151,1436,848,1751]" captionTargetPageId="6" captionText="FIGURE 3. Male genital: Phrynus operculatus (CASENT 9060512): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620718" httpUri="https://zenodo.org/record/4620718/files/figure.png" pageId="8" pageNumber="159">Fig. 3B</figureCitation>
,
<figureCitation id="B6F9B7EFEF5BD32AFDB6FEC50F3FFE0F" box="[558,592,367,393]" captionStart="FIGURE 9" captionStartId="16.[151,250,1626,1650]" captionTargetBox="[156,1431,816,1600]" captionTargetId="figure-348@16.[151,1435,813,1603]" captionTargetPageId="16" captionText="FIGURE 9. Male genitalia: Phrynus aliciae sp. nov. (CNAN-T01448): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620732" httpUri="https://zenodo.org/record/4620732/files/figure.png" pageId="8" pageNumber="159">9B</figureCitation>
) while in
<taxonomicName id="E9C2D0E9EF5BD32AFD5AFEC50E58FE0F" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[706,823,367,393]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF5BD32AFD5AFEC50E58FE0F" box="[706,823,367,393]" italics="true" pageId="8" pageNumber="159">P. alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF5BD32AFCA6FEC50EF7FE0F" bold="true" box="[830,920,367,393]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="0785CA03EF5BD32AFCA6FEC50EF7FE0F" box="[830,920,367,393]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the base of projection is wider. The LoD in
<taxonomicName id="E9C2D0E9EF5BD32AFA10FEC50C75FE2B" authorityName="Pocock" authorityYear="1902" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF5BD32AFA10FEC50C75FE2B" italics="true" pageId="8" pageNumber="159">P. operculatus</emphasis>
</taxonomicName>
is thicker than in
<taxonomicName id="E9C2D0E9EF5BD32AFE7CFE390F36FE2B" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[484,601,403,429]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF5BD32AFE7CFE390F36FE2B" box="[484,601,403,429]" italics="true" pageId="8" pageNumber="159">P. alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF5BD32AFDF8FE390FD5FE2B" bold="true" box="[608,698,403,429]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="0785CA03EF5BD32AFDF8FE390FD5FE2B" box="[608,698,403,429]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B6F9B7EFEF5BD32AFD51FE390E4FFE2B" box="[713,800,403,429]" captionStart="FIGURE 3" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[156,1432,852,1747]" captionTargetId="figure-368@6.[151,1436,848,1751]" captionTargetPageId="6" captionText="FIGURE 3. Male genital: Phrynus operculatus (CASENT 9060512): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620718" httpUri="https://zenodo.org/record/4620718/files/figure.png" pageId="8" pageNumber="159">Fig. 3C</figureCitation>
,
<figureCitation id="B6F9B7EFEF5BD32AFCB4FE390E20FE2B" box="[812,847,403,429]" captionStart="FIGURE 6" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[155,1432,855,1752]" captionTargetId="figure-323@11.[151,1437,849,1755]" captionTargetPageId="11" captionText="FIGURE 6. Male genitalia: Phrynus alfonsoi sp. nov. (CNAN-T01445): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620724" httpUri="https://zenodo.org/record/4620724/files/figure.png" pageId="8" pageNumber="159">6C</figureCitation>
,
<tableCitation id="63409ED1EF5BD32AFCC2FE390EC0FE28" box="[858,943,403,430]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="8" pageNumber="159">Table 6</tableCitation>
).
</paragraph>
<caption id="7ABDFBE2EF5BD32AFF0FF8060E6BF86E" ID-DOI="http://doi.org/10.5281/zenodo.4620720" ID-Zenodo-Dep="4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="8" pageNumber="159" startId="8.[151,250,1964,1988]" targetBox="[174,1415,459,1933]" targetPageId="8">
<paragraph id="2E7DAB6AEF5BD32AFF0FF8060E6BF86E" blockId="8.[151,1436,1964,2024]" pageId="8" pageNumber="159">
<emphasis id="1CB67778EF5BD32AFF0FF8060C7BF842" bold="true" box="[151,276,1964,1988]" pageId="8" pageNumber="159">FIGURE 4.</emphasis>
<taxonomicName id="E9C2D0E9EF5BD32AFE83F8060CA6F842" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[283,457,1964,1988]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF5BD32AFE83F8060CA6F842" box="[283,457,1964,1988]" italics="true" pageId="8" pageNumber="159">Phrynus alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF5BD32AFE48F8060F4DF842" bold="true" box="[464,546,1964,1988]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="0785CA03EF5BD32AFE48F8060F4DF842" box="[464,546,1964,1988]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view.
</paragraph>
</caption>
<paragraph id="2E7DAB6AEF5AD32BFF5FFF3D0EEDFF7F" blockId="9.[151,1437,150,501]" pageId="9" pageNumber="160">
The spine Pd1 is conspicuous, and frequently accompanied by a Pd1a and Pd1-a (
<figureCitation id="B6F9B7EFEF5AD32BFBF9FF3D09D6FF37" box="[1121,1209,150,177]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="9" pageNumber="160">Fig. 5D</figureCitation>
) that is similar in
<taxonomicName id="E9C2D0E9EF5AD32BFA10FF3D0D8CFF53" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="1CB67778EF5AD32BFA10FF3D0D8CFF53" italics="true" pageId="9" pageNumber="160">P. aliciae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0785CA03EF5AD32BFF72FF110C2CFF53" box="[234,323,187,213]" pageId="9" pageNumber="160" rank="species">
<emphasis id="1CB67778EF5AD32BFF72FF110C52FF53" bold="true" box="[234,317,187,213]" pageId="9" pageNumber="160">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="1CB67778EF5AD32BFEC9FF110CD2FF53" box="[337,445,187,213]" italics="true" pageId="9" pageNumber="160">
<taxonomicName id="E9C2D0E9EF5AD32BFEC9FF110CD6FF53" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[337,441,187,213]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="jalisco">P. jalisco</taxonomicName>
,
</emphasis>
and
<emphasis id="1CB67778EF5AD32BFE6AFF110FF8FF53" box="[498,663,187,213]" italics="true" pageId="9" pageNumber="160">
<taxonomicName id="E9C2D0E9EF5AD32BFE6AFF110FFBFF53" authorityName="Pocock" authorityYear="1902" box="[498,660,187,213]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="operculatus">P. operculatus</taxonomicName>
.
</emphasis>
Tv1 is longer than Tv3 (
<figureCitation id="B6F9B7EFEF5AD32BFC30FF110E93FF53" box="[936,1020,186,213]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="9" pageNumber="160">Fig. 5E</figureCitation>
) different from the other species here studied. Td1 is minute (
<figureCitation id="B6F9B7EFEF5AD32BFE39FF750C9BFF7F" box="[417,500,222,249]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="9" pageNumber="160">Fig. 5F</figureCitation>
) which differs from
<emphasis id="1CB67778EF5AD32BFD44FF750EEDFF7F" box="[732,898,223,249]" italics="true" pageId="9" pageNumber="160">
<taxonomicName id="E9C2D0E9EF5AD32BFD44FF750E11FF7F" authorityName="Pocock" authorityYear="1902" box="[732,894,223,249]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="operculatus">P. operculatus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF5AD32EFF5FFEA809E0FD48" lastPageId="12" lastPageNumber="163" pageId="9" pageNumber="160" type="description">
<paragraph id="2E7DAB6AEF5AD32BFF5FFEA80C98FEE3" blockId="9.[151,1437,150,501]" pageId="9" pageNumber="160">
<emphasis id="1CB67778EF5AD32BFF5FFEA80FE1FE9A" bold="true" box="[199,654,258,285]" pageId="9" pageNumber="160">Description. Female (CNAN-T01383):</emphasis>
Total length
<quantity id="E93A068FEF5AD32BFCBCFEA90EE4FE9B" box="[804,907,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="9" pageNumber="160" unit="mm" value="11.9">11.9 mm</quantity>
. Carapace and opisthosoma color brown, pedipalps and legs brown with red tonalities. Dimensions of prosoma, opisthosoma, pedipalps segments, and all leg femora are provided in
<tableCitation id="63409ED1EF5AD32BFE05FEE10C9CFEE3" box="[413,499,331,357]" captionStart="TABLE 2" captionStartId="9.[151,239,548,572]" captionText="TABLE 2. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus alfonsoi sp. nov. When the segment is missing, the cell is empty." pageId="9" pageNumber="160">Table 2</tableCitation>
.
</paragraph>
<paragraph id="2E7DAB6AEF5AD32BFF5FFEC50C25FE73" blockId="9.[151,1437,150,501]" pageId="9" pageNumber="160">
<emphasis id="1CB67778EF5AD32BFF5FFEC50C53FE0F" bold="true" box="[199,316,367,393]" italics="true" pageId="9" pageNumber="160">Carapace.</emphasis>
The frontal margin is almost straight, with one slightly pronounced lobe at each side, lobes are not joined in the middle area. Carapace presents a group of granules widely spaced; anterior, lateral, and posterior margins with a white border, posterior margin concave. Lateral and anteromedial eyes clearly visible, ocular tubercle black (
<figureCitation id="B6F9B7EFEF5AD32BFF7BFE710C51FE73" box="[227,318,475,501]" captionStart="FIGURE 4" captionStartId="8.[151,250,1964,1988]" captionTargetBox="[174,1415,459,1933]" captionTargetId="figure-192@8.[161,1427,453,1940]" captionTargetPageId="8" captionText="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="9" pageNumber="160">Fig. 4A</figureCitation>
).
</paragraph>
<caption id="7ABDFBE2EF5AD32BFF0FFD8E0F4AFDE7" pageId="9" pageNumber="160">
<paragraph id="2E7DAB6AEF5AD32BFF0FFD8E0F4AFDE7" blockId="9.[151,1437,548,609]" pageId="9" pageNumber="160">
<emphasis id="1CB67778EF5AD32BFF0FFD8E0C64FDBA" bold="true" box="[151,267,548,572]" pageId="9" pageNumber="160">TABLE 2.</emphasis>
Measurements (mm) of body, pedipalps segments, and femur of legs in
<taxonomicName id="E9C2D0E9EF5AD32BFB80FD8E09A1FDBA" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[1048,1230,548,572]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="9" pageNumber="160" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF5AD32BFB80FD8E09A1FDBA" box="[1048,1230,548,572]" italics="true" pageId="9" pageNumber="160">Phrynus alfonsoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0785CA03EF5AD32BFB4EFD8F0842FDBA" box="[1238,1325,549,573]" pageId="9" pageNumber="160" rank="species">
<emphasis id="1CB67778EF5AD32BFB4EFD8F0849FDBB" bold="true" box="[1238,1318,549,573]" pageId="9" pageNumber="160">sp. nov</emphasis>
.
</taxonomicNameLabel>
When the segment is missing, the cell is empty.
</paragraph>
</caption>
<paragraph id="2E7DAB6AEF5AD32BFD02FDDB085FF9CC" pageId="9" pageNumber="160">
<table id="5CC259CAEF5A2CDDFF3EFDDA0817F9CD" box="[166,1400,624,1611]" gridcols="4" gridrows="25" pageId="9" pageNumber="160">
<tr id="90F2A928EF5A2CDDFF3EFDDA0817FD0F" box="[166,1400,624,649]" gridrow="0" pageId="9" pageNumber="160" rowspan-0="1">
<th id="D323C054EF5A2CDDFDC2FDDA0E5FFD0F" box="[602,816,624,649]" gridcol="1" gridrow="0" pageId="9" pageNumber="160">Holotype</th>
<th id="D323C054EF5A2CDDFC3BFDDA0917FD0F" box="[931,1144,624,649]" gridcol="2" gridrow="0" pageId="9" pageNumber="160">CNAN-T01445</th>
<th id="D323C054EF5A2CDDFB42FDDA0817FD0F" box="[1242,1400,624,649]" gridcol="3" gridrow="0" pageId="9" pageNumber="160">CNAN-T01445</th>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFD3F0817FD2B" box="[166,1400,661,685]" gridrow="1" pageId="9" pageNumber="160" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="D323C054EF5A2CDDFDC2FD3F0E5FFD2B" box="[602,816,661,685]" gridcol="1" gridrow="1" pageId="9" pageNumber="160">CNAN-T01383</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFD170817FD53" box="[166,1400,701,725]" gridrow="2" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFD170CBFFD53" box="[166,464,701,725]" gridcol="0" gridrow="2" pageId="9" pageNumber="160">Sex</th>
<td id="D323C054EF5A2CDDFDC2FD170E5FFD53" box="[602,816,701,725]" gridcol="1" gridrow="2" pageId="9" pageNumber="160">Female</td>
<td id="D323C054EF5A2CDDFC3BFD170917FD53" box="[931,1144,701,725]" gridcol="2" gridrow="2" pageId="9" pageNumber="160">Male</td>
<td id="D323C054EF5A2CDDFB42FD170817FD53" box="[1242,1400,701,725]" gridcol="3" gridrow="2" pageId="9" pageNumber="160">Female</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFD4F0817FD7B" box="[166,1400,741,765]" gridrow="3" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFD4F0CBFFD7B" box="[166,464,741,765]" gridcol="0" gridrow="3" pageId="9" pageNumber="160">Total length (with chelicerae)</th>
<td id="D323C054EF5A2CDDFDC2FD4F0E5FFD7B" box="[602,816,741,765]" gridcol="1" gridrow="3" pageId="9" pageNumber="160">11.9</td>
<td id="D323C054EF5A2CDDFC3BFD4F0917FD7B" box="[931,1144,741,765]" gridcol="2" gridrow="3" pageId="9" pageNumber="160">12.7</td>
<td id="D323C054EF5A2CDDFB42FD4F0817FD7B" box="[1242,1400,741,765]" gridcol="3" gridrow="3" pageId="9" pageNumber="160">16.9</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFCA40817FCA3" box="[166,1400,782,805]" gridrow="4" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFCA40817FCA3" box="[166,1400,782,805]" colspan="4" colspanRight="3" gridcol="0" gridrow="4" pageId="9" pageNumber="160">Prosoma</th>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFC9C0817FCC8" box="[166,1400,822,846]" gridrow="5" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFC9C0CBFFCC8" box="[166,464,822,846]" gridcol="0" gridrow="5" pageId="9" pageNumber="160">Maximum width</th>
<td id="D323C054EF5A2CDDFDC2FC9C0E5FFCC8" box="[602,816,822,846]" gridcol="1" gridrow="5" pageId="9" pageNumber="160">6.3</td>
<td id="D323C054EF5A2CDDFC3BFC9C0917FCC8" box="[931,1144,822,846]" gridcol="2" gridrow="5" pageId="9" pageNumber="160">7.8</td>
<td id="D323C054EF5A2CDDFB42FC9C0817FCC8" box="[1242,1400,822,846]" gridcol="3" gridrow="5" pageId="9" pageNumber="160">8.4</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFCF40817FCF0" box="[166,1400,862,886]" gridrow="6" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFCF40CBFFCF0" box="[166,464,862,886]" gridcol="0" gridrow="6" pageId="9" pageNumber="160">Maximum length</th>
<td id="D323C054EF5A2CDDFDC2FCF40E5FFCF0" box="[602,816,862,886]" gridcol="1" gridrow="6" pageId="9" pageNumber="160">4.2</td>
<td id="D323C054EF5A2CDDFC3BFCF40917FCF0" box="[931,1144,862,886]" gridcol="2" gridrow="6" pageId="9" pageNumber="160">5.3</td>
<td id="D323C054EF5A2CDDFB42FCF40817FCF0" box="[1242,1400,862,886]" gridcol="3" gridrow="6" pageId="9" pageNumber="160">5.4</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFC2C0817FC18" box="[166,1400,902,926]" gridrow="7" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFC2C0817FC18" box="[166,1400,902,926]" colspan="4" colspanRight="3" gridcol="0" gridrow="7" pageId="9" pageNumber="160">Opisthosoma</th>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFC040817FC40" box="[166,1400,942,966]" gridrow="8" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFC040CBFFC40" box="[166,464,942,966]" gridcol="0" gridrow="8" pageId="9" pageNumber="160">Maximum width</th>
<td id="D323C054EF5A2CDDFDC2FC040E5FFC40" box="[602,816,942,966]" gridcol="1" gridrow="8" pageId="9" pageNumber="160">4.9</td>
<td id="D323C054EF5A2CDDFC3BFC040917FC40" box="[931,1144,942,966]" gridcol="2" gridrow="8" pageId="9" pageNumber="160">5.7</td>
<td id="D323C054EF5A2CDDFB42FC040817FC40" box="[1242,1400,942,966]" gridcol="3" gridrow="8" pageId="9" pageNumber="160">8</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFC7C0817FC69" box="[166,1400,982,1007]" gridrow="9" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFC7C0CBFFC69" box="[166,464,982,1007]" gridcol="0" gridrow="9" pageId="9" pageNumber="160">Maximum length</th>
<td id="D323C054EF5A2CDDFDC2FC7C0E5FFC69" box="[602,816,982,1007]" gridcol="1" gridrow="9" pageId="9" pageNumber="160">6.9</td>
<td id="D323C054EF5A2CDDFC3BFC7C0917FC69" box="[931,1144,982,1007]" gridcol="2" gridrow="9" pageId="9" pageNumber="160">7.2</td>
<td id="D323C054EF5A2CDDFB42FC7C0817FC69" box="[1242,1400,982,1007]" gridcol="3" gridrow="9" pageId="9" pageNumber="160">10.5</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFC550817FB91" box="[166,1400,1023,1047]" gridrow="10" pageId="9" pageNumber="160" rowspan-0="1">
<td id="D323C054EF5A2CDDFDC2FC550E5FFB91" box="[602,816,1023,1047]" gridcol="1" gridrow="10" pageId="9" pageNumber="160">Right Left</td>
<td id="D323C054EF5A2CDDFC3BFC550917FB91" box="[931,1144,1023,1047]" gridcol="2" gridrow="10" pageId="9" pageNumber="160">Right Left</td>
<td id="D323C054EF5A2CDDFB42FC550817FB91" box="[1242,1400,1023,1047]" gridcol="3" gridrow="10" pageId="9" pageNumber="160">Right</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFB8D0817FBB9" box="[166,1400,1063,1087]" gridrow="11" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFB8D0817FBB9" box="[166,1400,1063,1087]" colspan="4" colspanRight="3" gridcol="0" gridrow="11" pageId="9" pageNumber="160">Pedipalp lengths</th>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFBE50817FBE1" box="[166,1400,1103,1127]" gridrow="12" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFBE50CBFFBE1" box="[166,464,1103,1127]" gridcol="0" gridrow="12" pageId="9" pageNumber="160">Trochanter</th>
<td id="D323C054EF5A2CDDFDC2FBE50E5FFBE1" box="[602,816,1103,1127]" gridcol="1" gridrow="12" pageId="9" pageNumber="160">1.3 1.3</td>
<td id="D323C054EF5A2CDDFC3BFBE50917FBE1" box="[931,1144,1103,1127]" gridcol="2" gridrow="12" pageId="9" pageNumber="160">1.9 1.9</td>
<td id="D323C054EF5A2CDDFB42FBE50817FBE1" box="[1242,1400,1103,1127]" gridcol="3" gridrow="12" pageId="9" pageNumber="160">2.4</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFBDD0817FB16" box="[166,1400,1143,1168]" gridrow="13" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFBDD0CBFFB16" box="[166,464,1143,1168]" gridcol="0" gridrow="13" pageId="9" pageNumber="160">Femur</th>
<td id="D323C054EF5A2CDDFDC2FBDD0E5FFB16" box="[602,816,1143,1168]" gridcol="1" gridrow="13" pageId="9" pageNumber="160">3.7 5.3</td>
<td id="D323C054EF5A2CDDFC3BFBDD0917FB16" box="[931,1144,1143,1168]" gridcol="2" gridrow="13" pageId="9" pageNumber="160">5.3 5.6</td>
<td id="D323C054EF5A2CDDFB42FBDD0817FB16" box="[1242,1400,1143,1168]" gridcol="3" gridrow="13" pageId="9" pageNumber="160">5.5</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFB0A0817FB3E" box="[166,1400,1184,1208]" gridrow="14" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFB0A0CBFFB3E" box="[166,464,1184,1208]" gridcol="0" gridrow="14" pageId="9" pageNumber="160">
<taxonomicName id="E9C2D0E9EF5AD32BFF34FB0A0D9EFB3E" box="[172,241,1184,1208]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="9" pageNumber="176" phylum="Mollusca" rank="genus">Patella</taxonomicName>
</th>
<td id="D323C054EF5A2CDDFDC2FB0A0E5FFB3E" box="[602,816,1184,1208]" gridcol="1" gridrow="14" pageId="9" pageNumber="160">4.1 3.9</td>
<td id="D323C054EF5A2CDDFC3BFB0A0917FB3E" box="[931,1144,1184,1208]" gridcol="2" gridrow="14" pageId="9" pageNumber="160">5.3 5.3</td>
<td id="D323C054EF5A2CDDFB42FB0A0817FB3E" box="[1242,1400,1184,1208]" gridcol="3" gridrow="14" pageId="9" pageNumber="160">5.7</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFB620817FB66" box="[166,1400,1224,1248]" gridrow="15" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFB620CBFFB66" box="[166,464,1224,1248]" gridcol="0" gridrow="15" pageId="9" pageNumber="160">Tibia</th>
<td id="D323C054EF5A2CDDFDC2FB620E5FFB66" box="[602,816,1224,1248]" gridcol="1" gridrow="15" pageId="9" pageNumber="160">1.9 1.9</td>
<td id="D323C054EF5A2CDDFC3BFB620917FB66" box="[931,1144,1224,1248]" gridcol="2" gridrow="15" pageId="9" pageNumber="160">2.6 2.4</td>
<td id="D323C054EF5A2CDDFB42FB620817FB66" box="[1242,1400,1224,1248]" gridcol="3" gridrow="15" pageId="9" pageNumber="160">2.9</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFB5A0817FA8E" box="[166,1400,1264,1288]" gridrow="16" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFB5A0CBFFA8E" box="[166,464,1264,1288]" gridcol="0" gridrow="16" pageId="9" pageNumber="160">Tarsus-metatarsus</th>
<td id="D323C054EF5A2CDDFDC2FB5A0E5FFA8E" box="[602,816,1264,1288]" gridcol="1" gridrow="16" pageId="9" pageNumber="160">2.2 2.2</td>
<td id="D323C054EF5A2CDDFC3BFB5A0917FA8E" box="[931,1144,1264,1288]" gridcol="2" gridrow="16" pageId="9" pageNumber="160">2.3 2.3</td>
<td id="D323C054EF5A2CDDFB42FB5A0817FA8E" box="[1242,1400,1264,1288]" gridcol="3" gridrow="16" pageId="9" pageNumber="160">2.9</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFAB30817FAB7" box="[166,1400,1305,1329]" gridrow="17" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFAB30817FAB7" box="[166,1400,1305,1329]" colspan="4" colspanRight="3" gridcol="0" gridrow="17" pageId="9" pageNumber="160">Femur length</th>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFAEB0817FADF" box="[166,1400,1345,1369]" gridrow="18" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFAEB0CBFFADF" box="[166,464,1345,1369]" gridcol="0" gridrow="18" pageId="9" pageNumber="160">Leg I</th>
<td id="D323C054EF5A2CDDFDC2FAEB0E5FFADF" box="[602,816,1345,1369]" gridcol="1" gridrow="18" pageId="9" pageNumber="160">7.9 7.9</td>
<td id="D323C054EF5A2CDDFC3BFAEB0917FADF" box="[931,1144,1345,1369]" gridcol="2" gridrow="18" pageId="9" pageNumber="160">11.7 11.9</td>
<td id="D323C054EF5A2CDDFB42FAEB0817FADF" box="[1242,1400,1345,1369]" gridcol="3" gridrow="18" pageId="9" pageNumber="160">12.7</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFAC30817FA04" box="[166,1400,1385,1410]" gridrow="19" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFAC30CBFFA04" box="[166,464,1385,1410]" gridcol="0" gridrow="19" pageId="9" pageNumber="160">Leg II</th>
<td id="D323C054EF5A2CDDFDC2FAC30E5FFA04" box="[602,816,1385,1410]" gridcol="1" gridrow="19" pageId="9" pageNumber="160">6.4 6.4</td>
<td id="D323C054EF5A2CDDFC3BFAC30917FA04" box="[931,1144,1385,1410]" gridcol="2" gridrow="19" pageId="9" pageNumber="160">8.2 8.2</td>
<td id="D323C054EF5A2CDDFB42FAC30817FA04" box="[1242,1400,1385,1410]" gridcol="3" gridrow="19" pageId="9" pageNumber="160">8.1</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFA3B0817FA2C" box="[166,1400,1425,1450]" gridrow="20" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFA3B0CBFFA2C" box="[166,464,1425,1450]" gridcol="0" gridrow="20" pageId="9" pageNumber="160">Leg III</th>
<td id="D323C054EF5A2CDDFDC2FA3B0E5FFA2C" box="[602,816,1425,1450]" gridcol="1" gridrow="20" pageId="9" pageNumber="160">7.2 7</td>
<td id="D323C054EF5A2CDDFC3BFA3B0917FA2C" box="[931,1144,1425,1450]" gridcol="2" gridrow="20" pageId="9" pageNumber="160">9.1 9.1</td>
<td id="D323C054EF5A2CDDFB42FA3B0817FA2C" box="[1242,1400,1425,1450]" gridcol="3" gridrow="20" pageId="9" pageNumber="160">9.2</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFA130817FA54" box="[166,1400,1465,1490]" gridrow="21" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFA130CBFFA54" box="[166,464,1465,1490]" gridcol="0" gridrow="21" pageId="9" pageNumber="160">Leg IV</th>
<td id="D323C054EF5A2CDDFDC2FA130E5FFA54" box="[602,816,1465,1490]" gridcol="1" gridrow="21" pageId="9" pageNumber="160">6.1 6.1</td>
<td id="D323C054EF5A2CDDFC3BFA130917FA54" box="[931,1144,1465,1490]" gridcol="2" gridrow="21" pageId="9" pageNumber="160">8.2 8</td>
<td id="D323C054EF5A2CDDFB42FA130817FA54" box="[1242,1400,1465,1490]" gridcol="3" gridrow="21" pageId="9" pageNumber="160">1.9</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EFA480817FA7C" box="[166,1400,1506,1530]" gridrow="22" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EFA480CBFFA7C" box="[166,464,1506,1530]" gridcol="0" gridrow="22" pageId="9" pageNumber="160">No. segments tibia I</th>
<td id="D323C054EF5A2CDDFDC2FA480E5FFA7C" box="[602,816,1506,1530]" gridcol="1" gridrow="22" pageId="9" pageNumber="160">30</td>
<td id="D323C054EF5A2CDDFC3BFA480917FA7C" box="[931,1144,1506,1530]" gridcol="2" gridrow="22" pageId="9" pageNumber="160">30</td>
<td id="D323C054EF5A2CDDFB42FA480817FA7C" box="[1242,1400,1506,1530]" gridcol="3" gridrow="22" pageId="9" pageNumber="160">30</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EF9A00817F9A4" box="[166,1400,1546,1570]" gridrow="23" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EF9A00CBFF9A4" box="[166,464,1546,1570]" gridcol="0" gridrow="23" pageId="9" pageNumber="160">No. segments tarsus I</th>
<td id="D323C054EF5A2CDDFDC2F9A00E5FF9A4" box="[602,816,1546,1570]" gridcol="1" gridrow="23" pageId="9" pageNumber="160">58</td>
<td id="D323C054EF5A2CDDFC3BF9A00917F9A4" box="[931,1144,1546,1570]" gridcol="2" gridrow="23" pageId="9" pageNumber="160">64</td>
<td id="D323C054EF5A2CDDFB42F9A00817F9A4" box="[1242,1400,1546,1570]" gridcol="3" gridrow="23" pageId="9" pageNumber="160">64</td>
</tr>
<tr id="90F2A928EF5A2CDDFF3EF9980817F9CD" box="[166,1400,1586,1611]" gridrow="24" pageId="9" pageNumber="160">
<th id="D323C054EF5A2CDDFF3EF9980CBFF9CD" box="[166,464,1586,1611]" gridcol="0" gridrow="24" pageId="9" pageNumber="160">No. segments basitibia IV</th>
<td id="D323C054EF5A2CDDFDC2F9980E5FF9CD" box="[602,816,1586,1611]" gridcol="1" gridrow="24" pageId="9" pageNumber="160">3 3</td>
<td id="D323C054EF5A2CDDFC3BF9980917F9CD" box="[931,1144,1586,1611]" gridcol="2" gridrow="24" pageId="9" pageNumber="160">3 3</td>
<td id="D323C054EF5A2CDDFB42F9980817F9CD" box="[1242,1400,1586,1611]" gridcol="3" gridrow="24" pageId="9" pageNumber="160">3</td>
</tr>
</table>
</paragraph>
<paragraph id="2E7DAB6AEF5AD32BFF5FF9DE08E0F899" blockId="9.[151,1437,1652,2039]" pageId="9" pageNumber="160">
<emphasis id="1CB67778EF5AD32BFF5FF9DE0C5EF908" bold="true" box="[199,305,1652,1678]" italics="true" pageId="9" pageNumber="160">Sternum.</emphasis>
Tri-segmented; all segments clearly sclerotized, the area around segments is a little less sclerotized. Tritosternum projected anteriorly, elongated, conical, with six setae on the basal region, four on the medial region, three on the distal region, and two more on the apex. Second segment (tetrasternum) oval with two setae on the medial region. Third segment (pentasternum) oval, slightly smaller than the second segment, with two setae on the medial region. Metasternum longitudinally divided, with two setae on the posterior margin of each half (
<figureCitation id="B6F9B7EFEF5AD32BFAB2F8AE08EDF899" box="[1322,1410,1796,1823]" captionStart="FIGURE 4" captionStartId="8.[151,250,1964,1988]" captionTargetBox="[174,1415,459,1933]" captionTargetId="figure-192@8.[161,1427,453,1940]" captionTargetPageId="8" captionText="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="9" pageNumber="160">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF5AD32BFF5FF8820FF3F8E1" blockId="9.[151,1437,1652,2039]" pageId="9" pageNumber="160">
<emphasis id="1CB67778EF5AD32BFF5FF8820C53F8C5" bold="true" box="[199,316,1832,1859]" italics="true" pageId="9" pageNumber="160">Abdomen.</emphasis>
Oblong, color dark brown. Presents patches of light coloration on both sides of each tergite. The carapace is wider than the abdomen (
<figureCitation id="B6F9B7EFEF5AD32BFDAFF8E60FFFF8E1" box="[567,656,1868,1895]" captionStart="FIGURE 4" captionStartId="8.[151,250,1964,1988]" captionTargetBox="[174,1415,459,1933]" captionTargetId="figure-192@8.[161,1427,453,1940]" captionTargetPageId="8" captionText="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="9" pageNumber="160">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF5AD328FF5FF8DA0CDDFF53" blockId="9.[151,1437,1652,2039]" lastBlockId="10.[151,1436,151,213]" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160">
<emphasis id="1CB67778EF5AD32BFF5FF8DA0C51F80D" bold="true" box="[199,318,1904,1931]" italics="true" pageId="9" pageNumber="160">Chelicera.</emphasis>
The mesal row of the basal segment of chelicerae with three teeth, the first is bilobed, placed in the proximal portion, lobe 1b bigger than 1a, followed by one tooth shorter in the medial portion, and the third, bigger than the others, placed in the distal region (
<figureCitation id="B6F9B7EFEF5AD32BFD1EF8120F8CF855" box="[646,739,1976,2003]" captionStart="FIGURE 4" captionStartId="8.[151,250,1964,1988]" captionTargetBox="[174,1415,459,1933]" captionTargetId="figure-192@8.[161,1427,453,1940]" captionTargetPageId="8" captionText="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="9" pageNumber="160">Fig. 4D</figureCitation>
). The ectal row with one conspicuous tooth (tooth 2) with an acuminated apex, placed in the medial region. There is a small keel on the most proximal region (tooth 1) (
<figureCitation id="B6F9B7EFEF5AD32BFAA2F87608FFF871" box="[1338,1424,2012,2039]" captionStart="FIGURE 4" captionStartId="8.[151,250,1964,1988]" captionTargetBox="[174,1415,459,1933]" captionTargetId="figure-192@8.[161,1427,453,1940]" captionTargetPageId="8" captionText="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="9" pageNumber="160">Fig. 4F</figureCitation>
). The mobile segment of chelicerae with four teeth, the first is the biggest, the second and third are similar in size, and the fourth is the smallest.
</paragraph>
<caption id="7ABDFBE2EF59D328FF0FF9D80E32F928" ID-DOI="http://doi.org/10.5281/zenodo.4620722" ID-Zenodo-Dep="4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="10" pageNumber="161" startId="10.[151,250,1650,1674]" targetBox="[162,1425,246,1606]" targetPageId="10">
<paragraph id="2E7DAB6AEF59D328FF0FF9D80E32F928" blockId="10.[151,1437,1650,1710]" pageId="10" pageNumber="161">
<emphasis id="1CB67778EF59D328FF0FF9D80C7DF90C" bold="true" box="[151,274,1650,1674]" pageId="10" pageNumber="161">FIGURE 5.</emphasis>
<taxonomicName id="E9C2D0E9EF59D328FE80F9D80CABF90C" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[280,452,1650,1674]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF59D328FE80F9D80CABF90C" box="[280,452,1650,1674]" italics="true" pageId="10" pageNumber="161">Phrynus alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF59D328FE51F9D80F77F90C" bold="true" box="[457,536,1650,1674]" pageId="10" pageNumber="161">
<taxonomicNameLabel id="0785CA03EF59D328FE51F9D80F77F90C" box="[457,536,1650,1674]" pageId="10" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral
<taxonomicName id="E9C2D0E9EF59D328FAC1F9D80DA9F928" authority="(Pv)" baseAuthorityName="Pv" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="10" pageNumber="176" phylum="Mollusca" rank="species" species="patella">patella (Pv)</taxonomicName>
. D. Dorsal
<taxonomicName id="E9C2D0E9EF59D328FEA0F93C0CDEF928" authority="(Pd)" baseAuthorityName="Pd" box="[312,433,1686,1710]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="10" pageNumber="176" phylum="Mollusca" rank="species" species="patella">patella (Pd)</taxonomicName>
. E. Ventral tibia (Tv). F. Dorsal tibia (Td).
</paragraph>
</caption>
<paragraph id="2E7DAB6AEF59D329FF5FF9760FDFFD9F" blockId="10.[151,1437,1756,2035]" lastBlockId="11.[151,1437,151,826]" lastPageId="11" lastPageNumber="162" pageId="10" pageNumber="161">
<emphasis id="1CB67778EF59D328FF5FF9760CD7F970" box="[199,440,1756,1783]" italics="true" pageId="10" pageNumber="161">
<emphasis id="1CB67778EF59D328FF5FF9760C5CF971" bold="true" box="[199,307,1756,1783]" italics="true" pageId="10" pageNumber="161">Pedipalp.</emphasis>
Trochanter
</emphasis>
: Prolateral face with four spines; spines Tr1 and Tr3 placed in the medial region, Tr2 placed near the ventral margin. Tr3 and Tr4 subequal in length. Spine lengths: Tr2&gt;Tr1&gt;Tr4ŻTr3. Dorsal oblique series of five setiferous tubercles. Dorsomedial area without spines but with one big tubercle (
<figureCitation id="B6F9B7EFEF59D328FB42F88E085BF8B8" box="[1242,1332,1828,1854]" captionStart="FIGURE 4" captionStartId="8.[151,250,1964,1988]" captionTargetBox="[174,1415,459,1933]" captionTargetId="figure-192@8.[161,1427,453,1940]" captionTargetPageId="8" captionText="FIGURE 4. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620720" httpUri="https://zenodo.org/record/4620720/files/figure.png" pageId="10" pageNumber="161">Fig. 4E</figureCitation>
).
<emphasis id="1CB67778EF59D328FAD2F88F08FAF8B8" box="[1354,1429,1829,1854]" italics="true" pageId="10" pageNumber="161">Femur</emphasis>
: Ventral face with six major spines, Fv4, Fv3a, Fv4a, and Fv5a are small, and between Fv5 and Fv5a there are three tubercles, Fv8 small. Spine lengths: Fv1=Fv2&gt;Fv5&gt;Fv6&gt;Fv3&gt;Fv7&gt;Fv4&gt;Fv5a&gt;Fv3a=Fv4a=Fv8 (
<figureCitation id="B6F9B7EFEF59D328FB7BF8C60851F801" box="[1251,1342,1900,1927]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="10" pageNumber="161">Fig. 5A</figureCitation>
). Dorsal face with five major spines, Fd4 small with one tubercle on each side, there is a small spine Fd5a, one tubercle between Fd5Fd5a and Fd6Fd7, with a small spine Fd8, Fd1 and Fd2 share the same base, Fd3 is separated from Fd2. Spine lengths: Fd2&gt;Fd3&gt;Fd5&gt;Fd6&gt;Fd1&gt;Fd7&gt;Fd4=Fd5a&gt;Fd8 (
<figureCitation id="B6F9B7EFEF59D328FC00F8720E9FF875" box="[920,1008,2008,2035]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="10" pageNumber="161">Fig. 5B</figureCitation>
).
<taxonomicName id="E9C2D0E9EF59D328FB9CF87209DBF874" authority=": Ventral" authorityName="Ventral" box="[1028,1204,2008,2035]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="10" pageNumber="161" phylum="Mollusca" rank="genus">
<emphasis id="1CB67778EF59D328FB9CF872093AF874" box="[1028,1109,2008,2034]" italics="true" pageId="10" pageNumber="161">Patella</emphasis>
: Ventral
</taxonomicName>
face with five major spines. With a conspicuous spine Pv1-a and one very small spine at each side, one conspicuous spine Pv1a, one very small spine between Pv1Pv1a, there is a small spine between Pv2Pv3, Pv3Pv4, Pv4Pv5, and Pv5Pv6. Spine lengths: Pv2&gt;Pv5&gt;Pv1&gt;Pv4&gt;Pv6&gt;Pv7&gt;Pv1-a=Pv1a&gt;Pv3 (
<figureCitation id="B6F9B7EFEF58D329FCB6FF750EE8FF7F" box="[814,903,222,249]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="11" pageNumber="162">Fig. 5C</figureCitation>
). Dorsal face with seven major spines, with one small spine Pd1a and Pd1-a, Pd8 is absent. Spine lengths: Pd5&gt;Pd3&gt;Pd2&gt;Pd4&gt;Pd6&gt;Pd1&gt;Pd7&gt;Pd1-a=Pd1a (
<figureCitation id="B6F9B7EFEF58D329FAF7FEA90DD3FEC7" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="11" pageNumber="162">Fig. 5D</figureCitation>
).
<emphasis id="1CB67778EF58D329FF4AFE8D0C63FEC7" box="[210,268,295,321]" italics="true" pageId="11" pageNumber="162">Tibia</emphasis>
: Ventral face with three major spines, there is one tubercle between Tv1Tv2, and one small spine between Tv2Tv3. Spine lengths: Tv2&gt;Tv1&gt;Tv3 (
<figureCitation id="B6F9B7EFEF58D329FD58FEE10E76FEE3" box="[704,793,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="11" pageNumber="162">Fig. 5E</figureCitation>
). Dorsal face with two major spines, between Td2Td3 there is one tubercle, over Td3 there is one conspicuous spine, distal to Td3 there are two small spines, the most distal spine is a little bigger and here is named Td4, Td1 has less than a third of length of Td3, and is placed almost on the base of Td2. Spine lengths: Td2&gt;Td3&gt;Td1=Td4 (
<figureCitation id="B6F9B7EFEF58D329FC91FE1D0E31FE57" box="[777,862,438,465]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="11" pageNumber="162">Fig. 5F</figureCitation>
).
<emphasis id="1CB67778EF58D329FCE9FE1D0927FE56" box="[881,1096,439,464]" italics="true" pageId="11" pageNumber="162">Tarsus-metatarsus:</emphasis>
Internal face with two lines of dorsomedial bristles; suture between the tarsus and metatarsus is not visible. Tarsus without an inconspicuous spine on the proximal end of the dorsomedial surface.
</paragraph>
<paragraph id="2E7DAB6AEF58D329FF5FFD890921FD03" blockId="11.[151,1437,151,826]" pageId="11" pageNumber="162">
<emphasis id="1CB67778EF58D329FF5FFD890C6AFDBB" bold="true" box="[199,261,547,573]" pageId="11" pageNumber="162">
<emphasis id="1CB67778EF58D329FF5FFD890D92FDBB" bold="true" box="[199,253,547,573]" italics="true" pageId="11" pageNumber="162">Legs</emphasis>
.
</emphasis>
Femora brown. Femora lengths: I&gt;III&gt;II&gt;IV (
<tableCitation id="63409ED1EF58D329FC95FD890E0AFDBB" box="[781,869,547,573]" captionStart="TABLE 2" captionStartId="9.[151,239,548,572]" captionText="TABLE 2. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus alfonsoi sp. nov. When the segment is missing, the cell is empty." pageId="11" pageNumber="162">Table 2</tableCitation>
). Leg I: Tibia with 30 segments and tarsus with 58 segments in the right leg; the left leg is missing. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT1&gt;DT&gt;BT3&gt;BT2. Basitarsus and Telotarsus subequal in size. Tarsus tetramerous.
</paragraph>
<paragraph id="2E7DAB6AEF58D329FF5FFD240F9EFCBC" blockId="11.[151,1437,151,826]" pageId="11" pageNumber="162">
<emphasis id="1CB67778EF58D329FF5FFD240CFDFD2F" bold="true" box="[199,402,654,681]" italics="true" pageId="11" pageNumber="162">Female genitalia.</emphasis>
Genital operculum pentagonal, it extends to the second segment of the opisthosoma. Gonopod bases cylindrical, dorsal surface poorly sclerotized; on the most proximal area, there is a strongly sclerotized line, joining both bases, the coloration is brown on the most proximal region and white overall dorsal surface. The sclerites of gonopods claw-like, the base is more than three times wider than the medial and apical regions, apex curved. Sclerite coloration brown (
<figureCitation id="B6F9B7EFEF58D329FD85FCB50FECFCBF" box="[541,643,799,825]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="11" pageNumber="162">Fig. 17B</figureCitation>
,
<tableCitation id="63409ED1EF58D329FD16FCB50F8CFCBC" box="[654,739,799,826]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="11" pageNumber="162">Table 6</tableCitation>
).
</paragraph>
<caption id="7ABDFBE2EF58D329FF0FF9590E74F8F1" ID-DOI="http://doi.org/10.5281/zenodo.4620724" ID-Zenodo-Dep="4620724" httpUri="https://zenodo.org/record/4620724/files/figure.png" pageId="11" pageNumber="162" startId="11.[151,250,1779,1803]" targetBox="[155,1432,855,1752]" targetPageId="11">
<paragraph id="2E7DAB6AEF58D329FF0FF9590E74F8F1" blockId="11.[151,1437,1779,1911]" pageId="11" pageNumber="162">
<emphasis id="1CB67778EF58D329FF0FF9590C7CF88D" bold="true" box="[151,275,1779,1803]" pageId="11" pageNumber="162">FIGURE 6.</emphasis>
Male genitalia:
<taxonomicName id="E9C2D0E9EF58D329FE22F9590F08F88D" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[442,615,1779,1803]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="11" pageNumber="162" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF58D329FE22F9590F08F88D" box="[442,615,1779,1803]" italics="true" pageId="11" pageNumber="162">Phrynus alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF58D329FDF5F9590FD1F88D" bold="true" box="[621,702,1779,1803]" pageId="11" pageNumber="162">
<taxonomicNameLabel id="0785CA03EF58D329FDF5F9590FD1F88D" box="[621,702,1779,1803]" pageId="11" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(CNAN-T01445): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line).
</paragraph>
</caption>
<paragraph id="2E7DAB6AEF58D32EFF5FF80F0842FE73" blockId="11.[151,1437,1957,2020]" lastBlockId="12.[151,1437,150,718]" lastPageId="12" lastPageNumber="163" pageId="11" pageNumber="162">
<emphasis id="1CB67778EF58D329FF5FF80F0C50F839" bold="true" box="[199,319,1957,1983]" pageId="11" pageNumber="162">Variation.</emphasis>
Type series measurements and the number of segments in the basitibia IV, Tibia I and tarsus I, are summarized in
<tableCitation id="63409ED1EF58D329FEDCF8630CF5F865" box="[324,410,1993,2019]" captionStart="TABLE 2" captionStartId="9.[151,239,548,572]" captionText="TABLE 2. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus alfonsoi sp. nov. When the segment is missing, the cell is empty." pageId="11" pageNumber="162">Table 2</tableCitation>
.
<emphasis id="1CB67778EF58D329FE3DF8630F7DF865" bold="true" box="[421,530,1993,2019]" pageId="11" pageNumber="162">
<emphasis id="1CB67778EF58D329FE3DF8630F66F865" bold="true" box="[421,521,1993,2019]" italics="true" pageId="11" pageNumber="162">Sternum</emphasis>
:
</emphasis>
The number and position of setae from all segments of the sternum are variable.
<emphasis id="1CB67778EF5FD32EFF0FFF3C0CE8FF37" box="[151,391,150,177]" italics="true" pageId="12" pageNumber="163">
<emphasis id="1CB67778EF5FD32EFF0FFF3C0C69FF37" bold="true" box="[151,262,150,177]" italics="true" pageId="12" pageNumber="163">Pedipalp:</emphasis>
Trochanter
</emphasis>
: line of dorsal tubercles varies between four and six tubercles.
<emphasis id="1CB67778EF5FD32EFBD2FF3D09FAFF36" box="[1098,1173,151,176]" italics="true" pageId="12" pageNumber="163">Femur</emphasis>
: Ventral face: tubercles are constant in most individuals however the size is variable, as well as the size of spines Fv4, Fv3a, Fv4a, and Fv5a, which are always smaller than other spines but the relative sizes between them are variable. Dorsal face: the size and presence of tubercles mentioned in the
<typeStatus id="F17915C8EF5FD32EFDD2FEA90FC3FE9B" box="[586,684,259,285]" pageId="12" pageNumber="163" type="holotype">holotype</typeStatus>
are variable.
<taxonomicName id="E9C2D0E9EF5FD32EFCDAFEA90E9DFE9B" authority=": Ventral" authorityName="Ventral" box="[834,1010,259,285]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="12" pageNumber="163" phylum="Mollusca" rank="genus">
<emphasis id="1CB67778EF5FD32EFCDAFEA90EFCFE9B" box="[834,915,259,285]" italics="true" pageId="12" pageNumber="163">Patella</emphasis>
: Ventral
</taxonomicName>
face: Tubercles and small spines were more conspicuous in larger sized individuals. Dorsal face: tubercles can be more conspicuous, in some individuals can be present tubercles between each spine, in larger specimens can be present a small spine Pd8.
<emphasis id="1CB67778EF5FD32EFA9AFEE10853FEE3" box="[1282,1340,331,357]" italics="true" pageId="12" pageNumber="163">Tibia</emphasis>
: Ventral face: between Tv2Tv3 can be present two or three small spines. Dorsal face: tubercles between spines can be more conspicuous.
<emphasis id="1CB67778EF5FD32EFEB7FE390C02FE2B" bold="true" box="[303,365,403,429]" italics="true" pageId="12" pageNumber="163">Legs.</emphasis>
The number of segments of the tibia and tarsus of leg I, was variable among the individuals, the number of tibial segments fixed at 30 and the tarsal segments between 58 and 64. In the observed specimens, the number of segments did not depend on sex, there are more tarsal segments in larger specimens (
<tableCitation id="63409ED1EF5FD32EFB50FE71084FFE73" box="[1224,1312,475,501]" captionStart="TABLE 2" captionStartId="9.[151,239,548,572]" captionText="TABLE 2. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus alfonsoi sp. nov. When the segment is missing, the cell is empty." pageId="12" pageNumber="163">Table 2</tableCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF5FD32EFF5FFE550934FDBB" blockId="12.[151,1437,150,718]" pageId="12" pageNumber="163">
<emphasis id="1CB67778EF5FD32EFF5FFE550C64FD9F" bold="true" box="[199,267,511,537]" pageId="12" pageNumber="163">Male.</emphasis>
Observed males are similar in size to females (
<tableCitation id="63409ED1EF5FD32EFC8AFE550E08FD9F" box="[786,871,511,537]" captionStart="TABLE 2" captionStartId="9.[151,239,548,572]" captionText="TABLE 2. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus alfonsoi sp. nov. When the segment is missing, the cell is empty." pageId="12" pageNumber="163">Table 2</tableCitation>
) spination pattern as in females, genital operculum has the posterior margin oval and covers part of the third segment of the opisthosoma.
</paragraph>
<paragraph id="2E7DAB6AEF5FD32EFF5FFDEC09E0FD48" blockId="12.[151,1437,150,718]" pageId="12" pageNumber="163">
<emphasis id="1CB67778EF5FD32EFF5FFDEC0F24FDE6" bold="true" box="[199,587,582,609]" pageId="12" pageNumber="163">
<emphasis id="1CB67778EF5FD32EFF5FFDEC0C01FDE7" bold="true" box="[199,366,582,609]" italics="true" pageId="12" pageNumber="163">Male genitalia</emphasis>
. (CNAN-T01445):
</emphasis>
LaM longer than Pi. LoL 1 a little longer than LoL 2, both densely covered with minute projections, in LoL 1, projections are tubular, thin, and elongated, placed close together, the insertion base is not visible; projections in LoL 2 are similar to LoL 1, but with a slightly broader base, the insertion base is partially visible. In dorsal view, the LoD elongated but shorter than LoL1 (
<figureCitation id="B6F9B7EFEF5FD32EFC44FD19094EFD4B" box="[988,1057,691,717]" captionStart="FIGURE 6" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[155,1432,855,1752]" captionTargetId="figure-323@11.[151,1437,849,1755]" captionTargetPageId="11" captionText="FIGURE 6. Male genitalia: Phrynus alfonsoi sp. nov. (CNAN-T01445): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620724" httpUri="https://zenodo.org/record/4620724/files/figure.png" pageId="12" pageNumber="163">Fig. 6</figureCitation>
,
<tableCitation id="63409ED1EF5FD32EFBB4FD1909EEFD48" box="[1068,1153,691,718]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="12" pageNumber="163">Table 6</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>