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<document id="FC710BAD9691038229E69604B7CD6C3A" ID-CLB-Dataset="272018" ID-DOI="10.11646/zootaxa.5353.4.6" ID-GBIF-Dataset="ab136eda-cf71-4c24-9665-83fad102a156" ID-ISSN="1175-5326" ID-Zenodo-Dep="8430810" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1697020763370" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Song, Xiao-Bin &amp; Chen, Jia-Wei" docDate="2023" docId="A6632C6DF975140D149DFB7A0EBAFE7A" docLanguage="en" docName="zootaxa.5353.4.6.pdf" docOrigin="Zootaxa 5353 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colilodion tetramerus Lobl 1998" docType="treatment" docVersion="2" lastPageNumber="394" masterDocId="5A5A5415F975140C140AFFCA0C6BFFD6" masterDocTitle="A second specimen of Colilodion tetramerus Löbl (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="394" masterPageNumber="393" pageNumber="393" updateTime="1697021414249" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="44FE083C1288940DAC56D9BAD8B99F2C">A second specimen of Colilodion tetramerus Löbl (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="0B97D50A4CB3DD38124FF76506B684B9">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="1608B2F496C91A78B470567BEFA272A3">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China</mods:affiliation>
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<mods:namePart id="FA0ACDC32BBFB8ED95EF4E6D4ADD0360">Song, Xiao-Bin</mods:namePart>
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<mods:namePart id="991FAC5D588E66CA87AA67B97F791B92">Chen, Jia-Wei</mods:namePart>
<mods:affiliation id="3CDD493B846C9D82E891ED0EAE751A38">Diaoluoshan Branch, Hainan Tropical Rainforest National Park Administration, 572400 Lingshui County, Hainan, China</mods:affiliation>
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<mods:title id="B45B8AB0070737384A567B7871E5A58D">Zootaxa</mods:title>
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<mods:date id="1EE34BC13CA7C4966670B04F834612AE">2023</mods:date>
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<treatment id="A6632C6DF975140D149DFB7A0EBAFE7A" ID-DOI="http://doi.org/10.5281/zenodo.8430803" ID-Zenodo-Dep="8430803" LSID="urn:lsid:plazi:treatment:A6632C6DF975140D149DFB7A0EBAFE7A" httpUri="http://treatment.plazi.org/id/A6632C6DF975140D149DFB7A0EBAFE7A" lastPageId="1" lastPageNumber="394" pageId="0" pageNumber="393">
<subSubSection id="66D0CEF0F975140C149DFB7A0EE7FB1C" box="[151,652,1200,1227]" pageId="0" pageNumber="393" type="nomenclature">
<paragraph id="2E759D7BF975140C149DFB7A0EE7FB1C" blockId="0.[151,1165,1200,1258]" box="[151,652,1200,1227]" pageId="0" pageNumber="393">
<heading id="753D2A17F975140C149DFB7A0EE7FB1C" bold="true" box="[151,652,1200,1227]" fontSize="11" level="1" pageId="0" pageNumber="393" reason="1">
<emphasis id="1CBE4169F975140C149DFB7A0EE7FB1C" bold="true" box="[151,652,1200,1227]" pageId="0" pageNumber="393">
<taxonomicName id="E9CAE6F8F975140C149DFB7A0E47FB1C" ID-CoL="X2JM" authority="Lobl, 1998" authorityName="Lobl" authorityYear="1998" box="[151,556,1200,1227]" class="Insecta" family="Staphylinidae" genus="Colilodion" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="tetramerus">
<emphasis id="1CBE4169F975140C149DFB7A0DCBFB1D" bold="true" box="[151,416,1200,1227]" italics="true" pageId="0" pageNumber="393">Colilodion tetramerus</emphasis>
<bibRefCitation id="4A5BE08AF975140C15ADFB7B0E47FB1C" author="Lobl, I." box="[423,556,1200,1227]" pageId="0" pageNumber="393" pagination="467 - 470" refId="ref771" refString="Lobl, I. (1998) A new species of Colilodion (Coleoptera: Staphylinidae: Pselaphinae) from China. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71, 467 - 470." type="journal article" year="1998">Löbl, 1998</bibRefCitation>
</taxonomicName>
(
<figureCitation id="B6F181FEF975140C1631FB7A0EE9FB1C" box="[571,642,1200,1226]" captionStart="FIGURE 1" captionStartId="0.[151,250,1979,2003]" captionTargetBox="[341,1248,1278,1956]" captionTargetId="figure-334@0.[340,1248,1276,1956]" captionTargetPageId="0" captionText="FIGURE 1. Colilodion tetramerus. A. Habitus. B. Apex of antenna, showing sensillum. C. Distribution; circle—known distribution, square—new record. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8430812" httpUri="https://zenodo.org/record/8430812/files/figure.png" pageId="0" pageNumber="393">Fig. 1</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="66D0CEF0F975140C149DFB1B08E6FB3F" box="[151,1165,1233,1258]" pageId="0" pageNumber="393" type="reference_group">
<paragraph id="2E759D7BF975140C149DFB1B08E6FB3F" blockId="0.[151,1165,1200,1258]" box="[151,1165,1233,1258]" pageId="0" pageNumber="393">
<treatmentCitationGroup id="0EDABA55F975140C149DFB1B08E6FB3F" box="[151,1165,1233,1258]" pageId="0" pageNumber="393">
<treatmentCitation id="AF6BBB6AF975140C149DFB1B0E4FFB3F" author="Lobl, I." box="[151,548,1233,1257]" page="467" pageId="0" pageNumber="393" year="1998">
<taxonomicName id="E9CAE6F8F975140C149DFB1B0E4FFB3F" ID-CoL="X2JM" authority="Lobl, 1998: 467" authorityName="Lobl" authorityPageNumber="467" authorityYear="1998" box="[151,548,1233,1257]" class="Insecta" family="Staphylinidae" genus="Colilodion" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="tetramerus">
<emphasis id="1CBE4169F975140C149DFB1B0D13FB3F" box="[151,376,1233,1257]" italics="true" pageId="0" pageNumber="393">Colilodion tetramerus</emphasis>
<materialsCitation id="9EA29726F975140C1574FB1B0E4FFB3F" box="[382,548,1233,1257]" collectionCode="R" collectorName="Ba Wang Ling N." country="China" elevation="1000" location="Hainan" pageId="0" pageNumber="393" specimenCount="1" stateProvince="Hainan">
<bibRefCitation id="4A5BE08AF975140C1574FB1B0E4FFB3F" author="Lobl, I." box="[382,548,1233,1257]" pageId="0" pageNumber="393" pagination="467 - 470" refId="ref771" refString="Lobl, I. (1998) A new species of Colilodion (Coleoptera: Staphylinidae: Pselaphinae) from China. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 71, 467 - 470." type="journal article" year="1998">Löbl, 1998: 467</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="F17123D9F975140C1627FB180E34FB3C" box="[557,607,1234,1258]" pageId="0" pageNumber="393">Type</typeStatus>
locality:
<collectingCountry id="56DDDDEBF975140C16B5FB1B0E95FB3F" box="[703,766,1233,1257]" name="China" pageId="0" pageNumber="393">China</collectingCountry>
,
<collectingRegion id="EC0E5399F975140C1702FB1B0F3FFB3F" box="[776,852,1233,1257]" country="China" name="Hainan" pageId="0" pageNumber="393">Hainan</collectingRegion>
,
<collectorName id="833FF8ADF975140C1754FB18087BFB3F" box="[862,1040,1233,1258]" pageId="0" pageNumber="393">
Ba Wang Ling
<taxonomicNameLabel id="078DFC12F975140C17FDFB18087BFB3F" box="[1015,1040,1234,1257]" pageId="0" pageNumber="393" rank="subSpecies">N.</taxonomicNameLabel>
</collectorName>
R.,
<quantity id="E932309EF975140C1030FB1B08E1FB3F" box="[1082,1162,1233,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="393" unit="m" value="1000.0">
<elevation id="A5E77A48F975140C1030FB1B08E1FB3F" box="[1082,1162,1233,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="393" unit="m" value="1000.0">1000 m</elevation>
</quantity>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="7AB5CDF3F975140C149DF8710ED9F821" ID-DOI="http://doi.org/10.5281/zenodo.8430812" ID-Zenodo-Dep="8430812" httpUri="https://zenodo.org/record/8430812/files/figure.png" pageId="0" pageNumber="393" startId="0.[151,250,1979,2003]" targetBox="[341,1248,1278,1956]" targetPageId="0" targetType="figure">
<paragraph id="2E759D7BF975140C149DF8710ED9F821" blockId="0.[151,1437,1979,2040]" pageId="0" pageNumber="393">
<emphasis id="1CBE4169F975140C149DF8710D73F802" bold="true" box="[151,280,1979,2004]" pageId="0" pageNumber="393">FIGURE 1.</emphasis>
<taxonomicName id="E9CAE6F8F975140C1529F8760E62F805" authorityName="Lobl" authorityYear="1998" box="[291,521,1980,2004]" class="Insecta" family="Staphylinidae" genus="Colilodion" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="tetramerus">
<emphasis id="1CBE4169F975140C1529F8760E62F805" box="[291,521,1980,2004]" italics="true" pageId="0" pageNumber="393">Colilodion tetramerus</emphasis>
</taxonomicName>
.
<emphasis id="1CBE4169F975140C1610F8710E58F805" bold="true" box="[538,563,1979,2003]" pageId="0" pageNumber="393">A.</emphasis>
Habitus.
<emphasis id="1CBE4169F975140C1694F8760EDEF802" bold="true" box="[670,693,1980,2004]" pageId="0" pageNumber="393">B.</emphasis>
Apex of antenna, showing sensillum.
<emphasis id="1CBE4169F975140C105FF8710805F805" bold="true" box="[1109,1134,1979,2003]" pageId="0" pageNumber="393">C.</emphasis>
Distribution; circle—known distribution, square—new record. Scale bar: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="66D0CEF0F974140D149DFF5D0865FF02" pageId="1" pageNumber="394" type="materials_examined">
<paragraph id="2E759D7BF974140D149DFF5D0865FF02" blockId="1.[151,1437,151,428]" pageId="1" pageNumber="394">
<emphasis id="1CBE4169F974140D149DFF5D0D10FF79" bold="true" box="[151,379,151,175]" inLineHeading="true" pageId="1" pageNumber="394" reason="1">Material examined.</emphasis>
<materialsCitation id="9EA29726F974140D158DFF520D66FF05" collectingDate="2023-08-05" collectionCode="R" collectorName="Diaoluoshan N." country="China" county="Lingshui Co." elevation="950" latitude="18.7275" location="Lingshui Co." longLatPrecision="21" longitude="109.86527" pageId="1" pageNumber="394" specimenCount="1" stateProvince="Hainan">
<specimenCount id="38CC56F2F974140D158DFF520DA8FF79" box="[391,451,152,176]" pageId="1" pageNumber="394" type="generic">1 ex.</specimenCount>
, ‘
<collectingCountry id="56DDDDEBF974140D15D2FF5D0E4BFF79" box="[472,544,151,175]" name="China" pageId="1" pageNumber="394">China</collectingCountry>
:
<collectingRegion id="EC0E5399F974140D1625FF5D0E14FF79" box="[559,639,151,175]" country="China" name="Hainan" pageId="1" pageNumber="394">Hainan</collectingRegion>
,
<collectingCounty id="C714E5F7F974140D1685FF5D0F48FF66" box="[655,803,151,176]" pageId="1" pageNumber="394">Lingshui Co.</collectingCounty>
,
<collectorName id="833FF8ADF974140D1739FF5D0F8BFF79" box="[819,992,151,175]" pageId="1" pageNumber="394">
Diaoluoshan
<taxonomicNameLabel id="078DFC12F974140D17CCFF520F8BFF79" box="[966,992,152,175]" pageId="1" pageNumber="394" rank="subSpecies">N.</taxonomicNameLabel>
</collectorName>
<collectionCode id="48DB05BEF974140D17E6FF520869FF79" box="[1004,1026,152,175]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="394">R</collectionCode>
.,
<geoCoordinate id="4BFEFBBCF974140D101DFF5208F7FF66" box="[1047,1180,152,176]" degrees="18" direction="north" minutes="43" orientation="latitude" pageId="1" pageNumber="394" precision="15" seconds="39" value="18.7275">18°43’39”N</geoCoordinate>
,
<geoCoordinate id="4BFEFBBCF974140D10A7FF5D0957FF79" box="[1197,1340,151,175]" degrees="109" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="394" precision="15" seconds="55" value="109.86527">109°51’55”E</geoCoordinate>
,
<quantity id="E932309EF974140D1147FF5D09F3FF79" box="[1357,1432,151,175]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="394" unit="m" value="950.0">
<elevation id="A5E77A48F974140D1147FF5D09F3FF79" box="[1357,1432,151,175]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="394" unit="m" value="950.0">950 m</elevation>
</quantity>
,
<date id="5A74BBBBF974140D149DFF710D66FF05" box="[151,269,187,211]" pageId="1" pageNumber="394" value="2023-08-05">
<collectingDate id="4A304253F974140D149DFF710D66FF05" box="[151,269,187,211]" pageId="1" pageNumber="394" value="2023-08-05">5.viii.2023</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="9EA29726F974140D1512FF710860FF02" box="[280,1035,187,212]" collectionCode="SNUC" collectorName="Xiao-Bin Song" country="China" pageId="1" pageNumber="394" specimenCount="1" stateProvince="Hainan">
leaf litter,
<collectingMethod id="F78BE56CF974140D158FFF710E48FF05" box="[389,547,187,211]" pageId="1" pageNumber="394">Berlese funnel</collectingMethod>
,
<collectorName id="833FF8ADF974140D1627FF710EA5FF02" box="[557,718,187,212]" pageId="1" pageNumber="394">Xiao-Bin Song</collectorName>
leg., [DZẾBƔƜ]’ (
<collectionCode id="48DB05BEF974140D17BCFF760868FF02" box="[950,1027,188,212]" pageId="1" pageNumber="394">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66D0CEF0F974140D14CDFF150975FE51" pageId="1" pageNumber="394" type="discussion">
<paragraph id="2E759D7BF974140D14CDFF150975FE51" blockId="1.[151,1437,151,428]" pageId="1" pageNumber="394">
<emphasis id="1CBE4169F974140D14CDFF150D22FF21" bold="true" box="[199,329,223,247]" inLineHeading="true" pageId="1" pageNumber="394" reason="1">Comments.</emphasis>
The specimen (
<figureCitation id="B6F181FEF974140D15F6FF150E39FF2E" box="[508,594,223,248]" captionStart="FIGURE 1" captionStartId="0.[151,250,1979,2003]" captionTargetBox="[341,1248,1278,1956]" captionTargetId="figure-334@0.[340,1248,1276,1956]" captionTargetPageId="0" captionText="FIGURE 1. Colilodion tetramerus. A. Habitus. B. Apex of antenna, showing sensillum. C. Distribution; circle—known distribution, square—new record. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8430812" httpUri="https://zenodo.org/record/8430812/files/figure.png" pageId="1" pageNumber="394">Fig. 1A</figureCitation>
) is undoubtedly conspecific with the population of Bawanglin based on the identical four-segmented antennae, large sensilla of the antennomeres 4 (
<figureCitation id="B6F181FEF974140D1790FEC90F80FECA" box="[922,1003,259,284]" captionStart="FIGURE 1" captionStartId="0.[151,250,1979,2003]" captionTargetBox="[341,1248,1278,1956]" captionTargetId="figure-334@0.[340,1248,1276,1956]" captionTargetPageId="0" captionText="FIGURE 1. Colilodion tetramerus. A. Habitus. B. Apex of antenna, showing sensillum. C. Distribution; circle—known distribution, square—new record. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8430812" httpUri="https://zenodo.org/record/8430812/files/figure.png" pageId="1" pageNumber="394">Fig. 1B</figureCitation>
), and remaining external characters. The only other species showing a similar morphology is
<taxonomicName id="E9CAE6F8F974140D16CEFEE20863FE96" authority="Nomura &amp; Sugaya" authorityName="Nomura &amp; Sugaya" authorityYear="2007" box="[708,1032,295,320]" class="Insecta" family="Staphylinidae" genus="Colilodion" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="394" phylum="Arthropoda" rank="species" species="thienmu">
<emphasis id="1CBE4169F974140D16CEFEE20EBDFEE9" box="[708,726,296,319]" italics="true" pageId="1" pageNumber="394">C</emphasis>
.
<emphasis id="1CBE4169F974140D16E9FEED0F52FEE9" box="[739,825,295,319]" italics="true" pageId="1" pageNumber="394">thienmu</emphasis>
Nomura &amp; Sugaya
</taxonomicName>
from central
<collectingCountry id="56DDDDEBF974140D1092FEED089EFEE9" box="[1176,1269,295,319]" name="Vietnam" pageId="1" pageNumber="394">Vietnam</collectingCountry>
, which also has four segments on each antenna. However, the latter species has more rounded lateral margins of the head, the antennomeres 4 are much more curved near the bases, and the apical sensilla of antennomeres 4 are much less pronounced.
</paragraph>
</subSubSection>
<subSubSection id="66D0CEF0F974140D14CDFE590EBAFE7A" box="[199,721,403,428]" pageId="1" pageNumber="394" type="distribution">
<paragraph id="2E759D7BF974140D14CDFE590EBAFE7A" blockId="1.[151,1437,151,428]" box="[199,721,403,428]" pageId="1" pageNumber="394">
<emphasis id="1CBE4169F974140D14CDFE590D31FE7D" bold="true" box="[199,346,403,427]" inLineHeading="true" pageId="1" pageNumber="394" reason="1">Distribution.</emphasis>
Southern
<collectingCountry id="56DDDDEBF974140D15CDFE590E60FE7D" box="[455,523,403,427]" name="China" pageId="1" pageNumber="394">China</collectingCountry>
:
<collectingRegion id="EC0E5399F974140D161FFE590E09FE7D" box="[533,610,403,427]" country="China" name="Hainan" pageId="1" pageNumber="394">Hainan</collectingRegion>
(
<figureCitation id="B6F181FEF974140D167AFE590EAFFE7A" box="[624,708,403,428]" captionStart="FIGURE 1" captionStartId="0.[151,250,1979,2003]" captionTargetBox="[341,1248,1278,1956]" captionTargetId="figure-334@0.[340,1248,1276,1956]" captionTargetPageId="0" captionText="FIGURE 1. Colilodion tetramerus. A. Habitus. B. Apex of antenna, showing sensillum. C. Distribution; circle—known distribution, square—new record. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8430812" httpUri="https://zenodo.org/record/8430812/files/figure.png" pageId="1" pageNumber="394">Fig. 1C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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