359 lines
61 KiB
XML
359 lines
61 KiB
XML
<document id="819CDC0E54C7241A0C3F4116CCBCB367" ID-DOI="10.5281/zenodo.186314" ID-GBIF-Dataset="29bf32ba-5840-4c26-8af0-c7e8231a4821" ID-ISSN="1175-5326" ID-Zenodo-Dep="186314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461050178843" checkinUser="plazi" docAuthor="Solodovnikov, Alexey & Newton, Alfred F." docDate="2009" docId="A641F03DFFBC6705FF5FF951FA966072" docLanguage="en" docName="zt02033p057.pdf" docOrigin="Zootaxa 2033" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Australotarsius Solodovnikov & Newton" docType="treatment" docVersion="9" lastPageNumber="53" masterDocId="5A788845FFBD6701FFC8FF9FFFAC636E" masterDocTitle="Australotarsius — a new genus of the rove beetle tribe Staphylinini from Australia (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="57" masterPageNumber="49" pageNumber="50" updateTime="1698578517988" updateUser="plazi">
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<mods:titleInfo id="D840003DDF67BD7637410372F5D605B0">
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<mods:title id="43F9000E2D8DA69E973B59C0AF632A5B">Australotarsius — a new genus of the rove beetle tribe Staphylinini from Australia (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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</mods:titleInfo>
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<mods:name id="4C808AE888F74CF05AE35310E300154B" type="personal">
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<mods:namePart id="4F83C3C56ECF356D0FD1AA2E3FB6A2EB">Solodovnikov, Alexey</mods:namePart>
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<mods:namePart id="440B03CA87DA566E5B6033CAB5F870A8">Newton, Alfred F.</mods:namePart>
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<mods:typeOfResource id="9DEE0F195F1E23513E1C5EDB32EEA953">text</mods:typeOfResource>
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<mods:title id="DEF18B051C3161DBF67BBBA713447566">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B1477EC56747B46E7CDF0A358A6CAAC1">
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<mods:date id="8BC80CD220EF764EFB06E9FF7D8D824B">2009</mods:date>
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<mods:detail id="F71F658D5A45D862EDCC6218C54C3B92" type="volume">
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<mods:number id="FAA51C92D29DE782E006EE7D5B6D4D42">2033</mods:number>
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<mods:start id="5B87A61E5AADD84B001723E49646FF48">49</mods:start>
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<mods:end id="143E4AD3C33B9600F6406E3D6FB673D9">57</mods:end>
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<mods:classification id="6A50B71C11E325EF457B38202356C6F5">journal article</mods:classification>
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<mods:identifier id="934936D976A1EABF498BC36EB8A05989" type="DOI">10.5281/zenodo.186314</mods:identifier>
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<mods:identifier id="887F4D3124E12296D070468D2156E360" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F27E3148A28302A51C38840C5449ED20" type="Zenodo-Dep">186314</mods:identifier>
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<treatment id="A641F03DFFBC6705FF5FF951FA966072" ID-DOI="http://doi.org/10.5281/zenodo.6226722" ID-GBIF-Taxon="119611713" ID-Zenodo-Dep="6226722" LSID="urn:lsid:plazi:treatment:A641F03DFFBC6705FF5FF951FA966072" httpUri="http://treatment.plazi.org/id/A641F03DFFBC6705FF5FF951FA966072" lastPageId="4" lastPageNumber="53" pageId="1" pageNumber="50">
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<subSubSection id="66F212A0FFBC6700FF5FF951FE8A6465" pageId="1" pageNumber="50" type="nomenclature">
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<paragraph id="2E57412BFFBC6700FF5FF951FCB46586" blockId="1.[151,792,1742,1803]" box="[151,792,1742,1768]" pageId="1" pageNumber="50">
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<heading id="751FF647FFBC6700FF5FF951FCB46586" bold="true" box="[151,792,1742,1768]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
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<emphasis id="1C9C9D39FFBC6700FF5FF951FCB46586" bold="true" box="[151,792,1742,1768]" pageId="1" pageNumber="50">
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<taxonomicName id="E9E83AA8FFBC6700FF5FF951FD266586" ID-CoL="8XJTL" authority="Solodovnikov & Newton" authorityName="Solodovnikov & Newton" box="[151,650,1742,1768]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="1C9C9D39FFBC6700FF5FF951FEFC6586" bold="true" box="[151,336,1742,1768]" italics="true" pageId="1" pageNumber="50">Australotarsius</emphasis>
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Solodovnikov & Newton
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</taxonomicName>
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, new genus
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2E57412BFFBC6700FF5FF96EFE8A6465" blockId="1.[151,792,1742,1803]" box="[151,294,1777,1803]" pageId="1" pageNumber="50">
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(
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<figureCitation id="B6D35DAEFFBC6700FF57F96EFEB16465" box="[159,285,1777,1803]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 11" captionStart-2="FIGURES 12 – 19" captionStartId-0="3.[151,269,1125,1149]" captionStartId-1="6.[151,270,1874,1898]" captionStartId-2="7.[151,269,1291,1315]" captionTargetBox-0="[463,1131,387,1087]" captionTargetBox-1="[163,1422,212,1822]" captionTargetBox-2="[188,1418,191,1271]" captionTargetId-0="figure@3.[449,1139,351,1100]" captionTargetId-1="figure@6.[159,1434,197,1849]" captionTargetId-2="figure@7.[163,1448,183,1325]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 1 – 2. Australotarsius grandis (1) and A. tasmanicus (2), habitus. Scale bar 5 mm." captionText-1="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." captionText-2="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri-0="https://zenodo.org/record/186315/files/figure.png" httpUri-1="https://zenodo.org/record/186316/files/figure.png" httpUri-2="https://zenodo.org/record/186317/files/figure.png" pageId="1" pageNumber="50">Figs. 1–19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="66F212A0FFBC6700FF5FF8A1FB976436" box="[151,1083,1854,1880]" pageId="1" pageNumber="50" type="materials_examined">
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<paragraph id="2E57412BFFBC6700FF5FF8A1FB976436" blockId="1.[151,1437,1854,2035]" box="[151,1083,1854,1880]" pageId="1" pageNumber="50">
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<emphasis id="1C9C9D39FFBC6700FF5FF8A1FE956436" bold="true" box="[151,313,1854,1880]" pageId="1" pageNumber="50">
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<typeStatus id="F153FF89FFBC6700FF5FF8A1FF786436" box="[151,212,1854,1880]" pageId="1" pageNumber="50">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="E9E83AA8FFBC6700FE88F8A0FC366436" authority="Solodovnikov and Newton" authorityName="Solodovnikov and Newton" box="[320,922,1854,1880]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
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<emphasis id="1C9C9D39FFBC6700FE88F8A0FDF96436" box="[320,597,1855,1880]" italics="true" pageId="1" pageNumber="50">Australotarsius grandis</emphasis>
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Solodovnikov and Newton
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</taxonomicName>
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,
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<taxonomicNameLabel id="07AF2042FFBC6700FC6EF8A1FB9B6436" box="[934,1079,1854,1880]" pageId="1" pageNumber="50" rank="species">new species</taxonomicNameLabel>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="66F212A0FFBC6700FF0EF8FAFC3C649D" pageId="1" pageNumber="50" type="diagnosis">
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<paragraph id="2E57412BFFBC6700FF0EF8FAFC3C649D" blockId="1.[151,1437,1854,2035]" pageId="1" pageNumber="50">
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<emphasis id="1C9C9D39FFBC6700FF0EF8FAFEEA6411" bold="true" box="[198,326,1893,1919]" pageId="1" pageNumber="50">Diagnosis.</emphasis>
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From all other genera of
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<taxonomicName id="E9E83AA8FFBC6700FDB5F8FAFCA36411" box="[637,783,1893,1919]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
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,
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<taxonomicName id="E9E83AA8FFBC6700FCD4F8FAFC7C6410" box="[796,976,1893,1918]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
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<emphasis id="1C9C9D39FFBC6700FCD4F8FAFC7C6410" box="[796,976,1893,1918]" italics="true" pageId="1" pageNumber="50">Australotarsius</emphasis>
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</taxonomicName>
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can be distinguished by the following combination of characters: entire body evenly setose; apical segment of maxillary palpus setose; neck broad; infraorbital ridges well developed; anterior tarsi in both sexes very broad; male sternite VIII without secondary sexual modifications, of the same shape as in females.
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</paragraph>
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</subSubSection>
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<subSubSection id="66F212A0FFBF6702FF0EFF08FCFF63B6" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51" type="description">
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<paragraph id="2E57412BFFBF6703FF0EFF08FAC063DC" blockId="2.[151,1437,151,1995]" box="[198,1388,151,178]" pageId="2" pageNumber="51">
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<emphasis id="1C9C9D39FFBF6703FF0EFF08FEF363DF" bold="true" box="[198,351,151,177]" pageId="2" pageNumber="51">Description.</emphasis>
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Habitus as in
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<figureCitation id="B6D35DAEFFBF6703FDCEFF07FDC363DC" box="[518,623,152,178]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,269,1125,1149]" captionTargetBox="[463,1131,387,1087]" captionTargetId="figure@3.[449,1139,351,1100]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Australotarsius grandis (1) and A. tasmanicus (2), habitus. Scale bar 5 mm." httpUri="https://zenodo.org/record/186315/files/figure.png" pageId="2" pageNumber="51">Figs 1, 2</figureCitation>
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. Entire body covered by dense setation. Body length
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<quantity id="E910ECCEFFBF6703FB17FF07FAC563DC" box="[1247,1385,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.4" metricValueMin="0.74" pageId="2" pageNumber="51" unit="mm" value="10.7" valueMax="14.0" valueMin="7.4">7.4–14 mm</quantity>
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.
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</paragraph>
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<paragraph id="2E57412BFFBF6703FF0EFF21FB156198" blockId="2.[151,1437,151,1995]" pageId="2" pageNumber="51">
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Head capsule (
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<figureCitation id="B6D35DAEFFBF6703FEB5FF21FE4763B6" box="[381,491,190,216]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,269,1125,1149]" captionTargetBox="[463,1131,387,1087]" captionTargetId="figure@3.[449,1139,351,1100]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Australotarsius grandis (1) and A. tasmanicus (2), habitus. Scale bar 5 mm." httpUri="https://zenodo.org/record/186315/files/figure.png" pageId="2" pageNumber="51">Figs 1–2</figureCitation>
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,
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<figureCitation id="B6D35DAEFFBF6703FE35FF21FDA063B6" box="[509,524,190,216]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">8</figureCitation>
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) rounded with well developed, moderately large eyes; nuchal constriction indicated only laterally (in
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<taxonomicName id="E9E83AA8FFBF6703FE10FF7AFDF86390" box="[472,596,229,254]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="grandis">
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<emphasis id="1C9C9D39FFBF6703FE10FF7AFDF86390" box="[472,596,229,254]" italics="true" pageId="2" pageNumber="51">A. grandis</emphasis>
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</taxonomicName>
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) or laterally and dorsally (in
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<taxonomicName id="E9E83AA8FFBF6703FC79FF7AFBFA6390" box="[945,1110,229,254]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="tasmanicus">
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<emphasis id="1C9C9D39FFBF6703FC79FF7AFBFA6390" box="[945,1110,229,254]" italics="true" pageId="2" pageNumber="51">A. tasmanicus</emphasis>
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</taxonomicName>
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), in the former case nuchal ridge developed only laterally, in the latter—laterally and dorsally; epistomal suture absent; infraorbital ridges present, extending to about the middle of the distance from neck to base of mandibles; postgenal and ventral basal ridges well developed; postmandibular ridge and dorsal basal ridge absent. Antennal insertions at anterolateral margins of frons, anterior to eyes, not concealed from above; distance between them slightly longer than distance from either insertion to margin of eye. Antennae moderately long, with distinctly elongate first antennomere (as long as second and third antennomeres together); first three antennomeres setose, but without pubescence, remaining antennomeres setose and pubescent. Mandibles (
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<figureCitation id="B6D35DAEFFBF6703FB0BFE6BFAA46160" box="[1219,1288,500,526]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 6</figureCitation>
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) moderately long, symmetrical, each with teeth on inner side, without dorso-lateral mandibular groove; prostheca broadly attached near base of mandible, mola not developed. Labrum (
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<figureCitation id="B6D35DAEFFBF6703FCBDFDDEFC176135" box="[885,955,577,603]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 3</figureCitation>
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) strongly transverse, without transparent apical membrane, only slightly notched medio-apically. Maxilla (
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<figureCitation id="B6D35DAEFFBF6703FC53FDF7FC4F61EC" box="[923,995,616,642]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 4</figureCitation>
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) with lacinia and galea densely hairy at apex; stipes with clear sutures; maxillary palpus four-segmented, last segment fusiform, finely setose. Labium (
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<figureCitation id="B6D35DAEFFBF6703FEC0FD2AFEF961A1" box="[264,341,693,719]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 5</figureCitation>
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): ligula small, only slightly notched apically, paraglossae long with comb of stout setae internally; last segment of labial palpus fusiform with slightly truncate apex, finely setose.
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</paragraph>
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<paragraph id="2E57412BFFBF6703FF0EFC9DFEA966C0" blockId="2.[151,1437,151,1995]" pageId="2" pageNumber="51">
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Pronotum (
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<figureCitation id="B6D35DAEFFBF6703FE85FC9DFE3B6072" box="[333,407,770,796]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 7</figureCitation>
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) with anterior angles projecting anteriad over apical margin of prosternum; pronotal hypomera slightly inflected inwards, only slightly visible in lateral view; superior marginal line of pronotum well developed throughout its whole length, not deflexed ventrad; inferior marginal line shorter, not meeting with superior marginal line; pronotal hypomeron with weakly sclerotized postcoxal process. Pronotosternal sutures visible but not membranous. Prosternum with midlongitudinal carina, without large conspicuous macrosetae. Mesoventrite without large conspicuous macrosetae, with narrow, acute mesoventral intercoxal process. Meso-metaventral suture well developed, membranous. Mesoscutellum (
|
||
<figureCitation id="B6D35DAEFFBF6703FB9AFC75FB09676A" box="[1106,1189,1002,1028]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 11</figureCitation>
|
||
) with two transverse carinae: one very close to base, another close to middle. Elytra (
|
||
<figureCitation id="B6D35DAEFFBF6703FCB5FB8EFC7C6745" box="[893,976,1041,1067]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 11</figureCitation>
|
||
) relatively long, with sharp humeri and well developed sub-basal ridge immediately adjacent to elytral articulation and extending anteriad to anterior margin of elytron; epipleural part gradually deflexed, lacking epipleural ridge; sutural angle distinct.
|
||
<collectingCountry id="56FF01BBFFBF6703FAA8FBC1FA306716" box="[1376,1436,1118,1144]" name="India" pageId="2" pageNumber="51">Hind</collectingCountry>
|
||
wings (
|
||
<figureCitation id="B6D35DAEFFBF6703FF27FB1AFEE967F1" box="[239,325,1157,1183]" captionStart="FIGURES 12 – 19" captionStartId="7.[151,269,1291,1315]" captionTargetBox="[188,1418,191,1271]" captionTargetId="figure@7.[163,1448,183,1325]" captionTargetPageId="7" captionText="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri="https://zenodo.org/record/186317/files/figure.png" pageId="2" pageNumber="51">Fig. 12</figureCitation>
|
||
) fully developed, with veins CuA and MP4 completely separate. Metaventrite (
|
||
<figureCitation id="B6D35DAEFFBF6703FAC8FB1AFAF467F1" box="[1280,1368,1157,1183]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 10</figureCitation>
|
||
) well developed, with deep mesocoxal acetabuli delimited by carina. Legs moderately long and slender; anterior and middle (smaller) coxae large, conical, contiguous (
|
||
<figureCitation id="B6D35DAEFFBF6703FCFDFB4DFC106782" box="[821,956,1234,1260]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Figs. 7, 10</figureCitation>
|
||
).
|
||
<collectingCountry id="56FF01BBFFBF6703FC1AFB4DFBA26782" box="[978,1038,1234,1260]" name="India" pageId="2" pageNumber="51">Hind</collectingCountry>
|
||
coxae (
|
||
<figureCitation id="B6D35DAEFFBF6703FBA6FB4DFB6B6782" box="[1134,1223,1234,1260]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 10</figureCitation>
|
||
) as long as wide, almost contiguous at base. Anterior tibia densely setose, without spines externally; middle and posterior tibiae setose, with spines externally. Tarsal formula 5-5-5; claws moderately long, arcuate; pair of empodial setae on each tarsus, shorter on anterior tarsi and very long, longer than claws on middle and posterior tarsi. Anterior tarsi (
|
||
<figureCitation id="B6D35DAEFFBF6703FF12FAF2FE8D66E9" box="[218,289,1389,1415]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 9</figureCitation>
|
||
) in both sexes with tarsomeres I–IV strongly dilated and with dense, long whitish adhesive setae ventrally.
|
||
</paragraph>
|
||
<paragraph id="2E57412BFFBF6703FF0EFA25FC806439" blockId="2.[151,1437,151,1995]" pageId="2" pageNumber="51">
|
||
Abdomen more or less parallel-sided along most of its length, narrowed posteriad, slightly flattened dorsoventrally; abdominal tergite I with protergal glands externally manifested by deep indentation (acetabulum) at each side of tergite and adjacent groups of setae (
|
||
<figureCitation id="B6D35DAEFFBF6703FC79F997FBA6654C" box="[945,1034,1544,1570]" captionStart="FIGURES 3 – 11" captionStartId="6.[151,270,1874,1898]" captionTargetBox="[163,1422,212,1822]" captionTargetId="figure@6.[159,1434,197,1849]" captionTargetPageId="6" captionText="FIGURES 3 – 11. Australotarsius grandis, details of structure. 3, labrum, dorsally; 4, maxilla, left, ventrally; 5, labium, ventrally; 6, mandible, right, dorsally; 7, prothorax, ventrally, left leg removed; 8, head capsule, latero-ventrally; 9, anterior tarsus, left, dorsally; 10, meso- and metathorax, ventrally, left middle leg removed; 11, meso- and metathorax dorsally, right elytron removed. Scale bars 1 mm." httpUri="https://zenodo.org/record/186316/files/figure.png" pageId="2" pageNumber="51">Fig. 11</figureCitation>
|
||
). Segments III–VI each with two pairs of elongate paratergites, segment VII with two pairs of paratergites, basal short one and apical long one; segment VIII without paratergites; tergites III–VII only with one basal carina; sternite III with basal carina medially strongly angulate (
|
||
<figureCitation id="B6D35DAEFFBF6703FE2DF9E3FD9265F8" box="[485,574,1660,1686]" captionStart="FIGURES 12 – 19" captionStartId="7.[151,269,1291,1315]" captionTargetBox="[188,1418,191,1271]" captionTargetId="figure@7.[163,1448,183,1325]" captionTargetPageId="7" captionText="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri="https://zenodo.org/record/186317/files/figure.png" pageId="2" pageNumber="51">Fig. 13</figureCitation>
|
||
). Sternite VIII without apical emargination in males, with apical margin similarly rounded in both sexes. Abdominal intersegmental membranes attached preapically and having a pattern of small irregular rounded sclerites. Male (
|
||
<figureCitation id="B6D35DAEFFBF6703FD38F956FCEA658D" box="[752,838,1737,1763]" captionStart="FIGURES 12 – 19" captionStartId="7.[151,269,1291,1315]" captionTargetBox="[188,1418,191,1271]" captionTargetId="figure@7.[163,1448,183,1325]" captionTargetPageId="7" captionText="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri="https://zenodo.org/record/186317/files/figure.png" pageId="2" pageNumber="51">Fig. 14</figureCitation>
|
||
) and female (
|
||
<figureCitation id="B6D35DAEFFBF6703FC24F956FBEE658D" box="[1004,1090,1737,1763]" captionStart="FIGURES 12 – 19" captionStartId="7.[151,269,1291,1315]" captionTargetBox="[188,1418,191,1271]" captionTargetId="figure@7.[163,1448,183,1325]" captionTargetPageId="7" captionText="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri="https://zenodo.org/record/186317/files/figure.png" pageId="2" pageNumber="51">Fig. 15</figureCitation>
|
||
) tergite IX consisting of two elongate, inflated lateral sclerites, apically obtuse, widely separated dorsally by tergite X and ventrally by sternite IX in males or valvifers (first gonocoxites) in females. Male sternite IX (
|
||
<figureCitation id="B6D35DAEFFBF6703FBA9F889FB15645E" box="[1121,1209,1814,1840]" captionStart="FIGURES 12 – 19" captionStartId="7.[151,269,1291,1315]" captionTargetBox="[188,1418,191,1271]" captionTargetId="figure@7.[163,1448,183,1325]" captionTargetPageId="7" captionText="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri="https://zenodo.org/record/186317/files/figure.png" pageId="2" pageNumber="51">Fig. 14</figureCitation>
|
||
) elongate, more or less symmetrical, with short asymmetrical basal portion.
|
||
</paragraph>
|
||
<paragraph id="2E57412BFFBF6703FF0EF8FBFDEE64A5" blockId="2.[151,1437,151,1995]" pageId="2" pageNumber="51">
|
||
Aedeagus (
|
||
<figureCitation id="B6D35DAEFFBF6703FE82F8FBFE776410" box="[330,475,1892,1918]" captionStart="FIGURES 12 – 19" captionStartId="7.[151,269,1291,1315]" captionTargetBox="[188,1418,191,1271]" captionTargetId="figure@7.[163,1448,183,1325]" captionTargetPageId="7" captionText="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri="https://zenodo.org/record/186317/files/figure.png" pageId="2" pageNumber="51">Figs. 16–19</figureCitation>
|
||
): median lobe elongate, apically obtuse, symmetrical; paramere with peg setae on the underside, and four pairs of setae apically. Aedeagus rotated by 90° in the abdomen in repose: its dorsal (parameral) side facing left laterally.
|
||
</paragraph>
|
||
<paragraph id="2E57412BFFBE6702FF0EFF07FCFF63B6" blockId="3.[151,1436,152,294]" pageId="3" pageNumber="52">
|
||
Female external genitalia consisting of a pair of large basal valvifers, short coxites (second gonocoxites) and tiny styli; female tergite X symmetrical, large (
|
||
<figureCitation id="B6D35DAEFFBE6702FD26FF21FCE963B6" box="[750,837,190,216]" captionStart="FIGURES 12 – 19" captionStartId="7.[151,269,1291,1315]" captionTargetBox="[188,1418,191,1271]" captionTargetId="figure@7.[163,1448,183,1325]" captionTargetPageId="7" captionText="FIGURES 12 – 19. Australotarsius grandis (12 – 17) and A. tasmanicus (18 – 19), details of structure. 12, hind wing, right; 13, abdominal sternites II + III, ventrally; 14, male terminalia (abdominal segments IX, X), ventrally; 15, female terminalia (abdominal segments IX, X), ventrally; 16, 18, aedeagus, laterally; 17, 19, aedeagus, dorsally (parameral side). Scale bars 1 mm." httpUri="https://zenodo.org/record/186317/files/figure.png" pageId="3" pageNumber="52">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="66F212A0FFBE6705FF0EFF7AFA966072" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52" type="etymology">
|
||
<paragraph id="2E57412BFFBE6702FF0EFF7AFAFA6248" blockId="3.[151,1436,152,294]" pageId="3" pageNumber="52">
|
||
<emphasis id="1C9C9D39FFBE6702FF0EFF7AFEFF6391" bold="true" box="[198,339,229,255]" pageId="3" pageNumber="52">Etymology.</emphasis>
|
||
The genus name is derived from two words “
|
||
<collectingCountry id="56FF01BBFFBE6702FCB8FF7AFC4D6391" box="[880,993,229,255]" name="Australia" pageId="3" pageNumber="52">Australia</collectingCountry>
|
||
” and “tarsus”. It refers to the country of origin and spectacular wide anterior tarsi of this new taxon. It is Latinized noun of masculine gender.
|
||
</paragraph>
|
||
<caption id="7A9711A3FFBE6702FF5FFBFAFBDC6713" box="[151,1136,1125,1149]" httpUri="https://zenodo.org/record/186315/files/figure.png" pageId="3" pageNumber="52" targetBox="[463,1131,387,1087]" targetPageId="3">
|
||
<paragraph id="2E57412BFFBE6702FF5FFBFAFBDC6713" blockId="3.[151,1136,1125,1149]" box="[151,1136,1125,1149]" pageId="3" pageNumber="52">
|
||
<emphasis id="1C9C9D39FFBE6702FF5FFBFAFEEE6713" bold="true" box="[151,322,1125,1149]" pageId="3" pageNumber="52">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="E9E83AA8FFBE6702FE81FBF9FDEA6713" box="[329,582,1126,1149]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="1C9C9D39FFBE6702FE81FBF9FDEA6713" box="[329,582,1126,1149]" italics="true" pageId="3" pageNumber="52">Australotarsius grandis</emphasis>
|
||
</taxonomicName>
|
||
(1) and
|
||
<taxonomicName id="E9E83AA8FFBE6702FD57FBF9FC996713" box="[671,821,1126,1149]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="tasmanicus">
|
||
<emphasis id="1C9C9D39FFBE6702FD57FBF9FC996713" box="[671,821,1126,1149]" italics="true" pageId="3" pageNumber="52">A. tasmanicus</emphasis>
|
||
</taxonomicName>
|
||
(2), habitus. Scale bar 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2E57412BFFBE6702FF0EFB30FC436434" blockId="3.[151,1437,1199,2037]" pageId="3" pageNumber="52">
|
||
<emphasis id="1C9C9D39FFBE6702FF0EFB30FD5467A7" bold="true" box="[198,760,1199,1225]" pageId="3" pageNumber="52">Comparison and phylogenetic relationships.</emphasis>
|
||
Due to somewhat inflexed hypomera of pronotum and well developed infraorbital ridges
|
||
<taxonomicName id="E9E83AA8FFBE6702FDE4FB49FD4D6781" box="[556,737,1238,1263]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FDE4FB49FD4D6781" box="[556,737,1238,1263]" italics="true" pageId="3" pageNumber="52">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
looks like a member of the conventional subtribe Quediina (for its definition and list of included taxa see, for example,
|
||
<bibRefCitation id="4A793CDAFFBE6702FC4CFB63FB1C6678" author="Smetana" box="[900,1200,1276,1302]" pageId="3" pageNumber="57" refString="Smetana, A. & Davies, A. (2000) Reclassification of the north temperate taxa associated with Staphylinus sensu lato, including comments on relevant subtribes of Staphylinini (Coleoptera: Staphylinidae). American Museum Novitates, 3287, 1 - 88." type="journal article" year="2000">Smetana & Davies 2000</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="4A793CDAFFBE6702FB25FB63FA346678" author="Herman" box="[1261,1432,1276,1302]" pageId="3" pageNumber="57" refString="Herman, L. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Parts I - VII. Bulletin of the American Museum of Natural History, 265, 1 - 4218." type="journal article" year="2001">Herman 2001</bibRefCitation>
|
||
, respectively), which has been shown to be a polyphyletic group (
|
||
<bibRefCitation id="4A793CDAFFBE6702FC0EFABCFB6D6653" author="Solodovnikov" box="[966,1217,1315,1341]" pageId="3" pageNumber="57" refString="Solodovnikov, A. (2006) Revision and phylogenetic assessment of Afroquedius gen. nov. from South Africa: toward new concepts of the genus Quedius, subtribe Quediina and reclassification of the tribe Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Annals of the Entomological Society of America, 99 (6), 1064 - 1084." type="journal article" year="2006">Solodovnikov 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A793CDAFFBE6702FB06FABCFEF0660A" author="Solodovnikov" pageId="3" pageNumber="57" refString="Solodovnikov, A. & Schomann, A. (2009) Revised systematics and biogeography of " Quediina " of sub-Saharan Africa bring new insights about the phylogeny of the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology, (in press)." type="book" year="2009">Solodovnikov & Schomann 2009</bibRefCitation>
|
||
). Within the newly emerging phylogenetic framework for
|
||
<taxonomicName id="E9E83AA8FFBE6702FBDCFAD5FB04660A" box="[1044,1192,1354,1380]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9E83AA8FFBE6702FB7CFAD5FAC7660D" box="[1204,1387,1354,1379]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FB7CFAD5FAC7660D" box="[1204,1387,1354,1379]" italics="true" pageId="3" pageNumber="52">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
can be placed in the clade “Quediina propria”, which, according to
|
||
<bibRefCitation id="4A793CDAFFBE6702FC42FAEEFA3966E5" author="Solodovnikov" box="[906,1429,1393,1419]" pageId="3" pageNumber="57" refString="Solodovnikov, A. & Schomann, A. (2009) Revised systematics and biogeography of " Quediina " of sub-Saharan Africa bring new insights about the phylogeny of the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology, (in press)." type="book" year="2009">
|
||
Solodovnikov and Schomann (2009,
|
||
<figureCitation id="B6D35DAEFFBE6702FA8DFAEEFA2566E5" box="[1349,1417,1393,1419]" captionStart="FIGURES 1 – 2" captionStartId="3.[151,269,1125,1149]" captionTargetBox="[463,1131,387,1087]" captionTargetId="figure@3.[449,1139,351,1100]" captionTargetPageId="3" captionText="FIGURES 1 – 2. Australotarsius grandis (1) and A. tasmanicus (2), habitus. Scale bar 5 mm." httpUri="https://zenodo.org/record/186315/files/figure.png" pageId="3" pageNumber="52">Fig. 1</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
, includes two subclades: one with the north temperate members of the genus
|
||
<taxonomicName id="E9E83AA8FFBE6702FBE8FA07FA9266DF" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" box="[1056,1342,1431,1457]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FBE8FA07FB2E66DF" box="[1056,1154,1432,1457]" italics="true" pageId="3" pageNumber="52">Quedius</emphasis>
|
||
<bibRefCitation id="4A793CDAFFBE6702FB42FA08FA9266DF" author="Stephens" box="[1162,1342,1431,1457]" pageId="3" pageNumber="57" refString="Stephens, J. F. (1829) The nomenclature of British insects; being a compendious list of such species as are contained in the Systematic Catalogue of British Insects, and forming a guide to their classification. Baldwin and Cradock, London, 68 pp." type="book" year="1829">Stephens, 1829</bibRefCitation>
|
||
</taxonomicName>
|
||
and the genus
|
||
<taxonomicName id="E9E83AA8FFBE6702FF2DFA21FDCF66B6" authority="Blackwelder, 1952" authorityName="Blackwelder" authorityYear="1952" box="[229,611,1470,1496]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FF2DFA21FED566B9" box="[229,377,1470,1495]" italics="true" pageId="3" pageNumber="52">Indoquedius</emphasis>
|
||
<bibRefCitation id="4A793CDAFFBE6702FE49FA21FDCF66B6" author="Blackwelder" box="[385,611,1470,1496]" pageId="3" pageNumber="57" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. United States National Museum Bulletin, 200, i - iv, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
|
||
</taxonomicName>
|
||
and another with the genera
|
||
<taxonomicName id="E9E83AA8FFBE6702FC0BFA21FB5B66B6" authority="Fauvel, 1905" authorityName="Fauvel" authorityYear="1905" box="[963,1271,1470,1496]" class="Insecta" family="Staphylinidae" genus="Anchocerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FC0BFA21FBFC66B9" box="[963,1104,1470,1495]" italics="true" pageId="3" pageNumber="52">Anchocerus</emphasis>
|
||
<bibRefCitation id="4A793CDAFFBE6702FB90FA21FB5B66B6" author="Fauvel" box="[1112,1271,1470,1496]" pageId="3" pageNumber="57" refString="Fauvel, A. (1905) Staphylinides exotiques nouveaux (3 e partie). Revue d'Entomologie, 24, 113 - 147." type="journal article" year="1905">Fauvel, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9E83AA8FFBE6702FACDFA21FEC56690" authority="Nordmann, 1837" authorityName="Nordmann" authorityYear="1837" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FACDFA21FA3066B9" box="[1285,1436,1470,1495]" italics="true" pageId="3" pageNumber="52">Acylophorus</emphasis>
|
||
<bibRefCitation id="4A793CDAFFBE6702FF5FFA7BFEC56690" author="Nordmann" box="[151,361,1508,1534]" pageId="3" pageNumber="57" refString="Nordmann, A. (1837) Symbolae ad monographiam staphylinorum. Academiae Caesareae Scientiarum, Petropoli [St. Petersburg], 167 pp., 2 pls." type="book" year="1837">Nordmann, 1837</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9E83AA8FFBE6702FEBEFA7AFD166690" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[374,698,1508,1534]" class="Insecta" family="Staphylinidae" genus="Euryporus" higherTaxonomySource="CoL" kingdom="Fungi" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FEBEFA7AFE5B6690" box="[374,503,1509,1534]" italics="true" pageId="3" pageNumber="52">Euryporus</emphasis>
|
||
<bibRefCitation id="4A793CDAFFBE6702FDC8FA7BFD166690" author="Erichson" box="[512,698,1508,1534]" pageId="3" pageNumber="57" refString="Erichson, W. F. (1839) Die Kafer der Mark Brandenburg, 1 (2). F. H. Morin, Berlin, pp. 385 - 740." type="journal article" year="1839">Erichson, 1839</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E9E83AA8FFBE6702FD3FFA7AFB966690" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[759,1082,1508,1534]" class="Insecta" family="Staphylinidae" genus="Hemiquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FD3FFA7AFC346690" box="[759,920,1509,1534]" italics="true" pageId="3" pageNumber="52">Hemiquedius</emphasis>
|
||
<bibRefCitation id="4A793CDAFFBE6702FC68FA7BFB966690" author="Casey" box="[928,1082,1508,1534]" pageId="3" pageNumber="57" refString="Casey, T. L. (1915) Studies in some staphylinid genera of North America. In: Memoirs on the Coleoptera, VI. The New Era Printing Co., Lancaster, Pennsylvania, pp. 395 - 460." type="book chapter" year="1915">Casey, 1915</bibRefCitation>
|
||
</taxonomicName>
|
||
. Presumably
|
||
<taxonomicName id="E9E83AA8FFBE6702FB17FA7AFA306690" box="[1247,1436,1509,1534]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FB17FA7AFA306690" box="[1247,1436,1509,1534]" italics="true" pageId="3" pageNumber="52">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
belongs to that latter subclade of “Quediina propria”. The exact limits of this lineage within
|
||
<taxonomicName id="E9E83AA8FFBE6702FB28F994FADC654B" box="[1248,1392,1547,1573]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
|
||
are not finally clear yet: apparently it also includes the genus
|
||
<taxonomicName id="E9E83AA8FFBE6702FCAEF9ADFB346522" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[870,1176,1586,1612]" class="Insecta" family="Staphylinidae" genus="Anaquedius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FCAEF9ADFC5A6525" box="[870,1014,1586,1611]" italics="true" pageId="3" pageNumber="52">Anaquedius</emphasis>
|
||
<bibRefCitation id="4A793CDAFFBE6702FC37F9ADFB346522" author="Casey" box="[1023,1176,1586,1612]" pageId="3" pageNumber="57" refString="Casey, T. L. (1915) Studies in some staphylinid genera of North America. In: Memoirs on the Coleoptera, VI. The New Era Printing Co., Lancaster, Pennsylvania, pp. 395 - 460." type="book chapter" year="1915">Casey, 1915</bibRefCitation>
|
||
</taxonomicName>
|
||
, but may not include
|
||
<taxonomicName id="E9E83AA8FFBE6702FF5FF9C6FEB6651C" box="[151,282,1625,1650]" class="Insecta" family="Staphylinidae" genus="Euryporus" higherTaxonomySource="CoL" kingdom="Fungi" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FF5FF9C6FEB6651C" box="[151,282,1625,1650]" italics="true" pageId="3" pageNumber="52">Euryporus</emphasis>
|
||
</taxonomicName>
|
||
. All these genera here affiliated with
|
||
<taxonomicName id="E9E83AA8FFBE6702FD32F9C6FC17651C" box="[762,955,1625,1650]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FD32F9C6FC17651C" box="[762,955,1625,1650]" italics="true" pageId="3" pageNumber="52">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
share numerous character states (e.g., datamatrix in
|
||
<bibRefCitation id="4A793CDAFFBE6702FE98F9E0FCA465F7" author="Solodovnikov" box="[336,776,1663,1689]" pageId="3" pageNumber="57" refString="Solodovnikov, A. & Schomann, A. (2009) Revised systematics and biogeography of " Quediina " of sub-Saharan Africa bring new insights about the phylogeny of the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology, (in press)." type="book" year="2009">Solodovnikov & Schomann 2009</bibRefCitation>
|
||
; and work in progress) but only some of them are preliminarily thought to be synapomorphies: setose apical segments of the maxillary and labial palps; more or less distinctly elongate first antennomere; lack of conspicuous large macrosetae on mesoventrite; abdominal tergites III–VII with only one basal carina; and male sternite VIII without secondary sexual modifications, medially straight to very slightly concave. Within this tentative group, strongly dilated anterior tarsi and evenly setose head and pronotum are autapomorphies of
|
||
<emphasis id="1C9C9D39FFBE6702FCFAF8DEFC436434" box="[818,1007,1857,1882]" italics="true" pageId="3" pageNumber="52">
|
||
<taxonomicName id="E9E83AA8FFBE6702FCFAF8DEFC476434" box="[818,1003,1857,1882]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">Australotarsius</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2E57412BFFBE6705FF0EF8F8FE9A63B6" blockId="3.[151,1437,1199,2037]" lastBlockId="4.[151,1437,152,796]" lastPageId="4" lastPageNumber="53" pageId="3" pageNumber="52">
|
||
Of the listed genera, only one species of
|
||
<taxonomicName id="E9E83AA8FFBE6702FD3BF8F7FB3E64EF" authority="Solodovnikov 2008" authorityName="Solodovnikov" authorityYear="2008" box="[755,1170,1895,1921]" class="Insecta" family="Staphylinidae" genus="Anchocerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FD3BF8F7FCD264EF" box="[755,894,1896,1921]" italics="true" pageId="3" pageNumber="52">Anchocerus</emphasis>
|
||
(
|
||
<bibRefCitation id="4A793CDAFFBE6702FC5EF8F8FB2664EF" author="Solodovnikov" box="[918,1162,1895,1921]" pageId="3" pageNumber="57" refString="Solodovnikov, A. (2008) Review of the Oriental genus Anchocerus with the description of new species and new combinations (Coleoptera: Staphylinidae: Staphylininae). Insect Systematics and Evolution, 39 (3), 287 - 301." type="journal article" year="2008">Solodovnikov 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and five species of
|
||
<taxonomicName id="E9E83AA8FFBE6702FF5FF811FDC964C6" authority="Newton & Thayer 2005" authorityName="Newton & Thayer" authorityYear="2005" box="[151,613,1934,1960]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FF5FF811FE8164C9" box="[151,301,1934,1959]" italics="true" pageId="3" pageNumber="52">Acylophorus</emphasis>
|
||
(
|
||
<bibRefCitation id="4A793CDAFFBE6702FEF5F811FDF164C6" author="Newton" box="[317,605,1934,1960]" pageId="3" pageNumber="57" refString="Newton, A. F. & Thayer, M. K. (2005) Catalog of higher taxa of Staphyliniformia and genera and subgenera of Staphylinoidea. Field Museum of Natural History, Chicago. Available from: http: // www. fieldmuseum. org / peet _ staph / db _ 1 a. html (accessed 28. X. 2008)." type="journal article" year="2005">Newton & Thayer 2005</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
occur in
|
||
<collectingCountry id="56FF01BBFFBE6702FD1FF811FCE464C6" box="[727,840,1934,1960]" name="Australia" pageId="3" pageNumber="52">Australia</collectingCountry>
|
||
. But both species of
|
||
<taxonomicName id="E9E83AA8FFBE6702FB8AF811FB5564C9" box="[1090,1273,1934,1959]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FB8AF811FB5564C9" box="[1090,1273,1934,1959]" italics="true" pageId="3" pageNumber="52">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from the Australian
|
||
<taxonomicName id="E9E83AA8FFBE6702FDEEF82AFC0564A0" authority="Lea, 1929" authorityName="Lea" authorityYear="1929" box="[550,937,1972,1998]" class="Insecta" family="Staphylinidae" genus="Anchocerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="tenuipes">
|
||
<emphasis id="1C9C9D39FFBE6702FDEEF82AFCB664A0" box="[550,794,1973,1998]" italics="true" pageId="3" pageNumber="52">Anchocerus tenuipes</emphasis>
|
||
(
|
||
<bibRefCitation id="4A793CDAFFBE6702FCE1F82BFC0D64A0" author="Lea" box="[809,929,1972,1998]" pageId="3" pageNumber="57" refString="Lea, A. M. (1929) Notes on some miscellaneous Coleoptera, with descriptions of new species. Part VII. Transactions of the Royal Society of South Australia, 53, 203 - 244." type="journal article" year="1929">Lea, 1929</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
externally by very broad anterior tarsi and setose head and pronotum. The same characters also easily separate
|
||
<taxonomicName id="E9E83AA8FFBE6702FC02F844FB2D649A" box="[970,1153,2011,2036]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FC02F844FB2D649A" box="[970,1153,2011,2036]" italics="true" pageId="3" pageNumber="52">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="E9E83AA8FFBE6702FB03F844FACD649A" box="[1227,1377,2011,2036]" class="Insecta" family="Staphylinidae" genus="Acylophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFBE6702FB03F844FACD649A" box="[1227,1377,2011,2036]" italics="true" pageId="3" pageNumber="52">Acylophorus</emphasis>
|
||
</taxonomicName>
|
||
, and the latter genus, in addition, has a conspicuously narrow neck and a much more strongly elongate first antennomere.
|
||
</paragraph>
|
||
<paragraph id="2E57412BFFB96705FF0EFF7AFA966072" blockId="4.[151,1437,152,796]" pageId="4" pageNumber="53">
|
||
According to the preliminary phylogeny of
|
||
<taxonomicName id="E9E83AA8FFB96705FD07FF7AFCF26391" box="[719,862,229,255]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4A793CDAFFB96705FCA7FF7AFAAD6391" author="Solodovnikov" box="[879,1281,229,255]" pageId="4" pageNumber="57" refString="Solodovnikov, A. & Schomann, A. (2009) Revised systematics and biogeography of " Quediina " of sub-Saharan Africa bring new insights about the phylogeny of the rove beetle tribe Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology, (in press)." type="book" year="2009">Solodovnikov & Schomann 2009</bibRefCitation>
|
||
), a quediinelike
|
||
<taxonomicName id="E9E83AA8FFB96705FF1AFE93FE30624B" box="[210,412,268,293]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFB96705FF1AFE93FE30624B" box="[210,412,268,293]" italics="true" pageId="4" pageNumber="53">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
does not belong to “
|
||
<taxonomicName id="E9E83AA8FFB96705FD63FE93FCD46248" box="[683,888,268,294]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="subTribe" subTribe="Tanygnathinina">Tanygnathinina</taxonomicName>
|
||
sensu novo”, the lineage whose members superficially resemble “Quediina propria” and which is a predominant and the most abundant faunal element among Australian
|
||
<taxonomicName id="E9E83AA8FFB96705FEB9FEC6FDA8621D" box="[369,516,345,371]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
|
||
. Current Australian species of “
|
||
<taxonomicName id="E9E83AA8FFB96705FCB7FEC6FC4D621C" box="[895,993,345,370]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFB96705FCB7FEC6FC4D621C" box="[895,993,345,370]" italics="true" pageId="4" pageNumber="53">Quedius</emphasis>
|
||
</taxonomicName>
|
||
”, and “
|
||
<taxonomicName id="E9E83AA8FFB96705FBF5FEC6FB61621C" box="[1085,1229,345,370]" class="Insecta" family="Staphylinidae" genus="Heterothops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFB96705FBF5FEC6FB61621C" box="[1085,1229,345,370]" italics="true" pageId="4" pageNumber="53">Heterothops</emphasis>
|
||
</taxonomicName>
|
||
”, all members of “
|
||
<taxonomicName id="E9E83AA8FFB96705FF68FE1FFEF562F4" box="[160,345,384,410]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="subTribe" subTribe="Tanygnathinina">Tanygnathinina</taxonomicName>
|
||
sensu novo”, are not congeneric with the north temperate representatives of these genera; but some other, less species-rich Australian genera now formally in Quediina also belong to this group. From any species of “
|
||
<taxonomicName id="E9E83AA8FFB96705FED7FE52FE766289" box="[287,474,461,487]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="subTribe" subTribe="Tanygnathinina">Tanygnathinina</taxonomicName>
|
||
sensu novo”
|
||
<taxonomicName id="E9E83AA8FFB96705FDB5FE52FC986288" box="[637,820,461,486]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFB96705FDB5FE52FC986288" box="[637,820,461,486]" italics="true" pageId="4" pageNumber="53">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
differs in the structure of the mesoscutellum which has two (not one) transverse carinae, and in the shape of the paramere which is closely attached to the median lobe only at its base, and has distinct, strongly sclerotized and obtuse sensory peg setae (in “
|
||
<taxonomicName id="E9E83AA8FFB96705FB29FD85FA37615A" box="[1249,1435,538,564]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="subTribe" subTribe="Tanygnathinina">Tanygnathinina</taxonomicName>
|
||
sensu novo” the paramere is closely attached to the median lobe along the entire length of the latter, and lacks sensory peg setae, or, if those present, they are inconspicuous and sharp). By such combination of characters as very broad anterior tarsi, somewhat deflexed hypomera of the pronotum, evenly setose head and pronotum, strongly cuspidate teeth at the base of the mandibles, and well developed infraorbital ridges,
|
||
<taxonomicName id="E9E83AA8FFB96705FB2EFD2AFA3761A0" box="[1254,1435,693,718]" class="Insecta" family="Staphylinidae" genus="Australotarsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFB96705FB2EFD2AFA3761A0" box="[1254,1435,693,718]" italics="true" pageId="4" pageNumber="53">Australotarsius</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from any other genus of
|
||
<taxonomicName id="E9E83AA8FFB96705FCDFFD43FC076198" box="[791,939,732,758]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
|
||
recorded in
|
||
<collectingCountry id="56FF01BBFFB96705FB8EFD43FB196198" box="[1094,1205,732,758]" name="Australia" pageId="4" pageNumber="53">Australia</collectingCountry>
|
||
including the very poorly known and phylogenetically puzzling genera
|
||
<taxonomicName id="E9E83AA8FFB96705FD37FC9DFC4D6072" authority="Strand, 1943" authorityName="Strand" authorityYear="1943" box="[767,993,770,796]" class="Insecta" family="Staphylinidae" genus="Lonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFB96705FD37FC9DFCEF6075" box="[767,835,770,795]" italics="true" pageId="4" pageNumber="53">Lonia</emphasis>
|
||
<bibRefCitation id="4A793CDAFFB96705FC83FC9DFC4D6072" author="Strand" box="[843,993,770,796]" pageId="4" pageNumber="57" refString="Strand, E. (1943) Miscellanea nomenclatorica zoologica et paleontological, XI. Folia Zoologica et Hydrobiologica, 12, 94 - 114." type="journal article" year="1943">Strand, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E9E83AA8FFB96705FBD1FC9DFA9A6072" authority="Fauvel, 1878" authorityName="Fauvel" authorityYear="1878" box="[1049,1334,770,796]" class="Insecta" family="Staphylinidae" genus="Antimerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="53" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1C9C9D39FFB96705FBD1FC9DFB3D6075" box="[1049,1169,770,795]" italics="true" pageId="4" pageNumber="53">Antimerus</emphasis>
|
||
<bibRefCitation id="4A793CDAFFB96705FB51FC9DFA9A6072" author="Fauvel" box="[1177,1334,770,796]" pageId="4" pageNumber="57" refString="Fauvel, A. (1878) Les staphylinides de l'Australie et de la Polynesie. Annali del Museo Civico di Storia Naturale di Genova, 12, 171 - 315." type="journal article" year="1878">Fauvel, 1878</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |