245 lines
37 KiB
XML
245 lines
37 KiB
XML
<document id="7C16E9A773765292A6CD4CA51462B206" ID-CLB-Dataset="294590" ID-DOI="10.5194/fr-21-11-2018" ID-GBIF-Dataset="ab3dad8c-e248-4169-9ac2-79bbdca0377c" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965458" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.tables_requiresApprovalFor="GgImagineBatch" IM.treatments_approvedBy="diego" checkinTime="1712926397309" checkinUser="felipe" docAuthor="Ramassamy, Benjamin, Lambert, Olivier, Collareta, Alberto, Urbina, Mario & Bianucci, Giovanni" docDate="2018" docId="A63C8788FFC1493AFCC2F999FC35FEE6" docLanguage="en" docName="FossRec.21.1.11-32.pdf" docOrigin="Fossil Record 21 (1)" docSource="http://dx.doi.org/10.5194/fr-21-11-2018" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.1:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="1" docTitle="Messapicetus gregarius Bianucci, Lambert and Post 2011" docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="5A05FFF0FFC2493EFFABFFABFFABFFC1" masterDocTitle="Description of the skeleton of the fossil beaked whale Messapicetus gregarius: searching potential proxies for deep-diving abilities" masterLastPageNumber="32" masterPageNumber="11" pageNumber="14" updateTime="1713879443204" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="DC8B9F9C47345A24F5BFA0F0C0FD9E59">
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<mods:title id="679DF0978982A542F239642B591E768C">Description of the skeleton of the fossil beaked whale Messapicetus gregarius: searching potential proxies for deep-diving abilities</mods:title>
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<mods:name id="2A804E9C9298041F20DF7019EB6D6AB4" type="personal">
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<mods:namePart id="54DCDFC61D1E0A2882676E6B8886E1A2">Ramassamy, Benjamin</mods:namePart>
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<mods:namePart id="071FEAC2E54E8EDA5380A2E6EA4FAA3F">Lambert, Olivier</mods:namePart>
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<mods:roleTerm id="6EFC05B97EB9AF6368DDD0047A73D63F">Author</mods:roleTerm>
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<mods:namePart id="AB8DCD5385FC75B29B66F97839FBFC8B">Collareta, Alberto</mods:namePart>
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<mods:namePart id="BA57C2FB69A8268A6CAD4B053557EDBA">Urbina, Mario</mods:namePart>
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<mods:namePart id="4DDF2C71D25ED1C51DDF94898A8505B3">Bianucci, Giovanni</mods:namePart>
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<mods:title id="B2D87A184BA1605B79BACF393716E5BE">Fossil Record</mods:title>
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<mods:date id="693C97BCA7A69C30E9D3D2757B4BC740">2018</mods:date>
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<mods:number id="B165DB7DFD6530E2387ADD9DD8168C2D">2018-01-16</mods:number>
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<mods:number id="7E9C8C675675D06CFE4E9DEE465372D4">21</mods:number>
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<mods:start id="0AFE7FB6A2A5BF86B0AC813137271798">11</mods:start>
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<mods:identifier id="8F7A75ED8973B206D0C64319DE0C416E" type="DOI">10.5194/fr-21-11-2018</mods:identifier>
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<treatment id="A63C8788FFC1493AFCC2F999FC35FEE6" ID-DOI="http://doi.org/10.5281/zenodo.11047626" ID-Zenodo-Dep="11047626" LSID="urn:lsid:plazi:treatment:A63C8788FFC1493AFCC2F999FC35FEE6" httpUri="http://treatment.plazi.org/id/A63C8788FFC1493AFCC2F999FC35FEE6" lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14">
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<subSubSection id="668F6515FFC1493DFCC2F999FC72F9A8" pageId="3" pageNumber="14" type="nomenclature">
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<paragraph id="2E2A369EFFC1493DFCC2F999FC72F9A8" blockId="3.[873,1408,1586,1641]" pageId="3" pageNumber="14">
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<taxonomicName id="E9954D1DFFC1493DFCC2F999FC72F9A8" authority="Bianucci, Lambert and Post, 2011" authorityName="Bianucci, Lambert and Post" authorityYear="2011" class="Mammalia" family="Hyperoodontidae" genus="Messapicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="14" phylum="Chordata" rank="species" species="gregarius">
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<emphasis id="1CE1EA8CFFC1493DFCC2F999FBC7F98B" box="[873,1132,1586,1610]" italics="true" pageId="3" pageNumber="14">Messapicetus gregarius</emphasis>
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Bianucci, Lambert and Post, 2011
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="668F6515FFC1493AFC98F925FC35FEE6" lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14" type="materials_examined">
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<paragraph id="2E2A369EFFC1493DFC98F925FB90F967" blockId="3.[819,1083,1678,1702]" box="[819,1083,1678,1702]" pageId="3" pageNumber="14">
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<emphasis id="1CE1EA8CFFC1493DFC98F925FB90F967" bold="true" box="[819,1083,1678,1702]" pageId="3" pageNumber="14">3.1 Referred material</emphasis>
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</paragraph>
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<paragraph id="2E2A369EFFC1493AFC98F961FE8FFE86" blockId="3.[819,1462,1738,1921]" lastBlockId="4.[124,767,176,327]" lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14">
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Specimen
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<collectionCode id="4884AE5BFFC1493DFC01F961FC50F920" box="[938,1019,1738,1761]" pageId="3" pageNumber="14">MUSM</collectionCode>
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2548: three cervical vertebrae including the axis, three thoracic and two thoracic–post-thoracic vertebrae all lacking the neural spine (
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<figureCitation id="B6AE2A1BFFC1493DFB5FF8A1FAC9F8E0" box="[1268,1378,1802,1825]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStartId-0="5.[124,191,1483,1505]" captionStartId-1="6.[124,191,1508,1530]" captionTargetBox-0="[230,1348,197,1427]" captionTargetBox-1="[235,1345,232,1472]" captionTargetId-0="figure-17@5.[318,1348,-7,1383]" captionTargetId-1="figure-16@6.[257,1345,275,1424]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figure 2. Cervical vertebrae of the specimen MUSM 2548, Messapicetus gregarius. Axis in anterior (a), posterior (b), and ventral view (c); C5–C6 in anterior (d), posterior (e), and dorsal view (f); C7 in anterior (g), posterior (h), and dorsal view (i)." captionText-1="Figure 3. Thoracic and post-thoracic vertebrae of the specimen MUSM 2548, Messapicetus gregarius. Thoracic vertebra A in anterior (a), posterior (b), and dorsal view (c); thoracic B in posterior (d), left lateral (e), and ventral view (f); thoracic C in posterior (g), lateral (h), and ventral view (i); thoracic–post-thoracic D in posterior (j) and lateral view (k); thoracic–post-thoracic E in lateral view (l)." figureDoi-0="http://doi.org/10.5281/zenodo.10965462" figureDoi-1="http://doi.org/10.5281/zenodo.10965464" httpUri-0="https://zenodo.org/record/10965462/files/figure.png" httpUri-1="https://zenodo.org/record/10965464/files/figure.png" pageId="3" pageNumber="14">Figs. 2–3</figureCitation>
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), fused manubrium and left part of the second sternebra (
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<figureCitation id="B6AE2A1BFFC1493DFAEEF881FA2DF880" box="[1349,1414,1834,1857]" captionStart="Figure 4" captionStartId="7.[124,191,920,942]" captionTargetBox="[133,747,177,885]" captionTargetId="figure-16@7.[279,626,207,752]" captionText="Figure 4. Sternum of the specimen MUSM 2548, Messapicetus gregarius, in ventral view (a) and manubrium element in anterior view (b). Horizontal solid lines indicate major break. The dashed line represents a reconstruction of the second element of the sternebra 2." figureDoi="http://doi.org/10.5281/zenodo.11047628" httpUri="https://zenodo.org/record/11047628/files/figure.png" pageId="3" pageNumber="14">Fig. 4</figureCitation>
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), 11 complete to subcomplete ribs (
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<figureCitation id="B6AE2A1BFFC1493DFB28F8E1FB6DF8A0" box="[1155,1222,1866,1889]" captionStart="Figure 5" captionStartId="8.[124,191,780,802]" captionTargetBox="[233,1351,195,682]" captionTargetId="figure-17@8.[261,1314,197,677]" captionTargetPageId="8" captionText="Figure 5. Ribs of the specimen MUSM 2548, Messapicetus gregarius, in anterior view. Pair 1 (a) and (b); pair 2 (c) and (d); pair 3 (e) and (f); pair 4 (g) and (h); (i), (j), (k), and (l) cannot be precisely positioned." figureDoi="http://doi.org/10.5281/zenodo.10965466" httpUri="https://zenodo.org/record/10965466/files/figure.png" pageId="3" pageNumber="14">Fig. 5</figureCitation>
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); skull and mandibles are still in the field. Specimen
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<collectionCode id="4884AE5BFFC1493DFB3AF8C1FB49F840" box="[1169,1250,1898,1921]" pageId="3" pageNumber="14">MUSM</collectionCode>
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2542: partial right scapula lacking the acromion, the anterior part of the scapular blade, and the broken coracoid process (
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<figureCitation id="B6AE2A1BFFC6493AFDF4FF7BFD09FF26" box="[607,674,208,231]" captionStart="Figure 6" captionStartId="8.[124,191,1539,1561]" captionTargetBox="[149,746,927,1501]" captionTargetId="figure-118@8.[248,666,892,1488]" captionTargetPageId="8" captionText="Figure 6. Right scapula of the specimen MUSM 2542, Messapicetus gregarius, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10965468" httpUri="https://zenodo.org/record/10965468/files/figure.png" pageId="4" pageNumber="15">Fig. 6</figureCitation>
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); partial left and right humeri (
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<figureCitation id="B6AE2A1BFFC6493AFEC6FF5BFE1BFEC6" box="[365,432,240,263]" captionStart="Figure 7" captionStartId="9.[124,191,1765,1787]" captionTargetBox="[233,1350,286,1727]" captionTargetId="figure-38@9.[284,1350,344,1652]" captionTargetPageId="9" captionText="Figure 7. Humeri and left radius of MUSM 2542, Messapicetus gregarius. Right humerus in medial (a), lateral (b), ulnar (c), and radial view (d); left humerus in medial (e), lateral (f), ulnar (g), radial (h), and posterior view (i); left radius in lateral (j), medial (k), and ulnar view (l)." figureDoi="http://doi.org/10.5281/zenodo.10965470" httpUri="https://zenodo.org/record/10965470/files/figure.png" pageId="4" pageNumber="15">Fig. 7</figureCitation>
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), complete left radius (
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<figureCitation id="B6AE2A1BFFC6493AFD06FF5BFD5BFEC6" box="[685,752,240,263]" captionStart="Figure 8" captionStartId="12.[124,191,1789,1811]" captionTargetBox="[236,1353,239,1748]" captionTargetId="figure-17@12.[251,1232,318,1652]" captionTargetPageId="12" captionText="Figure 8. Comparison of muscular insertions along the atlas and axis in Messapicetus gregarius MUSM 2548 in ventral view (a) and posterior view (b); in Ninoziphius platyrostris MNHN SAS 941 in ventral view (c) and posterior view (d); in Mesoplodon densirostris USNM 593522 in ventral view (e) and posterior view (f); in Berardius sp. MNHN 1885-278." figureDoi="http://doi.org/10.5281/zenodo.10965472" httpUri="https://zenodo.org/record/10965472/files/figure.png" pageId="4" pageNumber="15">Fig. 8</figureCitation>
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); skull, mandibles, and other vertebrae of this individual are still in the field.
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</paragraph>
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<paragraph id="2E2A369EFFC6493AFFD7FEC2FE0BFE40" blockId="4.[124,416,361,385]" box="[124,416,361,385]" pageId="4" pageNumber="15">
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<emphasis id="1CE1EA8CFFC6493AFFD7FEC2FE0BFE40" bold="true" box="[124,416,361,385]" pageId="4" pageNumber="15">3.2 Horizon and locality</emphasis>
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</paragraph>
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<paragraph id="2E2A369EFFC6493AFFD7FE0EFD54FDDA" blockId="4.[124,767,421,1432]" pageId="4" pageNumber="15">
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<materialsCitation id="9EFD3CC3FFC6493AFFD7FE0EFD50FDDA" county="Pisco Formation" elevation="200" location="They" municipality="Cerro" pageId="4" pageNumber="15" specimenCount="1" stateProvince="Colorado">
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The specimens were discovered in the
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<quantity id="E96D9B7BFFC6493AFDBFFE0EFDFDFE7D" box="[532,598,421,444]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="15" unit="m" value="200.0">
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<elevation id="A5B8D1ADFFC6493AFDBFFE0EFDFDFE7D" box="[532,598,421,444]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="15" unit="m" value="200.0">200 m</elevation>
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</quantity>
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thick section of sediments from the
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<collectingCounty id="C74B4E12FFC6493AFEFEFE6EFDAEFE1D" box="[341,517,453,476]" pageId="4" pageNumber="15">Pisco Formation</collectingCounty>
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exposed at
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<collectingMunicipality id="CE4EACE4FFC6493AFD21FE6EFD6DFE1D" box="[650,710,453,476]" pageId="4" pageNumber="15">Cerro</collectingMunicipality>
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<collectingRegion id="EC51F87CFFC6493AFD64FE6EFF17FE3D" country="United States of America" name="Colorado" pageId="4" pageNumber="15">Colorado</collectingRegion>
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.
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<location id="2B4A6045FFC6493AFF60FE4EFEAAFE3D" LSID="urn:lsid:plazi:treatment:A63C8788FFC1493AFCC2F999FC35FEE6:2B4A6045FFC6493AFF60FE4EFEAAFE3D" box="[203,257,485,508]" county="Pisco Formation" municipality="Cerro" name="They" pageId="4" pageNumber="15" stateProvince="Colorado">They</location>
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were identified under the fieldwork numbers O37 and O
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<quantity id="E96D9B7BFFC6493AFF44FDAFFE8FFDDA" box="[239,292,516,539]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="4" pageNumber="15" unit="in" value="39.0">39 in</quantity>
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||
the map provided by
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<bibRefCitation id="4A044B6FFFC6493AFDACFDAFFD50FDDA" author="Bianucci, G. & Di Celma, C. & Landini, W. & Post, K. & Tinelli, C. & de Muizon, C. & Gariboldi, K. & Malinverno, E. & Cantalamessa, G. & Gioncada, A. & Collareta, A. & Salas-Gismondi, R. & Varas, R. & Stucchi, M. & Urbina, M. & Lambert, O." box="[519,763,516,539]" pageId="4" pageNumber="15" pagination="543 - 557" refId="ref12898" refString="Bianucci, G., Di Celma, C., Landini, W., Post, K., Tinelli, C., de Muizon, C., Gariboldi, K., Malinverno, E., Cantalamessa, G., Gioncada, A., Collareta, A., Salas-Gismondi, R., Varas, R., Stucchi, M., Urbina, M., and Lambert, O.: Distribution of fossil marine vertebrates in Cerro Colorado, the type locality of the giant raptorial sperm whale Livyatan melvillei (Miocene, Pisco Formation, Peru), Journal of Maps, 12, 543 - 557, 2016 b." type="journal article" year="2016">Bianucci et al. (2016b)</bibRefCitation>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="2E2A369EFFC6493AFF3CFD8FFE72FA59" blockId="4.[124,767,421,1432]" pageId="4" pageNumber="15">
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Two depositional sequences of the Pisco Formation, P1 and P2, are represented at Cerro
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<collectingRegion id="EC51F87CFFC6493AFE74FDEFFDE9FD9A" box="[479,578,580,603]" country="United States of America" name="Colorado" pageId="4" pageNumber="15">Colorado</collectingRegion>
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and separated by an angular unconformity (
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<bibRefCitation id="4A044B6FFFC6493AFE38FDCFFDD5FDBA" author="Di Celma, C. & Malinverno, E. & Gariboldi, K. & Gioncada, A. & Rustichelli, A. & Pierantoni, P. & Landini, W. & Bosio, G. & Tinelli, C. & Bianucci, G." box="[403,638,612,635]" pageId="4" pageNumber="15" pagination="1 - 15" refId="ref13898" refString="Di Celma, C., Malinverno, E., Gariboldi, K., Gioncada, A., Rustichelli, A., Pierantoni, P., Landini, W., Bosio, G., Tinelli, C., and Bianucci, G.: Stratigraphic framework of the late Miocene to Pliocene Pisco Formation at Cerro Colorado (Ica Desert, Peru), Journal of Maps, 12, 1 - 15, 2016." type="journal article" year="2016">Di Celma et al., 2016</bibRefCitation>
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, 2017). The specimens
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<collectionCode id="4884AE5BFFC6493AFF5DFD2FFEECFD5A" box="[246,327,644,667]" pageId="4" pageNumber="15">MUSM</collectionCode>
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2542 and
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<collectionCode id="4884AE5BFFC6493AFE6FFD2FFDBEFD5A" box="[452,533,644,667]" pageId="4" pageNumber="15">MUSM</collectionCode>
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2548 originate from the P1 sequence (the “lower allomember” of
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<bibRefCitation id="4A044B6FFFC6493AFDDDFD0FFF74FD1A" author="Di Celma, C. & Malinverno, E. & Gariboldi, K. & Gioncada, A. & Rustichelli, A. & Pierantoni, P. & Landini, W. & Bosio, G. & Tinelli, C. & Bianucci, G." pageId="4" pageNumber="15" pagination="1 - 15" refId="ref13898" refString="Di Celma, C., Malinverno, E., Gariboldi, K., Gioncada, A., Rustichelli, A., Pierantoni, P., Landini, W., Bosio, G., Tinelli, C., and Bianucci, G.: Stratigraphic framework of the late Miocene to Pliocene Pisco Formation at Cerro Colorado (Ica Desert, Peru), Journal of Maps, 12, 1 - 15, 2016." type="journal article" year="2016">Di Celma et al., 2016</bibRefCitation>
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), consisting of nearshore conglomerates and finegrained sandstones, bioturbated sandy siltstones, and mudstones (
|
||
<bibRefCitation id="4A044B6FFFC6493AFF79FCA8FE64FCDB" author="Di Celma, C. & Malinverno, E. & Gariboldi, K. & Gioncada, A. & Rustichelli, A. & Pierantoni, P. & Landini, W. & Bosio, G. & Tinelli, C. & Bianucci, G." box="[210,463,771,794]" pageId="4" pageNumber="15" pagination="1 - 15" refId="ref13898" refString="Di Celma, C., Malinverno, E., Gariboldi, K., Gioncada, A., Rustichelli, A., Pierantoni, P., Landini, W., Bosio, G., Tinelli, C., and Bianucci, G.: Stratigraphic framework of the late Miocene to Pliocene Pisco Formation at Cerro Colorado (Ica Desert, Peru), Journal of Maps, 12, 1 - 15, 2016." type="journal article" year="2016">Di Celma et al., 2016</bibRefCitation>
|
||
). The lower allomember is rich in marine fossil vertebrates, such as cartilaginous and bony fish, marine turtles, crocodiles, sea turtles, sea birds, and seals (
|
||
<bibRefCitation id="4A044B6FFFC6493AFF5AFCC8FE43FCBB" author="Bianucci, G. & Di Celma, C. & Landini, W. & Post, K. & Tinelli, C. & de Muizon, C. & Gariboldi, K. & Malinverno, E. & Cantalamessa, G. & Gioncada, A. & Collareta, A. & Salas-Gismondi, R. & Varas, R. & Stucchi, M. & Urbina, M. & Lambert, O." box="[241,488,867,890]" pageId="4" pageNumber="15" pagination="543 - 557" refId="ref12898" refString="Bianucci, G., Di Celma, C., Landini, W., Post, K., Tinelli, C., de Muizon, C., Gariboldi, K., Malinverno, E., Cantalamessa, G., Gioncada, A., Collareta, A., Salas-Gismondi, R., Varas, R., Stucchi, M., Urbina, M., and Lambert, O.: Distribution of fossil marine vertebrates in Cerro Colorado, the type locality of the giant raptorial sperm whale Livyatan melvillei (Miocene, Pisco Formation, Peru), Journal of Maps, 12, 543 - 557, 2016 b." type="journal article" year="2016">Bianucci et al., 2016b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A044B6FFFC6493AFE5EFCC8FD77FCBB" author="Landini, W. & Altamirano-Sierra, A. & Collareta, A. & Di Celma, C. & Urbina, M. & Bianucci, G." box="[501,732,867,890]" pageId="4" pageNumber="15" pagination="168 - 190" refId="ref15115" refString="Landini, W., Altamirano-Sierra, A., Collareta, A., Di Celma, C., Urbina, M., and Bianucci, G.: The late Miocene elasmobranch assemblage from Cerro Colorado (Pisco Formation, Peru), J. S. Am. Earth Sci., 73, 168 - 190, 2017 a." type="journal article" year="2017">Landini et al., 2017a</bibRefCitation>
|
||
, b;
|
||
<bibRefCitation id="4A044B6FFFC6493AFFD7FC28FE09FC5B" author="Parham, J. F. & Pyenson, N. D." box="[124,418,899,922]" pageId="4" pageNumber="15" pagination="231 - 247" refId="ref15847" refString="Parham, J. F. and Pyenson, N. D.: New sea turtle from the Miocene of Peru and the iterative evolution of feeding ecomorphologies since the Cretaceous, J. Paleontol., 84, 231 - 247, 2010." type="journal article" year="2010">Parham and Pyenson, 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A044B6FFFC6493AFE05FC28FD2AFC5B" author="Stucchi, M. & Varas-Malca, R. M. & Urbina-Schmitt, M." box="[430,641,899,922]" pageId="4" pageNumber="15" pagination="417 - 427" refId="ref16377" refString="Stucchi, M., Varas-Malca, R. M., and Urbina-Schmitt, M.: New Miocene sulid birds from Peru and considerations on their Neogene fossil record in the Eastern Pacific Ocean, Acta Palaeontologica Polonica, 61, 417 - 427, https: // doi. org / 10.4202 / app. 00170.2015, 2016." type="journal article" year="2016">Stucchi et al., 2016</bibRefCitation>
|
||
). Cetaceans are represented by diverse taxa: Physeteroidea (
|
||
<taxonomicName id="E9954D1DFFC6493AFD09FC09FF7CFC1B" authorityName="Lambert, Bianucci, Post, de Muizon, Salas-Gismondi, Urbina & Reumer" authorityYear="2010" class="Mammalia" genus="Livyatan" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="species" species="melvillei">
|
||
<emphasis id="1CE1EA8CFFC6493AFD09FC09FF7CFC1B" italics="true" pageId="4" pageNumber="15">Livyatan melvillei</emphasis>
|
||
</taxonomicName>
|
||
and aff.
|
||
<taxonomicName id="E9954D1DFFC6493AFEFBFC69FDA7FC1B" box="[336,524,962,986]" class="Mammalia" genus="Acrophyseter" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="1CE1EA8CFFC6493AFEFBFC69FE77FC1B" box="[336,476,962,986]" italics="true" pageId="4" pageNumber="15">Acrophyseter</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="E9954D1DFFC6493AFD82FC68FD24FC1B" authorityName="Gray" authorityYear="1850" box="[553,655,963,986]" family="Ziphiidae" pageId="4" pageNumber="15" rank="family">Ziphiidae</taxonomicName>
|
||
(
|
||
<taxonomicName id="E9954D1DFFC6493AFD0CFC69FF6BFC3B" authorityName="Bianucci, Lambert and Post" authorityYear="2011" class="Mammalia" family="Hyperoodontidae" genus="Messapicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="species" species="gregarius">
|
||
<emphasis id="1CE1EA8CFFC6493AFD0CFC69FF6BFC3B" italics="true" pageId="4" pageNumber="15">M. gregarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9954D1DFFC6493AFF7EFC49FE56FC3B" authorityName="Bianucci, Di Celma, Urbina & Lambert" authorityYear="2016" box="[213,509,994,1018]" class="Mammalia" family="Hyperoodontidae" genus="Chimuziphius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="species" species="coloradensis">
|
||
<emphasis id="1CE1EA8CFFC6493AFF7EFC49FE56FC3B" box="[213,509,994,1018]" italics="true" pageId="4" pageNumber="15">Chimuziphius coloradensis</emphasis>
|
||
</taxonomicName>
|
||
), Inioidea (
|
||
<taxonomicName id="E9954D1DFFC6493AFD22FC49FEA2FBDB" authorityName="Muizon" authorityYear="1988" class="Mammalia" family="Iniidae" genus="Brachydelphis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="species" species="mazeasi">
|
||
<emphasis id="1CE1EA8CFFC6493AFD22FC49FEA2FBDB" italics="true" pageId="4" pageNumber="15">Brachydelphis mazeasi</emphasis>
|
||
</taxonomicName>
|
||
and an undescribed taxon), two undescribed kentriodontid-like Delphinida, Balaenopteroidea, and
|
||
<taxonomicName id="E9954D1DFFC6493AFD73FB89FF45FB98" class="Mammalia" family="Cetotheriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="family">Cetotheriidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4A044B6FFFC6493AFF56FBE9FE4AFB98" author="Bianucci, G. & Lambert, O. & Post, K." box="[253,481,1090,1113]" pageId="4" pageNumber="15" pagination="1077 - 1098" refId="ref12692" refString="Bianucci, G., Lambert, O., and Post, K.: High concentration of long-snouted beaked whales (genus Messapicetus) from the Miocene of Peru, Palaeontology, 53, 1077 - 1098, https: // doi. org / 10.1111 / j. 1475 - 4983.2010.00995. x, 2010." type="journal article" year="2010">Bianucci et al., 2010</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4A044B6FFFC6493AFE46FBE9FD9AFB98" author="Bianucci, G. & Di Celma, C. & Landini, W. & Post, K. & Tinelli, C. & de Muizon, C. & Gariboldi, K. & Malinverno, E. & Cantalamessa, G. & Gioncada, A. & Collareta, A. & Salas-Gismondi, R. & Varas, R. & Stucchi, M. & Urbina, M. & Lambert, O." box="[493,561,1090,1113]" pageId="4" pageNumber="15" pagination="543 - 557" refId="ref12898" refString="Bianucci, G., Di Celma, C., Landini, W., Post, K., Tinelli, C., de Muizon, C., Gariboldi, K., Malinverno, E., Cantalamessa, G., Gioncada, A., Collareta, A., Salas-Gismondi, R., Varas, R., Stucchi, M., Urbina, M., and Lambert, O.: Distribution of fossil marine vertebrates in Cerro Colorado, the type locality of the giant raptorial sperm whale Livyatan melvillei (Miocene, Pisco Formation, Peru), Journal of Maps, 12, 543 - 557, 2016 b." type="journal article" year="2016">2016b</bibRefCitation>
|
||
, c;
|
||
<bibRefCitation id="4A044B6FFFC6493AFDF2FBE9FF1FFBB8" author="Collareta, A. & Landini, W. & Lambert, O. & Post, K. & Tinelli, C. & Di Celma, C. & Panetta, D. & Tripodi, M. & Salvadori, P. A. & Caramella, D. & Marchi, D. & Urbina, M. & Bianucci, G." pageId="4" pageNumber="15" pagination="1 - 12" refId="ref13174" refString="Collareta, A., Landini, W., Lambert, O., Post, K., Tinelli, C., Di Celma, C., Panetta, D., Tripodi, M., Salvadori, P. A., Caramella, D., Marchi, D., Urbina, M., and Bianucci, G.: Piscivory in a Miocene Cetotheriidae of Peru: first record of fossilized stomach content for an extinct baleen-bearing whale, The Science of Nature, 102, 1 - 12, 2015." type="journal article" year="2015">Collareta et al., 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A044B6FFFC6493AFF6BFBC9FE0DFBB8" author="Gioncada, A. & Collareta, A. & Gariboldi, K. & Lambert, O. & Di Celma, C. & Bonaccorsi, E. & Urbina, M. & Bianucci, G." box="[192,422,1122,1145]" pageId="4" pageNumber="15" pagination="839 - 842" refId="ref14449" refString="Gioncada, A., Collareta, A., Gariboldi, K., Lambert, O., Di Celma, C., Bonaccorsi, E., Urbina, M., and Bianucci, G.: Inside baleen: Exceptional microstructure preservation in a late Miocene whale skeleton from Peru, Geology, 44, 839 - 842, 2016." type="journal article" year="2016">Gioncada et al., 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A044B6FFFC6493AFE19FBC9FD31FBB8" author="Lambert, O. & Bianucci, G. & Post, K." box="[434,666,1122,1145]" pageId="4" pageNumber="15" pagination="19 - 26" refId="ref14837" refString="Lambert, O., Bianucci, G., and Post, K.: Tusk-bearing beaked whales from the Miocene of Peru: sexual dimorphism in fossil ziphiids?, J. Mammal., 91, 19 - 26, https: // doi. org / 10.1644 / 08 - MAMM-A- 388 R 1.1, 2010 a." type="journal article" year="2010">Lambert et al., 2010a</bibRefCitation>
|
||
). The geological age of the lower allomember is estimated as 9.9– 8.9 Ma (Tortonian, Late Miocene) based on the presence of the diatom species
|
||
<taxonomicName id="E9954D1DFFC6493AFEF9FB6AFDF0FB18" authorityName="Barron" authorityYear="1976" box="[338,603,1217,1241]" class="Bacillariophyceae" family="Lithodesmiaceae" genus="Lithodesmium" higherTaxonomySource="GBIF" kingdom="Chromista" order="Lithodesmiales" pageId="4" pageNumber="15" phylum="Ochrophyta" rank="species" species="reynoldsii">
|
||
<emphasis id="1CE1EA8CFFC6493AFEF9FB6AFDF0FB18" box="[338,603,1217,1241]" italics="true" pageId="4" pageNumber="15">Lithodesmium reynoldsii</emphasis>
|
||
</taxonomicName>
|
||
and radiometric dating of a local ash layer (
|
||
<bibRefCitation id="4A044B6FFFC6493AFE60FB49FD12FB38" author="Di Celma, C. & Malinverno, E. & Gariboldi, K. & Gioncada, A. & Rustichelli, A. & Pierantoni, P. & Landini, W. & Bosio, G. & Tinelli, C. & Bianucci, G." box="[459,697,1250,1273]" pageId="4" pageNumber="15" pagination="1 - 15" refId="ref13898" refString="Di Celma, C., Malinverno, E., Gariboldi, K., Gioncada, A., Rustichelli, A., Pierantoni, P., Landini, W., Bosio, G., Tinelli, C., and Bianucci, G.: Stratigraphic framework of the late Miocene to Pliocene Pisco Formation at Cerro Colorado (Ica Desert, Peru), Journal of Maps, 12, 1 - 15, 2016." type="journal article" year="2016">Di Celma et al., 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A044B6FFFC6493AFD6DFB49FE98FAD8" author="Gariboldi, K. & Bosio, G. & Malinverno, E. & Gioncada, A. & Di Celma, C. & Villa, I. M. & Urbina, M. & Bianucci, G." pageId="4" pageNumber="15" refId="ref14376" refString="Gariboldi, K., Bosio, G., Malinverno, E., Gioncada, A., Di Celma, C., Villa, I. M., Urbina, M., and Bianucci, G.: Biostratigraphy, geochronology and sedimentation rates of the upper Miocene Pisco Formation at two important marine vertebrate fossilbearing sites of southern Peru, Newsletters on Stratigraphy, 2017." type="book" year="2017">Gariboldi et al., 2017</bibRefCitation>
|
||
). Diatom genera found in the lower and upper allomembers (e.g.,
|
||
<taxonomicName id="E9954D1DFFC6493AFED0FA8AFE45FAF8" authorityName="G.W.Andrews" authorityYear="1977" box="[379,494,1313,1337]" class="Bacillariophyceae" family="Rhaphoneidaceae" genus="Delphineis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Rhaponeidales" pageId="4" pageNumber="15" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="1CE1EA8CFFC6493AFED0FA8AFE45FAF8" box="[379,494,1313,1337]" italics="true" pageId="4" pageNumber="15">Delphineis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9954D1DFFC6493AFE54FA8AFDC2FAF8" authorityName="Schaffer" authorityYear="1897" box="[511,617,1313,1337]" class="Collembola" family="Odontellidae" genus="Odontella" kingdom="Animalia" order="Poduromorpha" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1CE1EA8CFFC6493AFE54FA8AFDC2FAF8" box="[511,617,1313,1337]" italics="true" pageId="4" pageNumber="15">Odontella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9954D1DFFC6493AFDD1FA8AFD5CFAF8" authorityName="C.G.Ehrenberg" authorityYear="1844" box="[634,759,1313,1337]" class="Bacillariophyceae" family="Rhaphoneidaceae" genus="Rhaphoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Rhaponeidales" pageId="4" pageNumber="15" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="1CE1EA8CFFC6493AFDD1FA8AFD5CFAF8" box="[634,759,1313,1337]" italics="true" pageId="4" pageNumber="15">Rhaphoneis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9954D1DFFC6493AFFD7FAEAFEA2FA98" authorityName="K.Blazé" authorityYear="1984" box="[124,265,1345,1369]" class="Bacillariophyceae" family="Rhaphoneidaceae" genus="Diplomenora" higherTaxonomySource="GBIF" kingdom="Chromista" order="Rhaponeidales" pageId="4" pageNumber="15" phylum="Ochrophyta" rank="genus">
|
||
<emphasis id="1CE1EA8CFFC6493AFFD7FAEAFEA2FA98" box="[124,265,1345,1369]" italics="true" pageId="4" pageNumber="15">Diplomenora</emphasis>
|
||
</taxonomicName>
|
||
) are typical of a neritic environment, whereas open-ocean diatoms species are less frequent (
|
||
<bibRefCitation id="4A044B6FFFC6493AFDDCFACAFF70FA59" author="Di Celma, C. & Malinverno, E. & Gariboldi, K. & Gioncada, A. & Rustichelli, A. & Pierantoni, P. & Landini, W. & Bosio, G. & Tinelli, C. & Bianucci, G." pageId="4" pageNumber="15" pagination="1 - 15" refId="ref13898" refString="Di Celma, C., Malinverno, E., Gariboldi, K., Gioncada, A., Rustichelli, A., Pierantoni, P., Landini, W., Bosio, G., Tinelli, C., and Bianucci, G.: Stratigraphic framework of the late Miocene to Pliocene Pisco Formation at Cerro Colorado (Ica Desert, Peru), Journal of Maps, 12, 1 - 15, 2016." type="journal article" year="2016">Di Celma et al., 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A044B6FFFC6493AFF4DFA2AFE65FA59" author="Gariboldi, K. & Bosio, G. & Malinverno, E. & Gioncada, A. & Di Celma, C. & Villa, I. M. & Urbina, M. & Bianucci, G." box="[230,462,1409,1432]" pageId="4" pageNumber="15" refId="ref14376" refString="Gariboldi, K., Bosio, G., Malinverno, E., Gioncada, A., Di Celma, C., Villa, I. M., Urbina, M., and Bianucci, G.: Biostratigraphy, geochronology and sedimentation rates of the upper Miocene Pisco Formation at two important marine vertebrate fossilbearing sites of southern Peru, Newsletters on Stratigraphy, 2017." type="book" year="2017">Gariboldi et al., 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2E2A369EFFC6493AFFD7FA11FE1FFA13" blockId="4.[124,436,1466,1490]" box="[124,436,1466,1490]" pageId="4" pageNumber="15">
|
||
<emphasis id="1CE1EA8CFFC6493AFFD7FA11FE1FFA13" bold="true" box="[124,436,1466,1490]" pageId="4" pageNumber="15">3.3 Systematic attribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2E2A369EFFC6493AFFD7FA5DFE6BF88D" blockId="4.[124,767,1526,1868]" pageId="4" pageNumber="15">
|
||
Both specimens were identified based on complete skulls and mandibles that were left on the field. They are attributed to the family
|
||
<taxonomicName id="E9954D1DFFC6493AFF5BF99DFEFDF98C" authorityName="Gray" authorityYear="1850" box="[240,342,1590,1613]" family="Ziphiidae" pageId="4" pageNumber="15" rank="family">Ziphiidae</taxonomicName>
|
||
based on the enlargement of the hamular fossa of the pterygoid. In
|
||
<collectionCode id="4884AE5BFFC6493AFE6CF9FDFDB3F9AC" box="[455,536,1622,1645]" pageId="4" pageNumber="15">MUSM</collectionCode>
|
||
2548, this is further confirmed by the presence of an enlarged apical alveolus on the mandible. The two skulls have an extremely elongated rostrum, representing approximately 75 % of the total condylobasal length. Furthermore,
|
||
<collectionCode id="4884AE5BFFC6493AFE44F97EFDEBF92D" box="[495,576,1749,1772]" pageId="4" pageNumber="15">MUSM</collectionCode>
|
||
2548 displays the medial fusion of the premaxillae dorsal to the mesorostral groove along the rostrum. These two characteristics are typical of the genus
|
||
<taxonomicName id="E9954D1DFFC6493AFE81F89FFE12F88D" authorityName="Bianucci, Landini and Varola" authorityYear="1992" box="[298,441,1844,1868]" class="Mammalia" family="Hyperoodontidae" genus="Messapicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CE1EA8CFFC6493AFE81F89FFE12F88D" box="[298,441,1844,1868]" italics="true" pageId="4" pageNumber="15">Messapicetus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2E2A369EFFC6493AFCE5FF1BFC35FEE6" blockId="4.[819,1462,176,295]" pageId="4" pageNumber="15">
|
||
<materialsCitation id="9EFD3CC3FFC6493AFCE5FF1BFC31FEE6" location="Cerro" municipality="The" pageId="4" pageNumber="15" specimenCount="13" stateProvince="Colorado">
|
||
<collectingMunicipality id="CE4EACE4FFC6493AFCE5FF1BFCDCFF06" box="[846,887,176,199]" pageId="4" pageNumber="15">The</collectingMunicipality>
|
||
species
|
||
<taxonomicName id="E9954D1DFFC6493AFC75FF1BFBC6FF09" authorityName="Bianucci, Lambert and Post" authorityYear="2011" box="[990,1133,176,200]" class="Mammalia" family="Hyperoodontidae" genus="Messapicetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="15" phylum="Chordata" rank="species" species="gregarius">
|
||
<emphasis id="1CE1EA8CFFC6493AFC75FF1BFBC6FF09" box="[990,1133,176,200]" italics="true" pageId="4" pageNumber="15">M. gregarius</emphasis>
|
||
</taxonomicName>
|
||
is by far the most common ziphiid species in the lower allomember of
|
||
<location id="2B4A6045FFC6493AFAA3FF7BFAEFFF26" LSID="urn:lsid:plazi:treatment:A63C8788FFC1493AFCC2F999FC35FEE6:2B4A6045FFC6493AFAA3FF7BFAEFFF26" box="[1288,1348,208,231]" municipality="The" name="Cerro" pageId="4" pageNumber="15" stateProvince="Colorado">Cerro</location>
|
||
<collectingRegion id="EC51F87CFFC6493AFAE7FF7BFA19FF26" box="[1356,1458,208,231]" country="United States of America" name="Colorado" pageId="4" pageNumber="15">Colorado</collectingRegion>
|
||
, being represented by at least
|
||
<specimenCount id="3893FD17FFC6493AFBDAFF5BFAA9FEC6" box="[1137,1282,240,263]" count="13" pageId="4" pageNumber="15" type="generic">13 specimens</specimenCount>
|
||
(
|
||
<bibRefCitation id="4A044B6FFFC6493AFAB9FF5BFCD3FEE6" author="Bianucci, G. & Di Celma, C. & Landini, W. & Post, K. & Tinelli, C. & de Muizon, C. & Gariboldi, K. & Malinverno, E. & Cantalamessa, G. & Gioncada, A. & Collareta, A. & Salas-Gismondi, R. & Varas, R. & Stucchi, M. & Urbina, M. & Lambert, O." pageId="4" pageNumber="15" pagination="543 - 557" refId="ref12898" refString="Bianucci, G., Di Celma, C., Landini, W., Post, K., Tinelli, C., de Muizon, C., Gariboldi, K., Malinverno, E., Cantalamessa, G., Gioncada, A., Collareta, A., Salas-Gismondi, R., Varas, R., Stucchi, M., Urbina, M., and Lambert, O.: Distribution of fossil marine vertebrates in Cerro Colorado, the type locality of the giant raptorial sperm whale Livyatan melvillei (Miocene, Pisco Formation, Peru), Journal of Maps, 12, 543 - 557, 2016 b." type="journal article" year="2016">Bianucci et al., 2016b</bibRefCitation>
|
||
, c)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |