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<document id="44534E7E04DBAF358E076214A5E665CA" ID-DOI="10.11646/zootaxa.4758.2.6" ID-GBIF-Dataset="940e9590-f4cf-4960-8e23-76a1cfc6f88a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3734498" ID-ZooBank="A570EC50-061D-4DE5-8BEF-7A4A3B5C05D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585658104401" checkinUser="plazi" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2020" docId="A63287EF1A5C347394D907B0FCCEA6D7" docLanguage="en" docName="zootaxa.4758.2.6.pdf" docOrigin="Zootaxa 4758 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phrurolithus luppovae Spassky 1941" docType="treatment" docVersion="5" lastPageNumber="322" masterDocId="5A0BFF971A543478944E0115FFFCA570" masterDocTitle="A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia" masterLastPageNumber="329" masterPageNumber="311" pageNumber="319" updateTime="1698733622106" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="035ACAB51D768EF93138A2D958E461CE">A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia</mods:title>
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<mods:namePart id="6650EDF35840117117394191EBCBAE99">Zamani, Alireza</mods:namePart>
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<mods:namePart id="DDF453F713A5FB98ACE02144A0579B0C">Marusik, Yuri M.</mods:namePart>
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<treatment id="A63287EF1A5C347394D907B0FCCEA6D7" ID-DOI="http://doi.org/10.5281/zenodo.3812285" ID-GBIF-Taxon="163417986" ID-Zenodo-Dep="3812285" LSID="urn:lsid:plazi:treatment:A63287EF1A5C347394D907B0FCCEA6D7" httpUri="http://treatment.plazi.org/id/A63287EF1A5C347394D907B0FCCEA6D7" lastPageId="11" lastPageNumber="322" pageId="8" pageNumber="319">
<subSubSection id="668165721A5C347094D907B0FDA1A3CF" box="[151,605,1701,1728]" pageId="8" pageNumber="319" type="nomenclature">
<paragraph id="2E2436F91A5C347094D907B0FDA1A3CF" blockId="8.[151,605,1701,1764]" box="[151,605,1701,1728]" pageId="8" pageNumber="319">
<heading id="756C81951A5C347094D907B0FDA1A3CF" bold="true" box="[151,605,1701,1728]" fontSize="11" level="1" pageId="8" pageNumber="319" reason="1">
<taxonomicName id="E99B4D7A1A5C347094D907B0FDA1A3CF" authority="Spassky, 1941" authorityName="Spassky" authorityYear="1941" box="[151,605,1701,1728]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="319" phylum="Arthropoda" rank="species" species="luppovae">
<emphasis id="1CEFEAEB1A5C347094D907B0FDA1A3CF" bold="true" box="[151,605,1701,1728]" pageId="8" pageNumber="319">
<emphasis id="1CEFEAEB1A5C347094D907B0FE54A3B0" bold="true" box="[151,424,1701,1728]" italics="true" pageId="8" pageNumber="319">Phrurolithus luppovae</emphasis>
<bibRefCitation id="4A0A4B081A5C347095FE07B0FDA1A3CF" author="Spassky, S." box="[432,605,1701,1727]" pageId="8" pageNumber="319" pagination="12 - 26" refId="ref9417" refString="Spassky, S. (1941) Araneae palaearcticae novae VI. Folia Zoologica et Hydrobiologica, Riga, 11, 12 - 26." type="journal article" year="1941">Spassky, 1941</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165721A5C347094D907DCFE9CA394" box="[151,352,1737,1764]" pageId="8" pageNumber="319" type="description">
<paragraph id="2E2436F91A5C347094D907DCFE9CA394" blockId="8.[151,605,1701,1764]" box="[151,352,1737,1764]" pageId="8" pageNumber="319">
<figureCitation id="B6A02A7C1A5C347094D907DCFF27A394" box="[151,219,1737,1764]" captionStart="FIGURE 7" captionStartId="9.[151,250,1896,1921]" captionTargetBox="[236,1361,184,1866]" captionTargetId="figure@9.[219,1368,181,1870]" captionTargetPageId="9" captionText="FIGURE 7. General appearance and epigyne of Phrurolithus luppovae. A, C–D female habitus, dorsal, lateral and ventral; B, E male habitus, dorsal and ventral; F male prosoma, frontal; G macerated epigyne, dorsal; H intact epigyne, ventral. Scale=0.2 mm, if not otherwise indicated. Abbreviations: Co—copulatory opening, Fd—fertilization duct, Gr—receptacular gland, Pr— primary receptacle, Ro—receptacular outgrowth, Sr—secondary receptacle." figureDoi="http://doi.org/10.5281/zenodo.3734514" httpUri="https://zenodo.org/record/3734514/files/figure.png" pageId="8" pageNumber="319">Figs 7</figureCitation>
A–H, 8A–F
</paragraph>
</subSubSection>
<subSubSection id="668165721A5C347094D9061BFD1AA234" pageId="8" pageNumber="319" type="reference_group">
<paragraph id="2E2436F91A5C347094D9061BFD44A256" blockId="8.[151,742,1805,1861]" box="[151,696,1805,1830]" pageId="8" pageNumber="319">
<treatmentCitationGroup id="0E8B11D71A5C347094D9061BFD44A256" box="[151,696,1805,1830]" pageId="8" pageNumber="319">
<treatmentCitation id="AF3A10E81A5C347094D9061BFDC7A256" author="Spassky, S." box="[151,571,1805,1830]" page="24" pageId="8" pageNumber="319" year="1941">
<taxonomicName id="E99B4D7A1A5C347094D9061BFDC7A256" authority="Spassky, 1941: 24" authorityName="Spassky" authorityPageNumber="24" authorityYear="1941" box="[151,571,1805,1830]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="319" phylum="Arthropoda" rank="species" species="luppovae">
<emphasis id="1CEFEAEB1A5C347094D9061BFE87A256" box="[151,379,1806,1830]" italics="true" pageId="8" pageNumber="319">Phrurolithus luppovae</emphasis>
<bibRefCitation id="4A0A4B081A5C347095CF0618FDC7A256" author="Spassky, S." box="[385,571,1805,1830]" pageId="8" pageNumber="319" pagination="12 - 26" refId="ref9417" refString="Spassky, S. (1941) Araneae palaearcticae novae VI. Folia Zoologica et Hydrobiologica, Riga, 11, 12 - 26." type="journal article" year="1941">Spassky, 1941: 24</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 13 (
<specimenCount id="389DFD701A5C347096D9061BFD50A256" box="[663,684,1806,1830]" pageId="8" pageNumber="319" type="male">♂</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2E2436F91A5C347094D90639FD1AA234" blockId="8.[151,742,1805,1861]" box="[151,742,1836,1861]" pageId="8" pageNumber="319">
<treatmentCitationGroup id="0E8B11D71A5C347094D90639FD1AA234" box="[151,742,1836,1861]" pageId="8" pageNumber="319">
<treatmentCitation id="AF3A10E81A5C347094D90639FD48A234" author="Spassky, S. A. &amp; Luppova, E." box="[151,692,1836,1861]" page="49" pageId="8" pageNumber="319" year="1945">
<taxonomicName id="E99B4D7A1A5C347094D90639FD48A234" authority="Spassky &amp; Luppova, 1945: 49" authorityName="Spassky &amp; Luppova" authorityPageNumber="49" authorityYear="1945" box="[151,692,1836,1861]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="319" phylum="Arthropoda" rank="species" species="luppovae">
<emphasis id="1CEFEAEB1A5C347094D90639FE87A234" box="[151,379,1836,1860]" italics="true" pageId="8" pageNumber="319">Phrurolithus luppovae</emphasis>
<bibRefCitation id="4A0A4B081A5C347095CF0639FD48A234" author="Spassky, S. A. &amp; Luppova, E." box="[385,692,1836,1861]" pageId="8" pageNumber="319" pagination="43 - 55" refId="ref9442" refString="Spassky, S. A. &amp; Luppova, E. (1945) Materialy po faune paukov Tadjikistana [Materials on the spider fauna of Tajikistan]. Entomologicheskoe Obozrenie, 28 (1 - 2), 43 - 55. [in Russian]" type="journal article" year="1945">Spassky &amp; Luppova, 1945: 49</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<specimenCount id="389DFD701A5C3470968A0639FD25A234" box="[708,729,1836,1860]" pageId="8" pageNumber="319" type="male">♂</specimenCount>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="668165721A5C347094D90667FB06A2A5" pageId="8" pageNumber="319" type="materials_examined">
<paragraph id="2E2436F91A5C347094D90667FB06A2A5" blockId="8.[151,1436,1906,2005]" pageId="8" pageNumber="319">
<emphasis id="1CEFEAEB1A5C347094D90667FE78A2FD" bold="true" box="[151,388,1906,1933]" pageId="8" pageNumber="319">Material examined.</emphasis>
<materialsCitation id="9EF33CA41A5C347095C10666FBD2A2C1" ID-GBIF-Occurrence="2592928135" collectingDate="2015-04-29" collectionCode="ZMMU" collectorName="Y. M. Marusik" country="Tajikistan" county="Dangara Distr." elevation="1875" latitude="38.283333" location="Shar-Shar Pass" longLatPrecision="1176" longitude="69.21667" municipality="Sanglok Mt. Range" pageId="8" pageNumber="319" specimenCount="1" specimenCount-male="1" stateProvince="Khatlon">
<collectingCountry id="568C76691A5C347095C10666FDD3A2FD" box="[399,559,1907,1933]" name="Tajikistan" pageId="8" pageNumber="319">TAJIKISTAN</collectingCountry>
:
<emphasis id="1CEFEAEB1A5C347096730666FD0EA2FD" box="[573,754,1907,1933]" italics="true" pageId="8" pageNumber="319">
<collectingRegion id="EC5FF81B1A5C347096730666FD0EA2FD" box="[573,754,1907,1933]" country="Tajikistan" name="Khatlon" pageId="8" pageNumber="319">Khatlon Region</collectingRegion>
</emphasis>
:
<specimenCount id="389DFD701A5C3470974A0666FCDBA2FD" box="[772,807,1907,1933]" pageId="8" pageNumber="319" type="male">1♂</specimenCount>
(
<collectionCode id="488AAE3C1A5C347097740666FC6BA2FD" box="[826,919,1907,1933]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="8" pageNumber="319" type="Museum">ZMMU</collectionCode>
),
<collectingCounty id="C7454E751A5C347097E00666FBA4A2FD" box="[942,1112,1907,1933]" pageId="8" pageNumber="319">Dangara Distr.</collectingCounty>
,
<collectingMunicipality id="CE40AC831A5C347090290666FAB6A2FD" box="[1127,1354,1907,1933]" pageId="8" pageNumber="319">Sanglok Mt. Range</collectingMunicipality>
, above
<location id="2B4460221A5C347094D90682FEBBA2C1" LSID="urn:lsid:plazi:treatment:A63287EF1A5C347394D907B0FCCEA6D7:2B4460221A5C347094D90682FEBBA2C1" box="[151,327,1943,1969]" country="Tajikistan" county="Dangara Distr." latitude="38.283333" longLatPrecision="1176" longitude="69.21667" municipality="Sanglok Mt. Range" name="Shar-Shar Pass" pageId="8" pageNumber="319" stateProvince="Khatlon">Shar–Shar Pass</location>
,
<geoCoordinate id="4BAF503E1A5C3470951D0682FE49A2C1" box="[339,437,1943,1969]" degrees="38" direction="north" minutes="17" orientation="latitude" pageId="8" pageNumber="319" precision="925" value="38.283333">38°17’N</geoCoordinate>
,
<geoCoordinate id="4BAF503E1A5C3470958F0682FDDDA2C1" box="[449,545,1943,1969]" degrees="69" direction="east" minutes="13" orientation="longitude" pageId="8" pageNumber="319" precision="925" value="69.21667">69°13’E</geoCoordinate>
,
<quantity id="E9639B1C1A5C347096630683FD36A2C1" box="[557,714,1942,1969]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="2.05" metricValueMin="1.7" pageId="8" pageNumber="319" unit="m" value="1875.0" valueMax="2050.0" valueMin="1700.0">
<elevation id="A5B6D1CA1A5C347096630683FD36A2C1" box="[557,714,1942,1969]" metricMagnitude="3" metricUnit="m" metricValue="1.875" metricValueMax="2.05" metricValueMin="1.7" pageId="8" pageNumber="319" unit="m" value="1875.0" valueMax="2050.0" valueMin="1700.0">1700–2050 m</elevation>
</quantity>
,
<date id="5A2510391A5C3470969B0683FCA8A2C1" box="[725,852,1942,1969]" pageId="8" pageNumber="319" value="2015-04-29">
<collectingDate id="4A61E9D11A5C3470969B0683FCA8A2C1" box="[725,852,1942,1969]" pageId="8" pageNumber="319" value="2015-04-29">29.IV.2015</collectingDate>
</date>
, leg.
<collectorName id="836E532F1A5C347097C00682FBD2A2C1" box="[910,1070,1943,1969]" pageId="8" pageNumber="319">Y.M. Marusik</collectorName>
</materialsCitation>
;
<materialsCitation id="9EF33CA41A5C347090770682FB06A2A5" ID-GBIF-Occurrence="2592928162" collectingDate="2015-04-28" collectionCode="ZMMU" collectorName="Y. M. Marusik." country="Tajikistan" elevation="1579" latitude="38.383335" location="Darai-Mukhtor" longLatPrecision="1175" longitude="69.95" municipality="Khovaling Distr." pageId="8" pageNumber="319" specimenCount="1" specimenCount-female="1" stateProvince="Khatlon Region">
<specimenCount id="389DFD701A5C347090770682FBA0A2C1" box="[1081,1116,1943,1969]" pageId="8" pageNumber="319" type="female">1♀</specimenCount>
(
<collectionCode id="488AAE3C1A5C347090250682FB34A2C1" box="[1131,1224,1943,1969]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="8" pageNumber="319" type="Museum">ZMMU</collectionCode>
),
<collectingMunicipality id="CE40AC831A5C347090950682FA64A2C1" box="[1243,1432,1943,1969]" pageId="8" pageNumber="319">Khovaling Distr.</collectingMunicipality>
,
<location id="2B4460221A5C347094D906AEFEBDA2A5" LSID="urn:lsid:plazi:treatment:A63287EF1A5C347394D907B0FCCEA6D7:2B4460221A5C347094D906AEFEBDA2A5" box="[151,321,1979,2005]" country="Tajikistan" latitude="38.383335" longLatPrecision="1175" longitude="69.95" municipality="Khovaling Distr." name="Darai-Mukhtor" pageId="8" pageNumber="319" stateProvince="Khatlon Region">Darai-Mukhtor</location>
env. of “Vose Museum”,
<geoCoordinate id="4BAF503E1A5C3470962E06AEFD38A2A5" box="[608,708,1979,2005]" degrees="38" direction="north" minutes="23" orientation="latitude" pageId="8" pageNumber="319" precision="925" value="38.383335">38°23’N</geoCoordinate>
,
<geoCoordinate id="4BAF503E1A5C3470968106AFFCD3A2A5" box="[719,815,1978,2005]" degrees="69" direction="east" minutes="57" orientation="longitude" pageId="8" pageNumber="319" precision="925" value="69.95">69°57’E</geoCoordinate>
,
<quantity id="E9639B1C1A5C3470977506AFFC6EA2A5" box="[827,914,1978,2005]" metricMagnitude="3" metricUnit="m" metricValue="1.579" pageId="8" pageNumber="319" unit="m" value="1579.0">
<elevation id="A5B6D1CA1A5C3470977506AFFC6EA2A5" box="[827,914,1978,2005]" metricMagnitude="3" metricUnit="m" metricValue="1.579" pageId="8" pageNumber="319" unit="m" value="1579.0">1579 m</elevation>
</quantity>
,
<date id="5A2510391A5C347097D006AEFBE0A2A5" box="[926,1052,1978,2005]" pageId="8" pageNumber="319" value="2015-04-28">
<collectingDate id="4A61E9D11A5C347097D006AEFBE0A2A5" box="[926,1052,1978,2005]" pageId="8" pageNumber="319" value="2015-04-28">28.IV.2015</collectingDate>
</date>
, leg.
<collectorName id="836E532F1A5C3470901906AEFB06A2A5" box="[1111,1274,1979,2005]" pageId="8" pageNumber="319">Y.M. Marusik.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="7AE466711A5D347194D9067DFC6AA29D" ID-DOI="http://doi.org/10.5281/zenodo.3734514" ID-Zenodo-Dep="3734514" httpUri="https://zenodo.org/record/3734514/files/figure.png" pageId="9" pageNumber="320" startId="9.[151,250,1896,1921]" targetBox="[236,1361,184,1866]" targetPageId="9">
<paragraph id="2E2436F91A5D347194D9067DFC6AA29D" blockId="9.[151,1437,1896,2029]" pageId="9" pageNumber="320">
<emphasis id="1CEFEAEB1A5D347194D9067DFEE8A2F1" bold="true" box="[151,276,1896,1921]" pageId="9" pageNumber="320">FIGURE 7.</emphasis>
General appearance and epigyne of
<taxonomicName id="E99B4D7A1A5D347196C4067DFC92A2F0" authorityName="Spassky &amp; Luppova" authorityYear="1945" box="[650,878,1896,1920]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="320" phylum="Arthropoda" rank="species" species="luppovae">
<emphasis id="1CEFEAEB1A5D347196C4067DFC92A2F0" box="[650,878,1896,1920]" italics="true" pageId="9" pageNumber="320">Phrurolithus luppovae</emphasis>
</taxonomicName>
. A, C–D female habitus, dorsal, lateral and ventral; B, E male habitus, dorsal and ventral; F male prosoma, frontal; G macerated epigyne, dorsal; H intact epigyne, ventral. Scale=0.2 mm, if not otherwise indicated. Abbreviations:
<emphasis id="1CEFEAEB1A5D3471963506A4FD53A2B8" box="[635,687,1969,1992]" italics="true" pageId="9" pageNumber="320">Co—</emphasis>
copulatory opening,
<emphasis id="1CEFEAEB1A5D347197CC06A5FC49A2B8" box="[898,949,1968,1992]" italics="true" pageId="9" pageNumber="320">Fd—</emphasis>
fertilization duct,
<emphasis id="1CEFEAEB1A5D3471902506A4FB7BA2B8" box="[1131,1159,1969,1992]" italics="true" pageId="9" pageNumber="320">Gr</emphasis>
—receptacular gland,
<emphasis id="1CEFEAEB1A5D3471912406A4FA7FA2B8" box="[1386,1411,1969,1992]" italics="true" pageId="9" pageNumber="320">Pr</emphasis>
— primary receptacle,
<emphasis id="1CEFEAEB1A5D3471952F06C0FE68A29C" box="[353,404,2005,2028]" italics="true" pageId="9" pageNumber="320">Ro—</emphasis>
receptacular outgrowth,
<emphasis id="1CEFEAEB1A5D347196C506C0FD5EA29C" box="[651,674,2005,2028]" italics="true" pageId="9" pageNumber="320">Sr</emphasis>
—secondary receptacle.
</paragraph>
</caption>
<caption id="7AE466711A5E347294D904F6FC09A334" ID-DOI="http://doi.org/10.5281/zenodo.3734516" ID-Zenodo-Dep="3734516" httpUri="https://zenodo.org/record/3734516/files/figure.png" pageId="10" pageNumber="321" startId="10.[151,250,1507,1532]" targetBox="[151,1426,195,1482]" targetPageId="10">
<paragraph id="2E2436F91A5E347294D904F6FC09A334" blockId="10.[151,1437,1507,1604]" pageId="10" pageNumber="321">
<emphasis id="1CEFEAEB1A5E347294D904F6FEEFA08C" bold="true" box="[151,275,1507,1532]" pageId="10" pageNumber="321">FIGURE 8.</emphasis>
Male palp of
<taxonomicName id="E99B4D7A1A5E347295EC04F1FD7AA08C" authorityName="Spassky &amp; Luppova" authorityYear="1945" box="[418,646,1508,1532]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="321" phylum="Arthropoda" rank="species" species="luppovae">
<emphasis id="1CEFEAEB1A5E347295EC04F1FD7AA08C" box="[418,646,1508,1532]" italics="true" pageId="10" pageNumber="321">Phrurolithus luppovae</emphasis>
</taxonomicName>
. A ventral; B retrolateral; C dorso-retrolateral; D prolateral; E ventro-retrolateral; F dorsal. Scale=0.2 mm. Abbreviations:
<emphasis id="1CEFEAEB1A5E3472962C071CFD66A350" box="[610,666,1545,1568]" italics="true" pageId="10" pageNumber="321">Em—</emphasis>
embolus,
<emphasis id="1CEFEAEB1A5E347296B2071CFCD1A350" box="[764,813,1545,1568]" italics="true" pageId="10" pageNumber="321">Fe—</emphasis>
femoral extension,
<emphasis id="1CEFEAEB1A5E347297A1071CFBDEA350" box="[1007,1058,1545,1568]" italics="true" pageId="10" pageNumber="321">Ra—</emphasis>
retrolateral apophysis,
<emphasis id="1CEFEAEB1A5E34729146071DFADDA36F" box="[1288,1313,1544,1567]" italics="true" pageId="10" pageNumber="321">Sa</emphasis>
—subterminal apophysis,
<emphasis id="1CEFEAEB1A5E347295600739FEBBA334" box="[302,327,1580,1604]" italics="true" pageId="10" pageNumber="321">Sd</emphasis>
—sperm duct,
<emphasis id="1CEFEAEB1A5E347295900738FE0AA334" box="[478,502,1581,1604]" italics="true" pageId="10" pageNumber="321">Ta</emphasis>
—terminal apophysis,
<emphasis id="1CEFEAEB1A5E347296920738FCF0A334" box="[732,780,1581,1604]" italics="true" pageId="10" pageNumber="321">Va—</emphasis>
retroventral apophysis.
</paragraph>
</caption>
<subSubSection id="668165721A5E347294890767FB98A388" pageId="10" pageNumber="321" type="discussion">
<paragraph id="2E2436F91A5E347294890767FB98A388" blockId="10.[151,1437,1650,2037]" pageId="10" pageNumber="321">
<emphasis id="1CEFEAEB1A5E347294890767FEB3A3FD" bold="true" box="[199,335,1650,1677]" pageId="10" pageNumber="321">Comments.</emphasis>
This species was described twice, in two different publications.Apparently, the original manuscript was sent to the publisher in Riga prior to World War II, and Spassky was unaware that his paper was published during the war (in 1941) and resubmitted the manuscript again, in coauthorship with E.P. Luppova to the Soviet journal Entomologicheskoe Obozrenie, which appeared in 1945, although without any figures.
</paragraph>
</subSubSection>
<subSubSection id="668165721A5E347294890617FC22A285" pageId="10" pageNumber="321" type="diagnosis">
<paragraph id="2E2436F91A5E347294890617FC22A285" blockId="10.[151,1437,1650,2037]" pageId="10" pageNumber="321">
<emphasis id="1CEFEAEB1A5E347294890617FEBEA26D" bold="true" box="[199,322,1794,1821]" pageId="10" pageNumber="321">Diagnosis.</emphasis>
Males differ from most of its congeners by the sharply-pointed retroventral tibial apophysis. Only two other species have a similar apophysis,
<taxonomicName id="E99B4D7A1A5E347296C20632FBDDA231" authority="(Donitz &amp; Strand, 1906)" baseAuthorityName="Donitz &amp; Strand" baseAuthorityYear="1906" box="[652,1057,1830,1857]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="321" phylum="Arthropoda" rank="species" species="claripes">
<emphasis id="1CEFEAEB1A5E347296C20632FCFFA230" box="[652,771,1830,1856]" italics="true" pageId="10" pageNumber="321">P. claripes</emphasis>
(Dönitz &amp; Strand, 1906)
</taxonomicName>
and
<taxonomicName id="E99B4D7A1A5E347290140632FA64A231" authority="Zhu &amp; Mei, 1982" authorityName="Zhu &amp; Mei" authorityYear="1982" box="[1114,1432,1830,1857]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="321" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="1CEFEAEB1A5E347290140632FB39A230" box="[1114,1221,1831,1856]" italics="true" pageId="10" pageNumber="321">P. sinicus</emphasis>
Zhu &amp; Mei, 1982
</taxonomicName>
. It differs from
<taxonomicName id="E99B4D7A1A5E34729570065EFE54A214" authorityName="Zhu &amp; Mei" authorityYear="1982" box="[318,424,1867,1892]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="321" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="1CEFEAEB1A5E34729570065EFE54A214" box="[318,424,1867,1892]" italics="true" pageId="10" pageNumber="321">P. sinicus</emphasis>
</taxonomicName>
by its shorter apophysis (1.3 times longer than tibia
<emphasis id="1CEFEAEB1A5E347297B2065EFBEFA214" box="[1020,1043,1867,1892]" italics="true" pageId="10" pageNumber="321">vs</emphasis>
. 2.5), and by the absence of a proventral tibial apophysis.
<taxonomicName id="E99B4D7A1A5E347295E3067BFD61A2F8" baseAuthorityName="Donitz &amp; Strand" baseAuthorityYear="1906" box="[429,669,1902,1928]" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="321" phylum="Arthropoda" rank="species" species="claripes">
<emphasis id="1CEFEAEB1A5E347295E3067BFD61A2F8" box="[429,669,1902,1928]" italics="true" pageId="10" pageNumber="321">Phrurolithus claripes</emphasis>
</taxonomicName>
has a subtriangular terminal apophysis (
<emphasis id="1CEFEAEB1A5E34729029067AFB82A2F8" box="[1127,1150,1903,1928]" italics="true" pageId="10" pageNumber="321">vs</emphasis>
. square-shaped in
<taxonomicName id="E99B4D7A1A5E3472911D067AFF26A2DC" authorityName="Spassky &amp; Luppova" authorityYear="1945" class="Arachnida" family="Corinnidae" genus="Phrurolithus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="321" phylum="Arthropoda" rank="species" species="luppovae">
<emphasis id="1CEFEAEB1A5E3472911D067AFF26A2DC" italics="true" pageId="10" pageNumber="321">P. luppovae</emphasis>
</taxonomicName>
). The only known female differs from other female congeners by the diverging elongate subconical secondary receptacles spaced by more than 2 diameters, and the primary receptacles with a posterior outgrowth (
<emphasis id="1CEFEAEB1A5E3472914806A2FAE1A2A0" box="[1286,1309,1975,2000]" italics="true" pageId="10" pageNumber="321">vs</emphasis>
. secondary receptacles oval, parallel or converging, spaced by less than one diameter).
</paragraph>
</subSubSection>
<subSubSection id="668165721A5F34739489018DFC0AA62F" pageId="11" pageNumber="322" type="description">
<paragraph id="2E2436F91A5F34739489018DFE23A417" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="322">
<emphasis id="1CEFEAEB1A5F34739489018DFEA5A5C3" bold="true" box="[199,345,152,179]" pageId="11" pageNumber="322">Description.</emphasis>
Male. Habitus as in
<figureCitation id="B6A02A7C1A5F34739673018CFD2DA5C3" box="[573,721,153,179]" captionStart="FIGURE 7" captionStartId="9.[151,250,1896,1921]" captionTargetBox="[236,1361,184,1866]" captionTargetId="figure@9.[219,1368,181,1870]" captionTargetPageId="9" captionText="FIGURE 7. General appearance and epigyne of Phrurolithus luppovae. A, C–D female habitus, dorsal, lateral and ventral; B, E male habitus, dorsal and ventral; F male prosoma, frontal; G macerated epigyne, dorsal; H intact epigyne, ventral. Scale=0.2 mm, if not otherwise indicated. Abbreviations: Co—copulatory opening, Fd—fertilization duct, Gr—receptacular gland, Pr— primary receptacle, Ro—receptacular outgrowth, Sr—secondary receptacle." figureDoi="http://doi.org/10.5281/zenodo.3734514" httpUri="https://zenodo.org/record/3734514/files/figure.png" pageId="11" pageNumber="322">Figs 7B, E, F</figureCitation>
. Total length 2.02. Carapace 0.91 long, 0.80 wide. Eye sizes and interdistances: AME: 0.05, ALE: 0.06, PME: 0.05, PLE: 0.06, AME–AME: 0.04, PME–PME: 0.05. Carapace, sternum, labium, chelicerae and maxillae light reddish-brown; carapace with dark radiating patterns. Legs yellowishbrown, with darker annulation on femora. Abdomen dark, without any pattern, covered with entire dorsal scutum. Spinnerets uniformly pale yellow. Leg I measurements: 3.42 (0.90, 0.34, 0.96, 0.82, 0.40). Spination: I: Fe 1pl, Ti 12v, Mt 8v; II: Ti 10v, Mt 6v.
</paragraph>
<paragraph id="2E2436F91A5F347394890064FC5AA487" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="322">
Palp as in
<figureCitation id="B6A02A7C1A5F347395750064FE83A4FB" box="[315,383,369,395]" captionStart="FIGURE 8" captionStartId="10.[151,250,1507,1532]" captionTargetBox="[151,1426,195,1482]" captionTargetId="figure@10.[151,1435,181,1482]" captionTargetPageId="10" captionText="FIGURE 8. Male palp of Phrurolithus luppovae. A ventral; B retrolateral; C dorso-retrolateral; D prolateral; E ventro-retrolateral; F dorsal. Scale=0.2 mm.Abbreviations: Em—embolus, Fe—femoral extension, Ra—retrolateral apophysis, Sa—subterminal apophysis, Sd—sperm duct, Ta—terminal apophysis, Va—retroventral apophysis." figureDoi="http://doi.org/10.5281/zenodo.3734516" httpUri="https://zenodo.org/record/3734516/files/figure.png" pageId="11" pageNumber="322">Figs 8</figureCitation>
A–F; tibia with sharply pointed retroventral apophysis and retrolateral one also distinct, retro- ventral apophysis slightly longer than tibia; cymbium elongate, more than twice (ca. 2.25 times) as long as wide; sperm duct relatively thin; terminal apophysis wide, as wide as long; subterminal apophysis straight, finger-shaped; embolus with broad base, appears like thick spine, directed anteriorly.
</paragraph>
<paragraph id="2E2436F91A5F347394890314FC16A7DB" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="322">
Female. Habitus as in
<figureCitation id="B6A02A7C1A5F3473958C0314FD9EA76B" box="[450,610,513,539]" captionStart="FIGURE 7" captionStartId="9.[151,250,1896,1921]" captionTargetBox="[236,1361,184,1866]" captionTargetId="figure@9.[219,1368,181,1870]" captionTargetPageId="9" captionText="FIGURE 7. General appearance and epigyne of Phrurolithus luppovae. A, C–D female habitus, dorsal, lateral and ventral; B, E male habitus, dorsal and ventral; F male prosoma, frontal; G macerated epigyne, dorsal; H intact epigyne, ventral. Scale=0.2 mm, if not otherwise indicated. Abbreviations: Co—copulatory opening, Fd—fertilization duct, Gr—receptacular gland, Pr— primary receptacle, Ro—receptacular outgrowth, Sr—secondary receptacle." figureDoi="http://doi.org/10.5281/zenodo.3734514" httpUri="https://zenodo.org/record/3734514/files/figure.png" pageId="11" pageNumber="322">Figs 7A, C, D</figureCitation>
. Total length 2.31. Carapace 0.83 long, 0.80 wide. Eye sizes and interdistances: AME: 0.05, ALE: 0.07, PME: 0.07, PLE: 0.06, AME–AME: 0.03, PME–PME: 0.03. Colouration of carapace generally as in male, but with more intense scattered dark patches, light reddish-brown; abdomen with pair of large white spots on dorsum and postgastral light spot; spinnerets pale yellowish. Leg I measurements: 3.09 (0.88, 0.29, 0.80, 0.66, 0.46). Spination: I: Fe 1pl, Ti 10v, Mt 8v; II: Ti 10v, Mt 6v.
</paragraph>
<paragraph id="2E2436F91A5F3473948903A0FC0AA62F" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="322">
Epigyne as in
<figureCitation id="B6A02A7C1A5F3473952603A0FE51A7BF" box="[360,429,693,719]" captionStart="FIGURE 7" captionStartId="9.[151,250,1896,1921]" captionTargetBox="[236,1361,184,1866]" captionTargetId="figure@9.[219,1368,181,1870]" captionTargetPageId="9" captionText="FIGURE 7. General appearance and epigyne of Phrurolithus luppovae. A, C–D female habitus, dorsal, lateral and ventral; B, E male habitus, dorsal and ventral; F male prosoma, frontal; G macerated epigyne, dorsal; H intact epigyne, ventral. Scale=0.2 mm, if not otherwise indicated. Abbreviations: Co—copulatory opening, Fd—fertilization duct, Gr—receptacular gland, Pr— primary receptacle, Ro—receptacular outgrowth, Sr—secondary receptacle." figureDoi="http://doi.org/10.5281/zenodo.3734514" httpUri="https://zenodo.org/record/3734514/files/figure.png" pageId="11" pageNumber="322">Figs 7</figureCitation>
G–H; fovea weakly developed, copulatory openings widely separated (by about 10 diam- eters); primary receptacles visible through integument and located over copulatory openings; copulatory opening leads to elongate weakly sclerotized secondary receptacles, their anterior parts separated by about 2 diameters; primary receptacles globular, touching each other, each receptacle with anterior and posterior tuberculate outgrowth (
<emphasis id="1CEFEAEB1A5F347394EF0253FF3CA62F" box="[161,192,838,863]" italics="true" pageId="11" pageNumber="322">Ro</emphasis>
); fertilization ducts originate from anterior edge of receptacle, diverging.
</paragraph>
</subSubSection>
<subSubSection id="668165721A5F34739489027DFCCEA6D7" pageId="11" pageNumber="322" type="distribution">
<paragraph id="2E2436F91A5F34739489027DFCCEA6D7" blockId="11.[151,1437,152,935]" pageId="11" pageNumber="322">
<emphasis id="1CEFEAEB1A5F34739489027DFE9DA6F3" bold="true" box="[199,353,872,899]" pageId="11" pageNumber="322">Distribution.</emphasis>
Known only from the
<typeStatus id="F120885B1A5F3473962B027CFD69A6F3" box="[613,661,873,899]" pageId="11" pageNumber="322">type</typeStatus>
locality in Obigarm, and the new collection localities in
<collectingRegion id="EC5FF81B1A5F34739155027CFF37A6D7" country="Tajikistan" name="Khatlon" pageId="11" pageNumber="322">Khatlon region</collectingRegion>
, central-western and southwestern
<collectingCountry id="568C76691A5F3473961A0298FD3EA6D7" box="[596,706,909,935]" name="Tajikistan" pageId="11" pageNumber="322">Tajikistan</collectingCountry>
(
<figureCitation id="B6A02A7C1A5F3473969C0298FCD9A6D7" box="[722,805,909,935]" captionStart="FIGURE 14" captionStartId="17.[151,250,1567,1592]" captionTargetBox="[151,1436,899,1542]" captionTargetId="figure@17.[151,1436,899,1542]" captionTargetPageId="17" captionText="FIGURE 14. Distribution records of Phrurolithus spp. and Bosselaerius spp. in Azerbaijan, Iran and Tajikistan: Red circles—P. festivus, Pink stars—P. azarkinae sp. n., Blue squares—P. pullatus, Yellow crosses—P. luppovae, Cyan triangles—B. hyrcanicus sp. n., Black asterisk—B. tajikistanicus sp. n." figureDoi="http://doi.org/10.5281/zenodo.3734528" httpUri="https://zenodo.org/record/3734528/files/figure.png" pageId="11" pageNumber="322">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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