treatments-xml/data/A6/10/87/A61087FCFFEBF5344D44FDAFFADE63D5.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4759936" ID-GBIF-Dataset="8cc293b1-7429-40f0-a5a3-912f2730df83" ID-GBIF-Taxon="182075124" ID-ISSN="1854-0392" ID-Zenodo-Dep="4759936" checkinTime="1620961777533" checkinUser="felipe" docAuthor="Stark, Bill P. &amp; Baumann, Richard W." docDate="2011" docId="A61087FCFFEBF5344D44FDAFFADE63D5" docLanguage="en" docName="Illiesia.7.19.182-191.pdf" docOrigin="Illiesia 7 (19)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Anacroneuria ypane Stark &amp; Baumann 2011, sp. n." docType="treatment" docVersion="7" lastPageNumber="187" masterDocId="5A29FF84FFEEF5314C0BFFC8FF876155" masterDocTitle="Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species" masterLastPageNumber="191" masterPageNumber="182" pageId="5" pageNumber="187" updateTime="1632024467179" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Records Of Anacroneuria (Plecoptera: Perlidae) From Bolivia And Paraguay With Descriptions Of Three New Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumann, Richard W.</mods:namePart>
<mods:affiliation>Department of Biology and Monte L. Bean Life Science Museum, Brigham Young University, Provo, UT, U. S. A. 84602 E-mail: richard _ baumann @ byu. edu</mods:affiliation>
<mods:nameIdentifier type="email">richard_baumann@byu.edu</mods:nameIdentifier>
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<mods:title>Illiesia</mods:title>
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<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>7</mods:number>
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<mods:detail type="issue">
<mods:number>19</mods:number>
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<mods:start>182</mods:start>
<mods:end>191</mods:end>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760267</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4759936" ID-GBIF-Taxon="182075124" ID-Zenodo-Dep="4759936" LSID="urn:lsid:plazi:treatment:A61087FCFFEBF5344D44FDAFFADE63D5" httpUri="http://treatment.plazi.org/id/A61087FCFFEBF5344D44FDAFFADE63D5" lastPageNumber="187" pageId="5" pageNumber="187">
<subSubSection box="[335,651,615,640]" pageId="5" pageNumber="187" type="nomenclature">
<paragraph blockId="5.[335,651,615,640]" box="[335,651,615,640]" pageId="5" pageNumber="187">
<heading bold="true" box="[335,651,615,640]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="187" reason="2">
<emphasis bold="true" box="[335,651,615,640]" pageId="5" pageNumber="187">
<taxonomicName authority="Stark &amp; Baumann, 2011" authorityName="Stark &amp; Baumann" authorityYear="2011" box="[335,573,615,639]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="ypane" status="sp. n.">
<emphasis bold="true" box="[335,573,615,639]" italics="true" pageId="5" pageNumber="187">Anacroneuria ypane</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[587,651,616,640]" pageId="5" pageNumber="187" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[422,564,651,675]" pageId="5" pageNumber="187" type="description">
<paragraph blockId="5.[422,564,651,675]" box="[422,564,651,675]" pageId="5" pageNumber="187">
(
<figureCitation box="[430,556,651,675]" captionStart="Figs" captionStartId="6.[189,241,1575,1599]" captionTargetBox="[264,1383,215,1500]" captionTargetId="figure-60@6.[249,1383,215,1452]" captionTargetPageId="6" captionText="Figs. 11-15. Anacroneuria ypane structures. 11. Head and pronotum. 12. Male sternum 9. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760277" httpUri="https://zenodo.org/record/4760277/files/figure.png" pageId="5" pageNumber="187">Figs. 11-15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="187" type="materials_examined">
<paragraph blockId="5.[189,798,722,1921]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[189,437,722,746]" pageId="5" pageNumber="187">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111725336" collectingDate="1956-11-25" collectionCode="USNM" collectorName="C. J. D. Brown" country="Paraguay" county="Pedro Juan Caballero" location="Pedro Juan Caballero" municipality="Rio Ypane" pageId="5" pageNumber="187" specimenCount="1" specimenCount-male="1" stateProvince="Amambay Department" typeStatus="holotype">
<typeStatus box="[457,569,722,746]" pageId="5" pageNumber="187">Holotype</typeStatus>
<specimenCount box="[587,607,722,746]" pageId="5" pageNumber="187" type="male">♂</specimenCount>
, 
<emphasis bold="true" pageId="5" pageNumber="187">
<collectingCountry box="[633,793,722,746]" name="Paraguay" pageId="5" pageNumber="187">PARAGUAY</collectingCountry>
: 
<collectingRegion box="[189,467,758,782]" country="Paraguay" name="Amambay" pageId="5" pageNumber="187">Amambay Department</collectingRegion>
</emphasis>
, 
<collectingMunicipality box="[479,600,758,782]" pageId="5" pageNumber="187">Rio Ypane</collectingMunicipality>
, cerca 
<location LSID="urn:lsid:plazi:treatment:A61087FCFFEBF5344D44FDAFFADE63D5:2B666031FFEBF5344E94FD3EFEB66267" country="Paraguay" county="Pedro Juan Caballero" municipality="Rio Ypane" name="Pedro Juan Caballero" pageId="5" pageNumber="187" stateProvince="Amambay Department">Pedro Juan Caballero</location>
, 
<date box="[314,530,794,818]" pageId="5" pageNumber="187" value="1956-11-25">
<collectingDate box="[314,530,794,818]" pageId="5" pageNumber="187" value="1956-11-25">25 November 1956</collectingDate>
</date>
, 
<collectorName box="[539,687,794,818]" pageId="5" pageNumber="187">C.J.D. Brown</collectorName>
(
<collectionCode box="[700,786,794,818]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="5" pageNumber="187">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="187" type="description">
<paragraph blockId="5.[189,798,722,1921]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[189,365,829,853]" pageId="5" pageNumber="187">
<collectorName box="[189,260,829,853]" pageId="5" pageNumber="187">Adult</collectorName>
habitus.
</emphasis>
<collectorName box="[371,464,830,854]" pageId="5" pageNumber="187">General</collectorName>
color brown, patterned with areas of pale pigment. 
<collectorName box="[450,514,865,889]" pageId="5" pageNumber="187">Head</collectorName>
with a large area of dark pigment extending from ocellar region over central frons (
<figureCitation box="[263,341,936,961]" captionStart="Figs" captionStartId="6.[189,241,1575,1599]" captionTargetBox="[264,1383,215,1500]" captionTargetId="figure-60@6.[249,1383,215,1452]" captionTargetPageId="6" captionText="Figs. 11-15. Anacroneuria ypane structures. 11. Head and pronotum. 12. Male sternum 9. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760277" httpUri="https://zenodo.org/record/4760277/files/figure.png" pageId="5" pageNumber="187">Fig. 11</figureCitation>
); dark pigment area expanded laterally in central frons where a small oval pale area is located; lappets and antennae dark brown. 
<collectorName box="[680,797,1008,1031]" pageId="5" pageNumber="187">Pronotum</collectorName>
mostly dark but with a narrow pale median stripe and pale lateral areas. 
<collectorName box="[461,523,1079,1103]" pageId="5" pageNumber="187">Hind</collectorName>
femora dark brown in apical third but pale basally; hind tibia with small dark bands basally and apically but pale for most of length. 
<collectorName box="[276,341,1185,1209]" pageId="5" pageNumber="187">Wing</collectorName>
membrane brown, veins (particularly R vein) dark brown.
</paragraph>
<paragraph blockId="5.[189,798,722,1921]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[189,258,1256,1280]" pageId="5" pageNumber="187">Male.</emphasis>
Forewing length 
<quantity box="[484,573,1256,1280]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="187" unit="mm" value="10.0">10 mm</quantity>
. Hammer thimble shaped, apical diameter less than height (
<figureCitation box="[699,784,1292,1316]" captionStart="Figs" captionStartId="6.[189,241,1575,1599]" captionTargetBox="[264,1383,215,1500]" captionTargetId="figure-60@6.[249,1383,215,1452]" captionTargetPageId="6" captionText="Figs. 11-15. Anacroneuria ypane structures. 11. Head and pronotum. 12. Male sternum 9. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760277" httpUri="https://zenodo.org/record/4760277/files/figure.png" pageId="5" pageNumber="187">Fig. 12</figureCitation>
). Aedeagal apex a short, simple, truncate scoop bearing a ventral pair of prominent membranous lobes (
<figureCitation box="[264,343,1399,1423]" captionStart="Figs" captionStartId="6.[189,241,1575,1599]" captionTargetBox="[264,1383,215,1500]" captionTargetId="figure-60@6.[249,1383,215,1452]" captionTargetPageId="6" captionText="Figs. 11-15. Anacroneuria ypane structures. 11. Head and pronotum. 12. Male sternum 9. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760277" httpUri="https://zenodo.org/record/4760277/files/figure.png" pageId="5" pageNumber="187">Fig. 13</figureCitation>
); dorsal margin of apical area expanded into a pair of small rounded lateral lobes near point where hooks cross (
<figureCitation box="[419,499,1470,1494]" captionStart="Figs" captionStartId="6.[189,241,1575,1599]" captionTargetBox="[264,1383,215,1500]" captionTargetId="figure-60@6.[249,1383,215,1452]" captionTargetPageId="6" captionText="Figs. 11-15. Anacroneuria ypane structures. 11. Head and pronotum. 12. Male sternum 9. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760277" httpUri="https://zenodo.org/record/4760277/files/figure.png" pageId="5" pageNumber="187">Fig. 15</figureCitation>
); dorsal keel lines absent, apex bent strongly dorsad, giving a foot-like shape in lateral aspect (
<figureCitation box="[359,440,1541,1565]" captionStart="Figs" captionStartId="6.[189,241,1575,1599]" captionTargetBox="[264,1383,215,1500]" captionTargetId="figure-60@6.[249,1383,215,1452]" captionTargetPageId="6" captionText="Figs. 11-15. Anacroneuria ypane structures. 11. Head and pronotum. 12. Male sternum 9. 13. Aedeagus, ventral. 14. Aedeagus, lateral. 15. Aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760277" httpUri="https://zenodo.org/record/4760277/files/figure.png" pageId="5" pageNumber="187">Fig. 14</figureCitation>
). Hooks slender.
</paragraph>
<paragraph blockId="5.[189,798,722,1921]" box="[189,416,1576,1600]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[189,283,1576,1600]" pageId="5" pageNumber="187">Female.</emphasis>
Unknown.
</paragraph>
<paragraph blockId="5.[189,798,722,1921]" box="[189,396,1612,1636]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[189,264,1612,1636]" pageId="5" pageNumber="187">Larva.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="187" type="etymology">
<paragraph blockId="5.[189,798,722,1921]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[189,327,1648,1672]" pageId="5" pageNumber="187">Etymology.</emphasis>
The species name, used as a noun in apposition, is based on the 
<typeStatus box="[500,551,1684,1708]" pageId="5" pageNumber="187">type</typeStatus>
locality of Rio Ypane.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="187" type="diagnosis">
<paragraph blockId="5.[189,798,722,1921]" lastBlockId="5.[835,1443,225,640]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[189,319,1719,1743]" pageId="5" pageNumber="187">Diagnosis.</emphasis>
This species is related to the 
<taxonomicName authority="Klap" authorityName="Klap" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="atrifrons">
<emphasis italics="true" pageId="5" pageNumber="187">Anacroneuria atrifrons</emphasis>
Klap
</taxonomicName>
á lek group of species (
<bibRefCitation author="Froehlich, C. G." box="[614,789,1755,1779]" pageId="5" pageNumber="187" pagination="125 - 132" refId="ref4517" refString="Froehlich, C. G. 2008. Old species of Neotropical Plecoptera. pp. 125 - 132. In Hauer, F. R., J. A. Stanford &amp; R. L. Newell [eds.]. International advances in the ecology, zoogeography and systematics of mayflies and stoneflies. University of California Publications in Entomology. Volume 128. University of California Press, Berkeley. 412 pp." type="book chapter" year="2008">Froehlich 2008</bibRefCitation>
) which includes at least 
<taxonomicName authority="Rojas &amp; Baena" authorityName="Rojas &amp; Baena" authorityYear="1999" box="[466,720,1791,1815]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="azul">
<emphasis box="[466,545,1791,1815]" italics="true" pageId="5" pageNumber="187">A. azul</emphasis>
Rojas &amp; Baena
</taxonomicName>
(
<bibRefCitation author="Stark, B. P. &amp; Zuniga, A. M." pageId="5" pageNumber="187" pagination="13 - 46" refId="ref4822" refString="Stark, B. P., M. del C. Zuniga, A. M. Rojas, &amp; M. L. Baena. 1999. Colombian Anacroneuria: Descriptions of new and old species (Insecta, Plecoptera, Perlidae). Spixiana, 22: 13 - 46." type="journal article" year="1999">Stark et al. 1999</bibRefCitation>
), 
<taxonomicName authority="Stark &amp; Sivec" authorityName="Stark &amp; Sivec" authorityYear="1998" box="[338,615,1826,1850]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="canchi">
<emphasis box="[338,442,1826,1850]" italics="true" pageId="5" pageNumber="187">A. canchi</emphasis>
Stark &amp; Sivec
</taxonomicName>
(Stark &amp; Sivec 1998) and 
<taxonomicName authority="Stark" authorityName="Stark" authorityYear="2001" box="[323,516,1862,1886]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="pastaza">
<emphasis box="[323,442,1862,1886]" italics="true" pageId="5" pageNumber="187">A. pastaza</emphasis>
Stark
</taxonomicName>
(
<bibRefCitation author="Stark, B. P." box="[536,669,1862,1886]" pageId="5" pageNumber="187" pagination="1 - 42" refId="ref4730" refString="Stark, B. P. 2001. Records and descriptions of Anacroneuria from Ecuador (Plecoptera: Perlidae). Scopolia, 46: 1 - 42." type="journal article" year="2001">Stark 2001</bibRefCitation>
). Of these species, the aedeagus of 
<taxonomicName authorityName="Stark &amp; Baumann" authorityYear="2011" box="[484,582,1898,1921]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="ypane">
<emphasis box="[484,582,1898,1921]" italics="true" pageId="5" pageNumber="187">A. ypane</emphasis>
</taxonomicName>
is most similar to that of the Colombian species, 
<emphasis box="[1242,1336,225,249]" italics="true" pageId="5" pageNumber="187">
<taxonomicName authorityName="Rojas &amp; Baena" authorityYear="1999" box="[1242,1331,225,249]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="azul">A. azul</taxonomicName>
,
</emphasis>
but the membranous lobes in that species are smaller and a small transverse dorsal keel is present. The latter species also differs from 
<taxonomicName authorityName="Stark &amp; Baumann" authorityYear="2011" box="[1131,1229,332,355]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="ypane">
<emphasis box="[1131,1229,332,355]" italics="true" pageId="5" pageNumber="187">A. ypane</emphasis>
</taxonomicName>
in having a much smaller area of dark head pigment.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="187" type="discussion">
<paragraph blockId="5.[835,1443,225,640]" pageId="5" pageNumber="187">
<emphasis bold="true" box="[835,973,402,426]" pageId="5" pageNumber="187">Comments.</emphasis>
<bibRefCitation author="Froehlich, C. G." box="[992,1192,402,426]" pageId="5" pageNumber="187" pagination="75 - 107" refId="ref4481" refString="Froehlich, C. G. 2002. Anacroneuria mainly from southern Brazil and northeastern Argentina (Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 115: 75 - 107." type="journal article" year="2002">Froehlich (2002)</bibRefCitation>
reported 
<taxonomicName authority="(Pictet)" baseAuthorityName="Pictet" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="debilis">
<emphasis box="[1331,1443,402,426]" italics="true" pageId="5" pageNumber="187">A. debilis</emphasis>
(Pictet)
</taxonomicName>
from a single male collected in Parque Nacional Ybycui and 
<taxonomicName authority="Jewett" authorityName="Jewett" authorityYear="1959" box="[1112,1344,474,498]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="trimacula">
<emphasis box="[1112,1256,474,498]" italics="true" pageId="5" pageNumber="187">A. trimacula</emphasis>
Jewett
</taxonomicName>
from a single female collected at the same locality. Thus, 
<taxonomicName authorityName="Stark &amp; Baumann" authorityYear="2011" box="[835,1052,545,569]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="187" phylum="Arthropoda" rank="species" species="ypane">
<emphasis box="[835,1052,545,569]" italics="true" pageId="5" pageNumber="187">Anacroneuria ypane</emphasis>
</taxonomicName>
becomes only the third member of the genus to be confirmed for 
<collectingCountry box="[1180,1294,581,605]" name="Paraguay" pageId="5" pageNumber="187">Paraguay</collectingCountry>
, and the first to be described from a Paraguayan specimen.
</paragraph>
</subSubSection>
</treatment>
</document>