218 lines
35 KiB
XML
218 lines
35 KiB
XML
<document id="023FA37C3560C8DA4628277E33A34D21" ID-DOI="10.1111/j.1096-3642.2011.00706.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5441392" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630451174621" checkinUser="valdenar" docAuthor="Mendes, Amanda Cruz" docDate="2011" docId="A60A87F2FFBBFFA1D92AE9095D774366" docLanguage="en" docName="j.1096-3642.2011.00706.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (2)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00706.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heteropachylus crassicalcanei Mendes 2011" docType="treatment" docVersion="3" lastPageNumber="474" masterDocId="5A33FF8AFF9FFF84DA1BEF345E0A4645" masterDocTitle="Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae)" masterLastPageNumber="483" masterPageNumber="437" pageNumber="473" updateTime="1699275278278" updateUser="plazi">
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<mods:title id="279644C39705823C4FF89200706A2D3E">Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae)</mods:title>
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<mods:namePart id="9C987D5E387EF60F2A62649C9E78CA6E">Mendes, Amanda Cruz</mods:namePart>
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<mods:title id="AAF0D98C56F884FDD6636896CC33DEF9">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="ECA7E505F89CDBD9E016E34EBE8687FF">
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<mods:date id="CA270E4B0A8AFC98F65F4227C2015EB8">2011</mods:date>
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<mods:number id="52F3071CEF063B3A04F5A01B2202171F">2011-09-27</mods:number>
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<mods:number id="480858D9714C63DD4BDB87520B078F2E">163</mods:number>
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<paragraph id="2E1C36E4FFBBFFA0D92AE9095B894017" blockId="36.[817,1411,1596,1650]" box="[817,1411,1596,1618]" pageId="36" pageNumber="473">
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<heading id="75548188FFBBFFA0D92AE9095B894017" box="[817,1411,1596,1618]" centered="true" fontSize="8" level="2" pageId="36" pageNumber="473" reason="8">
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<taxonomicName id="E9A34D67FFBBFFA0D92AE9095B894017" ID-CoL="7WLPJ" authority="(SOARES, 1977)" authorityName="Mendes" authorityYear="2011" baseAuthorityName="H. Soares" baseAuthorityYear="1977" box="[817,1411,1596,1618]" class="Arachnida" family="Gonyleptidae" genus="Heteropachylus" kingdom="Animalia" order="Opiliones" pageId="36" pageNumber="473" phylum="Arthropoda" rank="species" species="crassicalcanei" status="comb. nov.">
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<emphasis id="1CD7EAF6FFBBFFA0D92AE9095ACD4015" box="[817,1223,1597,1618]" italics="true" pageId="36" pageNumber="473">HETEROPACHYLUS CRASSICALCANEI</emphasis>
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(
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<bibRefCitation id="4A324B15FFBBFFA0DEC1E9085B714017" author="Soares HEM" box="[1242,1403,1596,1618]" pageId="36" pageNumber="473" pagination="251 - 260" refId="ref32399" refString="Soares HEM. 1977. Opera Opiliologica Varia VI (Opiliones, Gonyleptidae). Physis, Seccion C 36: 251 - 260." type="journal article" year="1977">SOARES, 1977</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="2E1C36E4FFBBFFA0D965E96B5B3C4037" blockId="36.[817,1411,1596,1650]" box="[894,1334,1628,1650]" pageId="36" pageNumber="473">
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<heading id="75548188FFBBFFA0D965E96B5B3C4037" box="[894,1334,1628,1650]" centered="true" fontSize="9" level="2" pageId="36" pageNumber="473" reason="4">
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<emphasis id="1CD7EAF6FFBBFFA0D965E96B5A034034" bold="true" box="[894,1033,1628,1649]" pageId="36" pageNumber="473">COMB. NOV.</emphasis>
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(
|
||
<figureCitation id="B6982A61FFBBFFA0DE07E9685A7C4034" box="[1052,1142,1628,1649]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="36" pageNumber="473">FIGS 17</figureCitation>
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,
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<figureCitation id="B6982A61FFBBFFA0DE9EE9685AA94034" box="[1157,1187,1628,1649]" captionStart="Figure 18" captionStartId="27.[164,243,1135,1154]" captionTargetBox="[164,777,655,1048]" captionTargetPageId="27" captionText="Figure 18. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. Right trochanter, femur and patella IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441426" httpUri="https://zenodo.org/record/5441426/files/figure.png" pageId="36" pageNumber="473">18</figureCitation>
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,
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<figureCitation id="B6982A61FFBBFFA0DEA9E9685B094034" box="[1202,1283,1628,1650]" captionStart="Figure 25" captionStartId="34.[144,223,1747,1766]" captionTargetBox="[206,1359,196,1716]" captionTargetPageId="34" captionText="Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441440" httpUri="https://zenodo.org/record/5441440/files/figure.png" pageId="36" pageNumber="473">25E, F</figureCitation>
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,
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<figureCitation id="B6982A61FFBBFFA0DF0AE9685B244037" box="[1297,1326,1628,1650]" captionStart="Figure 28" captionStartId="36.[144,223,927,946]" captionTargetBox="[177,1389,200,897]" captionTargetId="graphics-419@36.[220,813,219,871]" captionTargetPageId="36" captionText="Figure 28. Geographical distribution of the species of Heteropachylus." figureDoi="http://doi.org/10.5281/zenodo.5441446" httpUri="https://zenodo.org/record/5441446/files/figure.png" pageId="36" pageNumber="473">28</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="66B9656FFFBBFFA0D93DE9B75D90412B" pageId="36" pageNumber="473" type="reference_group">
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<paragraph id="2E1C36E4FFBBFFA0D93DE9B75D5C4093" blockId="36.[806,1422,1666,1902]" pageId="36" pageNumber="473">
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<treatmentCitationGroup id="0EB311CAFFBBFFA0D93DE9B75D5C4093" pageId="36" pageNumber="473">
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<treatmentCitation id="AF0210F5FFBBFFA0D93DE9B75B8340DD" author="Soares HEM" box="[806,1417,1666,1688]" page="252" pageId="36" pageNumber="473" year="1977">
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<taxonomicName id="E9A34D67FFBBFFA0D93DE9B75B8340DD" ID-CoL="7QMXM" authority="Soares, 1977: 252" authorityName="H. Soares" authorityPageNumber="252" authorityYear="1977" box="[806,1417,1666,1688]" class="Arachnida" family="Gonyleptidae" genus="Aesotrinoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="36" pageNumber="473" phylum="Arthropoda" rank="species" species="crassicalcanei">
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<emphasis id="1CD7EAF6FFBBFFA0D93DE9B75A7840DD" box="[806,1138,1667,1688]" italics="true" pageId="36" pageNumber="473">Aesotrinoma crassicalcanei</emphasis>
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<bibRefCitation id="4A324B15FFBBFFA0DE8BE9B75B8340DD" author="Soares HEM" box="[1168,1417,1666,1688]" pageId="36" pageNumber="473" pagination="251 - 260" refId="ref32399" refString="Soares HEM. 1977. Opera Opiliologica Varia VI (Opiliones, Gonyleptidae). Physis, Seccion C 36: 251 - 260." type="journal article" year="1977">Soares, 1977: 252</bibRefCitation>
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</taxonomicName>
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, figures 1–6.
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||
<treatmentCitation id="AF0210F5FFBBFFA0D9AFE9965A7640F3" author="Kury AB" box="[948,1148,1697,1719]" page="142" pageId="36" pageNumber="473" year="2003">
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<bibRefCitation id="4A324B15FFBBFFA0D9AFE9965A7640F3" author="Kury AB" box="[948,1148,1697,1719]" pageId="36" pageNumber="473" pagination="1 - 337" refId="ref31027" refString="Kury AB. 2003 a. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia especial monografico: 1 - 337." type="book chapter" year="2003">Kury, 2003a: 142</bibRefCitation>
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</treatmentCitation>
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.
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<treatmentCitation id="AF0210F5FFBBFFA0DE92E9955D584093" author="Mendes AC & Kury AB" page="151" pageId="36" pageNumber="473" year="2003">
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<bibRefCitation id="4A324B15FFBBFFA0DE92E9955D584093" author="Mendes AC & Kury AB" pageId="36" pageNumber="473" pagination="151 - 156" refId="ref31553" refString="Mendes AC, Kury AB. 2003. A review of Thaumatoleptes Roewer (Arachnida, Opiliones, Gonyleptidae). Revista Iberica de Aracnologia 7: 151 - 156." type="journal article" year="2003">Mendes & Kury, 2003: 151</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="2E1C36E4FFBBFFA0D93DE9EA5D90412B" blockId="36.[806,1422,1666,1902]" pageId="36" pageNumber="473">
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<treatmentCitationGroup id="0EB311CAFFBBFFA0D93DE9EA5D90412B" pageId="36" pageNumber="473">
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||
<treatmentCitation id="AF0210F5FFBBFFA0D93DE9EA5D874157" author="Sigrist MS & Carvalho CJB" page="39" pageId="36" pageNumber="473" year="2008">
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||
<taxonomicName id="E9A34D67FFBBFFA0D93DE9EA5D874157" authority=": Sigrist & Carvalho, 2008: 39" authorityName=": Sigrist & Carvalho" authorityPageNumber="39" authorityYear="2008" class="Arachnida" family="Gonyleptidae" genus="Chavesincola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="36" pageNumber="473" phylum="Arthropoda" rank="species" species="crassicalcanei">
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<emphasis id="1CD7EAF6FFBBFFA0D93DE9EA5A7940B6" box="[806,1139,1758,1779]" italics="true" pageId="36" pageNumber="473">Chavesincola crassicalcanei</emphasis>
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:
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<bibRefCitation id="4A324B15FFBBFFA0DE8BE9EA5D874157" author="Sigrist MS & Carvalho CJB" pageId="36" pageNumber="473" refId="ref32145" refString="Sigrist MS, Carvalho CJB. 2008. Detection of areas of endemism on two spatial scales using parsimony analysis of endemicity (PAE): the Neotropical region and the Atlantic Forest. Biota Neotropica 8: 033 - 042." type="journal volume" year="2008">Sigrist & Carvalho, 2008: 39</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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. [This combination was never formally proposed; the authors used data from an unpublished MSc thesis (
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||
<bibRefCitation id="4A324B15FFBBFFA0D9A0E80E5A6A4115" author="Mendes AC" box="[955,1120,1850,1872]" pageId="36" pageNumber="473" refId="ref31519" refString="Mendes AC. 2005. Revisao sistematica e analise filogenetica de Heteropachylinae Kury, 1994 (Arachnida, Opiliones, Gonyleptidae). Unpublished MSc Thesis, Universidade Federal do Rio de Janeiro." type="book" year="2005">Mendes, 2005</bibRefCitation>
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), not valid for taxonomic purposes.]
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="66B9656FFFBAFFA1DABFEFF25D774366" pageId="37" pageNumber="474" type="distribution">
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<paragraph id="2E1C36E4FFBAFFA1DABFEFF25CDC46BF" blockId="37.[164,779,198,250]" pageId="37" pageNumber="474">
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<emphasis id="1CD7EAF6FFBAFFA1DABFEFF25F3F469E" box="[164,309,198,219]" italics="true" pageId="37" pageNumber="474">Distribution:</emphasis>
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Species known only from
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<typeStatus id="F1188846FFBAFFA1D86CEFF25CA2469E" box="[631,680,198,219]" pageId="37" pageNumber="474">type</typeStatus>
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||
locality. WWF Ecoregion: NT0103,
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||
<collectingRegion id="EC67F806FFBAFFA1DBC0EFD05C1546BF" box="[475,543,228,250]" country="Brazil" name="Bahia" pageId="37" pageNumber="474">Bahia</collectingRegion>
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coastal forests.
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</paragraph>
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<paragraph id="2E1C36E4FFBAFFA1DABFEE165F0647D6" blockId="37.[164,779,289,403]" pageId="37" pageNumber="474">
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<emphasis id="1CD7EAF6FFBAFFA1DABFEE165F904772" box="[164,410,290,311]" italics="true" pageId="37" pageNumber="474">Material examined:</emphasis>
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m
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||
<materialsCitation id="9ECB3CB9FFBAFFA1DBEAEE165F344731" collectingDate="1955-06" collectionCode="MNRJ" collectorName="Becker J" country="Brazil" location="Rio Vermelho." pageId="37" pageNumber="474" specimenCode="MNRJ5216" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
|
||
<typeStatus id="F1188846FFBAFFA1DBEAEE165C58477D" box="[497,594,290,312]" pageId="37" pageNumber="474" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCode id="7E059E9FFFBAFFA1D862EE155CF54772" box="[633,767,289,311]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="37" pageNumber="474">MNRJ5216</specimenCode>
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||
),
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||
<collectingCountry id="56B47674FFBAFFA1DABFEE745F0C4710" box="[164,262,320,341]" name="Brazil" pageId="37" pageNumber="474">BRAZIL</collectingCountry>
|
||
,
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||
<collectingRegion id="EC67F806FFBAFFA1DB00EE745F6B4713" box="[283,353,320,342]" country="Brazil" name="Bahia" pageId="37" pageNumber="474">Bahia</collectingRegion>
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, Salvador.
|
||
<location id="2B7C603FFFBAFFA1DBEFEE745CAA4713" LSID="urn:lsid:plazi:treatment:A60A87F2FFBBFFA1D92AE9095D774366:2B7C603FFFBAFFA1DBEFEE745CAA4713" box="[500,672,320,342]" country="Brazil" name="Rio Vermelho." pageId="37" pageNumber="474" stateProvince="Bahia">Rio Vermelho.</location>
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||
<date id="5A1D1024FFBAFFA1D8AAEE745D0D4713" box="[689,775,320,342]" pageId="37" pageNumber="474" value="1955-06">
|
||
<collectingDate id="4A59E9CCFFBAFFA1D8AAEE745D0D4713" box="[689,775,320,342]" pageId="37" pageNumber="474" value="1955-06">vi.1955</collectingDate>
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||
</date>
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||
.
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||
<collectorName id="83565332FFBAFFA1DABFEE6B5F184731" box="[164,274,351,373]" pageId="37" pageNumber="474">Becker J</collectorName>
|
||
; m
|
||
</materialsCitation>
|
||
<materialsCitation id="9ECB3CB9FFBAFFA1DB55EE6B5F0247D6" collectingDate="1955-06" collectionCode="HEMS" collectorName="Becker J" country="Brazil" location="Rio Vermelho." pageId="37" pageNumber="474" specimenCode="HEMS382" specimenCount="1" stateProvince="Bahia" typeStatus="paratype">
|
||
<typeStatus id="F1188846FFBAFFA1DB55EE6B5FBE4731" box="[334,436,351,372]" pageId="37" pageNumber="474" type="paratype">paratype</typeStatus>
|
||
(
|
||
<specimenCode id="7E059E9FFFBAFFA1DBD7EE6B5C4C4730" box="[460,582,351,373]" collectionCode="HEMS" country="United Kingdom" httpUri="http://grbio.org/cool/35t9-4j66" name="Haslemere Educational Museum" pageId="37" pageNumber="474">HEMS382</specimenCode>
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||
), same data as holotype
|
||
</materialsCitation>
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||
.
|
||
</paragraph>
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||
<paragraph id="2E1C36E4FFBAFFA1DABFEE8F5C424538" blockId="37.[164,779,443,893]" pageId="37" pageNumber="474">
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||
<emphasis id="1CD7EAF6FFBAFFA1DABFEE8F5F114795" box="[164,283,443,464]" italics="true" pageId="37" pageNumber="474">Diagnosis:</emphasis>
|
||
Three mesotergal areas (
|
||
<figureCitation id="B6982A61FFBAFFA1D880EE8F5CF54794" box="[667,767,443,465]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A</figureCitation>
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||
). Mesotergal area I divided by longitudinal groove; complete groove between mesotergal areas II and III (
|
||
<figureCitation id="B6982A61FFBAFFA1DAB7ED225F054468" box="[172,271,534,557]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A</figureCitation>
|
||
); mesotergal area III at the same level as the other areas and with unpaired armature (
|
||
<figureCitation id="B6982A61FFBAFFA1DAB7ED605F37442C" box="[172,317,596,618]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A, D</figureCitation>
|
||
); free tergites II–III with projected corners and median blunt conical apophysis (
|
||
<figureCitation id="B6982A61FFBAFFA1D8B8ED465ED744E3" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A, B, D</figureCitation>
|
||
); free tergite III without longitudinal keel, not fused to the dorsal scutum in males (
|
||
<figureCitation id="B6982A61FFBAFFA1D86EED9B5CF44480" box="[629,766,687,709]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A, B</figureCitation>
|
||
). Dorsal anal operculum covered by tubercles, elevated in the middle (
|
||
<figureCitation id="B6982A61FFBAFFA1DB4FEDD95FD14547" box="[340,475,749,771]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17B, D</figureCitation>
|
||
). Femur IV (
|
||
<figureCitation id="B6982A61FFBAFFA1D86DEDD95CCE4546" box="[630,708,749,771]" captionStart="Figure 18" captionStartId="27.[164,243,1135,1154]" captionTargetBox="[164,777,655,1048]" captionTargetPageId="27" captionText="Figure 18. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. Right trochanter, femur and patella IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441426" httpUri="https://zenodo.org/record/5441426/files/figure.png" pageId="37" pageNumber="474">Fig. 18</figureCitation>
|
||
) with bifid dorsobasal apophysis, lacking dorsodistal one.
|
||
<emphasis id="1CD7EAF6FFBAFFA1DABFEC1E5F58457A" box="[164,338,810,831]" italics="true" pageId="37" pageNumber="474">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="B6982A61FFBAFFA1DB70EC1E5FFC457A" box="[363,502,810,832]" captionStart="Figure 25" captionStartId="34.[144,223,1747,1766]" captionTargetBox="[206,1359,196,1716]" captionTargetPageId="34" captionText="Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441440" httpUri="https://zenodo.org/record/5441440/files/figure.png" pageId="37" pageNumber="474">Fig. 25E, F</figureCitation>
|
||
): fourth pair of setae larger than the first three and dislocated to the median portion of the ventral plate.
|
||
</paragraph>
|
||
<paragraph id="2E1C36E4FFBAFFA1DABFEC915C8045B2" blockId="37.[164,779,932,1015]" pageId="37" pageNumber="474">
|
||
<emphasis id="1CD7EAF6FFBAFFA1DABFEC915F4445FF" box="[164,334,933,954]" italics="true" pageId="37" pageNumber="474">Measurements:</emphasis>
|
||
Male
|
||
<typeStatus id="F1188846FFBAFFA1DBBEEC905C0345FF" box="[421,521,932,954]" pageId="37" pageNumber="474" type="holotype">holotype</typeStatus>
|
||
. CL: 1.6; MCW: 2.3; ASL: 3.4; MASW: 5.0; FL: 3.8. Male
|
||
<typeStatus id="F1188846FFBAFFA1D825ECF75CAD459D" box="[574,679,963,984]" pageId="37" pageNumber="474" type="paratype">paratype</typeStatus>
|
||
. CL: 1.8; MCW: 2.3; ASL: 3.1; MASW: 4.9; FL: 3.1.
|
||
</paragraph>
|
||
<paragraph id="2E1C36E4FFBAFFA1DABFEB2B5F2F4271" blockId="37.[164,779,1055,1903]" box="[164,293,1055,1076]" pageId="37" pageNumber="474">
|
||
<emphasis id="1CD7EAF6FFBAFFA1DABFEB2B5F2F4271" box="[164,293,1055,1076]" italics="true" pageId="37" pageNumber="474">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2E1C36E4FFBAFFA1DABFEB095AA242D5" blockId="37.[164,779,1055,1903]" lastBlockId="37.[826,1442,198,1168]" pageId="37" pageNumber="474">
|
||
<emphasis id="1CD7EAF6FFBAFFA1DABFEB095FB14216" box="[164,443,1085,1107]" italics="true" pageId="37" pageNumber="474">
|
||
Male
|
||
<typeStatus id="F1188846FFBAFFA1DAF7EB095F414217" box="[236,331,1085,1106]" pageId="37" pageNumber="474" type="holotype">holotype</typeStatus>
|
||
: Dorsum
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="B6982A61FFBAFFA1DBC8EB095C3D4216" box="[467,567,1085,1107]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A</figureCitation>
|
||
). Scutum outline pyriform, widest at mesotergal area II. Anterior margin of carapace with three small tubercles on each side. Frontal hump covered with granules. Eye mound with pair of paramedian tubercles (
|
||
<figureCitation id="B6982A61FFBAFFA1D8B8EB8C5EBC42A9" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A, C</figureCitation>
|
||
) and some smaller granules near the eyes. Carapace smooth. Mesotergum divided into three areas by shallow grooves, areas not well marked; mesotergal area I divided into left and right halves by median longitudinal groove; mesotergal areas I–II with pair of small paramedian granules each; mesotergal area III bearing median spiniform apophysis pointed upwards. Posterior margin smooth. Free tergites I–II fused to the scutum, with row of small granules each; free tergite II with a blunt apophysis on each corner and median blunt apophysis dorsally directed with two rounded tubercles on its base, one on each side; free tergite III smooth, with posterior border substraight, projected corners and median blunt apophysis dorsally pointed (
|
||
<figureCitation id="B6982A61FFBAFFA1DB8EE9B75C3740DD" box="[405,573,1667,1689]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A, B, D</figureCitation>
|
||
). Lateral areas of mesotergum each with row of tubercles, the sizes of which increase towards the posterior margin of the dorsal scutum, reaching the groove between areas II and III (
|
||
<figureCitation id="B6982A61FFBAFFA1DB1CE9C95F844157" box="[263,398,1789,1811]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A, D</figureCitation>
|
||
). Dorsal anal operculum covered by tubercles concentrated in the middle, forming an elevation (
|
||
<figureCitation id="B6982A61FFBAFFA1DB2BE80E5FC34115" box="[304,457,1850,1872]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17B, D</figureCitation>
|
||
).
|
||
<emphasis id="1CD7EAF6FFBAFFA1DBEAE80E5C73410A" box="[497,633,1850,1871]" italics="true" pageId="37" pageNumber="474">Appendages</emphasis>
|
||
. Pedipalps (
|
||
<figureCitation id="B6982A61FFBAFFA1DAB7E86D5F25412B" box="[172,303,1881,1903]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17A, D</figureCitation>
|
||
) – trochanter with two ventral setiferous granules. Femur with ventral row of setiferous granules, subapical mesal spine reduced. Tibia with four mesal (Ii + [Ii]) and two ectal ([Ii]) spines. Tarsus with four mesal (IiIi) and four ectal (IiIi) spines. Legs – calcaneus I length about half length of metatarsus I, lighter in colour and conspicuously thicker than the astragalus. Coxa IV armed with apical prolateral blunt apophysis. Trochanter IV armed with apical prolateral blunt apophysis and retrolateral distal curved short apophysis. Femur IV (
|
||
<figureCitation id="B6982A61FFBAFFA1DEE2EEED5B4D47AA" box="[1273,1351,473,495]" captionStart="Figure 18" captionStartId="27.[164,243,1135,1154]" captionTargetBox="[164,777,655,1048]" captionTargetPageId="27" captionText="Figure 18. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. Right trochanter, femur and patella IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441426" httpUri="https://zenodo.org/record/5441426/files/figure.png" pageId="37" pageNumber="474">Fig. 18</figureCitation>
|
||
) short, distinctly curved in dorsal view and with two curvatures (‘S-shaped’) in lateral view, bearing rows of tubercles; with bifid dorsobasal apophysis preceded by a basal rounded tubercle and granules and followed by medial row of rounded tubercles, the sizes of which decrease towards the apex; retrolateral surface with row of rounded tubercles equally sized, prolateral surface with row of about five large rounded tubercles located on the distal half, larger at the median portion of the row; apex (
|
||
<figureCitation id="B6982A61FFBAFFA1DE70EC3F5AC64564" box="[1131,1228,779,801]" captionStart="Figure 17" captionStartId="26.[144,223,1630,1649]" captionTargetBox="[143,1426,195,1600]" captionTargetId="graphics-82@26.[145,1426,196,1599]" captionTargetPageId="26" captionText="Figure 17. Heteropachylus crassicalcanei comb. nov. Male holotype MNRJ 5216. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, patella and apex of femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441424" httpUri="https://zenodo.org/record/5441424/files/figure.png" pageId="37" pageNumber="474">Fig. 17E</figureCitation>
|
||
) with retrolateral conical apophysis, pointed upwards in apical view, and prolateral rounded tubercle with acuminated apex. Tarsal counts: 6/11/6/6.
|
||
<emphasis id="1CD7EAF6FFBAFFA1DEFDEC535BAB4539" box="[1254,1441,871,892]" italics="true" pageId="37" pageNumber="474">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="B6982A61FFBAFFA1D959ECB25DC345DE" box="[834,969,901,924]" captionStart="Figure 25" captionStartId="34.[144,223,1747,1766]" captionTargetBox="[206,1359,196,1716]" captionTargetPageId="34" captionText="Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441440" httpUri="https://zenodo.org/record/5441440/files/figure.png" pageId="37" pageNumber="474">Fig. 25E, F</figureCitation>
|
||
). Ventral plate with cleft ‘U’-shaped at the distal border; lateral apexes of ventral plate curved. Group of three distal short setae; two basal long setae.
|
||
<taxonomicName id="E9A34D67FFBAFFA1D9D7ECD55A0545B2" authorityName="Megerle von Muhlfeld" authorityYear="1811" box="[972,1039,993,1015]" class="Bivalvia" family="Carditidae" genus="Glans" kingdom="Animalia" order="Carditida" pageId="37" pageNumber="444" phylum="Mollusca" rank="genus">Glans</taxonomicName>
|
||
with thumb-like dorsal process.
|
||
<taxonomicName id="E9A34D67FFBAFFA1D921EB345D884253" box="[826,898,1024,1046]" class="Pilidiophora" family="Lineidae" genus="Stylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Heteronemertea" pageId="37" pageNumber="444" phylum="Nemertea" rank="genus">Stylus</taxonomicName>
|
||
with apical setae subequally sized.
|
||
<emphasis id="1CD7EAF6FFBAFFA1DF30EB345D914271" italics="true" pageId="37" pageNumber="474">Colour (in alcohol).</emphasis>
|
||
Background colour of body and coxa, trochanter, and femur IV dark brown; Legs I–III light brown, except the tarsi, which are yellow and calcaneus I pale.
|
||
<emphasis id="1CD7EAF6FFBAFFA1D9D5EB4E5A2242CA" box="[974,1064,1146,1167]" italics="true" pageId="37" pageNumber="474">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="2E1C36E4FFBAFFA1D921EBE55D774366" blockId="37.[826,1441,1232,1315]" pageId="37" pageNumber="474">
|
||
<emphasis id="1CD7EAF6FFBAFFA1D921EBE55D7F42A3" box="[826,885,1233,1254]" italics="true" pageId="37" pageNumber="474">Note:</emphasis>
|
||
Male genitalia in bad state of conservation, probably because of presumed past treatment with KOH.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |