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<document id="D59EFB2B30A908CF8653268A109322AA" ID-DOI="10.1111/j.1096-3642.2011.00706.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5441392" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630451174621" checkinUser="valdenar" docAuthor="Mendes, Amanda Cruz" docDate="2011" docId="A60A87F2FFB9FFADD926E9E25CFB4795" docLanguage="en" docName="j.1096-3642.2011.00706.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (2)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00706.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Heteropachylus peracchii Soares &amp; Soares 1974" docType="treatment" docVersion="3" lastPageNumber="478" masterDocId="5A33FF8AFF9FFF84DA1BEF345E0A4645" masterDocTitle="Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae)" masterLastPageNumber="483" masterPageNumber="437" pageNumber="475" updateTime="1699275278278" updateUser="plazi">
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<mods:title id="3F266FC150E74BD6F0FF85964EBEED3D">Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae)</mods:title>
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<mods:namePart id="FE51409AB327C28B58F574CE992498D3">Mendes, Amanda Cruz</mods:namePart>
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<mods:date id="FF1890A6D247B6D5D1D32123B99A6B14">2011</mods:date>
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<mods:identifier id="8C9A2EB412E55EB7B38F504E31298791" type="DOI">10.1111/j.1096-3642.2011.00706.x</mods:identifier>
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<treatment id="A60A87F2FFB9FFADD926E9E25CFB4795" LSID="urn:lsid:plazi:treatment:A60A87F2FFB9FFADD926E9E25CFB4795" httpUri="http://treatment.plazi.org/id/A60A87F2FFB9FFADD926E9E25CFB4795" lastPageId="41" lastPageNumber="478" pageId="38" pageNumber="475">
<subSubSection id="66B9656FFFB9FFA2D926E9E25B02414E" pageId="38" pageNumber="475" type="nomenclature">
<paragraph id="2E1C36E4FFB9FFA2D926E9E25B02414E" blockId="38.[829,1398,1749,1803]" pageId="38" pageNumber="475">
<heading id="75548188FFB9FFA2D926E9E25B02414E" centered="true" fontSize="8" level="2" pageId="38" pageNumber="475" reason="8">
<taxonomicName id="E9A34D67FFB9FFA2D926E9E25DE9414F" ID-CoL="7WLPM" authority="SOARES &amp; SOARES, 1974" authorityName="Soares &amp; Soares" authorityYear="1974" class="Arachnida" family="Gonyleptidae" genus="Heteropachylus" kingdom="Animalia" order="Opiliones" pageId="38" pageNumber="475" phylum="Arthropoda" rank="species" species="peracchii">
<emphasis id="1CD7EAF6FFB9FFA2D926E9E25A8440AC" box="[829,1166,1750,1771]" italics="true" pageId="38" pageNumber="475">HETEROPACHYLUS PERACCHII</emphasis>
<bibRefCitation id="4A324B15FFB9FFA2DE83E9E15DE9414F" author="Soares BAM &amp; Soares HEM" pageId="38" pageNumber="475" pagination="599 - 614" refId="ref32314" refString="Soares BAM, Soares HEM. 1974. Opera Opiliologica Varia IV (Opiliones, Gonyleptidae, Stygnidae). Revista brasileira de Biologia 34: 599 - 614." type="journal article" year="1974">SOARES &amp; SOARES, 1974</bibRefCitation>
</taxonomicName>
(
<figureCitation id="B6982A61FFB9FFA2D9EDE9C15A5A414F" box="[1014,1104,1781,1802]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="38" pageNumber="475">FIGS 21</figureCitation>
,
<figureCitation id="B6982A61FFB9FFA2DE44E9C15A74414F" box="[1119,1150,1781,1802]" captionStart="Figure 22" captionStartId="31.[164,243,1310,1329]" captionTargetBox="[165,782,196,1245]" captionTargetId="graphics-515@31.[165,781,196,1244]" captionTargetPageId="31" captionText="Figure 22. Heteropachylus peracchii comb. nov. Male holotype HEMS 485. Left trochanter and femur IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441434" httpUri="https://zenodo.org/record/5441434/files/figure.png" pageId="38" pageNumber="475">22</figureCitation>
,
<figureCitation id="B6982A61FFB9FFA2DE96E9C15AD9414F" box="[1165,1235,1781,1803]" captionStart="Figure 25" captionStartId="34.[144,223,1747,1766]" captionTargetBox="[206,1359,196,1716]" captionTargetPageId="34" captionText="Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441440" httpUri="https://zenodo.org/record/5441440/files/figure.png" pageId="38" pageNumber="475">25I, J</figureCitation>
,
<figureCitation id="B6982A61FFB9FFA2DEF8E9C15B0A414E" box="[1251,1280,1781,1803]" captionStart="Figure 28" captionStartId="36.[144,223,927,946]" captionTargetBox="[177,1389,200,897]" captionTargetId="graphics-419@36.[220,813,219,871]" captionTargetPageId="36" captionText="Figure 28. Geographical distribution of the species of Heteropachylus." figureDoi="http://doi.org/10.5281/zenodo.5441446" httpUri="https://zenodo.org/record/5441446/files/figure.png" pageId="38" pageNumber="475">28</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="66B9656FFFB9FFA2D93DE8285D9A412B" pageId="38" pageNumber="475" type="reference_group">
<paragraph id="2E1C36E4FFB9FFA2D93DE8285D9A412B" blockId="38.[806,1421,1820,1902]" pageId="38" pageNumber="475">
<treatmentCitationGroup id="0EB311CAFFB9FFA2D93DE8285D9A412B" pageId="38" pageNumber="475">
<treatmentCitation id="AF0210F5FFB9FFA2D93DE8285B834177" author="Soares BAM &amp; Soares HEM" box="[806,1417,1820,1842]" page="609" pageId="38" pageNumber="475" year="1974">
<taxonomicName id="E9A34D67FFB9FFA2D93DE8285B834177" ID-CoL="7WLPM" authority="Soares &amp; Soares, 1974: 609" authorityName="Soares &amp; Soares" authorityPageNumber="609" authorityYear="1974" box="[806,1417,1820,1842]" class="Arachnida" family="Gonyleptidae" genus="Heteropachylus" kingdom="Animalia" order="Opiliones" pageId="38" pageNumber="475" phylum="Arthropoda" rank="species" species="peracchii">
<emphasis id="1CD7EAF6FFB9FFA2D93DE8285A4E4174" box="[806,1092,1820,1841]" italics="true" pageId="38" pageNumber="475">Heteropachylus peracchii</emphasis>
<bibRefCitation id="4A324B15FFB9FFA2DE56E8285B834177" author="Soares BAM &amp; Soares HEM" box="[1101,1417,1820,1842]" pageId="38" pageNumber="475" pagination="599 - 614" refId="ref32314" refString="Soares BAM, Soares HEM. 1974. Opera Opiliologica Varia IV (Opiliones, Gonyleptidae, Stygnidae). Revista brasileira de Biologia 34: 599 - 614." type="journal article" year="1974">Soares &amp; Soares, 1974: 609</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figures 44–51.
<treatmentCitation id="AF0210F5FFB9FFA2D9CAE80F5A90410A" author="Kury AB" box="[977,1178,1850,1872]" page="142" pageId="38" pageNumber="475" year="2003">
<bibRefCitation id="4A324B15FFB9FFA2D9CAE80F5A90410A" author="Kury AB" box="[977,1178,1850,1872]" pageId="38" pageNumber="475" pagination="1 - 337" refId="ref31027" refString="Kury AB. 2003 a. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia especial monografico: 1 - 337." type="book chapter" year="2003">Kury, 2003a: 142</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF0210F5FFB9FFA2DEB2E80E5D87412B" author="Sigrist MS &amp; Carvalho CJB" page="39" pageId="38" pageNumber="475" year="2008">
<bibRefCitation id="4A324B15FFB9FFA2DEB2E80E5D87412B" author="Sigrist MS &amp; Carvalho CJB" pageId="38" pageNumber="475" refId="ref32145" refString="Sigrist MS, Carvalho CJB. 2008. Detection of areas of endemism on two spatial scales using parsimony analysis of endemicity (PAE): the Neotropical region and the Atlantic Forest. Biota Neotropica 8: 033 - 042." type="journal volume" year="2008">Sigrist &amp; Carvalho, 2008: 39</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="66B9656FFFB8FFACDABFEFF25C5C46B9" lastPageId="40" lastPageNumber="477" pageId="39" pageNumber="476" type="distribution">
<paragraph id="2E1C36E4FFB8FFA3DABFEFF25CDD475C" blockId="39.[164,779,198,281]" pageId="39" pageNumber="476">
<emphasis id="1CD7EAF6FFB8FFA3DABFEFF25F3F469E" box="[164,309,198,219]" italics="true" pageId="39" pageNumber="476">Distribution:</emphasis>
<collectingCountry id="56B47674FFB8FFA3DB5CEFF25FA0469E" box="[327,426,198,219]" name="Brazil" pageId="39" pageNumber="476">BRAZIL</collectingCountry>
.
<collectingRegion id="EC67F806FFB8FFA3DBA5EFF25C084699" box="[446,514,198,220]" country="Brazil" name="Bahia" pageId="39" pageNumber="476">Bahia</collectingRegion>
(Ilhéus and Itajibá). WWF Ecoregions: NT0103 and NT0104, respectively,
<collectingRegion id="EC67F806FFB8FFA3DABFEE375EE2475C" box="[164,232,259,281]" country="Brazil" name="Bahia" pageId="39" pageNumber="476">Bahia</collectingRegion>
coastal forests and
<collectingRegion id="EC67F806FFB8FFA3DBCEEE375C13475C" box="[469,537,259,281]" country="Brazil" name="Bahia" pageId="39" pageNumber="476">Bahia</collectingRegion>
interior forests.
</paragraph>
<paragraph id="2E1C36E4FFB8FFA3DABFEE745EC84795" blockId="39.[164,779,320,464]" pageId="39" pageNumber="476">
<emphasis id="1CD7EAF6FFB8FFA3DABFEE745F8E4710" box="[164,388,320,341]" italics="true" pageId="39" pageNumber="476">Material examined:</emphasis>
<materialsCitation id="9ECB3CB9FFB8FFA3DB94EE755EC24730" collectionCode="HEMS" pageId="39" pageNumber="476" specimenCode="HEMS 485" specimenCount="1" typeStatus="holotype">
m
<typeStatus id="F1188846FFB8FFA3DBABEE745C1B4713" box="[432,529,320,342]" pageId="39" pageNumber="476" type="holotype">holotype</typeStatus>
(
<specimenCode id="7E059E9FFFB8FFA3D838EE745CA24713" box="[547,680,320,342]" collectionCode="HEMS" country="United Kingdom" httpUri="http://grbio.org/cool/35t9-4j66" name="Haslemere Educational Museum" pageId="39" pageNumber="476">HEMS 485</specimenCode>
), m and juv
</materialsCitation>
<materialsCitation id="9ECB3CB9FFB8FFA3DAC9EE6B5EB54795" collectingDate="1972-01-13" collectingDateMax="1977-05-20" collectingDateMin="1972-01-13" collectionCode="HEMS" collectorName="Jim J. &amp; de Menezes M" country="Brazil" elevation="2" location="Bahia" pageId="39" pageNumber="476" specimenCode="HEMS 486, HEMS 665" specimenCount="1" stateProvince="Bahia" typeStatus="paratype">
<typeStatus id="F1188846FFB8FFA3DAC9EE6B5F494731" box="[210,323,351,372]" pageId="39" pageNumber="476" type="paratype">paratypes</typeStatus>
(
<specimenCode id="7E059E9FFFB8FFA3DB4DEE6B5FD14730" box="[342,475,351,373]" collectionCode="HEMS" country="United Kingdom" httpUri="http://grbio.org/cool/35t9-4j66" name="Haslemere Educational Museum" pageId="39" pageNumber="476">HEMS 486</specimenCode>
),
<collectingCountry id="56B47674FFB8FFA3DBE9EE6B5C5F4731" box="[498,597,351,372]" name="Brazil" pageId="39" pageNumber="476">BRAZIL</collectingCountry>
.
<collectingRegion id="EC67F806FFB8FFA3D879EE6B5CA34730" box="[610,681,351,373]" country="Brazil" name="Bahia" pageId="39" pageNumber="476">Bahia</collectingRegion>
. Itajibá. Fazenda Pedra Branca.
<date id="5A1D1024FFB8FFA3DBD9EE495C2647D6" box="[450,556,381,403]" pageId="39" pageNumber="476" value="1972-01-13">
<collectingDate id="4A59E9CCFFB8FFA3DBD9EE495C2647D6" box="[450,556,381,403]" pageId="39" pageNumber="476" value="1972-01-13">13.i.1972</collectingDate>
</date>
.
<collectorName id="83565332FFB8FFA3D820EE495C8C47D6" box="[571,646,381,403]" pageId="39" pageNumber="476">Jim J.</collectorName>
<quantity id="E95B9B01FFB8FFA3D88AEE4A5CBD47D6" box="[657,695,382,403]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="476" unit="m" value="2.0">
<elevation id="A58ED1D7FFB8FFA3D88AEE4A5CBD47D6" box="[657,695,382,403]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="476" unit="m" value="2.0">2m</elevation>
</quantity>
, f, and juv (
<specimenCode id="7E059E9FFFB8FFA3DACCEEA85F5347F7" box="[215,345,412,434]" collectionCode="HEMS" country="United Kingdom" httpUri="http://grbio.org/cool/35t9-4j66" name="Haslemere Educational Museum" pageId="39" pageNumber="476">HEMS 665</specimenCode>
), CEPLAC.
<date id="5A1D1024FFB8FFA3DBF8EEA85C7347F4" box="[483,633,412,433]" pageId="39" pageNumber="476" value="1977-05-19" valueMax="1977-05-20" valueMin="1977-05-19">
<collectingDate id="4A59E9CCFFB8FFA3DBF8EEA85C7347F4" box="[483,633,412,433]" pageId="39" pageNumber="476" value="1977-05-19" valueMax="1977-05-20" valueMin="1977-05-19">19–20.v.1977</collectingDate>
</date>
,
<collectorName id="83565332FFB8FFA3D89DEEA85EB54795" pageId="39" pageNumber="476">de Menezes M</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="2E1C36E4FFB8FFA3DABFEECC5FAE45B2" blockId="39.[164,779,504,1015]" pageId="39" pageNumber="476">
<emphasis id="1CD7EAF6FFB8FFA3DABFEECC5F114448" box="[164,283,504,525]" italics="true" pageId="39" pageNumber="476">Diagnosis:</emphasis>
Four mesotergal areas; mesotergal area I divided by longitudinal groove (
<figureCitation id="B6982A61FFB8FFA3D833ED225C814468" box="[552,651,534,557]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A</figureCitation>
); complete groove between mesotergal areas II and III; mesotergal area III bearing row of granules with pair of paramedian tubercles and at the same level of the other areas (
<figureCitation id="B6982A61FFB8FFA3DB90EDA55C1B44E3" box="[395,529,657,679]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A, E</figureCitation>
); mesotergal area IV entire, without a longitudinal groove (
<figureCitation id="B6982A61FFB8FFA3D880ED9B5CF54483" box="[667,767,687,710]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A</figureCitation>
). Free tergites I–III with a small tubercle on each corner, free tergites II–III bearing median conical apophysis (
<figureCitation id="B6982A61FFB8FFA3DB2AEC3F5FCA4564" box="[305,448,779,801]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A, E</figureCitation>
); free tergite III without keel and not fused to the dorsal scutum in males (
<figureCitation id="B6982A61FFB8FFA3DAB7EC7C5F1A451A" box="[172,272,840,863]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A</figureCitation>
). Dorsal anal operculum covered by tubercles and bearing median apophysis (
<figureCitation id="B6982A61FFB8FFA3D8B8EC535EBD45DE" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21E, F</figureCitation>
). Femur IV (
<figureCitation id="B6982A61FFB8FFA3DB4FECB25FA945D9" box="[340,419,902,924]" captionStart="Figure 22" captionStartId="31.[164,243,1310,1329]" captionTargetBox="[165,782,196,1245]" captionTargetId="graphics-515@31.[165,781,196,1244]" captionTargetPageId="31" captionText="Figure 22. Heteropachylus peracchii comb. nov. Male holotype HEMS 485. Left trochanter and femur IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441434" httpUri="https://zenodo.org/record/5441434/files/figure.png" pageId="39" pageNumber="476">Fig. 22</figureCitation>
) with anvil-shaped dorsobasal apophysis and robust dorsodistal apophysis. Male genitalia: the four pairs of distal setae equal sized and forming a group.
</paragraph>
<paragraph id="2E1C36E4FFB8FFA3DABFEB2B5EC74288" blockId="39.[164,779,1054,1229]" pageId="39" pageNumber="476">
<emphasis id="1CD7EAF6FFB8FFA3DABFEB2B5F444271" box="[164,334,1055,1076]" italics="true" pageId="39" pageNumber="476">Measurements:</emphasis>
Male
<typeStatus id="F1188846FFB8FFA3DBBEEB2B5C034270" box="[421,521,1055,1077]" pageId="39" pageNumber="476" type="holotype">holotype</typeStatus>
. CL: 1.5; MCW: 2.1; ASL: 2.8; MASW: 4.0. FL: 2.4. Males (
<emphasis id="1CD7EAF6FFB8FFA3D87AEB0A5C7F4216" box="[609,629,1086,1107]" italics="true" pageId="39" pageNumber="476">N</emphasis>
= 3). CL: 1.5 (1.5–1.5); MCW: 2.1 (2.0–2.1); ASL: 2.9 (2.8–2.9); MASW: 4.0 (3.8–4.2). FL: 2.4 (2.2–2.5). Female (
<emphasis id="1CD7EAF6FFB8FFA3DAB7EBAE5ECA42EA" box="[172,192,1178,1199]" italics="true" pageId="39" pageNumber="476">N</emphasis>
= 1): CL: 1.5; MCW: 2.1; ASL: 2.5; MASW: 3.8. FL: 2.4.
</paragraph>
<paragraph id="2E1C36E4FFB8FFA3DABFEBC15F2F434F" blockId="39.[164,779,1269,1903]" box="[164,293,1269,1290]" pageId="39" pageNumber="476">
<emphasis id="1CD7EAF6FFB8FFA3DABFEBC15F2F434F" box="[164,293,1269,1290]" italics="true" pageId="39" pageNumber="476">Description</emphasis>
</paragraph>
<paragraph id="2E1C36E4FFB8FFA3DABFEA275B144079" blockId="39.[164,779,1269,1903]" lastBlockId="39.[826,1442,198,1597]" pageId="39" pageNumber="476">
<emphasis id="1CD7EAF6FFB8FFA3DABFEA275FB1436C" box="[164,443,1299,1321]" italics="true" pageId="39" pageNumber="476">
Male
<typeStatus id="F1188846FFB8FFA3DAF7EA275F41436D" box="[236,331,1299,1320]" pageId="39" pageNumber="476" type="holotype">holotype</typeStatus>
: Dorsum
</emphasis>
(
<figureCitation id="B6982A61FFB8FFA3DBC8EA275C32436F" box="[467,568,1299,1322]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A</figureCitation>
). Scutum outline pyriform, widest at groove between mesotergal areas I and II. Anterior margin of carapace with three granules at each side. Frontal hump covered with granules. Eye mound with pair of small paramedian tubercles (
<figureCitation id="B6982A61FFB8FFA3DB3BEA985FAD4386" box="[288,423,1452,1475]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A, C</figureCitation>
). Carapace with sparse small granules. Mesotergum divided into four areas; mesotergal area I divided into left and right halves by median longitudinal groove; mesotergal areas I–IV with row of granules each, the paramedian of areas I–III larger than the remaining; mesotergal area IV entire. Posterior margin with row of granules. Free tergites I–II fused to the scutum, each with transversal row of granules and one tubercle on each corner (
<figureCitation id="B6982A61FFB8FFA3DAB7E9F45F244090" box="[172,302,1728,1750]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A, E</figureCitation>
); free tergite II bearing median spiniform apophysis pointed backwards (
<figureCitation id="B6982A61FFB8FFA3D837E9EA5CB740B1" box="[556,701,1758,1780]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A, E</figureCitation>
); free tergite III with transversal row of rounded tubercles, with posterior border slightly convex, one tubercle on each corner and median blunt apophysis (
<figureCitation id="B6982A61FFB8FFA3D8B8E80E5EB2412B" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A, E</figureCitation>
). Lateral areas of mesotergum each with row of tubercles larger at area II reaching area III (
<figureCitation id="B6982A61FFB8FFA3DF21EFF25D4446BF" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A, E</figureCitation>
). Dorsal anal operculum covered by tubercles and bearing median apophysis with broad basis (
<figureCitation id="B6982A61FFB8FFA3DF21EE375D474772" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21E, F</figureCitation>
).
<emphasis id="1CD7EAF6FFB8FFA3D971EE165DF34772" box="[874,1017,290,311]" italics="true" pageId="39" pageNumber="476">Appendages.</emphasis>
Pedipalps (
<figureCitation id="B6982A61FFB8FFA3DE89EE165AF8477D" box="[1170,1266,290,312]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21E</figureCitation>
) – trochanter with two ventral setiferous granules. Femur with ventral row of setiferous granules and with subapical mesal spine. Tibia with four mesal (IiIi) and two ectal ([Ii]) spines. Tarsus with four mesal (IiIi) and four ectal (IiIi) spines. Legs – calcaneus I less than half the length of metatarsus I, lighter in colour, and thicker than astragalus. Coxa IV (
<figureCitation id="B6982A61FFB8FFA3DEF9EECC5B48444B" box="[1250,1346,504,526]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A</figureCitation>
) armed with apical prolateral blunt apophysis. Trochanter IV (
<figureCitation id="B6982A61FFB8FFA3D959ED015DA9440E" box="[834,931,565,587]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21A</figureCitation>
) armed with apical prolateral blunt apophysis and retrolateral curved and acuminate distal apophysis. Femur IV (
<figureCitation id="B6982A61FFB8FFA3DE0DED465A6F44CD" box="[1046,1125,626,648]" captionStart="Figure 22" captionStartId="31.[164,243,1310,1329]" captionTargetBox="[165,782,196,1245]" captionTargetId="graphics-515@31.[165,781,196,1244]" captionTargetPageId="31" captionText="Figure 22. Heteropachylus peracchii comb. nov. Male holotype HEMS 485. Left trochanter and femur IV. A, dorsal view. B, ventral view. C, prolateral view. D, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441434" httpUri="https://zenodo.org/record/5441434/files/figure.png" pageId="39" pageNumber="476">Fig. 22</figureCitation>
) short, distinctly curved in dorsal view and with two curvatures (‘S-shaped’) in lateral view, bearing rows of tubercles; with anvilshaped dorsobasal apophysis preceded by a basal rounded tubercle; dorsal surface with three rows of rounded tubercles equal sized, the distal tubercles forming a circle around a large conical distal apophysis; retrolateral surface with row of rounded tubercles, two subdistal larger than the remaining; prolateral of femur IV with row of blunt apophysis located at the distal two-thirds, larger at the median portion of the row; apex (
<figureCitation id="B6982A61FFB8FFA3DE71ECF75AC6459C" box="[1130,1228,963,985]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21D</figureCitation>
) with retrolateral conical apophysis, pointed upwards in apical view, and prolateral rounded apophysis of acuminate apex. Tarsal counts: 5/8/6/6.
<emphasis id="1CD7EAF6FFB8FFA3DE50EB2B5AFE4271" box="[1099,1268,1055,1076]" italics="true" pageId="39" pageNumber="476">Male genitalia</emphasis>
(
<figureCitation id="B6982A61FFB8FFA3DF12EB2B5B9E4271" box="[1289,1428,1054,1077]" captionStart="Figure 25" captionStartId="34.[144,223,1747,1766]" captionTargetBox="[206,1359,196,1716]" captionTargetPageId="34" captionText="Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441440" httpUri="https://zenodo.org/record/5441440/files/figure.png" pageId="39" pageNumber="476">Fig. 25G, H</figureCitation>
). Ventral plate with distal border distinctly concave. Group of four distal short setae on the dorsal portion, two short distal blunt setae on the ventral portion of ventral plate, one large seta after group of distal setae.
<taxonomicName id="E9A34D67FFB8FFA3D995EB8C5DDB428B" authorityName="Megerle von Muhlfeld" authorityYear="1811" box="[910,977,1208,1230]" class="Bivalvia" family="Carditidae" genus="Glans" kingdom="Animalia" order="Carditida" pageId="39" pageNumber="444" phylum="Mollusca" rank="genus">Glans</taxonomicName>
without dorsal process.
<taxonomicName id="E9A34D67FFB8FFA3DF08EB8C5B51428B" box="[1299,1371,1208,1230]" class="Pilidiophora" family="Lineidae" genus="Stylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Heteronemertea" pageId="39" pageNumber="444" phylum="Nemertea" rank="genus">Stylus</taxonomicName>
with apical setae of subequal size.
<emphasis id="1CD7EAF6FFB8FFA3DEAEEBE25BA842AE" box="[1205,1442,1238,1259]" italics="true" pageId="39" pageNumber="476">Colour (in alcohol).</emphasis>
Background colour of body, and coxa, trochanter, and femur IV dark brown. Chelicerae and pedipalps light brown, reticulated by dark brown. Trochanters I–III orange-light brown. Metatarsus I–IV with dark rings. Tarsi orange-yellow.
<emphasis id="1CD7EAF6FFB8FFA3DE36EA5B5A7543C1" box="[1069,1151,1391,1412]" italics="true" pageId="39" pageNumber="476">Female</emphasis>
(
<figureCitation id="B6982A61FFB8FFA3DE89EA5B5AFF43C0" box="[1170,1269,1391,1413]" captionStart="Figure 21" captionStartId="30.[144,223,1475,1494]" captionTargetBox="[144,1425,195,1445]" captionTargetId="graphics-108@30.[144,1416,195,1444]" captionTargetPageId="30" captionText="Figure 21. Heteropachylus peracchii comb. nov. A, C–F, male holotype HEMS 485. A, habitus, dorsal view; C, frontal view; D, patella and apex of femur IV; E, habitus, lateral view; F, ventroposterior view. B, female HEMS 665. Habitus, dorsal view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441432" httpUri="https://zenodo.org/record/5441432/files/figure.png" pageId="39" pageNumber="476">Fig. 21B</figureCitation>
). Free tergites not fused to the dorsal scutum, free tergite II bearing small median spine. Dorsal anal operculum granulous, unarmed. Apical apophysis of coxa IV reduced in relation to the male and spiniform. Trochanter IV narrow, armed with a subdistal retrolateral spine. Femur IV without conspicuous armature.
</paragraph>
<paragraph id="2E1C36E4FFB8FFA3D921E9B75B6A412A" blockId="39.[826,1441,1666,1903]" pageId="39" pageNumber="476">
<emphasis id="1CD7EAF6FFB8FFA3D921E9B75DA140DD" box="[826,939,1667,1688]" italics="true" pageId="39" pageNumber="476">Variation:</emphasis>
The description was based on a not fully developed male (the
<typeStatus id="F1188846FFB8FFA3DE24E9955AAE40F2" box="[1087,1188,1697,1719]" pageId="39" pageNumber="476" type="holotype">holotype</typeStatus>
). In more developed exemplars, the ratio between the maximum width of the abdominal scutum and the carapace is greater and the apophyses of the free tergites and leg IV are more developed. Exemplars less developed than the
<typeStatus id="F1188846FFB8FFA3D921E80E5D914115" box="[826,923,1850,1872]" pageId="39" pageNumber="476" type="holotype">holotype</typeStatus>
were also detected. The armature of their coxa, trochanter, and femur IV is even shorter.
</paragraph>
<paragraph id="2E1C36E4FFB7FFACDAD3EFF35CB4469E" blockId="40.[200,702,198,252]" box="[200,702,198,220]" pageId="40" pageNumber="477">
<taxonomicName id="E9A34D67FFB7FFACDAD3EFF35CB4469E" authority="ROEWER, 1913" authorityName="Roewer" authorityYear="1913" box="[200,702,198,220]" class="Arachnida" family="Gonyleptidae" genus="Heteropachylus" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="477" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="1CD7EAF6FFB7FFACDAD3EFF35C01469F" box="[200,523,199,220]" italics="true" pageId="40" pageNumber="477">HETEROPACHYLUS SPINIGER</emphasis>
ROEWER, 1913
</taxonomicName>
</paragraph>
<paragraph id="2E1C36E4FFB7FFACDB2BEFD25C5C46B9" blockId="40.[200,702,198,252]" box="[304,598,230,252]" pageId="40" pageNumber="477">
(
<figureCitation id="B6982A61FFB7FFACDB23EFD25F9846BE" box="[312,402,230,251]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">FIGS 23</figureCitation>
,
<figureCitation id="B6982A61FFB7FFACDBB9EFD25FCB46BE" box="[418,449,230,251]" captionStart="Figure 24" captionStartId="33.[164,243,1369,1388]" captionTargetBox="[176,757,778,1339]" captionTargetId="graphics-488@33.[176,750,1056,1339]" captionTargetPageId="33" captionText="Figure 24. Heteropachylus spiniger. Male lectotype SMF RI 798. A, right trochanter, femur, and patella IV. Dorsal view. B, right trochanter and femur. Ventral view. C, right femur and patella IV. Prolateral view. The arrow indicates the anvil-shaped dorsobasal apophysis. D, right femur and patella IV. Retrolateral view. The arrow indicates the two subdistal tubercles. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441438" httpUri="https://zenodo.org/record/5441438/files/figure.png" pageId="40" pageNumber="477">24</figureCitation>
,
<figureCitation id="B6982A61FFB7FFACDBD4EFD25C2946BE" box="[463,547,230,252]" captionStart="Figure 25" captionStartId="34.[144,223,1747,1766]" captionTargetBox="[206,1359,196,1716]" captionTargetPageId="34" captionText="Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441440" httpUri="https://zenodo.org/record/5441440/files/figure.png" pageId="40" pageNumber="477">25K, L</figureCitation>
,
<figureCitation id="B6982A61FFB7FFACD829EFD25C4546B9" box="[562,591,230,252]" captionStart="Figure 28" captionStartId="36.[144,223,927,946]" captionTargetBox="[177,1389,200,897]" captionTargetId="graphics-419@36.[220,813,219,871]" captionTargetPageId="36" captionText="Figure 28. Geographical distribution of the species of Heteropachylus." figureDoi="http://doi.org/10.5281/zenodo.5441446" httpUri="https://zenodo.org/record/5441446/files/figure.png" pageId="40" pageNumber="477">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="66B9656FFFB7FFACDA8BEE385C2A47D9" pageId="40" pageNumber="477" type="reference_group">
<paragraph id="2E1C36E4FFB7FFACDA8BEE385C2A47D9" blockId="40.[144,759,268,413]" pageId="40" pageNumber="477">
<treatmentCitationGroup id="0EB311CAFFB7FFACDA8BEE385C2A47D9" pageId="40" pageNumber="477">
<treatmentCitation id="AF0210F5FFB7FFACDA8BEE385C744767" author="Roewer CF" box="[144,638,268,290]" page="76" pageId="40" pageNumber="477" year="1913">
<taxonomicName id="E9A34D67FFB7FFACDA8BEE385C744767" ID-CoL="7WLPP" authority="Roewer, 1913 a: 76" authorityName="Roewer" authorityPageNumber="76" authorityYear="1913" box="[144,638,268,290]" class="Arachnida" family="Gonyleptidae" genus="Heteropachylus" kingdom="Animalia" order="Opiliones" pageId="40" pageNumber="477" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="1CD7EAF6FFB7FFACDA8BEE385FA94767" box="[144,419,268,290]" italics="true" pageId="40" pageNumber="477">Heteropachylus spiniger</emphasis>
<bibRefCitation id="4A324B15FFB7FFACDBB0EE395C744767" author="Roewer CF" box="[427,638,268,290]" pageId="40" pageNumber="477" pagination="1 - 256" refId="ref31985" refString="Roewer CF. 1913 a. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte 4: 1 - 256." type="journal article" year="1913">Roewer, 1913a: 76</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figure 34, 1923: 421, figure 525;
<treatmentCitation id="AF0210F5FFB7FFACDB8DEE1F5CA34705" author="Mello-Leitao CF" box="[406,681,299,321]" page="123" pageId="40" pageNumber="477" year="1923">
<bibRefCitation id="4A324B15FFB7FFACDB8DEE1F5CA34705" author="Mello-Leitao CF" box="[406,681,299,321]" pageId="40" pageNumber="477" pagination="107 - 197" refId="ref31311" refString="Mello-Leitao CF. 1923. Opilioes Laniatores do Brasil. Archivos do Museu. Nacional do Rio de Janeiro 24: 107 - 197." type="journal article" year="1923">Mello-Leitão, 1923: 123</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="AF0210F5FFB7FFACD8A2EE1F5EB1471A" author="Mello-Leitao CF" page="186" pageId="40" pageNumber="477" year="1932">
<bibRefCitation id="4A324B15FFB7FFACD8A2EE1F5EB1471A" author="Mello-Leitao CF" pageId="40" pageNumber="477" pagination="1 - 505" refId="ref31402" refString="Mello-Leitao CF. 1932. Opilioes do Brasil. Revista do Museu Paulista 17: 1 - 505." type="journal article" year="1932">1932: 186</bibRefCitation>
</treatmentCitation>
, figure 113;
<treatmentCitation id="AF0210F5FFB7FFACDB5CEE7E5C89471A" author="Soares BAM &amp; Soares HEM" box="[327,643,329,351]" page="267" pageId="40" pageNumber="477" year="1954">
<bibRefCitation id="4A324B15FFB7FFACDB5CEE7E5C89471A" author="Soares BAM &amp; Soares HEM" box="[327,643,329,351]" pageId="40" pageNumber="477" pagination="225 - 302" refId="ref32284" refString="Soares BAM, Soares HEM. 1954. Monografia dos generos de opilioes neotropicos III. Archivos de Zoologia do Estado de Sao Paulo 8: 225 - 302." type="journal article" year="1954">Soares &amp; Soares, 1954: 267</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF0210F5FFB7FFACD895EE7E5F6F473B" author="Soares BAM &amp; Soares HEM" page="611" pageId="40" pageNumber="477" year="1974">
<bibRefCitation id="4A324B15FFB7FFACD895EE7E5F6F473B" author="Soares BAM &amp; Soares HEM" pageId="40" pageNumber="477" pagination="599 - 614" refId="ref32314" refString="Soares BAM, Soares HEM. 1974. Opera Opiliologica Varia IV (Opiliones, Gonyleptidae, Stygnidae). Revista brasileira de Biologia 34: 599 - 614." type="journal article" year="1974">Soares &amp; Soares, 1974: 611</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF0210F5FFB7FFACDB6CEE5C5C444738" author="Acosta LE" box="[375,590,360,382]" page="217" pageId="40" pageNumber="477" year="1996">
<bibRefCitation id="4A324B15FFB7FFACDB6CEE5C5C444738" author="Acosta LE" box="[375,590,360,382]" pageId="40" pageNumber="477" pagination="209 - 225" refId="ref30352" refString="Acosta LE. 1996. Die Typus-Exemplare der von Carl- Friedrich Roewer beschriebenen Pachylinae (Arachnida: Opiliones: Gonyleptidae). Senckenbergiana Biologica 76: 209 - 225." type="journal article" year="1996">Acosta, 1996: 217</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="AF0210F5FFB7FFACD844EE5D5EB147D9" author="Kury AB" page="142" pageId="40" pageNumber="477" year="2003">
<bibRefCitation id="4A324B15FFB7FFACD844EE5D5EB147D9" author="Kury AB" pageId="40" pageNumber="477" pagination="1 - 337" refId="ref31027" refString="Kury AB. 2003 a. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia especial monografico: 1 - 337." type="book chapter" year="2003">Kury, 2003a: 142</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF0210F5FFB7FFACDAD2EEB35C1747D9" author="Sigrist MS &amp; Carvalho CJB" box="[201,541,391,413]" page="39" pageId="40" pageNumber="477" year="2008">
<bibRefCitation id="4A324B15FFB7FFACDAD2EEB35C1747D9" author="Sigrist MS &amp; Carvalho CJB" box="[201,541,391,413]" pageId="40" pageNumber="477" refId="ref32145" refString="Sigrist MS, Carvalho CJB. 2008. Detection of areas of endemism on two spatial scales using parsimony analysis of endemicity (PAE): the Neotropical region and the Atlantic Forest. Biota Neotropica 8: 033 - 042." type="journal volume" year="2008">Sigrist &amp; Carvalho, 2008: 39</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="66B9656FFFB7FFADDA8BEEF35CFB4795" lastPageId="41" lastPageNumber="478" pageId="40" pageNumber="477" type="distribution">
<paragraph id="2E1C36E4FFB7FFACDA8BEEF35C9C44B4" blockId="40.[144,759,455,753]" pageId="40" pageNumber="477">
<emphasis id="1CD7EAF6FFB7FFACDA8BEEF35F854798" box="[144,399,455,477]" italics="true" pageId="40" pageNumber="477">
Distribution (
<figureCitation id="B6982A61FFB7FFACDB2AEEFC5F8B4798" box="[305,385,456,477]" captionStart="Figure 28" captionStartId="36.[144,223,927,946]" captionTargetBox="[177,1389,200,897]" captionTargetId="graphics-419@36.[220,813,219,871]" captionTargetPageId="36" captionText="Figure 28. Geographical distribution of the species of Heteropachylus." figureDoi="http://doi.org/10.5281/zenodo.5441446" httpUri="https://zenodo.org/record/5441446/files/figure.png" pageId="40" pageNumber="477">Fig. 28</figureCitation>
):
</emphasis>
Species known only from its type locality, in the original description Santos,
<collectingCountry id="56B47674FFB7FFACD8AAEED25CFD47B9" box="[689,759,486,508]" name="Brazil" pageId="40" pageNumber="477">Brazil</collectingCountry>
(
<bibRefCitation id="4A324B15FFB7FFACDA8CED315F7B445F" author="Roewer CF" box="[151,369,517,538]" pageId="40" pageNumber="477" pagination="1 - 256" refId="ref31985" refString="Roewer CF. 1913 a. Die Familie der Gonyleptiden der Opiliones-Laniatores. Archiv fur Naturgeschichte 4: 1 - 256." type="journal article" year="1913">Roewer, 1913a: 77</bibRefCitation>
). However, on the
<typeStatus id="F1188846FFB7FFACD84FED315CBF445E" box="[596,693,517,539]" pageId="40" pageNumber="477" type="holotype">holotype</typeStatus>
label only ‘Brasilien’ is given, without mention of the exact locality. It could be a mistake or ‘Santos’ in this case may correspond to ‘
<collectingRegion id="EC67F806FFB7FFACDB77ED545FBB4433" box="[364,433,608,630]" country="Brazil" name="Bahia" pageId="40" pageNumber="477">Bahia</collectingRegion>
de Todos os Santos’, and not to the town Santos in
<collectingRegion id="EC67F806FFB7FFACDB94ED4B5C0844D0" box="[399,514,639,661]" country="Brazil" name="Sao Paulo" pageId="40" pageNumber="477">São Paulo</collectingRegion>
Brazilian state. This would be reasonable considering the distribution of the genus. In the map (
<figureCitation id="B6982A61FFB7FFACDBD8ED885C1E4497" box="[451,532,700,722]" captionStart="Figure 28" captionStartId="36.[144,223,927,946]" captionTargetBox="[177,1389,200,897]" captionTargetId="graphics-419@36.[220,813,219,871]" captionTargetPageId="36" captionText="Figure 28. Geographical distribution of the species of Heteropachylus." figureDoi="http://doi.org/10.5281/zenodo.5441446" httpUri="https://zenodo.org/record/5441446/files/figure.png" pageId="40" pageNumber="477">Fig. 28</figureCitation>
), this species was plotted in Santos but with a question mark.
</paragraph>
<paragraph id="2E1C36E4FFB7FFACDA8BEC285FB4450A" blockId="40.[144,759,796,848]" pageId="40" pageNumber="477">
<emphasis id="1CD7EAF6FFB7FFACDA8BEC285F7D4574" box="[144,375,796,817]" italics="true" pageId="40" pageNumber="477">Material examined:</emphasis>
<materialsCitation id="9ECB3CB9FFB7FFACDB93EC285FB1450A" collectionCode="SMF" country="Brazil" location="BRAZIL" pageId="40" pageNumber="477" specimenCount="1" typeStatus="lectotype">
m
<typeStatus id="F1188846FFB7FFACDBB4EC285C1F4577" box="[431,533,796,818]" pageId="40" pageNumber="477" type="lectotype">lectotype</typeStatus>
(here designated) (
<collectionCode id="48B2AE21FFB7FFACDA8CEC0E5EC5450A" box="[151,207,826,847]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="40" pageNumber="477">SMF</collectionCode>
RI – 798),
<collectingCountry id="56B47674FFB7FFACDB43EC0E5FB1450A" box="[344,443,826,847]" name="Brazil" pageId="40" pageNumber="477">BRAZIL</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="2E1C36E4FFB7FFACDA8BEC4F5F2545F5" blockId="40.[144,758,891,944]" pageId="40" pageNumber="477">
<materialsCitation id="9ECB3CB9FFB7FFACDA8BEC4F5F2145F5" collectionCode="SMF" country="Brazil" location="BRAZIL" pageId="40" pageNumber="477" specimenCount="2" typeStatus="paralectotype">
<emphasis id="1CD7EAF6FFB7FFACDA8BEC4F5F4045D5" box="[144,330,891,912]" italics="true" pageId="40" pageNumber="477">Other material:</emphasis>
f
<typeStatus id="F1188846FFB7FFACDB6EEC4F5C1A45D4" box="[373,528,891,913]" pageId="40" pageNumber="477" type="paralectotype">paralectotype</typeStatus>
(
<collectionCode id="48B2AE21FFB7FFACD833EC4F5C6A45D5" box="[552,608,891,912]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="40" pageNumber="477">SMF</collectionCode>
RI – 798),
<collectingCountry id="56B47674FFB7FFACDA8BECAE5EF845EA" box="[144,242,922,943]" name="Brazil" pageId="40" pageNumber="477">BRAZIL</collectingCountry>
, lost
</materialsCitation>
.
</paragraph>
<paragraph id="2E1C36E4FFB7FFACDA8BECEF5FC143FE" blockId="40.[144,759,986,1467]" pageId="40" pageNumber="477">
<emphasis id="1CD7EAF6FFB7FFACDA8BECEF5F0C45B5" box="[144,262,987,1008]" italics="true" pageId="40" pageNumber="477">Diagnosis:</emphasis>
Four mesotergal areas (
<figureCitation id="B6982A61FFB7FFACD806ECEE5C8B45B5" box="[541,641,986,1008]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
); mesotergal area I divided by longitudinal groove; complete groove between mesotergal areas II and III (
<figureCitation id="B6982A61FFB7FFACD89CEB235CE0426B" box="[647,746,1047,1070]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
); mesotergal area III bearing pair of paramedian tubercles and at the same level as the other areas (
<figureCitation id="B6982A61FFB7FFACDA83EB475F1442CC" box="[152,286,1139,1161]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, D</figureCitation>
); mesotergal area IV divided by longitudinal groove (
<figureCitation id="B6982A61FFB7FFACDB21EBA65F9442ED" box="[314,414,1170,1192]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
). Free tergites I–III with a tubercle on each corner (
<figureCitation id="B6982A61FFB7FFACDBB7EB845C1A4282" box="[428,528,1200,1223]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
); free tergites II–III bearing median conical apophysis (
<figureCitation id="B6982A61FFB7FFACD822EBFB5CE042A0" box="[569,746,1231,1253]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, B, D</figureCitation>
); free tergite III without keel and not fused to the dorsal scutum in males (
<figureCitation id="B6982A61FFB7FFACDBAAEA385C504367" box="[433,602,1292,1314]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, B, D</figureCitation>
). Dorsal anal operculum bearing median apophysis (
<figureCitation id="B6982A61FFB7FFACD87BEA1F5CE04305" box="[608,746,1323,1345]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23B, D</figureCitation>
). Femur IV (
<figureCitation id="B6982A61FFB7FFACDB0BEA7D5F554325" box="[272,351,1353,1376]" captionStart="Figure 24" captionStartId="33.[164,243,1369,1388]" captionTargetBox="[176,757,778,1339]" captionTargetId="graphics-488@33.[176,750,1056,1339]" captionTargetPageId="33" captionText="Figure 24. Heteropachylus spiniger. Male lectotype SMF RI 798. A, right trochanter, femur, and patella IV. Dorsal view. B, right trochanter and femur. Ventral view. C, right femur and patella IV. Prolateral view. The arrow indicates the anvil-shaped dorsobasal apophysis. D, right femur and patella IV. Retrolateral view. The arrow indicates the two subdistal tubercles. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441438" httpUri="https://zenodo.org/record/5441438/files/figure.png" pageId="40" pageNumber="477">Fig. 24</figureCitation>
) with anvil-shaped dorsobasal apophysis and lacking dorsodistal one.
<emphasis id="1CD7EAF6FFB7FFACD850EA5C5CFD4338" box="[587,759,1384,1405]" italics="true" pageId="40" pageNumber="477">Male genitalia</emphasis>
(
<figureCitation id="B6982A61FFB7FFACDA83EAB35F2843D9" box="[152,290,1414,1437]" captionStart="Figure 25" captionStartId="34.[144,223,1747,1766]" captionTargetBox="[206,1359,196,1716]" captionTargetPageId="34" captionText="Figure 25. Male genitalia of species of Heteropachylus. A, B, Heteropachylus ramphonotus sp. nov. holotype IBSP 994. A, dorsal view; B, lateral view. C, D, Heteropachylus inexpectabilis comb. nov. MNRJ. C, dorsal view; D, lateral view. E, F, Heteropachylus crassicalcanei comb. nov. MNRJ 5216. E, dorsal view; F, lateral view. G, H. Heteropachylus gracilis sp. nov. paratype IBSP 450. G, dorsal view; H, lateral view. I, J, Heteropachylus peracchii HEMS 665. I, dorsal view; J, lateral view. K, L, Heteropachylus spiniger lectotype SMF RI 798. K, dorsal view; L, lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441440" httpUri="https://zenodo.org/record/5441440/files/figure.png" pageId="40" pageNumber="477">Fig. 25K, L</figureCitation>
). The four pairs of distal setae equal sized and forming a group.
</paragraph>
<paragraph id="2E1C36E4FFB7FFACDA8BEAD35FF9405F" blockId="40.[144,758,1510,1562]" pageId="40" pageNumber="477">
<emphasis id="1CD7EAF6FFB7FFACDA8BEAD35F3043B9" box="[144,314,1511,1532]" italics="true" pageId="40" pageNumber="477">Measurements:</emphasis>
Male
<typeStatus id="F1188846FFB7FFACDB8AEAD25FFF43B9" box="[401,501,1510,1532]" pageId="40" pageNumber="477" type="holotype">holotype</typeStatus>
. CL: 1.7; MCW: 2.4; ASL: 3.2; MASW: 4.7. FL: 2.9.
</paragraph>
<paragraph id="2E1C36E4FFB7FFACDA8BE9725F1B401E" blockId="40.[144,759,1606,1903]" box="[144,273,1606,1627]" pageId="40" pageNumber="477">
<emphasis id="1CD7EAF6FFB7FFACDA8BE9725F1B401E" box="[144,273,1606,1627]" italics="true" pageId="40" pageNumber="477">Description</emphasis>
</paragraph>
<paragraph id="2E1C36E4FFB7FFADDA8BE9505CFE477D" blockId="40.[144,759,1606,1903]" lastBlockId="41.[164,779,198,312]" lastPageId="41" lastPageNumber="478" pageId="40" pageNumber="477">
<emphasis id="1CD7EAF6FFB7FFACDA8BE9505FAD403F" box="[144,423,1636,1658]" italics="true" pageId="40" pageNumber="477">
Male
<typeStatus id="F1188846FFB7FFACDAC3E9505F3D403C" box="[216,311,1636,1657]" pageId="40" pageNumber="477" type="holotype">holotype</typeStatus>
: Dorsum
</emphasis>
(
<figureCitation id="B6982A61FFB7FFACDBA4E9505C29403F" box="[447,547,1636,1658]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
). Scutum outline pyriform, widest at mesotergal area II. Anterior margin of carapace with three to four granules at each side; frontal hump covered with granules. Eye mound with pair of large paramedian tubercles (
<figureCitation id="B6982A61FFB7FFACDA83E9C95F154156" box="[152,287,1789,1811]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, C</figureCitation>
), with some smaller granules near the eyes. Carapace smooth. Mesotergum divided into four areas; mesotergal area I divided into left and right halves by median longitudinal groove; mesotergal area III with pair of paramedian tubercles; mesotergal area IV divided into left and right halves by longitudinal groove. Posterior margin with transversal row of granules. Free tergites I–II fused to the scutum, each with a transversal row of granules and with one tubercle on each corner (
<figureCitation id="B6982A61FFB7FFACDEDAEE6B5B404731" box="[1217,1354,351,373]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, D</figureCitation>
); free tergite II bearing median curved apophysis, pointed backwards (
<figureCitation id="B6982A61FFB7FFACD9AFEEA85A6847F4" box="[948,1122,412,434]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, B, D</figureCitation>
); free tergite III smooth, with posterior border slightly convex, bearing one tubercle on each corner and a median curved apophysis pointed backwards (
<figureCitation id="B6982A61FFB7FFACDE23EECC5AB44448" box="[1080,1214,504,526]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, D</figureCitation>
). Lateral areas of mesotergum each with row of tubercles that increase in size towards the posterior margin of dorsal scutum, reaching mesotergal area II (
<figureCitation id="B6982A61FFB7FFACDE9CED605AE1442F" box="[1159,1259,596,618]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
). Dorsal anal operculum covered with rounded tubercles, bearing median apophysis pointed backwards (
<figureCitation id="B6982A61FFB7FFACDEECEDA55B8B44E3" box="[1271,1409,657,679]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23B, D</figureCitation>
).
<emphasis id="1CD7EAF6FFB7FFACD93DED9B5DBC4481" box="[806,950,687,708]" italics="true" pageId="40" pageNumber="477">Appendages.</emphasis>
Pedipalps (
<figureCitation id="B6982A61FFB7FFACDE4BED9B5AD74480" box="[1104,1245,687,709]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A, D</figureCitation>
) – trochanter with two ventral setiferous granules. Femur with ventral row of setiferous granules and with subapical mesal spine. Tibia with four mesal (IiIi) and three ectal (I + Ii) spines. Tarsus with four mesal (IiIi) and four ectal (IiIi) spines. Legs – calcaneus I less than half the length of metatarsus I, lighter in colour, and thicker than astragalus. Coxa IV (
<figureCitation id="B6982A61FFB7FFACDED5ECB25B2545D9" box="[1230,1327,902,924]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
) armed with apical prolateral blunt apophysis. Trochanter IV (
<figureCitation id="B6982A61FFB7FFACD935ECF75D9A459C" box="[814,912,963,985]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23A</figureCitation>
) armed with apical prolateral blunt apophysis and distal retrolateral curved apophysis. Femur IV (
<figureCitation id="B6982A61FFB7FFACD94AEB345DAA4253" box="[849,928,1024,1046]" captionStart="Figure 24" captionStartId="33.[164,243,1369,1388]" captionTargetBox="[176,757,778,1339]" captionTargetId="graphics-488@33.[176,750,1056,1339]" captionTargetPageId="33" captionText="Figure 24. Heteropachylus spiniger. Male lectotype SMF RI 798. A, right trochanter, femur, and patella IV. Dorsal view. B, right trochanter and femur. Ventral view. C, right femur and patella IV. Prolateral view. The arrow indicates the anvil-shaped dorsobasal apophysis. D, right femur and patella IV. Retrolateral view. The arrow indicates the two subdistal tubercles. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441438" httpUri="https://zenodo.org/record/5441438/files/figure.png" pageId="40" pageNumber="477">Fig. 24</figureCitation>
) short, distinctly curved in dorsal view and with two curvatures (‘S-shaped’) in lateral view, bearing rows of tubercles; with anvil-shaped dorsbasal apophysis preceded by a basal rounded tubercle; dorsal surface with three rows of rounded tubercles equal sized, the distal tubercles forming a circle; retrolateral surface with two subdistal rounded large tubercles; prolateral surface with row of blunt apophysis located at the distal two-thirds, larger at the median portion of the row; apex (
<figureCitation id="B6982A61FFB7FFACDEA8EA275B1E436C" box="[1203,1300,1299,1321]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23E</figureCitation>
) with retrolateral conical apophysis, pointed upwards in apical view, and prolateral rounded apophysis of acuminate apex. Tarsal counts: 6/8–9/6/6.
<emphasis id="1CD7EAF6FFB7FFACDECEEA5B5B8743C1" box="[1237,1421,1391,1412]" italics="true" pageId="40" pageNumber="477">Male genitalia</emphasis>
(
<figureCitation id="B6982A61FFB7FFACD935EABA5DB943E6" box="[814,947,1422,1444]" captionStart="Figure 23" captionStartId="32.[144,223,1558,1577]" captionTargetBox="[143,1424,196,1528]" captionTargetId="graphics-129@32.[145,1368,197,1528]" captionTargetPageId="32" captionText="Figure 23. Heteropachylus spiniger. Male lectotype SMF RI 798. A, habitus, dorsal view. B, ventroposterior view. C, frontal view. D, habitus, lateral view. E, apical view of right femur IV. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5441436" httpUri="https://zenodo.org/record/5441436/files/figure.png" pageId="40" pageNumber="477">Fig. 23K, L</figureCitation>
). Ventral plate subrectangular, narrower in basal portion, with distal border distinctly concave; dorsolateral surface excavated, seeming to be divided into ventral and dorsal layers. Group of four distal short setae on the dorsal portion, two short distal blunt setae on the ventral portion of ventral plate, one large seta after group of distal setae.
<taxonomicName id="E9A34D67FFB7FFACDF51E9715B87401E" authorityName="Megerle von Muhlfeld" authorityYear="1811" box="[1354,1421,1605,1627]" class="Bivalvia" family="Carditidae" genus="Glans" kingdom="Animalia" order="Carditida" pageId="40" pageNumber="444" phylum="Mollusca" rank="genus">Glans</taxonomicName>
without dorsal process.
<taxonomicName id="E9A34D67FFB7FFACDE27E9505A8E403F" box="[1084,1156,1636,1658]" class="Pilidiophora" family="Lineidae" genus="Stylus" higherTaxonomySource="CoL" kingdom="Animalia" order="Heteronemertea" pageId="40" pageNumber="444" phylum="Nemertea" rank="genus">Stylus</taxonomicName>
with apical setae subequally sized.
<emphasis id="1CD7EAF6FFB7FFACD9CBE9B65AB840D2" box="[976,1202,1666,1688]" italics="true" pageId="40" pageNumber="477">Colour (in alcohol).</emphasis>
As it is old material, the examined exemplar has lost its colour and is entirely dark yellow. The original description says that the species body and all the appendages are rusty-brown (‘Färbung des Körpers und sämtlicher Gliedmaben einfarbig rostbraun’).
<emphasis id="1CD7EAF6FFB7FFACDEAAE82F5B014175" box="[1201,1291,1819,1840]" italics="true" pageId="40" pageNumber="477">Female.</emphasis>
Translated from original description: apical apophysis of coxa IV reduced and spiniform. Trochanter IV narrow, armed with a distal spiniform apophysis without conspicuous armature on femur IV. Free tergites not fused to the dorsal scutum, free tergite II bearing a small median spine and free tergite III a small tubercle.
</paragraph>
<paragraph id="2E1C36E4FFB6FFADDABFEE6B5CFB4795" blockId="41.[164,779,351,464]" pageId="41" pageNumber="478">
<emphasis id="1CD7EAF6FFB6FFADDABFEE6B5EE34731" box="[164,233,351,372]" italics="true" pageId="41" pageNumber="478">Notes:</emphasis>
The original description was based on
<typeStatus id="F1188846FFB6FFADD8C0EE6B5EE947D6" pageId="41" pageNumber="478" type="syntype">syntypes</typeStatus>
, a male and a female. The male was here designed as
<typeStatus id="F1188846FFB6FFADDB5DEEA85FA547F7" box="[326,431,412,434]" pageId="41" pageNumber="478" type="lectotype">lectotype</typeStatus>
, although the female, that would have been the unique
<typeStatus id="F1188846FFB6FFADDBE2EE8E5C9C4795" box="[505,662,442,464]" pageId="41" pageNumber="478" type="paralectotype">paralectotype</typeStatus>
, is lost.
</paragraph>
</subSubSection>
</treatment>
</document>
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